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<document id="0A45666119DC2F40726C97B0CA7EF5BA" ID-DOI="10.11865/zs.20150303" ID-ISSN="2095-6827" ID-Zenodo-Dep="7176340" ID-ZooBank="6BBCCA0D-1141-4204-BFFA-DB70EFB62202" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665296370487" checkinUser="felipe" docAuthor="Luo, Ying &amp; Chen, Jian" docDate="2015" docId="03FC87B5FFCE6557FF2DFED4FB73FABD" docLanguage="en" docName="ZoolSyst.40.3.258–267.pdf" docOrigin="Zoological Systematics 40 (3)" docSource="http://zoobank.org/6bbcca0d-1141-4204-bffa-db70efb62202" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Coelotes tiantangensis Luo &amp; Chen 2015, sp. nov." docType="treatment" docVersion="3" lastPageNumber="267" masterDocId="FFC5FFCDFFC8655EFFA5FFECFFE7FF91" masterDocTitle="Two new Coelotes species from China (Araneae: Agelenidae: Coelotinae)" masterLastPageNumber="267" masterPageNumber="258" pageNumber="264" updateTime="1699381651166" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="8A15E5B8F92F7316CAD66088B88D963D">
<mods:title id="2D9110F942A75CFD3353C439693253ED">Two new Coelotes species from China (Araneae: Agelenidae: Coelotinae)</mods:title>
</mods:titleInfo>
<mods:name id="5F8C2C6D2FAC33CC351B93F4863BF52F" type="personal">
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<mods:namePart id="6B5D09107D5623D5947E8173841AD317">Luo, Ying</mods:namePart>
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<mods:namePart id="C850EAC7ACE5D059E2432C54E7DBAEDA">Chen, Jian</mods:namePart>
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<mods:title id="F8F1A44045ADD3DA28DF7C7D4129C7CB">Zoological Systematics</mods:title>
</mods:titleInfo>
<mods:part id="43FAB18F7EF5E7E03F0107BB5BB31F39">
<mods:date id="6A38A07BE6089948B772D50C94EA0C84">2015</mods:date>
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<mods:number id="514EDD99EE4B71E2FEBBDC2A5E7CB376">40</mods:number>
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<mods:number id="0A3710A9DBA7DD25133042E1CB0C78C4">3</mods:number>
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<mods:classification id="2A8732C01220F220BCB5B3DA7E5D9769">journal article</mods:classification>
<mods:identifier id="870B499E8506003E70C48D01CC0AE720" type="DOI">10.11865/zs.20150303</mods:identifier>
<mods:identifier id="97574BBF357A5B52FE9822DE26D25737" type="ISSN">2095-6827</mods:identifier>
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<treatment id="03FC87B5FFCE6557FF2DFED4FB73FABD" LSID="urn:lsid:plazi:treatment:03FC87B5FFCE6557FF2DFED4FB73FABD" httpUri="http://treatment.plazi.org/id/03FC87B5FFCE6557FF2DFED4FB73FABD" lastPageId="9" lastPageNumber="267" pageId="6" pageNumber="264">
<subSubSection id="C34F6528FFCE6558FF2DFED4FD98FEC1" box="[136,639,312,337]" pageId="6" pageNumber="264" type="nomenclature">
<paragraph id="8BEA36A3FFCE6558FF2DFED4FD98FEC1" blockId="6.[136,1452,312,635]" box="[136,639,312,337]" pageId="6" pageNumber="264">
<heading id="D0A281CFFFCE6558FF2DFED4FD98FEC1" box="[136,639,312,337]" fontSize="10" level="2" pageId="6" pageNumber="264" reason="2">
<emphasis id="B921EAB1FFCE6558FF2DFED4FE3CFEC0" bold="true" box="[136,475,312,337]" pageId="6" pageNumber="264">
<taxonomicName id="4C554D20FFCE6558FF2DFED4FE98FEC1" ID-CoL="WTSD" authority="Luo &amp; Chen, 2015" authorityName="Luo &amp; Chen" authorityYear="2015" box="[136,383,312,336]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="264" phylum="Arthropoda" rank="species" species="tiantangensis" status="sp. nov.">
<emphasis id="B921EAB1FFCE6558FF2DFED4FE98FEC1" bold="true" box="[136,383,312,336]" italics="true" pageId="6" pageNumber="264">Coelotes tiantangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257CAFFCE6558FE23FED5FE3CFEC0" box="[390,475,313,337]" pageId="6" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="136E2A26FFCE6558FE58FED5FD90FEC1" box="[509,631,312,337]" captionStart-0="Figs 15–16" captionStart-1="Figs 17–19" captionStart-2="Figs 20–24" captionStart-3="Figs 25–28" captionStart-4="Fig" captionStartId-0="5.[155,196,1876,1898]" captionStartId-1="6.[155,196,1875,1897]" captionStartId-2="7.[155,196,1930,1952]" captionStartId-3="8.[155,196,1244,1266]" captionStartId-4="9.[347,382,994,1016]" captionTargetBox-0="[230,1356,739,1821]" captionTargetBox-1="[204,1389,686,1818]" captionTargetBox-2="[360,1237,243,1890]" captionTargetBox-3="[206,1385,275,1197]" captionTargetBox-4="[342,1249,266,951]" captionTargetId-0="figure-305@5.