523 lines
81 KiB
XML
523 lines
81 KiB
XML
<document id="557B89673B832B58EE9DDBAE3A778B0D" ID-DOI="10.5281/zenodo.282163" ID-GBIF-Dataset="478af170-cc64-4fc1-9425-8f2cdf3f3ea5" ID-ISSN="1175-5326" ID-Zenodo-Dep="282163" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460515836557" checkinUser="plazi" docAuthor="Benzoni, Francesca & Stefani, Fabrizio" docDate="2012" docId="03FC87ADFFBD9B15FF0AC60FB1DCF81B" docLanguage="en" docName="zt03447p068.pdf" docOrigin="Zootaxa 3447" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Porites fontanesii Benzoni & Stefani, 2012, sp. nov." docType="treatment" docVersion="10" lastPageNumber="61" masterDocId="FFC5FFD5FFBF9B10FF9DC13DB32FFFC5" masterDocTitle="Porites fontanesii, a new species of hard coral (Scleractinia, Poritidae) from the southern Red Sea, the Gulf of Tadjoura, and the Gulf of Aden, and its phylogenetic relationships within the genus" masterLastPageNumber="68" masterPageNumber="56" pageNumber="58" updateTime="1698313974647" updateUser="plazi">
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<mods:titleInfo id="89AE98ED4AED46C50A9919B4D9E58405">
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<mods:title id="833EF96A6ABDC1900241A36CBB89B8A3">Porites fontanesii, a new species of hard coral (Scleractinia, Poritidae) from the southern Red Sea, the Gulf of Tadjoura, and the Gulf of Aden, and its phylogenetic relationships within the genus</mods:title>
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</mods:titleInfo>
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<mods:name id="7FE1B86A131C5B509AFBF4412E463ADD" type="personal">
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<mods:roleTerm id="E7027DF72938C577D2F1CF9220318D4E">Author</mods:roleTerm>
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<mods:namePart id="B4F32BD35408BA58F78E1AC122D1C6BE">Benzoni, Francesca</mods:namePart>
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</mods:name>
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<mods:name id="D5AF78D6DC8EE1D9FD26402DF81A840A" type="personal">
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<mods:role id="752957B12255F51E0102096FA214B913">
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<mods:roleTerm id="5F2F50D747369C9DAFDCF794D3D595A5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4BAF8B2756874E6D8AE8F3650BCF5289">Stefani, Fabrizio</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A5C7BA730E29ED457F988E07CED7B823">text</mods:typeOfResource>
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<mods:title id="FB04EFA2B8989B939DC2A835C1ED38E9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="4189E5F5AFFA0C7E0DCA882740FC5CD9">
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<mods:date id="3E0751759B26DEEFC4805087ED041A2B">2012</mods:date>
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<mods:detail id="29B6BC21085C9E70E4F13FC4771CC740" type="volume">
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<mods:number id="8B9F4B9E071C5610835E1EDDF1ECBE70">3447</mods:number>
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<mods:start id="6315826ABAFD3D899B8D385F6DCB6126">56</mods:start>
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<mods:classification id="A9242D6F07DC3BD5924C34CA18F2D29B">journal article</mods:classification>
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<mods:identifier id="5FB4418FAE03281CAFA2478FEC23FF8F" type="DOI">10.5281/zenodo.282163</mods:identifier>
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<mods:identifier id="1BCE5C0EA3BD56111B42A42C94906717" type="GBIF-Dataset">478af170-cc64-4fc1-9425-8f2cdf3f3ea5</mods:identifier>
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<mods:identifier id="DE7686A4DAD2A348E78C283449957FFA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A508922E28F9C098FAE0AD1C408E1C17" type="Zenodo-Dep">282163</mods:identifier>
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<treatment id="03FC87ADFFBD9B15FF0AC60FB1DCF81B" ID-DOI="http://doi.org/10.5281/zenodo.6173752" ID-GBIF-Taxon="119523935" ID-Zenodo-Dep="6173752" LSID="urn:lsid:zoobank.org:act:9A01A99B-5160-4A7B-B968-CAA4CD28B0AC" httpUri="http://treatment.plazi.org/id/03FC87ADFFBD9B15FF0AC60FB1DCF81B" lastPageId="5" lastPageNumber="61" pageId="2" pageNumber="58">
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<subSubSection id="C34F6530FFBD9B12FF0AC60FB218F8A8" pageId="2" pageNumber="58" type="nomenclature">
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<paragraph id="8BEA36BBFFBD9B12FF0AC60FB2E0F889" blockId="2.[151,463,1842,1901]" box="[151,463,1842,1868]" pageId="2" pageNumber="58">
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<heading id="D0A281D7FFBD9B12FF0AC60FB2E0F889" bold="true" box="[151,463,1842,1868]" fontSize="11" level="1" pageId="2" pageNumber="58" reason="1">
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<emphasis id="B921EAA9FFBD9B12FF0AC60FB2E0F889" bold="true" box="[151,463,1842,1868]" pageId="2" pageNumber="58">
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<taxonomicName id="4C554D38FFBD9B12FF0AC60FB244F889" ID-CoL="4LXSZ" box="[151,363,1842,1868]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="58" phylum="Cnidaria" rank="species" species="fontanesii" status="sp. nov.">
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<emphasis id="B921EAA9FFBD9B12FF0AC60FB244F889" bold="true" box="[151,363,1842,1868]" italics="true" pageId="2" pageNumber="58">Porites fontanesii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21257D2FFBD9B12FEEFC60FB2E0F889" box="[370,463,1842,1868]" pageId="2" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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<paragraph id="8BEA36BBFFBD9B12FF0AC668B218F8A8" blockId="2.[151,463,1842,1901]" box="[151,311,1877,1901]" pageId="2" pageNumber="58">
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<figureCitation id="136E2A3EFFBD9B12FF0AC668B3CCF8A8" box="[151,227,1877,1901]" captionStart="FIGURE 1" captionStartId="4.[151,250,1233,1256]" captionTargetBox="[151,1435,190,1211]" captionTargetId="figure@4.[151,1436,190,1212]" captionTargetPageId="4" captionText="FIGURE 1 Holotype of Porites fontanesii sp. nov. (MNHN-IK. 2009 - 834) (A) In vivo image of the whole colony; (B) The holotype; (C) Detail of a digitation of the holotype typical of the species; (D) Close up of C showing smooth colony surface, regular corallite arrangement and coenosteum; (E) Detail of D showing the corallite septal plan." httpUri="https://zenodo.org/record/282164/files/figure.png" pageId="2" pageNumber="58">Figs. 1</figureCitation>
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,
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<figureCitation id="136E2A3EFFBD9B12FF6CC668B22EF8A8" box="[241,257,1877,1901]" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="2" pageNumber="58">2</figureCitation>
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,
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<figureCitation id="136E2A3EFFBD9B12FE90C668B233F8A8" box="[269,284,1877,1901]" captionStart="FIGURE 3" captionStartId="6.[151,250,1867,1890]" captionTargetBox="[176,1411,195,1822]" captionTargetId="figure@6.[176,1413,195,1844]" captionTargetPageId="6" captionText="FIGURE 3 Porites fontanesii sp. nov. in situ (A) A large branching colony at Balhaf, Yemen (depth 5 m); (B) Branching colony at Araher, Socotra Island (depth 15 m); (C) Colony at Tiqfash Island, Yemen (depth 3 m); (D) A beige colony with thick digitations at Musha Island, Djibouti (depth 4 m); (E) Detail of a digitation tip showing the smooth appearance of the colony surface once the polyps are retracted; (F) Close up of a colony at Tiqfash Island, Yemen, showing the polyps usually expanded during the day (depth 7 m); (G) Close-up of E showing living polyps with retracted tentacles." httpUri="https://zenodo.org/record/282166/files/figure.png" pageId="2" pageNumber="58">3</figureCitation>
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,
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<figureCitation id="136E2A3EFFBD9B12FEB4C668B218F8A8" box="[297,311,1877,1901]" captionStart="FIGURE 4" captionStartId="7.[151,250,1914,1937]" captionTargetBox="[175,1398,193,1888]" captionTargetId="figure@7.[175,1399,193,1889]" captionTargetPageId="7" captionText="FIGURE 4 (A – C, E – H, I) Specimens of Porites fontanesii sp. nov. showing the range of variability of the corallum shape and of its dimensions; (D) detail of C showing the usual surface appearance of this species; (J) detail of I showing the flower-like appearance of corallites in specimens with well developed denticles. Specimen registration code close to the scale bar. White scale bar = 1 cm." httpUri="https://zenodo.org/record/282167/files/figure.png" pageId="2" pageNumber="58">4</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34F6530FFBD9B14FF0AC6A0B16CF99E" lastPageId="4" lastPageNumber="60" pageId="2" pageNumber="58" type="materials_examined">
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<paragraph id="8BEA36BBFFBD9B12FF0AC6A0B06CF81C" blockId="2.[151,1437,1949,2009]" pageId="2" pageNumber="58">
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<emphasis id="B921EAA9FFBD9B12FF0AC6A0B2ADF873" bold="true" box="[151,386,1949,1974]" pageId="2" pageNumber="58">Material examined.</emphasis>
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<typeStatus id="54EE8819FFBD9B12FE16C6A0B2D8F870" box="[395,503,1949,1973]" pageId="2" pageNumber="58" type="holotype">Holotype</typeStatus>
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: MNHN-IK.2009-834 Balhaf,
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<collectingCountry id="F342762BFFBD9B12FCC8C6A3B08CF870" box="[853,931,1950,1973]" name="Yemen" pageId="2" pageNumber="58">Yemen</collectingCountry>
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(
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<geoCoordinate id="EE61507CFFBD9B12FC28C6A0B77CF870" box="[949,1107,1949,1973]" direction="north" orientation="latitude" pageId="2" pageNumber="58" precision="1" value="13.970467">13°58.228’ N</geoCoordinate>
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;
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<geoCoordinate id="EE61507CFFBD9B12FBFCC6A0B7D4F870" box="[1121,1275,1949,1973]" direction="east" orientation="longitude" pageId="2" pageNumber="58" precision="1" value="48.179317">48°10.759’ E</geoCoordinate>
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),
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<quantity id="4CAD9B5EFFBD9B12FA8CC6A0B66FF870" box="[1297,1344,1949,1973]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="58" unit="m" value="5.0">5 m</quantity>
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, coll. F. Benzoni & M. Pichon,
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<date id="FFEB107BFFBD9B12FE06C6FCB132F81C" box="[411,541,1985,2009]" pageId="2" pageNumber="58" value="2007-09-20">20/09/2007</date>
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, collection code BAL085.