[229,1363,739,1828]" captionTargetId-1="figure-227@6.[191,1401,680,1844]" captionTargetId-2="figure-52@7.[217,1374,237,1900]" captionTargetId-3="figure-327@8.[206,1385,275,1197]" captionTargetId-4="figure-312@9.[342,1249,266,955]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="Figs 15–16. Coelotes tiantangensis sp. nov., male, holotype. 15. Left male palp, prolateral view. 16. Ditto, retrolateral view. Scale bars = 0.2 mm. Abbreviations: C — conductor; CDA— conductor dorsal apophysis; CF — cymbial furrow; E — embolus; LTA— lateral tibial apophysis; MA—median apophysis; PA—patellar apophysis; RTA—retrolateral tibial apophysis; ST—subtegulum; T —tegulum; TS—tegular sclerite." captionText-1="Figs 17–19. Coelotes tiantangensis sp. nov. 17. Left male palp, ventral view, holotype. 18. Epigynum, ventral view, female, paratype. 19. Vulva, dorsal, female, paratype. Scale bars = 0.2 mm. Abbreviations: A— atrium; C—conductor; CD— copulatory duct; E—embolus; ET— epigynal teeth; FD— fertilization duct; H —hood; MA—median apophysis; S— spermathecae; SH— spermathecal head; TS—tegular sclerite." captionText-2="Figs 20–24. Coelotes tiantangensis sp. nov. 20. Left male palp, prolateral. 21. Ditto, retrolateral. 22. Left male palp, ventral view, holotype. 23. Epigynum, ventral view, female, paratype. 24. Vulva, dorsal view, female, paratype. Scale bars =0.2mm." captionText-3="Figs 25–28. Coelotes tiantangensis sp. nov. 25. Male, dorsal view, holotype. 26. Ditto, ventral view. 27. Female, dorsal view, paratype. 28. Ditto, ventral view. Scale bars=1 mm." captionText-4="Fig. 29. Loaction map of the Tiantangzhai National Forest Park." figureDoi-0="http://doi.org/10.5281/zenodo.7176350" figureDoi-1="http://doi.org/10.5281/zenodo.7176354" figureDoi-2="http://doi.org/10.5281/zenodo.7176356" figureDoi-3="http://doi.org/10.5281/zenodo.7176358" figureDoi-4="http://doi.org/10.5281/zenodo.7176360" httpUri-0="https://zenodo.org/record/7176350/files/figure.png" httpUri-1="https://zenodo.org/record/7176354/files/figure.png" httpUri-2="https://zenodo.org/record/7176356/files/figure.png" httpUri-3="https://zenodo.org/record/7176358/files/figure.png" httpUri-4="https://zenodo.org/record/7176360/files/figure.png" pageId="6" pageNumber="264">Figs 15–29</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F6528FFCE6557FF18FE9CFB73FABD" lastPageId="9" lastPageNumber="267" pageId="6" pageNumber="264" type="materials_examined">
<paragraph id="8BEA36A3FFCE6558FF18FE9CFBC0FE5C" blockId="6.[136,1452,312,635]" pageId="6" pageNumber="264">
Material examined.
<materialsCitation id="3B3D3CFEFFCE6558FE3DFE9CFE9FFE3A" collectingDate="2012-09-27" collectorName="Ying Luo &amp; Jie Liu." country="China" location="Tiantangzhai National Forest Park" municipality="Tiantangzhai National Forest" pageId="6" pageNumber="264" specimenCount="1" specimenCount-male="1" stateProvince="Hubei" typeStatus="holotype">
<typeStatus id="54EE8801FFCE6558FE3DFE9CFE1CFE19" box="[408,507,368,392]" pageId="6" pageNumber="264" type="holotype">Holotype</typeStatus>
<specimenCount id="9D53FD2AFFCE6558FDA7FE81FDF1FE19" box="[514,534,365,392]" pageId="6" pageNumber="264" type="male">♂</specimenCount>
,
<location id="8E8A6078FFCE6558FD81FE9CFC70FE19" LSID="urn:lsid:plazi:treatment:03FC87B5FFCE6557FF2DFED4FB73FABD:8E8A6078FFCE6558FD81FE9CFC70FE19" box="[548,919,368,392]" country="China" municipality="Tiantangzhai National Forest" name="Tiantangzhai National Forest Park" pageId="6" pageNumber="264" stateProvince="Hubei">
<collectingMunicipality id="6B8EACD9FFCE6558FD81FE9CFCBBFE19" box="[548,860,368,392]" pageId="6" pageNumber="264">Tiantangzhai National Forest</collectingMunicipality>
Park
</location>
,
<collectingRegion id="4991F841FFCE6558FC07FE9CFBAAFE19" box="[930,1101,368,392]" country="China" name="Hubei" pageId="6" pageNumber="264">Hubei Province</collectingRegion>
,
<collectingCountry id="F3427633FFCE6558FBFDFE9CFB7DFE19" box="[1112,1178,368,392]" name="China" pageId="6" pageNumber="264">China</collectingCountry>
,
<date id="FFEB1063FFCE6558FB00FE9CFA9EFE19" box="[1189,1401,368,392]" pageId="6" pageNumber="264" value="2012-09-27">
<collectingDate id="EFAFE98BFFCE6558FB00FE9CFA9EFE19" box="[1189,1401,368,392]" pageId="6" pageNumber="264" value="2012-09-27">27 September 2012</collectingDate>
</date>
, leg.
<collectorName id="26A05375FFCE6558FF2DFE7FFF08FE3B" box="[136,239,403,427]" pageId="6" pageNumber="264">Ying Luo</collectorName>
and
<collectorName id="26A05375FFCE6558FE83FE7FFE9FFE3A" box="[294,376,403,427]" pageId="6" pageNumber="264">Jie Liu.</collectorName>
</materialsCitation>
<materialsCitation id="3B3D3CFEFFCE6558FE25FE7FFCFCFE3A" box="[384,795,400,427]" collectingDate="2012-09-27" collectorName="Ying Luo &amp; Jie Liu." country="China" location="Tiantangzhai National Forest Park" municipality="Tiantangzhai National Forest" pageId="6" pageNumber="264" specimenCount="8" specimenCount-female="8" stateProvince="Hubei" typeStatus="paratype">
<typeStatus id="54EE8801FFCE6558FE25FE7FFE0BFE3A" box="[384,492,403,427]" pageId="6" pageNumber="264" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D53FD2AFFCE6558FE52FE7FFDFEFE3A" box="[503,537,400,427]" pageId="6" pageNumber="264" type="female">8♀</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="3B3D3CFEFFCE6558FC83FE7FFBC3FE5C" collectingDate="2009-09-28" collectionCode="CBEE" collectorName="Hai-Juan Xie &amp; Xin Xu. All" country="China" location="Tiantangzhai National Forest Park" municipality="Tiantangzhai National Forest" pageId="6" pageNumber="264" specimenCount="4" specimenCount-male="4" stateProvince="Anhui" typeStatus="paratype">
<specimenCount id="9D53FD2AFFCE6558FC83FE7FFCAFFE3A" box="[806,840,400,427]" pageId="6" pageNumber="264" type="male">4♂</specimenCount>
,
<location id="8E8A6078FFCE6558FCF2FE7FFB2BFE3A" LSID="urn:lsid:plazi:treatment:03FC87B5FFCE6557FF2DFED4FB73FABD:8E8A6078FFCE6558FCF2FE7FFB2BFE3A" box="[855,1228,403,427]" country="China" municipality="Tiantangzhai National Forest" name="Tiantangzhai National Forest Park" pageId="6" pageNumber="264" stateProvince="Anhui">
<collectingMunicipality id="6B8EACD9FFCE6558FCF2FE7FFB77FE3B" box="[855,1168,403,427]" pageId="6" pageNumber="264">Tiantangzhai National Forest</collectingMunicipality>
Park
</location>
,
<collectingRegion id="4991F841FFCE6558FB7DFE7FFA62FE3A" box="[1240,1413,403,427]" country="China" name="Anhui" pageId="6" pageNumber="264">Anhui Province</collectingRegion>
,
<date id="FFEB1063FFCE6558FA34FE7FFEDEFE5C" pageId="6" pageNumber="264" value="2009-09-28">
<collectingDate id="EFAFE98BFFCE6558FA34FE7FFEDEFE5C" pageId="6" pageNumber="264" value="2009-09-28">28 September 2009</collectingDate>
</date>
, leg.
<collectorName id="26A05375FFCE6558FED4FE59FE18FE5C" box="[369,511,437,461]" pageId="6" pageNumber="264">Hai-Juan Xie</collectorName>
and
<collectorName id="26A05375FFCE6558FD96FE59FD57FE5C" box="[563,688,437,461]" pageId="6" pageNumber="264">Xin Xu. All</collectorName>
specimens are deposited in
<collectionCode id="ED44AE66FFCE6558FC79FE59FBC3FE5C" box="[988,1060,437,461]" pageId="6" pageNumber="264">CBEE</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BEA36A3FFCE6558FF18FE34FD94FD82" blockId="6.[136,1452,312,635]" pageId="6" pageNumber="264">
Etymology. The specific name is an adjective, referring to the
<typeStatus id="54EE8801FFCE6558FCF9FE35FC6DFE60" box="[860,906,473,497]" pageId="6" pageNumber="264">type</typeStatus>
locality, Tiantangzhai, in which “Tiantang” means paradise, and “zhai” means stockaded village.