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</paragraph>
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<paragraph id="8BEA36BBFFBC9B13FF58C1A5B1F9FF32" blockId="3.[153,1437,151,2015]" pageId="3" pageNumber="59">
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<typeStatus id="54EE8819FFBC9B13FF58C1A5B219FF75" box="[197,310,152,176]" pageId="3" pageNumber="59" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="ED44AE7EFFBC9B13FEDDC1AAB2BAFF6A" box="[320,405,151,175]" pageId="3" pageNumber="59">MSNM</collectionCode>
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COE332 Al Mukallah (
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<geoCoordinate id="EE61507CFFBC9B13FD3EC1AAB069FF6A" box="[675,838,151,175]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="14.51765">14° 31.059’ N</geoCoordinate>
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;
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||
<geoCoordinate id="EE61507CFFBC9B13FCCFC1AAB0DEFF6A" box="[850,1009,151,175]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="49.171383">49° 10.283’ E</geoCoordinate>
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),
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<quantity id="4CAD9B5EFFBC9B13FB9BC1AAB769FF6A" box="[1030,1094,151,175]" metricMagnitude="0" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="59" unit="m" value="3.5">3.5 m</quantity>
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coll. M. Pichon & F. Benzoni,
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<date id="FFEB107BFFBC9B13FF04C181B234FF11" box="[153,283,188,212]" pageId="3" pageNumber="59" value="2007-03-17">17/03/2007</date>
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||
, collection code MU003; UNIMIB Y359 Balhaf (
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<geoCoordinate id="EE61507CFFBC9B13FCCDC181B0C2FF11" box="[848,1005,188,212]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="13.969383">13°58.163’ N</geoCoordinate>
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||
;
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||
<geoCoordinate id="EE61507CFFBC9B13FC67C181B7BCFF11" box="[1018,1171,188,212]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="48.182133">48°10.928’ E</geoCoordinate>
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),
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<quantity id="4CAD9B5EFFBC9B13FB35C181B7F9FF11" box="[1192,1238,188,212]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="59" unit="m" value="4.0">4 m</quantity>
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, coll. F. Benzoni,
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<date id="FFEB107BFFBC9B13FF04C1E2B3D3FF32" box="[153,252,223,247]" pageId="3" pageNumber="59" value="2005-09-08">08/09/05</date>
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(GenBank accession number
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<accessionNumber id="9406AB58FFBC9B13FDD1C1E2B1E4FF32" box="[588,715,223,247]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HE585991" pageId="3" pageNumber="59">HE585991</accessionNumber>
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).
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</paragraph>
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||
<paragraph id="8BEA36BBFFBC9B13FF58C03EB648FC61" blockId="3.[153,1437,151,2015]" pageId="3" pageNumber="59">
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<emphasis id="B921EAA9FFBC9B13FF58C03EB16CFED9" bold="true" box="[197,579,259,284]" pageId="3" pageNumber="59">
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Other material:
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<collectingCountry id="F342762BFFBC9B13FE14C039B2F5FED9" box="[393,474,260,284]" name="Yemen" pageId="3" pageNumber="59">Yemen</collectingCountry>
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Red Sea
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</emphasis>
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UNIMIB KA022 Kamaran Island (
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<geoCoordinate id="EE61507CFFBC9B13FC49C039B75FFED9" box="[980,1136,260,284]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="15.3664665">15°21.988’ N</geoCoordinate>
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;
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<geoCoordinate id="EE61507CFFBC9B13FBE0C039B63CFED9" box="[1149,1299,260,284]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="42.625668">42°37.540’ E</geoCoordinate>
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||
) F. Benzoni & A. Caragnano,
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<date id="FFEB107BFFBC9B13FEFEC01AB2C9FEFA" box="[355,486,295,319]" pageId="3" pageNumber="59" value="2009-09-28">28/09/2009</date>
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||
; KA170 Al Badi' (15°29.573’;
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<geoCoordinate id="EE61507CFFBC9B13FCD4C01AB0CFFEFA" box="[841,992,295,319]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="42.53515">42°32.109’ E</geoCoordinate>
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||
)
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||
<date id="FFEB107BFFBC9B13FC6CC01AB75BFEFA" box="[1009,1140,295,319]" pageId="3" pageNumber="59" value="2009-10-02">02/10/2009</date>
|
||
;
|
||
<emphasis id="B921EAA9FFBC9B13FB1DC01AB7CBFE85" bold="true" box="[1152,1252,295,320]" pageId="3" pageNumber="59">
|
||
<collectingCountry id="F342762BFFBC9B13FB1DC01AB7CBFE85" box="[1152,1252,295,320]" name="Djibouti" pageId="3" pageNumber="59">Djibouti</collectingCountry>
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||
</emphasis>
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TO-DJ030, Ras Ali, Gulf of Tadjoura, (
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<geoCoordinate id="EE61507CFFBC9B13FE00C071B117FEA1" box="[413,568,332,356]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="11.772567">11°46.354’ N</geoCoordinate>
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||
;
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||
<geoCoordinate id="EE61507CFFBC9B13FDD8C071B1F2FEA1" box="[581,733,332,356]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="42.954765">42°57.286’ E</geoCoordinate>
|
||
),
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||
<quantity id="4CAD9B5EFFBC9B13FD6FC070B030FEA1" box="[754,799,333,356]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="59" unit="m" value="7.0">7 m</quantity>
|
||
, coll. F. Benzoni,
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||
<date id="FFEB107BFFBC9B13FC77C071B742FEA1" box="[1002,1133,332,356]" pageId="3" pageNumber="59" value="2010-01-28">28/01/2010</date>
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||
; TO-DJ079, Oblal, Gulf of Tadjoura, (
|
||
<geoCoordinate id="EE61507CFFBC9B13FE89C052B29EFE42" box="[276,433,367,391]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="11.811283">11°48.677’ N</geoCoordinate>
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||
;
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||
<geoCoordinate id="EE61507CFFBC9B13FE22C052B164FE42" box="[447,587,367,391]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="43.057117">43°3.427’ E</geoCoordinate>
|
||
),
|
||
<quantity id="4CAD9B5EFFBC9B13FDFFC052B1BEFE42" box="[610,657,367,391]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="59" unit="m" value="8.0">8 m</quantity>
|
||
, coll. F. Benzoni,
|
||
<date id="FFEB107BFFBC9B13FCFFC052B0C9FE42" box="[866,998,367,391]" pageId="3" pageNumber="59" value="2010-01-29">29/01/2010</date>
|
||
; TO-DJ134, Musha, Gulf of Tadjoura, (
|
||
<geoCoordinate id="EE61507CFFBC9B13FF3CC0A9B210FE69" box="[161,319,404,428]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="11.743716">11°44.623’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FED1C0A9B2C9FE69" box="[332,486,404,428]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="43.1686">43°10.116’ E</geoCoordinate>
|
||
),
|
||
<quantity id="4CAD9B5EFFBC9B13FE61C0A9B117FE69" box="[508,568,404,428]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="59" unit="m" value="10.0">10 m</quantity>
|
||
, coll. F. Benzoni,
|
||
<date id="FFEB107BFFBC9B13FC97C0A9B0A2FE69" box="[778,909,404,428]" pageId="3" pageNumber="59" value="2010-01-31">31/01/2010</date>
|
||
;
|
||
<emphasis id="B921EAA9FFBC9B13FC07C0A9B7A0FE69" bold="true" box="[922,1167,403,428]" pageId="3" pageNumber="59">Ye m e n G u lf o f Ade n</emphasis>
|
||
UNIMIB Y358 Balhaf (
|
||
<geoCoordinate id="EE61507CFFBC9B13FF3CC08AB26FFE0A" box="[161,320,439,463]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="13.969383">13°58.163’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FED3C08AB2C9FE0A" box="[334,486,439,463]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="48.182133">48°10.928’ E</geoCoordinate>
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) coll. F. Benzoni,
|
||
<date id="FFEB107BFFBC9B13FD26C08AB030FE0A" box="[699,799,439,463]" pageId="3" pageNumber="59" value="2005-09-08">08/09/05</date>
|
||
(GenBank accession number
|
||
<accessionNumber id="9406AB58FFBC9B13FBEAC08AB7DAFE0A" box="[1143,1269,439,463]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HE585990" pageId="3" pageNumber="59">HE585990</accessionNumber>
|
||
); Y360 Balhaf (
|
||
<geoCoordinate id="EE61507CFFBC9B13FF3CC0E1B26FFE31" box="[161,320,476,500]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="13.969383">13°58.163’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FED3C0E1B2C9FE31" box="[334,486,476,500]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="48.182133">48°10.928’ E</geoCoordinate>
|
||
) coll. F. Benzoni,
|
||
<date id="FFEB107BFFBC9B13FD26C0E1B00FFE31" box="[699,800,476,500]" pageId="3" pageNumber="59" value="2006-04-08">08/04/06</date>
|
||
(GenBank accession number
|
||
<accessionNumber id="9406AB58FFBC9B13FBEAC0E1B7DAFE31" box="[1143,1269,476,500]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HE585992" pageId="3" pageNumber="59">HE585992</accessionNumber>
|
||
); Y694 Balhaf (
|
||
<geoCoordinate id="EE61507CFFBC9B13FF3CC0C2B210FDD2" box="[161,319,511,535]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="13.973367">13°58.402’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FED1C0C2B2CBFDD2" box="[332,484,511,535]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="48.206833">48°12.410’ E</geoCoordinate>
|
||
) coll. F. Benzoni,
|
||
<date id="FFEB107BFFBC9B13FD2AC0C2B031FDD2" box="[695,798,511,535]" pageId="3" pageNumber="59" value="2008-03-10">10/03/08</date>
|
||
; BAL015 Balhaf (
|
||
<geoCoordinate id="EE61507CFFBC9B13FC6DC0C2B7A1FDD2" box="[1008,1166,511,535]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="13.972183">13°58.331’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FB06C0C2B61DFDD2" box="[1179,1330,511,535]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="48.186066">48°11.164’ E</geoCoordinate>
|
||
) coll. M. Pichon & F. Benzoni,
|
||
<date id="FFEB107BFFBC9B13FE0AC319B134FDF9" box="[407,539,548,572]" pageId="3" pageNumber="59" value="2007-03-04">04/03/2007</date>
|
||
; BAL042 (
|
||
<geoCoordinate id="EE61507CFFBC9B13FD07C319B017FDF9" box="[666,824,548,572]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="13.981083">13°58.865’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FCDBC319B0F2FDF9" box="[838,989,548,572]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="48.17685">48°10.611’ E</geoCoordinate>
|
||
) coll. F. Benzoni
|
||
<date id="FFEB107BFFBC9B13FB37C319B601FDF9" box="[1194,1326,548,572]" pageId="3" pageNumber="59" value="2007-05-20">20/05/2007</date>
|
||
; BAL044 (
|
||
<geoCoordinate id="EE61507CFFBC9B13FF3CC37AB26AFD9A" box="[161,325,583,607]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="13.981083">13°58.865’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FEC4C37AB2D9FD9A" box="[345,502,583,607]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="48.17685">48°10.611’ E</geoCoordinate>
|
||
) coll. M. Pichon & F. Benzoni
|
||
<date id="FFEB107BFFBC9B13FC16C37AB722FD9A" box="[907,1037,583,607]" pageId="3" pageNumber="59" value="2007-09-16">16/09/2007</date>
|
||
; BAL069 Balhaf (
|
||
<geoCoordinate id="EE61507CFFBC9B13FB6EC37AB6B8FD9A" box="[1267,1431,583,607]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="13.970467">13°58.228’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FF04C351B21FFD41" box="[153,304,620,644]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="48.179317">48°10.759’ E</geoCoordinate>
|
||
) coll. M. Pichon & F. Benzoni,
|
||
<date id="FFEB107BFFBC9B13FD00C351B00FFD41" box="[669,800,620,644]" pageId="3" pageNumber="59" value="2007-09-20">20/09/2007</date>
|
||
; BAL108 Balhaf (
|
||
<geoCoordinate id="EE61507CFFBC9B13FC6CC351B7A0FD41" box="[1009,1167,620,644]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="13.970467">13°58.228’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FB06C351B61CFD41" box="[1179,1331,620,644]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="48.179317">48°10.759’ E</geoCoordinate>
|
||
) coll. M. Pichon & F. Benzoni,
|
||
<date id="FFEB107BFFBC9B13FE09C3B2B138FD62" box="[404,535,655,679]" pageId="3" pageNumber="59" value="2007-09-20">20/09/2007</date>
|
||
; BAL175b Balhaf (
|
||
<geoCoordinate id="EE61507CFFBC9B13FD6BC3B2B0BCFD62" box="[758,915,655,679]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="13.969383">13°58.163’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FC3DC3B2B717FD62" box="[928,1080,655,679]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="48.182133">48°10.928’ E</geoCoordinate>
|
||
) coll. M. Pichon & F. Benzoni,
|
||
<date id="FFEB107BFFBC9B13FF04C389B234FD09" box="[153,283,692,716]" pageId="3" pageNumber="59" value="2007-09-22">22/09/2007</date>
|
||
; BAL250 Balhaf (
|
||
<geoCoordinate id="EE61507CFFBC9B13FE76C389B1A7FD09" box="[491,648,692,716]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="13.974067">13°58.444’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FD09C389B005FD09" box="[660,810,692,716]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="48.192665">48°11.560’ E</geoCoordinate>
|
||
) coll. M. Pichon & F. Benzoni,
|
||
<date id="FFEB107BFFBC9B13FB0EC389B639FD09" box="[1171,1302,692,716]" pageId="3" pageNumber="59" value="2007-09-23">23/09/2007</date>
|
||
; MU027 Al Mukallah (
|
||
<geoCoordinate id="EE61507CFFBC9B13FE8EC3EAB29AFD2A" box="[275,437,727,751]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="14.51765">14° 31.059’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FE5CC3EAB172FD2A" box="[449,605,727,751]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="49.171383">49° 10.283’ E</geoCoordinate>
|
||
) coll. M. Pichon, F. Benzoni & C. Riva,
|
||
<date id="FFEB107BFFBC9B13FBBEC3EAB789FD2A" box="[1059,1190,727,751]" pageId="3" pageNumber="59" value="2007-03-17">17/03/2007</date>
|
||
; MU091 Al Mukallah (14°
|
||
<geoCoordinate id="EE61507CFFBC9B13FF48C3C1B26AFCD1" box="[213,325,764,788]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="55" value="30.923">30.923 N</geoCoordinate>
|
||
; 49°
|
||
<geoCoordinate id="EE61507CFFBC9B13FE15C3C1B2CAFCD1" box="[392,485,764,788]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="55" value="9.254">9.254 E</geoCoordinate>
|
||
) coll. M. Pichon & F. Benzoni,
|
||
<date id="FFEB107BFFBC9B13FCF1C3C1B0C0FCD1" box="[876,1007,764,788]" pageId="3" pageNumber="59" value="2007-03-17">17/03/2007</date>
|
||
; BA092 Sikha Island (13°
|
||
<geoCoordinate id="EE61507CFFBC9B13FABCC3C1B6B8FCD1" box="[1313,1431,764,788]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="55" value="55.775">55,775 N</geoCoordinate>
|
||
; 48°
|
||
<geoCoordinate id="EE61507CFFBC9B13FF20C222B208FCF2" box="[189,295,799,823]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="55" value="23.219">23,219 E</geoCoordinate>
|
||
)
|
||
<quantity id="4CAD9B5EFFBC9B13FEAAC222B24AFCFD" box="[311,357,799,824]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="59" unit="m" value="8.0">8 m</quantity>
|
||
, coll. F. Benzoni & S. Montano,
|
||
<date id="FFEB107BFFBC9B13FD41C222B075FCF2" box="[732,858,799,823]" pageId="3" pageNumber="59" value="2008-11-21">21/11/2008</date>
|
||
(GenBank accession number
|
||
<accessionNumber id="9406AB58FFBC9B13FB33C222B603FCF2" box="[1198,1324,799,823]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HE585993" pageId="3" pageNumber="59">HE585993</accessionNumber>
|
||
);
|
||
<emphasis id="B921EAA9FFBC9B13FADDC222B3CBFC99" bold="true" pageId="3" pageNumber="59">
|
||
<collectingCountry id="F342762BFFBC9B13FADDC222B6B3FCFD" box="[1344,1436,799,824]" name="Yemen" pageId="3" pageNumber="59">Socotra</collectingCountry>
|
||
Island
|
||
</emphasis>
|
||
UNIMIB SO057 Araher (
|
||
<geoCoordinate id="EE61507CFFBC9B13FD8BC279B19AFC99" box="[534,693,836,860]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="12.582367">12°34.942’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FD59C279B04FFC99" box="[708,864,836,860]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="54.4334">54°26.004’ E</geoCoordinate>
|
||
),
|
||
<quantity id="4CAD9B5EFFBC9B13FCEAC279B09AFC99" box="[887,949,836,860]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="59" unit="m" value="18.0">18 m</quantity>
|
||
, coll. F. Benzoni & A. Caragnano,
|
||
<date id="FFEB107BFFBC9B13FAC9C279B3FAFCBA" pageId="3" pageNumber="59" value="2010-03-13">13/03/ 2010</date>
|
||
; SO114 Ras Qadamah (
|
||
<geoCoordinate id="EE61507CFFBC9B13FE7AC25AB1ADFC45" box="[487,642,871,896]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="1" value="12.698367">12°41.902’ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE61507CFFBC9B13FD00C25AB01AFCBA" box="[669,821,871,895]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="1" value="53.661366">53°39.682’ E</geoCoordinate>
|
||
)
|
||
<quantity id="4CAD9B5EFFBC9B13FCDAC25AB0ABFC45" box="[839,900,871,896]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="59" unit="m" value="10.0">10 m</quantity>
|
||
, coll. F. Benzoni & A. Caragnano
|
||
<date id="FFEB107BFFBC9B13FA88C25AB6B7FCBA" box="[1301,1432,871,895]" pageId="3" pageNumber="59" value="2010-03-17">17/03/2010</date>
|
||
;
|
||
<collectionCode id="ED44AE7EFFBC9B13FF04C2B1B3FEFC61" LSID="urn:lsid:biocol.org:col:34838" box="[153,209,908,932]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="59">SMF</collectionCode>
|
||
(housed at EPA
|
||
<collectingCountry id="F342762BFFBC9B13FE11C2B1B2CCFC61" box="[396,483,908,932]" name="Yemen" pageId="3" pageNumber="59">Socotra</collectingCountry>
|
||
) C90S, ST-145 W Ras Momi,
|
||
<date id="FFEB107BFFBC9B13FCA6C2B1B091FC61" box="[827,958,908,932]" pageId="3" pageNumber="59" value="1999-04-02">02/04/1999</date>
|
||
; C325S, ST-738 Medina,
|
||
<date id="FFEB107BFFBC9B13FB42C2B1B64CFC61" box="[1247,1379,908,932]" pageId="3" pageNumber="59" value="2000-04-18">18/04/2000</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEA36BBFFBC9B13FF58C292B207F9CF" blockId="3.[153,1437,151,2015]" pageId="3" pageNumber="59">