</paragraph>
<paragraph id="8BEA36A3FFCE6558FF18FDF1FB42FDEA" blockId="6.[136,1452,312,635]" pageId="6" pageNumber="264">
Diagnosis. This new species is similar to
<taxonomicName id="4C554D20FFCE6558FD2FFDF2FCDDFDA4" authority="Luo &amp; Chen, 2015" authorityName="Luo &amp; Chen" authorityYear="2015" box="[650,826,541,565]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="264" phylum="Arthropoda" rank="species" species="obtusangulus" status="sp. nov.">
<emphasis id="B921EAB1FFCE6558FD2FFDF2FCDDFDA4" box="[650,826,541,565]" italics="true" pageId="6" pageNumber="264">C. obtusangulus</emphasis>
</taxonomicName>
<emphasis id="B921EAB1FFCE6558FCE6FDF1FC7BFDA4" bold="true" box="[835,924,541,565]" pageId="6" pageNumber="264">
<taxonomicNameLabel id="A21257CAFFCE6558FCE6FDF1FC7BFDA4" box="[835,924,541,565]" pageId="6" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but can be distinguished from the latter by the shorter patellar apophysis, the RTA with distal end not extending beyond tibia, the large epigynal teeth closed with each other (
<figureCitation id="136E2A26FFCE6558FF35FD8FFF32FDEA" box="[144,213,611,635]" captionStart="Figs 1–2" captionStartId="1.[155,196,1875,1897]" captionTargetBox="[229,1356,736,1831]" captionTargetId="figure-183@1.[229,1363,730,1838]" captionTargetPageId="1" captionText="Figs 1–2. Coelotes obtusangulus sp. nov., male, holotype. 1. Left male palp, prolateral view. 2. Ditto, retrolateral view. Scale bars= 0.2 mm. Abbreviations: C — conductor; CDA—conductor dorsal apophysis; CF — cymbial furrow; E— embolus; LTA— lateral tibial apophysis; MA— median apophysis; PA—patellar apophysis; RTA—retrolateral tibial apophysis; ST— subtegulum; T— tegulum; TS—tegular sclerite." figureDoi="http://doi.org/10.5281/zenodo.7176342" httpUri="https://zenodo.org/record/7176342/files/figure.png" pageId="6" pageNumber="264">Figs 2</figureCitation>
,
<figureCitation id="136E2A26FFCE6558FF45FD8FFF17FDEA" box="[224,240,611,635]" captionStart="Figs 3–5" captionStartId="2.[155,196,1856,1878]" captionTargetBox="[203,1387,764,1798]" captionTargetId="figure-181@2.[191,1401,756,1814]" captionTargetPageId="2" captionText="Figs 3–5. Coelotes obtusangulus sp. nov. 3. Left male palp, ventral view, holotype. 4. Epigynum, ventral view, female, paratype. 5. Vulva, dorsal view, female, paratype. Scale bars=0.2 mm. Abbreviations: A— atrium; C—conductor; CDA—conductor dorsal apophysis; E — embolus; ET —epigynal teeth; MA — median apophysis; PA — patellar apophysis; RTA — retrolateral tibial apophysis; S— spermathecae; SH—spermathecal head; TS— tegular sclerite." figureDoi="http://doi.org/10.5281/zenodo.7176344" httpUri="https://zenodo.org/record/7176344/files/figure.png" pageId="6" pageNumber="264">4</figureCitation>
,
<figureCitation id="136E2A26FFCE6558FF5EFD8FFEFFFDEA" box="[251,280,611,635]" captionStart="Figs 15–16" captionStartId="5.[155,196,1876,1898]" captionTargetBox="[230,1356,739,1821]" captionTargetId="figure-305@5.[229,1363,739,1828]" captionTargetPageId="5" captionText="Figs 15–16. Coelotes tiantangensis sp. nov., male, holotype. 15. Left male palp, prolateral view. 16. Ditto, retrolateral view. Scale bars = 0.2 mm. Abbreviations: C — conductor; CDA— conductor dorsal apophysis; CF — cymbial furrow; E — embolus; LTA— lateral tibial apophysis; MA—median apophysis; PA—patellar apophysis; RTA—retrolateral tibial apophysis; ST—subtegulum; T —tegulum; TS—tegular sclerite." figureDoi="http://doi.org/10.5281/zenodo.7176350" httpUri="https://zenodo.org/record/7176350/files/figure.png" pageId="6" pageNumber="264">16</figureCitation>
,
<figureCitation id="136E2A26FFCE6558FE86FD8FFEA7FDEA" box="[291,320,611,635]" captionStart="Figs 17–19" captionStartId="6.[155,196,1875,1897]" captionTargetBox="[204,1389,686,1818]" captionTargetId="figure-227@6.[191,1401,680,1844]" captionTargetPageId="6" captionText="Figs 17–19. Coelotes tiantangensis sp. nov. 17. Left male palp, ventral view, holotype. 18. Epigynum, ventral view, female, paratype. 19. Vulva, dorsal, female, paratype. Scale bars = 0.2 mm. Abbreviations: A— atrium; C—conductor; CD— copulatory duct; E—embolus; ET— epigynal teeth; FD— fertilization duct; H —hood; MA—median apophysis; S— spermathecae; SH— spermathecal head; TS—tegular sclerite." figureDoi="http://doi.org/10.5281/zenodo.7176354" httpUri="https://zenodo.org/record/7176354/files/figure.png" pageId="6" pageNumber="264">18</figureCitation>
). More details can be seen under the above diagnosis of
<taxonomicName id="4C554D20FFCE6558FC3EFD8FFBAEFDEA" authority="Luo &amp; Chen, 2015" authorityName="Luo &amp; Chen" authorityYear="2015" box="[923,1097,611,635]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="264" phylum="Arthropoda" rank="species" species="obtusangulus" status="sp. nov.">
<emphasis id="B921EAB1FFCE6558FC3EFD8FFBAEFDEA" box="[923,1097,611,635]" italics="true" pageId="6" pageNumber="264">C. obtusangulus</emphasis>
</taxonomicName>
<emphasis id="B921EAB1FFCE6558FBF5FD8FFB42FDEA" bold="true" box="[1104,1189,611,635]" pageId="6" pageNumber="264">
<taxonomicNameLabel id="A21257CAFFCE6558FBF5FD8FFB42FDEA" box="[1104,1189,611,635]" pageId="6" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<caption id="DF2A662BFFCE6558FF3EF8BFFDCFF858" ID-DOI="http://doi.org/10.5281/zenodo.7176354" ID-Zenodo-Dep="7176354" httpUri="https://zenodo.org/record/7176354/files/figure.png" pageId="6" pageNumber="264" startId="6.[155,196,1875,1897]" targetBox="[204,1389,686,1818]" targetPageId="6">
<paragraph id="8BEA36A3FFCE6558FF3EF8BFFDCFF858" blockId="6.[155,1433,1875,1993]" pageId="6" pageNumber="264">
Figs 17–19.