|
||
<emphasis id="B921EAA9FFBC9B13FF58C292B181FC0D" bold="true" box="[197,686,943,968]" pageId="3" pageNumber="59">
|
||
Skeletal characteristics of the
|
||
<typeStatus id="54EE8819FFBC9B13FDDCC28DB185FC0D" box="[577,682,944,968]" pageId="3" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
The
|
||
<typeStatus id="54EE8819FFBC9B13FD6FC292B07BFC02" box="[754,852,943,967]" pageId="3" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="136E2A3EFFBC9B13FCF4C292B080FC02" box="[873,943,943,967]" captionStart="FIGURE 1" captionStartId="4.[151,250,1233,1256]" captionTargetBox="[151,1435,190,1211]" captionTargetId="figure@4.[151,1436,190,1212]" captionTargetPageId="4" captionText="FIGURE 1 Holotype of Porites fontanesii sp. nov. (MNHN-IK. 2009 - 834) (A) In vivo image of the whole colony; (B) The holotype; (C) Detail of a digitation of the holotype typical of the species; (D) Close up of C showing smooth colony surface, regular corallite arrangement and coenosteum; (E) Detail of D showing the corallite septal plan." httpUri="https://zenodo.org/record/282164/files/figure.png" pageId="3" pageNumber="59">Fig. 1</figureCitation>
|
||
) is part of a large colony (
|
||
<figureCitation id="136E2A3EFFBC9B13FB62C292B66BFC02" box="[1279,1348,943,967]" captionStart="FIGURE 1" captionStartId="4.[151,250,1233,1256]" captionTargetBox="[151,1435,190,1211]" captionTargetId="figure@4.[151,1436,190,1212]" captionTargetPageId="4" captionText="FIGURE 1 Holotype of Porites fontanesii sp. nov. (MNHN-IK. 2009 - 834) (A) In vivo image of the whole colony; (B) The holotype; (C) Detail of a digitation of the holotype typical of the species; (D) Close up of C showing smooth colony surface, regular corallite arrangement and coenosteum; (E) Detail of D showing the corallite septal plan." httpUri="https://zenodo.org/record/282164/files/figure.png" pageId="3" pageNumber="59">Fig. 1</figureCitation>
|
||
a). The fragment is
|
||
<quantity id="4CAD9B5EFFBC9B13FEBCC2E9B276FC29" box="[289,345,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="59" unit="cm" value="9.0">9 cm</quantity>
|
||
long,
|
||
<quantity id="4CAD9B5EFFBC9B13FE3FC2E9B2C0FC29" box="[418,495,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="59" unit="cm" value="4.5">4.5 cm</quantity>
|
||
wide, and
|
||
<quantity id="4CAD9B5EFFBC9B13FDF3C2E9B195FC29" box="[622,698,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="59" unit="cm" value="2.5">2.5 cm</quantity>
|
||
thick. It is characterized by an encrusting base from which three taller digitations and one shorter digitiform non-anastomosing projections develop. Digitations tips are rounded (
|
||
<figureCitation id="136E2A3EFFBC9B13FF3CC522B3CEFBF2" box="[161,225,1055,1079]" captionStart="FIGURE 1" captionStartId="4.[151,250,1233,1256]" captionTargetBox="[151,1435,190,1211]" captionTargetId="figure@4.[151,1436,190,1212]" captionTargetPageId="4" captionText="FIGURE 1 Holotype of Porites fontanesii sp. nov. (MNHN-IK. 2009 - 834) (A) In vivo image of the whole colony; (B) The holotype; (C) Detail of a digitation of the holotype typical of the species; (D) Close up of C showing smooth colony surface, regular corallite arrangement and coenosteum; (E) Detail of D showing the corallite septal plan." httpUri="https://zenodo.org/record/282164/files/figure.png" pageId="3" pageNumber="59">Fig. 1</figureCitation>
|
||
A–C). Corallite density over the corallum surface is 50–66 corallites/cm2 (
|
||
<figureCitation id="136E2A3EFFBC9B13FBAFC522B75DFBF2" box="[1074,1138,1055,1079]" captionStart="FIGURE 1" captionStartId="4.[151,250,1233,1256]" captionTargetBox="[151,1435,190,1211]" captionTargetId="figure@4.[151,1436,190,1212]" captionTargetPageId="4" captionText="FIGURE 1 Holotype of Porites fontanesii sp. nov. (MNHN-IK. 2009 - 834) (A) In vivo image of the whole colony; (B) The holotype; (C) Detail of a digitation of the holotype typical of the species; (D) Close up of C showing smooth colony surface, regular corallite arrangement and coenosteum; (E) Detail of D showing the corallite septal plan." httpUri="https://zenodo.org/record/282164/files/figure.png" pageId="3" pageNumber="59">Fig. 1</figureCitation>
|
||
C–D). Corallites are flush with the coenosteum, although they tend be slightly sunken at the base of the digitations. They are separated by coarsely ornamented coenosteum and can be locally arranged in short series 3 to 6 corallites long (
|
||
<figureCitation id="136E2A3EFFBC9B13FAACC55AB65BFBBA" box="[1329,1396,1127,1151]" captionStart="FIGURE 1" captionStartId="4.[151,250,1233,1256]" captionTargetBox="[151,1435,190,1211]" captionTargetId="figure@4.[151,1436,190,1212]" captionTargetPageId="4" captionText="FIGURE 1 Holotype of Porites fontanesii sp. nov. (MNHN-IK. 2009 - 834) (A) In vivo image of the whole colony; (B) The holotype; (C) Detail of a digitation of the holotype typical of the species; (D) Close up of C showing smooth colony surface, regular corallite arrangement and coenosteum; (E) Detail of D showing the corallite septal plan." httpUri="https://zenodo.org/record/282164/files/figure.png" pageId="3" pageNumber="59">Fig. 1</figureCitation>
|
||
D). Collines, or ridges, are not present. Calice diameter ranges from
|
||
<quantity id="4CAD9B5EFFBC9B13FCE6C5B7B724FB67" box="[891,1035,1162,1186]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="3" pageNumber="59" unit="mm" value="0.95" valueMax="1.0" valueMin="0.9">0.9 to 1 mm</quantity>
|
||
. A columella is absent. The typical septa arrangement of the genus
|
||
<taxonomicName id="4C554D38FFBC9B13FD9DC58DB17EFB02" box="[512,593,1200,1223]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="3" pageNumber="59" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B921EAA9FFBC9B13FD9DC58DB17EFB02" box="[512,593,1200,1223]" italics="true" pageId="3" pageNumber="59">Porites</emphasis>
|
||
</taxonomicName>
|
||
is observed with 12 septa arranged in four lateral pairs, one dorsal septum and three ventral septa (
|
||
<figureCitation id="136E2A3EFFBC9B13FE36C5EFB2C3FB2F" box="[427,492,1234,1258]" captionStart="FIGURE 1" captionStartId="4.[151,250,1233,1256]" captionTargetBox="[151,1435,190,1211]" captionTargetId="figure@4.[151,1436,190,1212]" captionTargetPageId="4" captionText="FIGURE 1 Holotype of Porites fontanesii sp. nov. (MNHN-IK. 2009 - 834) (A) In vivo image of the whole colony; (B) The holotype; (C) Detail of a digitation of the holotype typical of the species; (D) Close up of C showing smooth colony surface, regular corallite arrangement and coenosteum; (E) Detail of D showing the corallite septal plan." httpUri="https://zenodo.org/record/282164/files/figure.png" pageId="3" pageNumber="59">Fig. 1</figureCitation>
|
||
E;
|
||
<figureCitation id="136E2A3EFFBC9B13FD8DC5EFB17CFB2F" box="[528,595,1234,1258]" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="3" pageNumber="59">Fig. 2</figureCitation>
|
||
A–B). The dorsal septum is shorter than other septa (larger arrows in
|
||
<figureCitation id="136E2A3EFFBC9B13FAEDC5EFB387FACA" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="3" pageNumber="59">Fig. 2</figureCitation>
|
||
B) and the synapticulae connecting it to one of the two lateral pairs, or both, can be more or less visible from the top (smaller arrows in
|
||
<figureCitation id="136E2A3EFFBC9B13FE3BC427B2C1FAF7" box="[422,494,1306,1330]" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="3" pageNumber="59">Fig. 2</figureCitation>
|
||
B). The ventral triplet is fused (
|
||
<figureCitation id="136E2A3EFFBC9B13FCF9C427B082FAF7" box="[868,941,1306,1330]" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="3" pageNumber="59">Fig. 2</figureCitation>
|
||
B). Five large pali are normally present in corallites, four in front of the four lateral pairs of septa, and one in front of the ventral triplet (
|
||
<figureCitation id="136E2A3EFFBC9B13FB2FC402B7D9FA92" box="[1202,1270,1343,1367]" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="3" pageNumber="59">Fig. 2</figureCitation>
|
||
B). No palus is found in front of the short dorsal septum. Upper septal margins carry 1 finely ornamented denticle (
|
||
<figureCitation id="136E2A3EFFBC9B13FA8EC45FB675FABF" box="[1299,1370,1378,1402]" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="3" pageNumber="59">Fig. 2</figureCitation>
|
||
). The innermost circle of synapticulae connecting the pali is visible and can be thin (
|
||
<figureCitation id="136E2A3EFFBC9B13FB96C4BAB762FA5A" box="[1035,1101,1415,1439]" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="3" pageNumber="59">Fig. 2</figureCitation>
|
||
A–C) or thick (
|
||
<figureCitation id="136E2A3EFFBC9B13FB64C4BAB614FA5A" box="[1273,1339,1415,1439]" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="3" pageNumber="59">Fig. 2</figureCitation>
|
||
D–F). In some corallites it can be so thick to resemble a hollow columellar structure (
|
||
<figureCitation id="136E2A3EFFBC9B13FB86C497B74EFA07" box="[1051,1121,1450,1474]" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="3" pageNumber="59">Fig. 2</figureCitation>
|
||
E–F). In these corallites an irregular fusion of the ventral triplet can be observed (
|
||
<figureCitation id="136E2A3EFFBC9B13FD6BC4F2B016FA22" box="[758,825,1487,1511]" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="3" pageNumber="59">Fig. 2</figureCitation>
|
||
F), and two smaller pali or no palus at all can be found in front of it.
|
||
</paragraph>
|
||
<paragraph id="8BEA36BBFFBC9B13FF58C72BB2DEF81A" blockId="3.[153,1437,151,2015]" pageId="3" pageNumber="59">
|
||
<emphasis id="B921EAA9FFBC9B13FF58C72BB265F9EA" bold="true" box="[197,330,1558,1583]" pageId="3" pageNumber="59">Variability.</emphasis>
|
||
The study of the
|
||
<typeStatus id="54EE8819FFBC9B13FD89C725B16BF9F5" box="[532,580,1560,1584]" pageId="3" pageNumber="59">type</typeStatus>
|
||
series and of the other collected specimens showed that although a digitate colony developing from an encrusting base is typical of
|
||
<taxonomicName id="4C554D38FFBC9B13FC8BC701B0F1F996" box="[790,990,1595,1619]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="3" pageNumber="59" phylum="Cnidaria" rank="species" species="fontanesii" status="sp. nov.">
|
||
<emphasis id="B921EAA9FFBC9B13FC8BC701B0F1F996" box="[790,990,1595,1619]" italics="true" pageId="3" pageNumber="59">Porites fontanesii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B921EAA9FFBC9B13FC78C706B76FF996" bold="true" box="[997,1088,1595,1619]" pageId="3" pageNumber="59">
|
||
<taxonomicNameLabel id="A21257D2FFBC9B13FC78C706B76FF996" box="[997,1088,1595,1619]" pageId="3" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, a certain degree of variability of the digitations upper ends diameter and shape is observed. These can be circular to oval in section (
|
||
<figureCitation id="136E2A3EFFBC9B13FA8AC762B674F9B2" box="[1303,1371,1631,1655]" captionStart="FIGURE 4" captionStartId="7.[151,250,1914,1937]" captionTargetBox="[175,1398,193,1888]" captionTargetId="figure@7.[175,1399,193,1889]" captionTargetPageId="7" captionText="FIGURE 4 (A – C, E – H, I) Specimens of Porites fontanesii sp. nov. showing the range of variability of the corallum shape and of its dimensions; (D) detail of C showing the usual surface appearance of this species; (J) detail of I showing the flower-like appearance of corallites in specimens with well developed denticles. Specimen registration code close to the scale bar. White scale bar = 1 cm." httpUri="https://zenodo.org/record/282167/files/figure.png" pageId="3" pageNumber="59">Fig. 4</figureCitation>
|
||
) with thickness ranging from
|
||
<quantity id="4CAD9B5EFFBC9B13FE3DC7BFB125F95F" box="[416,522,1666,1690]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="3" pageNumber="59" unit="cm" value="2.5" valueMax="4.0" valueMin="1.0">1 to 4 cm</quantity>
|
||
. In larger colonies digitations tend to fan out and occasionally branch (
|
||
<figureCitation id="136E2A3EFFBC9B13FA80C7BFB673F95F" box="[1309,1372,1666,1690]" captionStart="FIGURE 3" captionStartId="6.[151,250,1867,1890]" captionTargetBox="[176,1411,195,1822]" captionTargetId="figure@6.[176,1413,195,1844]" captionTargetPageId="6" captionText="FIGURE 3 Porites fontanesii sp. nov. in situ (A) A large branching colony at Balhaf, Yemen (depth 5 m); (B) Branching colony at Araher, Socotra Island (depth 15 m); (C) Colony at Tiqfash Island, Yemen (depth 3 m); (D) A beige colony with thick digitations at Musha Island, Djibouti (depth 4 m); (E) Detail of a digitation tip showing the smooth appearance of the colony surface once the polyps are retracted; (F) Close up of a colony at Tiqfash Island, Yemen, showing the polyps usually expanded during the day (depth 7 m); (G) Close-up of E showing living polyps with retracted tentacles." httpUri="https://zenodo.org/record/282166/files/figure.png" pageId="3" pageNumber="59">Fig. 3</figureCitation>
|
||
A–D,
|
||
<figureCitation id="136E2A3EFFBC9B13FF04C79AB3F3F97A" box="[153,220,1703,1727]" captionStart="FIGURE 4" captionStartId="7.[151,250,1914,1937]" captionTargetBox="[175,1398,193,1888]" captionTargetId="figure@7.[175,1399,193,1889]" captionTargetPageId="7" captionText="FIGURE 4 (A – C, E – H, I) Specimens of Porites fontanesii sp. nov. showing the range of variability of the corallum shape and of its dimensions; (D) detail of C showing the usual surface appearance of this species; (J) detail of I showing the flower-like appearance of corallites in specimens with well developed denticles. Specimen registration code close to the scale bar. White scale bar = 1 cm." httpUri="https://zenodo.org/record/282167/files/figure.png" pageId="3" pageNumber="59">Fig. 4</figureCitation>
|
||
C–G). Corallite organization over the colony surface is the same as the
|
||
<typeStatus id="54EE8819FFBC9B13FB90C795B712F905" box="[1037,1085,1704,1728]" pageId="3" pageNumber="59">type</typeStatus>
|
||
material throughout the whole collection. However, although corallites are generally flush with the coenosteum, in certain specimens they can be more sunken in certain parts of the corallum, typically towards the base of the digitations. At the corallite level, calice diameter, septal plan, and number of pali and denticles are consistent with the
|
||
<typeStatus id="54EE8819FFBC9B13FBD9C62EB75BF8EE" box="[1092,1140,1811,1835]" pageId="3" pageNumber="59">type</typeStatus>
|
||
series. In some specimens the denticles can be more visible (
|
||
<figureCitation id="136E2A3EFFBC9B13FD8FC60AB17AF88A" box="[530,597,1847,1871]" captionStart="FIGURE 4" captionStartId="7.[151,250,1914,1937]" captionTargetBox="[175,1398,193,1888]" captionTargetId="figure@7.[175,1399,193,1889]" captionTargetPageId="7" captionText="FIGURE 4 (A – C, E – H, I) Specimens of Porites fontanesii sp. nov. showing the range of variability of the corallum shape and of its dimensions; (D) detail of C showing the usual surface appearance of this species; (J) detail of I showing the flower-like appearance of corallites in specimens with well developed denticles. Specimen registration code close to the scale bar. White scale bar = 1 cm." httpUri="https://zenodo.org/record/282167/files/figure.png" pageId="3" pageNumber="59">Fig. 4</figureCitation>
|
||
J) than in others (
|
||
<figureCitation id="136E2A3EFFBC9B13FC89C60AB077F88A" box="[788,856,1847,1871]" captionStart="FIGURE 4" captionStartId="7.[151,250,1914,1937]" captionTargetBox="[175,1398,193,1888]" captionTargetId="figure@7.[175,1399,193,1889]" captionTargetPageId="7" captionText="FIGURE 4 (A – C, E – H, I) Specimens of Porites fontanesii sp. nov. showing the range of variability of the corallum shape and of its dimensions; (D) detail of C showing the usual surface appearance of this species; (J) detail of I showing the flower-like appearance of corallites in specimens with well developed denticles. Specimen registration code close to the scale bar. White scale bar = 1 cm." httpUri="https://zenodo.org/record/282167/files/figure.png" pageId="3" pageNumber="59">Fig. 4</figureCitation>
|
||
D). When this happens, the small corallites are more easily visible to the naked eye and attain a flower-like appearance (
|
||
<figureCitation id="136E2A3EFFBC9B13FC3AC667B0C1F8B7" box="[935,1006,1882,1906]" captionStart="FIGURE 4" captionStartId="7.[151,250,1914,1937]" captionTargetBox="[175,1398,193,1888]" captionTargetId="figure@7.[175,1399,193,1889]" captionTargetPageId="7" captionText="FIGURE 4 (A – C, E – H, I) Specimens of Porites fontanesii sp. nov. showing the range of variability of the corallum shape and of its dimensions; (D) detail of C showing the usual surface appearance of this species; (J) detail of I showing the flower-like appearance of corallites in specimens with well developed denticles. Specimen registration code close to the scale bar. White scale bar = 1 cm." httpUri="https://zenodo.org/record/282167/files/figure.png" pageId="3" pageNumber="59">Fig. 4</figureCitation>
|
||
J). The character showing the highest within and between specimens variability is the thickness of the innermost cycle of synapticulae (
|
||
<figureCitation id="136E2A3EFFBC9B13FB4AC642B633F852" box="[1239,1308,1919,1943]" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="3" pageNumber="59">Fig. 2</figureCitation>
|
||
). When the synapticulae connecting proximal ends of the septa are thicker the species typical pattern of septal fusion can be less easily visible (
|
||
<figureCitation id="136E2A3EFFBC9B13FEF1C6FAB280F81A" box="[364,431,1991,2015]" captionStart="FIGURE 2" captionStartId="5.[151,250,1059,1082]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@5.[151,1436,193,1038]" captionTargetPageId="5" captionText="FIGURE 2 SEM images of Porites fontanesii sp. nov. (A) Holotype (MNHN-IK. 2009 - 834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU 027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL 151 (Fig. 4 E – F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs." httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="3" pageNumber="59">Fig. 2</figureCitation>
|
||
D–F).