<taxonomicName id="4C554D20FFCE6558FEB8F8BFFE1BF8F9" authority="Luo &amp; Chen, 2015" authorityName="Luo &amp; Chen" authorityYear="2015" box="[285,508,1875,1896]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="264" phylum="Arthropoda" rank="species" species="tiantangensis" status="sp. nov.">
<emphasis id="B921EAB1FFCE6558FEB8F8BFFE1BF8F9" box="[285,508,1875,1896]" italics="true" pageId="6" pageNumber="264">Coelotes tiantangensis</emphasis>
</taxonomicName>
<emphasis id="B921EAB1FFCE6558FDA2F8BFFDBEF8F9" bold="true" box="[519,601,1875,1896]" pageId="6" pageNumber="264">
<taxonomicNameLabel id="A21257CAFFCE6558FDA2F8BFFDBEF8F9" box="[519,601,1875,1896]" pageId="6" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
17. Left male palp, ventral view, holotype. 18. Epigynum, ventral view, female, paratype. 19. Vulva, dorsal, female, paratype. Scale bars = 0.2 mm. Abbreviations: A— atrium; C — conductor; CD — copulatory duct; E — embolus; ET — epigynal teeth; FD — fertilization duct; H — hood; MA— median apophysis; S — spermathecae; SH — spermathecal head; TS — tegular sclerite.
</paragraph>
</caption>
<paragraph id="8BEA36A3FFCE6558FBB1F7C6FA4BF7D0" blockId="6.[1044,1452,2090,2113]" box="[1044,1452,2090,2113]" pageId="6" pageNumber="264">
©
<emphasis id="B921EAB1FFCE6558FB88F7C7FAECF7D1" box="[1069,1291,2091,2112]" italics="true" pageId="6" pageNumber="264">Zoological Systematics</emphasis>
, 40(3): 258–267
</paragraph>
<caption id="DF2A662BFFCF6559FF3EF866FAE8F851" ID-DOI="http://doi.org/10.5281/zenodo.7176356" ID-Zenodo-Dep="7176356" httpUri="https://zenodo.org/record/7176356/files/figure.png" pageId="7" pageNumber="265" startId="7.[155,196,1930,1952]" targetBox="[360,1237,243,1890]" targetPageId="7">
<paragraph id="8BEA36A3FFCF6559FF3EF866FAE8F851" blockId="7.[155,1433,1930,1984]" pageId="7" pageNumber="265">
Figs 20–24.
<taxonomicName id="4C554D20FFCF6559FEB3F866FE15F831" authority="Luo &amp; Chen, 2015" authorityName="Luo &amp; Chen" authorityYear="2015" box="[278,498,1930,1952]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="265" phylum="Arthropoda" rank="species" species="tiantangensis" status="sp. nov.">
<emphasis id="B921EAB1FFCF6559FEB3F866FE15F831" box="[278,498,1930,1952]" italics="true" pageId="7" pageNumber="265">Coelotes tiantangensis</emphasis>
</taxonomicName>
<emphasis id="B921EAB1FFCF6559FE5FF867FDAFF831" bold="true" box="[506,584,1931,1952]" pageId="7" pageNumber="265">
<taxonomicNameLabel id="A21257CAFFCF6559FE5FF867FDAFF831" box="[506,584,1931,1952]" pageId="7" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
20. Left male palp, prolateral. 21. Ditto, retrolateral. 22. Left male palp, ventral view, holotype. 23. Epigynum, ventral view, female, paratype. 24. Vulva, dorsal view, female, paratype. Scale bars = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BEA36A3FFCF6559FF2DF7C6FDF8F7D0" blockId="7.[136,543,2090,2113]" box="[136,543,2090,2113]" pageId="7" pageNumber="265">
©
<emphasis id="B921EAB1FFCF6559FF05F7C7FE98F7D1" box="[160,383,2091,2112]" italics="true" pageId="7" pageNumber="265">Zoological Systematics</emphasis>
, 40(3): 258–267
</paragraph>
<caption id="DF2A662BFFC06556FF3EFB30FD68FA80" ID-DOI="http://doi.org/10.5281/zenodo.7176358" ID-Zenodo-Dep="7176358" httpUri="https://zenodo.org/record/7176358/files/figure.png" pageId="8" pageNumber="266" startId="8.[155,196,1244,1266]" targetBox="[206,1385,275,1197]" targetPageId="8">
<paragraph id="8BEA36A3FFC06556FF3EFB30FD68FA80" blockId="8.[155,1433,1244,1298]" pageId="8" pageNumber="266">
Figs 25–28.
<taxonomicName id="4C554D20FFC06556FEBEFB30FE1EFB63" authority="Luo &amp; Chen, 2015" authorityName="Luo &amp; Chen" authorityYear="2015" box="[283,505,1244,1266]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="266" phylum="Arthropoda" rank="species" species="tiantangensis" status="sp. nov.">
<emphasis id="B921EAB1FFC06556FEBEFB30FE1EFB63" box="[283,505,1244,1266]" italics="true" pageId="8" pageNumber="266">Coelotes tiantangensis</emphasis>
</taxonomicName>
<emphasis id="B921EAB1FFC06556FDA6FB30FDB3FB60" bold="true" box="[515,596,1244,1265]" pageId="8" pageNumber="266">
<taxonomicNameLabel id="A21257CAFFC06556FDA6FB30FDB3FB60" box="[515,596,1244,1265]" pageId="8" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
25. Male, dorsal view, holotype. 26. Ditto, ventral view. 27. Female, dorsal view, paratype. 28. Ditto, ventral view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BEA36A3FFC06556FF18FAB6FB2CF9F4" blockId="8.[136,1452,1370,1949]" pageId="8" pageNumber="266">
Description. Male (
<typeStatus id="54EE8801FFC06556FE29FAB6FE17FAE3" box="[396,496,1370,1394]" pageId="8" pageNumber="266" type="holotype">holotype</typeStatus>
). Total length 5.21. Prosoma 2.53 long, 1.94 wide; opisthosoma 2.68 long, 1.73 wide. Eye AME 0.08, ALE 0.13, PME 0.13, PLE 0.13, AME–AME 0.05, AME– ALE 0.05, ALE–PLE 0.04, PME–PME 0.08, PME–PLE 0.10. Leg formula
<collectionCode id="ED44AE66FFC06556FE69FA4CFE00FA26" box="[460,487,1440,1463]" pageId="8" pageNumber="266">IV</collectionCode>
, I,
<collectionCode id="ED44AE66FFC06556FDACFA4CFDFAFA26" box="[521,541,1440,1463]" pageId="8" pageNumber="266">II</collectionCode>
,
<collectionCode id="ED44AE66FFC06556FD8DFA4CFDA1FA26" box="[552,582,1440,1463]" pageId="8" pageNumber="266">III</collectionCode>
; leg I 7.50 (2.15, 2.55, 1.55, 1.25);
<collectionCode id="ED44AE66FFC06556FC66FA4CFC32FA26" box="[963,981,1440,1463]" pageId="8" pageNumber="266">II</collectionCode>
6.40 (1.70, 2.15, 1.50, 1.05);
<collectionCode id="ED44AE66FFC06556FAB3FA4CFAD7FA26" box="[1302,1328,1440,1463]" pageId="8" pageNumber="266">III</collectionCode>
6.20 (1.75, 1.85, 1.60, 1.00);
<collectionCode id="ED44AE66FFC06556FEF0FA2FFE97FA4B" box="[341,368,1475,1498]" pageId="8" pageNumber="266">IV</collectionCode>
8.70 (2.30, 2.75, 2.40, 1.25). Chelicerae with 3 promarginal and 2 retromarginal teeth. Patellar apophysis short, bifurcated; retolateral tibial apophysis large and broad, with end not extending beyond tibia; lateral tibial apophysis slightly broad, closed to retolateral tibial apophysis; cymbial furrow short, no more than half the length of the cymbium; conductor broad and complex, significantly curved, almost &quot;
<collectionCode id="ED44AE66FFC06556FC04F9C7FC5EF9D2" box="[929,953,1579,1603]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="266" type="Museum">V</collectionCode>
&quot; shaped; conductor dorsal apophysis absent; median apophysis large and round; embolus filiform, originating prolaterally (
<figureCitation id="136E2A26FFC06556FC6FF9A1FBFAF9F4" box="[970,1053,1613,1637]" captionStart="Figs 15–16" captionStartId="5.[155,196,1876,1898]" captionTargetBox="[230,1356,739,1821]" captionTargetId="figure-305@5.[229,1363,739,1828]" captionTargetPageId="5" captionText="Figs 15–16. Coelotes tiantangensis sp. nov., male, holotype. 15. Left male palp, prolateral view. 16. Ditto, retrolateral view. Scale bars = 0.2 mm. Abbreviations: C — conductor; CDA— conductor dorsal apophysis; CF — cymbial furrow; E — embolus; LTA— lateral tibial apophysis; MA—median apophysis; PA—patellar apophysis; RTA—retrolateral tibial apophysis; ST—subtegulum; T —tegulum; TS—tegular sclerite." figureDoi="http://doi.org/10.5281/zenodo.7176350" httpUri="https://zenodo.org/record/7176350/files/figure.png" pageId="8" pageNumber="266">Figs 15</figureCitation>
,
<figureCitation id="136E2A26FFC06556FB8DF9A1FBA2F9F4" box="[1064,1093,1613,1637]" captionStart="Figs 17–19" captionStartId="6.[155,196,1875,1897]" captionTargetBox="[204,1389,686,1818]" captionTargetId="figure-227@6.[191,1401,680,1844]" captionTargetPageId="6" captionText="Figs 17–19. Coelotes tiantangensis sp. nov. 17. Left male palp, ventral view, holotype. 18. Epigynum, ventral view, female, paratype. 19. Vulva, dorsal, female, paratype. Scale bars = 0.2 mm. Abbreviations: A— atrium; C—conductor; CD— copulatory duct; E—embolus; ET— epigynal teeth; FD— fertilization duct; H —hood; MA—median apophysis; S— spermathecae; SH— spermathecal head; TS—tegular sclerite." figureDoi="http://doi.org/10.5281/zenodo.7176354" httpUri="https://zenodo.org/record/7176354/files/figure.png" pageId="8" pageNumber="266">17</figureCitation>
,
<figureCitation id="136E2A26FFC06556FBF5F9A1FB8AF9F4" box="[1104,1133,1613,1637]" captionStart="Figs 20–24" captionStartId="7.[155,196,1930,1952]" captionTargetBox="[360,1237,243,1890]" captionTargetId="figure-52@7.[217,1374,237,1900]" captionTargetPageId="7" captionText="Figs 20–24. Coelotes tiantangensis sp. nov. 20. Left male palp, prolateral. 21. Ditto, retrolateral. 22. Left male palp, ventral view, holotype. 23. Epigynum, ventral view, female, paratype. 24. Vulva, dorsal view, female, paratype. Scale bars =0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7176356" httpUri="https://zenodo.org/record/7176356/files/figure.png" pageId="8" pageNumber="266">22</figureCitation>
,
<figureCitation id="136E2A26FFC06556FBDDF9A1FB59F9F4" box="[1144,1214,1613,1637]" captionStart="Figs 25–28" captionStartId="8.[155,196,1244,1266]" captionTargetBox="[206,1385,275,1197]" captionTargetId="figure-327@8.[206,1385,275,1197]" captionTargetPageId="8" captionText="Figs 25–28. Coelotes tiantangensis sp. nov. 25. Male, dorsal view, holotype. 26. Ditto, ventral view. 27. Female, dorsal view, paratype. 28. Ditto, ventral view. Scale bars=1 mm." figureDoi="http://doi.org/10.5281/zenodo.7176358" httpUri="https://zenodo.org/record/7176358/files/figure.png" pageId="8" pageNumber="266">25, 26</figureCitation>
).