|
||
</paragraph>
|
||
<caption id="DF2A6633FFBB9B14FF0AC5ECB74DFAE0" httpUri="https://zenodo.org/record/282164/files/figure.png" pageId="4" pageNumber="60" targetBox="[151,1435,190,1211]" targetPageId="4">
|
||
<paragraph id="8BEA36BBFFBB9B14FF0AC5ECB74DFAE0" blockId="4.[151,1436,1233,1318]" pageId="4" pageNumber="60">
|
||
<emphasis id="B921EAA9FFBB9B14FF0AC5ECB220FB22" bold="true" box="[151,271,1233,1256]" pageId="4" pageNumber="60">FIGURE 1</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C554D38FFBB9B14FE01C5EEB17EFB2D" box="[412,593,1234,1256]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="4" pageNumber="60" phylum="Cnidaria" rank="species" species="fontanesii" status="sp. nov.">
|
||
<emphasis id="B921EAA9FFBB9B14FE01C5EEB17EFB2D" box="[412,593,1234,1256]" italics="true" pageId="4" pageNumber="60">Porites fontanesii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B921EAA9FFBB9B14FDC7C5EFB182FB22" bold="true" box="[602,685,1234,1255]" pageId="4" pageNumber="60">
|
||
<taxonomicNameLabel id="A21257D2FFBB9B14FDC7C5EFB182FB22" box="[602,685,1234,1255]" pageId="4" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(MNHN-IK.2009-834) (A)
|
||
<emphasis id="B921EAA9FFBB9B14FC4EC5EEB734FB2D" box="[979,1051,1235,1256]" italics="true" pageId="4" pageNumber="60">In vivo</emphasis>
|
||
image of the whole colony; (B) The holotype; (C) Detail of a digitation of the holotype typical of the species; (D) Close up of C showing smooth colony surface, regular corallite arrangement and coenosteum; (E) Detail of D showing the corallite septal plan.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEA36BBFFBB9B14FF58C456B16CF99E" blockId="4.[153,1436,1387,2023]" pageId="4" pageNumber="60">
|
||
<emphasis id="B921EAA9FFBB9B14FF58C456B186FA41" bold="true" box="[197,681,1387,1412]" pageId="4" pageNumber="60">Field characteristics and identification.</emphasis>
|
||
<taxonomicName id="4C554D38FFBB9B14FD2BC451B0ADFA46" box="[694,898,1387,1411]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="4" pageNumber="60" phylum="Cnidaria" rank="species" species="fontanesii" status="sp. nov.">
|
||
<emphasis id="B921EAA9FFBB9B14FD2BC451B0ADFA46" box="[694,898,1387,1411]" italics="true" pageId="4" pageNumber="60">Porites fontanesii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257D2FFBB9B14FC12C451B0C1FA46" box="[911,1006,1388,1412]" pageId="4" pageNumber="60" rank="species">
|
||
<emphasis id="B921EAA9FFBB9B14FC12C451B0C8FA41" bold="true" box="[911,999,1388,1412]" pageId="4" pageNumber="60">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
forms colonies ranging from a few centimeters to
|
||
<quantity id="4CAD9B5EFFBB9B14FEA2C4B2B2ABFA62" box="[319,388,1423,1447]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="60" unit="cm" value="50.0">50 cm</quantity>
|
||
in diameter and is found between 5 and
|
||
<quantity id="4CAD9B5EFFBB9B14FCCDC4B2B0A6FA62" box="[848,905,1423,1447]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="60" unit="m" value="15.0">15 m</quantity>
|
||
depth. The corallum base is usually encrusting and from it digitations of variable thickness and height develop (
|
||
<figureCitation id="136E2A3EFFBB9B14FCE4C48EB093FA0E" box="[889,956,1459,1483]" captionStart="FIGURE 3" captionStartId="6.[151,250,1867,1890]" captionTargetBox="[176,1411,195,1822]" captionTargetId="figure@6.[176,1413,195,1844]" captionTargetPageId="6" captionText="FIGURE 3 Porites fontanesii sp. nov. in situ (A) A large branching colony at Balhaf, Yemen (depth 5 m); (B) Branching colony at Araher, Socotra Island (depth 15 m); (C) Colony at Tiqfash Island, Yemen (depth 3 m); (D) A beige colony with thick digitations at Musha Island, Djibouti (depth 4 m); (E) Detail of a digitation tip showing the smooth appearance of the colony surface once the polyps are retracted; (F) Close up of a colony at Tiqfash Island, Yemen, showing the polyps usually expanded during the day (depth 7 m); (G) Close-up of E showing living polyps with retracted tentacles." httpUri="https://zenodo.org/record/282166/files/figure.png" pageId="4" pageNumber="60">Fig. 3</figureCitation>
|
||
A–D). In larger colonies branching can be observed. Digitations and branches tend to fan out (
|
||
<figureCitation id="136E2A3EFFBB9B14FD79C4EAB009FA2A" box="[740,806,1495,1519]" captionStart="FIGURE 3" captionStartId="6.[151,250,1867,1890]" captionTargetBox="[176,1411,195,1822]" captionTargetId="figure@6.[176,1413,195,1844]" captionTargetPageId="6" captionText="FIGURE 3 Porites fontanesii sp. nov. in situ (A) A large branching colony at Balhaf, Yemen (depth 5 m); (B) Branching colony at Araher, Socotra Island (depth 15 m); (C) Colony at Tiqfash Island, Yemen (depth 3 m); (D) A beige colony with thick digitations at Musha Island, Djibouti (depth 4 m); (E) Detail of a digitation tip showing the smooth appearance of the colony surface once the polyps are retracted; (F) Close up of a colony at Tiqfash Island, Yemen, showing the polyps usually expanded during the day (depth 7 m); (G) Close-up of E showing living polyps with retracted tentacles." httpUri="https://zenodo.org/record/282166/files/figure.png" pageId="4" pageNumber="60">Fig. 3</figureCitation>
|
||
A–D). Polyps are usually extended during the day (
|
||
<figureCitation id="136E2A3EFFBB9B14FAF2C4EAB389F9D6" captionStart="FIGURE 3" captionStartId="6.[151,250,1867,1890]" captionTargetBox="[176,1411,195,1822]" captionTargetId="figure@6.[176,1413,195,1844]" captionTargetPageId="6" captionText="FIGURE 3 Porites fontanesii sp. nov. in situ (A) A large branching colony at Balhaf, Yemen (depth 5 m); (B) Branching colony at Araher, Socotra Island (depth 15 m); (C) Colony at Tiqfash Island, Yemen (depth 3 m); (D) A beige colony with thick digitations at Musha Island, Djibouti (depth 4 m); (E) Detail of a digitation tip showing the smooth appearance of the colony surface once the polyps are retracted; (F) Close up of a colony at Tiqfash Island, Yemen, showing the polyps usually expanded during the day (depth 7 m); (G) Close-up of E showing living polyps with retracted tentacles." httpUri="https://zenodo.org/record/282166/files/figure.png" pageId="4" pageNumber="60">Fig. 3</figureCitation>
|
||
D, F). When these are retracted the smooth surface of the corallum (
|
||
<figureCitation id="136E2A3EFFBB9B14FC2BC4C6B0DCF9D6" box="[950,1011,1531,1555]" captionStart="FIGURE 3" captionStartId="6.[151,250,1867,1890]" captionTargetBox="[176,1411,195,1822]" captionTargetId="figure@6.[176,1413,195,1844]" captionTargetPageId="6" captionText="FIGURE 3 Porites fontanesii sp. nov. in situ (A) A large branching colony at Balhaf, Yemen (depth 5 m); (B) Branching colony at Araher, Socotra Island (depth 15 m); (C) Colony at Tiqfash Island, Yemen (depth 3 m); (D) A beige colony with thick digitations at Musha Island, Djibouti (depth 4 m); (E) Detail of a digitation tip showing the smooth appearance of the colony surface once the polyps are retracted; (F) Close up of a colony at Tiqfash Island, Yemen, showing the polyps usually expanded during the day (depth 7 m); (G) Close-up of E showing living polyps with retracted tentacles." httpUri="https://zenodo.org/record/282166/files/figure.png" pageId="4" pageNumber="60">Fig 3</figureCitation>
|
||
E) and the corallites separated by the coenosteum (
|
||