</paragraph>
<paragraph id="8BEA36A3FFC06556FF18F99CFCCCF8EB" blockId="8.[136,1452,1370,1949]" pageId="8" pageNumber="266">
Female (
<typeStatus id="54EE8801FFC06556FEBCF99CFE9CF919" box="[281,379,1648,1672]" pageId="8" pageNumber="266" type="paratype">paratype</typeStatus>
). Total length 4.96. Prosoma 2.58 long, 1.75 wide; opisthosoma 2.38 long, 1.65 wide. Eye AME 0.08, ALE 0.14, PME 0.13, PLE 0.14, AME–AME 0.04, AME–ALE 0.05, ALE–PLE 0.03, PME–PME 0.08, PME–PLE 0.11. Leg formula
<collectionCode id="ED44AE66FFC06556FF41F95AFEE7F95C" box="[228,256,1718,1741]" pageId="8" pageNumber="266">IV</collectionCode>
, I,
<collectionCode id="ED44AE66FFC06556FE87F95AFED1F95C" box="[290,310,1718,1741]" pageId="8" pageNumber="266">II</collectionCode>
,
<collectionCode id="ED44AE66FFC06556FEE7F95AFE87F95C" box="[322,352,1718,1741]" pageId="8" pageNumber="266">III</collectionCode>
; leg I 6.30 (1.60, 2.35, 1.35, 1.00);
<collectionCode id="ED44AE66FFC06556FD7BF95AFD17F95C" box="[734,752,1718,1741]" pageId="8" pageNumber="266">II</collectionCode>
5.65 (1.60, 1.95, 1.15, 0.95);
<collectionCode id="ED44AE66FFC06556FB96F95AFBA9F95C" box="[1075,1102,1718,1741]" pageId="8" pageNumber="266">III</collectionCode>
5.38 (1.50, 1.65, 1.35, 0.88);
<collectionCode id="ED44AE66FFC06556FA35F95AFA4CF95C" box="[1424,1451,1718,1741]" pageId="8" pageNumber="266">IV</collectionCode>
7.47 (1.94, 2.47, 2.01, 1.05). Chelicerae with 3 promarginal and 2 retromarginal teeth. Epigynal teeth lamella large, broad, slightly longer than wide, situated anteriorly and medially, closed to each other basically; the epigynal hoods present, situated anteriorly and laterally; atrium large, with a broad septum, atrium extending into epigynum, forming 2 round copulatory duct openings; copulatory duct short; spermathecae oval, slightly separated; spermathecal heads short, situated anteriorly laterad of spermathecae (
<figureCitation id="136E2A26FFC06556FDA7F88EFD9AF8EB" box="[514,637,1890,1914]" captionStart="Figs 17–19" captionStartId="6.[155,196,1875,1897]" captionTargetBox="[204,1389,686,1818]" captionTargetId="figure-227@6.[191,1401,680,1844]" captionTargetPageId="6" captionText="Figs 17–19. Coelotes tiantangensis sp. nov. 17. Left male palp, ventral view, holotype. 18. Epigynum, ventral view, female, paratype. 19. Vulva, dorsal, female, paratype. Scale bars = 0.2 mm. Abbreviations: A— atrium; C—conductor; CD— copulatory duct; E—embolus; ET— epigynal teeth; FD— fertilization duct; H —hood; MA—median apophysis; S— spermathecae; SH— spermathecal head; TS—tegular sclerite." figureDoi="http://doi.org/10.5281/zenodo.7176354" httpUri="https://zenodo.org/record/7176354/files/figure.png" pageId="8" pageNumber="266">Figs 18–19</figureCitation>
,
<figureCitation id="136E2A26FFC06556FD2DF88EFD2AF8EB" box="[648,717,1890,1914]" captionStart="Figs 20–24" captionStartId="7.[155,196,1930,1952]" captionTargetBox="[360,1237,243,1890]" captionTargetId="figure-52@7.[217,1374,237,1900]" captionTargetPageId="7" captionText="Figs 20–24. Coelotes tiantangensis sp. nov. 20. Left male palp, prolateral. 21. Ditto, retrolateral. 22. Left male palp, ventral view, holotype. 23. Epigynum, ventral view, female, paratype. 24. Vulva, dorsal view, female, paratype. Scale bars =0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7176356" httpUri="https://zenodo.org/record/7176356/files/figure.png" pageId="8" pageNumber="266">23–24</figureCitation>
,
<figureCitation id="136E2A26FFC06556FD7DF88EFCF8F8EB" box="[728,799,1890,1914]" captionStart="Figs 25–28" captionStartId="8.[155,196,1244,1266]" captionTargetBox="[206,1385,275,1197]" captionTargetId="figure-327@8.[206,1385,275,1197]" captionTargetPageId="8" captionText="Figs 25–28. Coelotes tiantangensis sp. nov. 25. Male, dorsal view, holotype. 26. Ditto, ventral view. 27. Female, dorsal view, paratype. 28. Ditto, ventral view. Scale bars=1 mm." figureDoi="http://doi.org/10.5281/zenodo.7176358" httpUri="https://zenodo.org/record/7176358/files/figure.png" pageId="8" pageNumber="266">27–28</figureCitation>
).
</paragraph>
<paragraph id="8BEA36A3FFC06556FF18F869FD7EF80C" blockId="8.[136,1452,1370,1949]" box="[189,665,1925,1949]" pageId="8" pageNumber="266">
Distribution.
<collectingCountry id="F3427633FFC06556FEE9F869FE6BF80C" box="[332,396,1925,1949]" name="China" pageId="8" pageNumber="266">China</collectingCountry>
(
<collectingRegion id="4991F841FFC06556FE3CF869FE07F80C" box="[409,480,1925,1949]" country="China" name="Anhui" pageId="8" pageNumber="266">Anhui</collectingRegion>
,
<collectingRegion id="4991F841FFC06556FE4EF869FDC9F80C" box="[491,558,1925,1949]" country="China" name="Hubei" pageId="8" pageNumber="266">Hubei</collectingRegion>
) (
<figureCitation id="136E2A26FFC06556FDE6F869FD6BF80C" box="[579,652,1925,1949]" captionStart="Fig" captionStartId="9.[347,382,994,1016]" captionTargetBox="[342,1249,266,951]" captionTargetId="figure-312@9.[342,1249,266,955]" captionTargetPageId="9" captionText="Fig. 29. Loaction map of the Tiantangzhai National Forest Park." figureDoi="http://doi.org/10.5281/zenodo.7176360" httpUri="https://zenodo.org/record/7176360/files/figure.png" pageId="8" pageNumber="266">Fig 29</figureCitation>
).
</paragraph>
<paragraph id="8BEA36A3FFC06557FBB1F7C6FEAFFB0A" blockId="8.[1044,1452,2090,2113]" lastBlockId="9.[136,1452,1085,1179]" lastPageId="9" lastPageNumber="267" pageId="8" pageNumber="266">
©
<emphasis id="B921EAB1FFC06556FB88F7C7FAECF7D1" box="[1069,1291,2091,2112]" italics="true" pageId="8" pageNumber="266">Zoological Systematics</emphasis>
, 40(3): 258–267
<emphasis id="B921EAB1FFC16557FF2DFBD1FF0FFBC4" bold="true" box="[136,232,1085,1109]" inLineHeading="true" pageId="9" pageNumber="267" reason="1">Funding</emphasis>
This study was financially supported by the National Natural Sciences Foundation of
<collectingCountry id="F3427633FFC16557FB42FBD1FAC0FBC4" box="[1255,1319,1085,1109]" name="China" pageId="9" pageNumber="267">China</collectingCountry>
(31172113, 31272268) and the Special Foundation of the Ministry of Science and Technology of the
<collectingCountry id="F3427633FFC16557FBD9FB8CFA4BFBE9" box="[1148,1452,1120,1144]" name="China" pageId="9" pageNumber="267">People's Republic of China</collectingCountry>
(2014FY110100).
</paragraph>
<caption id="DF2A662BFFC16557FEFEFC0EFC24FC69" ID-DOI="http://doi.org/10.5281/zenodo.7176360" ID-Zenodo-Dep="7176360" box="[347,963,994,1016]" httpUri="https://zenodo.org/record/7176360/files/figure.png" pageId="9" pageNumber="267" startId="9.[347,382,994,1016]" targetBox="[342,1249,266,951]" targetPageId="9">
<paragraph id="8BEA36A3FFC16557FEFEFC0EFC24FC69" blockId="9.[347,963,994,1016]" box="[347,963,994,1016]" pageId="9" pageNumber="267">Fig. 29. Loaction map of the Tiantangzhai National Forest Park.</paragraph>
</caption>
<paragraph id="8BEA36A3FFC16557FF2DFB23FB73FABD" blockId="9.[136,1452,1231,1325]" pageId="9" pageNumber="267">
<emphasis id="B921EAB1FFC16557FF2DFB23FE82FB76" bold="true" box="[136,357,1231,1255]" inLineHeading="true" pageId="9" pageNumber="267" reason="1">Acknowledgements</emphasis>
We thank Hai-Juan Xie and Xin Xu (
<collectionCode id="ED44AE66FFC16557FCA7FB23FCAEFB76" box="[770,841,1231,1255]" pageId="9" pageNumber="267">CBEE</collectionCode>
) for providing specimens. The manuscript benefited from comments by Jie Liu (
<collectionCode id="ED44AE66FFC16557FE28FB1DFE33FA98" box="[397,468,1265,1289]" pageId="9" pageNumber="267">CBEE</collectionCode>
) and Jeremy Miller (Department of Terrestrial Zoology, Naturalis Biodiversity Center,
<collectingCountry id="F3427633FFC16557FF2DFAF8FEE9FABD" box="[136,270,1300,1324]" name="Netherlands" pageId="9" pageNumber="267">Netherlands</collectingCountry>
). We are grateful to the anonymous reviewers for their comments on the manuscript.
</paragraph>
</subSubSection>
</treatment>
</document>
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