<figureCitation id="136E2A3EFFBB9B14FEB2C722B243F9F2" box="[303,364,1567,1591]" captionStart="FIGURE 3" captionStartId="6.[151,250,1867,1890]" captionTargetBox="[176,1411,195,1822]" captionTargetId="figure@6.[176,1413,195,1844]" captionTargetPageId="6" captionText="FIGURE 3 Porites fontanesii sp. nov. in situ (A) A large branching colony at Balhaf, Yemen (depth 5 m); (B) Branching colony at Araher, Socotra Island (depth 15 m); (C) Colony at Tiqfash Island, Yemen (depth 3 m); (D) A beige colony with thick digitations at Musha Island, Djibouti (depth 4 m); (E) Detail of a digitation tip showing the smooth appearance of the colony surface once the polyps are retracted; (F) Close up of a colony at Tiqfash Island, Yemen, showing the polyps usually expanded during the day (depth 7 m); (G) Close-up of E showing living polyps with retracted tentacles." httpUri="https://zenodo.org/record/282166/files/figure.png" pageId="4" pageNumber="60">Fig 3</figureCitation>
|
||
G) are visible. Colour ranges from grey or pale beige to light brown, and polyps are typically extended during the day. (
|
||
<figureCitation id="136E2A3EFFBB9B14FE26C77EB2D3F99E" box="[443,508,1603,1627]" captionStart="FIGURE 3" captionStartId="6.[151,250,1867,1890]" captionTargetBox="[176,1411,195,1822]" captionTargetId="figure@6.[176,1413,195,1844]" captionTargetPageId="6" captionText="FIGURE 3 Porites fontanesii sp. nov. in situ (A) A large branching colony at Balhaf, Yemen (depth 5 m); (B) Branching colony at Araher, Socotra Island (depth 15 m); (C) Colony at Tiqfash Island, Yemen (depth 3 m); (D) A beige colony with thick digitations at Musha Island, Djibouti (depth 4 m); (E) Detail of a digitation tip showing the smooth appearance of the colony surface once the polyps are retracted; (F) Close up of a colony at Tiqfash Island, Yemen, showing the polyps usually expanded during the day (depth 7 m); (G) Close-up of E showing living polyps with retracted tentacles." httpUri="https://zenodo.org/record/282166/files/figure.png" pageId="4" pageNumber="60">Fig. 3</figureCitation>
|
||
C–D).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F6530FFBB9B14FF58C755B27AF966" pageId="4" pageNumber="60" type="etymology">
|
||
<paragraph id="8BEA36BBFFBB9B14FF58C755B27AF966" blockId="4.[153,1436,1387,2023]" pageId="4" pageNumber="60">
|
||
<emphasis id="B921EAA9FFBB9B14FF58C755B264F945" bold="true" box="[197,331,1640,1664]" pageId="4" pageNumber="60">Etymology.</emphasis>
|
||
This species is named after Prof. Marcello Fontanesi, Magnifico Rettore of the University of Milano-Bicocca.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F6530FFBB9B15FF58C792B1DCF81B" lastPageId="5" lastPageNumber="61" pageId="4" pageNumber="60" type="distribution">
|
||
<paragraph id="8BEA36BBFFBB9B14FF58C792B785F892" blockId="4.[153,1436,1387,2023]" pageId="4" pageNumber="60">
|
||
<emphasis id="B921EAA9FFBB9B14FF58C792B13EF90D" bold="true" box="[197,529,1711,1736]" pageId="4" pageNumber="60">Distribution and frequency.</emphasis>
|
||
<taxonomicName id="4C554D38FFBB9B14FD84C78DB18BF902" box="[537,676,1711,1735]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="4" pageNumber="60" phylum="Cnidaria" rank="species" species="fontanesii" status="sp. nov.">
|
||
<emphasis id="B921EAA9FFBB9B14FD84C78DB18BF902" box="[537,676,1711,1735]" italics="true" pageId="4" pageNumber="60">P. fontanesii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B921EAA9FFBB9B14FD31C78DB029F90D" bold="true" box="[684,774,1712,1736]" pageId="4" pageNumber="60">
|
||
<taxonomicNameLabel id="A21257D2FFBB9B14FD31C78DB029F90D" box="[684,774,1712,1736]" pageId="4" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has been recorded from the southern Red Sea, the Gulf of Tadjoura, the Gulf of Aden and the north-western Indian Ocean (
|
||
<figureCitation id="136E2A3EFFBB9B14FC0CC7EEB0F7F92E" box="[913,984,1747,1771]" captionStart="FIGURE 5" captionStartId="8.[151,250,631,654]" captionTargetBox="[151,1436,193,610]" captionTargetId="figure@8.[151,1436,193,610]" captionTargetPageId="8" captionText="FIGURE 5 (A) Map of the Indian Ocean, the inset indicates the position of (B); (B) Map showing the sites where Porites fontanesii sp. nov. was collected (blue dots) in the southern Red Sea, Gulf of Tadjoura, and Gulf of Aden. The red star indicates the type locality in Balhaf, Yemen." httpUri="https://zenodo.org/record/282168/files/figure.png" pageId="4" pageNumber="60">Fig. 5</figureCitation>
|
||
). Southern Red Sea (
|
||
<collectingCountry id="F342762BFFBB9B14FB4CC7EEB60CF92E" box="[1233,1315,1747,1771]" name="Yemen" pageId="4" pageNumber="60">Yemen</collectingCountry>
|
||
) Kamaran Island, Kutama, Tiqfash, Uqban, Al Badi, Gulf of Tadjoura (
|
||
<collectingCountry id="F342762BFFBB9B14FCD2C7CAB09FF8CA" box="[847,944,1783,1807]" name="Djibouti" pageId="4" pageNumber="60">Djibouti</collectingCountry>
|
||
) Tadjoura, Obock, Musha Island, Northern Gulf of Aden (
|
||
<collectingCountry id="F342762BFFBB9B14FED8C626B2B8F8F6" box="[325,407,1819,1843]" name="Yemen" pageId="4" pageNumber="60">Yemen</collectingCountry>
|
||
) Aden, Balhaf, Bir Ali, Sikha Island, Burum, Al Mukallah, and from
|
||
<collectingCountry id="F342762BFFBB9B14FB58C626B635F8F6" box="[1221,1306,1819,1843]" name="Yemen" pageId="4" pageNumber="60">Socotra</collectingCountry>
|
||
Island. No additional records from published illustrations or museum specimens have been found so far.
|
||
</paragraph>
|
||
<paragraph id="8BEA36BBFFBB9B14FF58C659B619F822" blockId="4.[153,1436,1387,2023]" pageId="4" pageNumber="60">
|
||
<taxonomicName id="4C554D38FFBB9B14FF58C659B2A2F8BE" box="[197,397,1891,1915]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="4" pageNumber="60" phylum="Cnidaria" rank="species" species="fontanesii" status="sp. nov.">
|
||
<emphasis id="B921EAA9FFBB9B14FF58C659B2A2F8BE" box="[197,397,1891,1915]" italics="true" pageId="4" pageNumber="60">Porites fontanesii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B921EAA9FFBB9B14FE08C659B2DFF8B9" bold="true" box="[405,496,1892,1916]" pageId="4" pageNumber="60">
|
||
<taxonomicNameLabel id="A21257D2FFBB9B14FE08C659B2DFF8B9" box="[405,496,1892,1916]" pageId="4" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was most frequently observed in the
|
||
<collectingCountry id="F342762BFFBB9B14FC01C659B0C5F8BE" box="[924,1002,1892,1915]" name="Yemen" pageId="4" pageNumber="60">Yemen</collectingCountry>
|
||
Red Sea islands (
|
||
<figureCitation id="136E2A3EFFBB9B14FB2AC65EB7D4F8BE" box="[1207,1275,1891,1915]" captionStart="FIGURE 6" captionStartId="8.[151,250,1399,1422]" captionTargetBox="[151,1436,765,1378]" captionTargetId="figure@8.[151,1436,765,1378]" captionTargetPageId="8" captionText="FIGURE 6 Map of the southern Red Sea, Gulf of Tadjoura, and of the Gulf of Aden showing the frequency of Porites fontanesii sp. nov. in the different sampled areas. Radius of the blue circle is proportional to the species frequency, numbers above each circle indicate number of sites where Porites fontanesii sp. nov. was recorded / total number of sampled sites in each area. Area codes: a = southern Red Sea, Yemen; b = Gulf of Tadjoura, Djibouti; c = Aden, Yemen; d = Balhaf-Bir Ali, Yemen; e = Burum-Al Mukallah, Yemen; f = Socotra Island." httpUri="https://zenodo.org/record/282169/files/figure.png" pageId="4" pageNumber="60">Fig. 6</figureCitation>
|
||
) where it was observed in 88% of the sampled sites (n = 16). It was also frequently observed along the Gulf of Aden coast of
|
||
<collectingCountry id="F342762BFFBB9B14FF04C691B3C8F806" box="[153,231,1964,1987]" name="Yemen" pageId="4" pageNumber="60">Yemen</collectingCountry>
|
||
at Aden (60%, n = 5), Balhaf-Bir Ali (72%, n = 32) and Burum-Mukallah (67%, n = 18). Finally, the species was recorded at 20% and 11% of the sites in
|
||
<collectingCountry id="F342762BFFBB9B14FD7AC6F2B06BF822" box="[743,836,1999,2023]" name="Djibouti" pageId="4" pageNumber="60">Djibouti</collectingCountry>
|
||
(n = 27) and
|
||
<collectingCountry id="F342762BFFBB9B14FC79C6F2B715F822" box="[996,1082,1999,2023]" name="Yemen" pageId="4" pageNumber="60">Socotra</collectingCountry>
|
||
(n = 15), respectively.
|
||
</paragraph>
|
||
<caption id="DF2A6633FFBA9B15FF0AC51EB74BFB17" httpUri="https://zenodo.org/record/282165/files/figure.png" pageId="5" pageNumber="61" targetBox="[151,1435,193,1037]" targetPageId="5">
|
||
<paragraph id="8BEA36BBFFBA9B15FF0AC51EB74BFB17" blockId="5.[151,1437,1059,1234]" pageId="5" pageNumber="61">
|
||
<emphasis id="B921EAA9FFBA9B15FF0AC51EB222FBFD" bold="true" box="[151,269,1059,1082]" pageId="5" pageNumber="61">FIGURE 2</emphasis>
|
||
SEM images of
|
||
<taxonomicName id="4C554D38FFBA9B15FE2BC519B145FBFC" box="[438,618,1059,1081]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="5" pageNumber="61" phylum="Cnidaria" rank="species" species="fontanesii" status="sp. nov.">
|
||
<emphasis id="B921EAA9FFBA9B15FE2BC519B145FBFC" box="[438,618,1059,1081]" italics="true" pageId="5" pageNumber="61">Porites fontanesii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B921EAA9FFBA9B15FDEDC519B1EFFBFC" bold="true" box="[624,704,1060,1081]" pageId="5" pageNumber="61">
|
||
<taxonomicNameLabel id="A21257D2FFBA9B15FDEDC519B1EFFBFC" box="[624,704,1060,1081]" pageId="5" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A) Holotype (MNHN-IK.2009-834) corallite arrangement; The typical septal plan with a fused vetral triplet, a short dorsal septum, and five pali in a corallite of specimen UNIMIB MU027 (B), and in the holotype C); (D) Corallite arrangement at the base of specimen UNIMIB BAL151 (Fig. 4E–F); (E) A corallite in the same specimen with a typically thickened inner cycle of synapticulae connecting the proximal end of the septa resembling a hollow columella; (F) Detail of another corallite with a very thickened inner cycle of synapticulae. Larger arrows indicate the dorsal septum, smaller arrows the synapticulae connecting the dorsal septum to the lateral pairs.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEA36BBFFBA9B15FF58C42AB27DFA7A" blockId="5.[153,1436,1303,2014]" pageId="5" pageNumber="61">
|
||
<emphasis id="B921EAA9FFBA9B15FF58C42AB2E4FAF5" bold="true" box="[197,459,1303,1328]" pageId="5" pageNumber="61">Phylogenetic analysis.</emphasis>
|
||
A total of 48 sequences were aligned and subjected to phylogenetic analysis. Overall, 572 positions (194 showing gaps) characterized the alignment after excluding two hypervariable and unaligneable regions (about 200 bp in total, situated at the beginning of ITS1 and at the end of ITS2). Among 85 variable sites, 69 were parsimony informative, while a GTR+G+I evolutionary models was selected as the most appropriate fro the data by AIC.
|
||
</paragraph>
|
||
<paragraph id="8BEA36BBFFBA9B15FF58C4F6B7A6F91A" blockId="5.[153,1436,1303,2014]" pageId="5" pageNumber="61">
|
||
The phylogenetic trees obtained according to the two different criteria showed concordant topologies (
|
||
<figureCitation id="136E2A3EFFBA9B15FAD5C4F6B6A2FA26" box="[1352,1421,1483,1507]" captionStart="FIGURE 7" captionStartId="9.[151,250,1594,1617]" captionTargetBox="[280,1320,291,1573]" captionTargetId="figure@9.[240,1347,265,1581]" captionTargetPageId="9" captionText="FIGURE 7 Phylogenetic relationships based on rDNA among sequences of P. fontanesii and other homolog sequences of the Poritidae. Numbers at each node represent a posteriori probabilities obtained from a Bayesian analysis (above) and bootstrap values obtained from a Maximum Likelihood analysis (below). The clades described by Forsman et al. (2009) are indicated." httpUri="https://zenodo.org/record/282170/files/figure.png" pageId="5" pageNumber="61">Fig. 7</figureCitation>
|
||
). Two main clades were found, one relating to Central American
|
||
<taxonomicName id="4C554D38FFBA9B15FCEAC4CDB0E7F9C2" box="[887,968,1520,1543]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="5" pageNumber="61" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B921EAA9FFBA9B15FCEAC4CDB0E7F9C2" box="[887,968,1520,1543]" italics="true" pageId="5" pageNumber="61">Porites</emphasis>
|
||
</taxonomicName>
|
||
species, (subclades X, XI and XII
|
||
<emphasis id="B921EAA9FFBA9B15FAC3C4CDB6B3F9C2" box="[1374,1436,1520,1543]" italics="true" pageId="5" pageNumber="61">sensu</emphasis>
|
||
Forsman
|
||
<emphasis id="B921EAA9FFBA9B15FE9FC729B214F9EE" box="[258,315,1555,1579]" italics="true" pageId="5" pageNumber="61">et al.</emphasis>
|
||
2009) and the other, a large, polytomic clade including both Atlantic and Pacific species (subclades IX, VIII, VII, VI, V, III, II, I
|
||
<emphasis id="B921EAA9FFBA9B15FE7BC705B10BF98A" box="[486,548,1592,1615]" italics="true" pageId="5" pageNumber="61">sensu</emphasis>
|
||
Forsman
|
||
<emphasis id="B921EAA9FFBA9B15FD0AC705B1FEF98A" box="[663,721,1591,1615]" italics="true" pageId="5" pageNumber="61">et al.</emphasis>
|
||
2009). In both cases, the internal relationships agreed with the phylogeny obtained by Forsman
|
||
<emphasis id="B921EAA9FFBA9B15FDBEC761B14DF9B6" box="[547,610,1627,1651]" italics="true" pageId="5" pageNumber="61">et al.</emphasis>
|
||
(2009) from which the sequences were taken. Specimens of
|
||
<taxonomicName id="4C554D38FFBA9B15FAD6C761B227F952" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="5" pageNumber="61" phylum="Cnidaria" rank="species" species="fontanesii" status="sp. nov.">
|
||
<emphasis id="B921EAA9FFBA9B15FAD6C761B227F952" italics="true" pageId="5" pageNumber="61">Porites fontanesii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257D2FFBA9B15FE93C7BDB247F952" box="[270,360,1664,1688]" pageId="5" pageNumber="61" rank="species">
|
||
<emphasis id="B921EAA9FFBA9B15FE93C7BDB24EF95D" bold="true" box="[270,353,1664,1688]" pageId="5" pageNumber="61">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
are found in a subclade basal to, and well divergent from, the second main clade. A mean value of 6.4% (±
|
||
<geoCoordinate id="EE61507CFFBA9B15FEB5C79EB24CF97E" box="[296,355,1699,1723]" direction="south" orientation="latitude" pageId="5" pageNumber="61" precision="5555" value="-1.3">1.3 s</geoCoordinate>
|
||
.d.) p-distance separates
|
||
<taxonomicName id="4C554D38FFBA9B15FD18C799B03CF97E" box="[645,787,1699,1723]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="5" pageNumber="61" phylum="Cnidaria" rank="species" species="fontanesii" status="sp. nov.">
|
||
<emphasis id="B921EAA9FFBA9B15FD18C799B03CF97E" box="[645,787,1699,1723]" italics="true" pageId="5" pageNumber="61">P. fontanesii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B921EAA9FFBA9B15FCBDC799B051F979" bold="true" box="[800,894,1700,1724]" pageId="5" pageNumber="61">
|
||
<taxonomicNameLabel id="A21257D2FFBA9B15FCBDC799B051F979" box="[800,894,1700,1724]" pageId="5" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from the other species of the genus. Internal divergence within
|
||
<taxonomicName id="4C554D38FFBA9B15FEF4C7F5B2DDF91A" box="[361,498,1735,1759]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="5" pageNumber="61" phylum="Cnidaria" rank="species" species="fontanesii" status="sp. nov.">
|
||
<emphasis id="B921EAA9FFBA9B15FEF4C7F5B2DDF91A" box="[361,498,1735,1759]" italics="true" pageId="5" pageNumber="61">P. fontanesii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B921EAA9FFBA9B15FE67C7F5B17CF925" bold="true" box="[506,595,1736,1760]" pageId="5" pageNumber="61">
|
||
<taxonomicNameLabel id="A21257D2FFBA9B15FE67C7F5B17CF925" box="[506,595,1736,1760]" pageId="5" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is one order of magnitude lower (0.5 % ±
|
||
<geoCoordinate id="EE61507CFFBA9B15FBB2C7FAB74AF91A" box="[1071,1125,1735,1759]" direction="south" orientation="latitude" pageId="5" pageNumber="61" precision="5555" value="-0.3">0.3 s</geoCoordinate>
|
||
.d).
|
||
</paragraph>
|
||
<paragraph id="8BEA36BBFFBA9B15FF58C7D6B1DCF81B" blockId="5.[153,1436,1303,2014]" pageId="5" pageNumber="61">
|
||
Bayesian analysis performed with the software BEAST showed signals of convergence to the stationary distributions of parameters after 10.000.0 0 0 generations, since all ESS were largely> 200 and parameters distribution were smooth. The analysis was repeated three times obtaining the same inferences. An evolutionary pattern congruent with a molecular clock was not rejected (ucld.stdev = 0.3, 95% HPD 0.0–0.6) and a mean rate of 2.4
|
||
<date id="FFEB107BFFBA9B15FF20C643B3DDF84C" box="[189,242,1915,1942]" pageId="5" pageNumber="61" value="2010-10-02">x10-2</date>
|
||
(95% HPD 1.3
|
||
<date id="FFEB107BFFBA9B15FE3DC643B2F8F84C" box="[416,471,1915,1942]" pageId="5" pageNumber="61" value="2010-10-02">x10-2</date>
|
||
–3.7
|
||
<date id="FFEB107BFFBA9B15FD97C643B110F84C" box="[522,575,1915,1942]" pageId="5" pageNumber="61" value="2010-10-02">x10-2</date>
|
||
) subst/site*My was estimated. On these basis, the main divergence between
|
||
<taxonomicName id="4C554D38FFBA9B15FF04C69EB208F87F" box="[153,295,1954,1978]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="5" pageNumber="61" phylum="Cnidaria" rank="species" species="fontanesii" status="sp. nov.">
|
||
<emphasis id="B921EAA9FFBA9B15FF04C69EB208F87F" box="[153,295,1954,1978]" italics="true" pageId="5" pageNumber="61">P. fontanesii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257D2FFBA9B15FEA9C69EB2BDF87E" box="[308,402,1955,1979]" pageId="5" pageNumber="61" rank="species">
|
||
<emphasis id="B921EAA9FFBA9B15FEA9C69EB2BDF87E" bold="true" box="[308,402,1955,1979]" pageId="5" pageNumber="61">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and the other polytomic clade is dated at 8.53 MYA (95% HPD 4.0–14.0 MYA), thus indicating a likely late Miocene origin of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |