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<document id="2444D417714F3EE208C5CCC7926FE520" ID-DOI="10.11646/phytotaxa.323.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13696786" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725566605671" checkinUser="felipe" docAuthor="Corrêa, André Luis, Scudeller, Veridiana Vizoni &amp; Goldenberg, Renato" docDate="2017" docId="03FC87A2FFADFFE674C4F949FE17FDCE" docLanguage="en" docName="phytotaxa.323.2.1.pdf" docOrigin="Phytotaxa 323 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.323.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Miconia alata De Candolle 1828" docType="treatment" docVersion="2" lastPageNumber="115" masterDocId="FFC5FFDAFFA0FFE8744CFFE5FFA2FFEF" masterDocTitle="Melastomataceae in the Reserva de Desenvolvimento Sustentável do Tupé, Amazonas, Brazil" masterLastPageNumber="127" masterPageNumber="101" pageNumber="114" updateTime="1725568089067" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="83CEE94D895369FF4C871B4E551094E9">Melastomataceae in the Reserva de Desenvolvimento Sustentável do Tupé, Amazonas, Brazil</mods:title>
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<mods:namePart id="EFE001452F27E63B393A51D30A2D7DBD">Corrêa, André Luis</mods:namePart>
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<mods:namePart id="7C4978055775C2136D34C1E933DAEB41">Scudeller, Veridiana Vizoni</mods:namePart>
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<mods:namePart id="960179780D6D7405C13C83DE594089CF">Goldenberg, Renato</mods:namePart>
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<mods:title id="B220D79C7FD9AABC2C6BEEF6363DE874">Phytotaxa</mods:title>
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<mods:date id="EFB0E1A4DCD752395F0AE5BFE0E60336">2017</mods:date>
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<mods:number id="8E624DD68A6D70F74ACF786147D6B17A">2017-09-29</mods:number>
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<mods:identifier id="45D9298471362CE42E4876C3462A145C" type="DOI">10.11646/phytotaxa.323.2.1</mods:identifier>
<mods:identifier id="158B928E11BF8FB6E387F60C0D43D993" type="ISSN">1179-3163</mods:identifier>
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<treatment id="03FC87A2FFADFFE674C4F949FE17FDCE" ID-DOI="http://doi.org/10.5281/zenodo.13696830" ID-Zenodo-Dep="13696830" LSID="urn:lsid:plazi:treatment:03FC87A2FFADFFE674C4F949FE17FDCE" httpUri="http://treatment.plazi.org/id/03FC87A2FFADFFE674C4F949FE17FDCE" lastPageId="14" lastPageNumber="115" pageId="13" pageNumber="114">
<subSubSection id="C34F653FFFADFFE574C4F949FCF4F92A" box="[136,854,1708,1733]" pageId="13" pageNumber="114" type="nomenclature">
<paragraph id="8BEA36B4FFADFFE574C4F949FCF4F92A" blockId="13.[136,854,1708,1733]" box="[136,854,1708,1733]" pageId="13" pageNumber="114">
<heading id="D0A281D8FFADFFE574C4F949FCF4F92A" box="[136,854,1708,1733]" fontSize="11" level="2" pageId="13" pageNumber="114" reason="4">
<emphasis id="B921EAA6FFADFFE574C4F949FF09F92A" bold="true" box="[136,171,1708,1733]" pageId="13" pageNumber="114">17.</emphasis>
<taxonomicName id="4C554D37FFADFFE574FEF949FCF1F92A" ID-CoL="42ML9" authority="(Aublet 1829: 410) De Candolle (1828: 184)" authorityName="De Candolle" authorityPageNumber="184" authorityYear="1828" baseAuthorityName="Aublet" baseAuthorityPageNumber="410" baseAuthorityYear="1829" box="[178,851,1708,1733]" class="Magnoliopsida" family="Melastomataceae" genus="Miconia" kingdom="Plantae" order="Myrtales" pageId="13" pageNumber="114" phylum="Tracheophyta" rank="species" species="alata">
<emphasis id="B921EAA6FFADFFE574FEF949FEF6F92A" bold="true" box="[178,340,1708,1733]" italics="true" pageId="13" pageNumber="114">Miconia alata</emphasis>
(Aublet 1829: 410)
<bibRefCitation id="EFC44B45FFADFFE57674F949FCF1F92A" author="De Candolle, A. P." box="[568,851,1708,1733]" pageId="13" pageNumber="114" refId="ref30239" refString="De Candolle, A. P. (1828) Prodromus Systematis Naturalis Regni Vegetabilis. Tom. I et II. Paris." type="book" year="1828">De Candolle (1828: 184)</bibRefCitation>
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F653FFFADFFE674C4F911FE1EFE82" lastPageId="14" lastPageNumber="115" pageId="13" pageNumber="114" type="description">
<paragraph id="8BEA36B4FFADFFE674C4F911FADAFEEE" blockId="13.[136,1452,1780,2093]" lastBlockId="14.[136,1452,160,545]" lastPageId="14" lastPageNumber="115" pageId="13" pageNumber="114">
Shrubs ca.
<quantity id="4CAD9B51FFADFFE57547F911FEECF8E2" box="[267,334,1780,1805]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="114" unit="m" value="1.2">1.2 m</quantity>
tall. Light gray and smooth rhytidome. Branches erect, quadrangular, winged, rufous; branches, leaves both side, inflorescences, bracts, bracteoles, hypanthium and fruits with stellate-stipitate persistent trichomes. Leaves opposite isophyllous, sessile; leaf blades 10.515.2 ×
<quantity id="4CAD9B51FFADFFE57716F8D9FC7CF8BA" box="[858,990,1852,1877]" metricMagnitude="-2" metricUnit="m" metricValue="5.55" metricValueMax="6.9" metricValueMin="4.2" pageId="13" pageNumber="114" unit="cm" value="5.55" valueMax="6.9" valueMin="4.2">4.26.9 cm</quantity>
, coriaceous, discolored, abaxial surface rufous, symmetrical, oval, base attenuate, apex acute, margins slightly denticulate and ciliate, adaxial side with stellate-stipitate caducous trichomes only at the central veins, leaf veins 7, suprabasal; ant domatia absent. Paniculate 8.415.3 ×
<quantity id="4CAD9B51FFADFFE57541F84DFE2DF82E" box="[269,399,1960,1985]" metricMagnitude="-2" metricUnit="m" metricValue="7.05" metricValueMax="7.2" metricValueMin="6.9" pageId="13" pageNumber="114" unit="cm" value="7.05" valueMax="7.2" valueMin="6.9">6.97.2 cm</quantity>
, terminal; bracts
<quantity id="4CAD9B51FFADFFE57614F84DFD13F82E" box="[600,689,1960,1985]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="114" unit="mm" value="1.1">1.1 mm</quantity>
long, elliptical, caducous; bracteoles ca.
<quantity id="4CAD9B51FFADFFE570C1F84DFB47F82E" box="[1165,1253,1960,1985]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="114" unit="mm" value="0.3">0.3 mm</quantity>
long. Flowers 5- merous; pedicel ca.
<quantity id="4CAD9B51FFADFFE5752BF829FE1EF80A" box="[359,444,1996,2021]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="114" unit="mm" value="0.3">0.3 mm</quantity>
long; hypanthium ca.
<quantity id="4CAD9B51FFADFFE576F9F829FCA9F80A" box="[693,779,1996,2021]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="13" pageNumber="114" unit="mm" value="2.3">2.3 mm</quantity>
long, campanulate, smooth; calyx simple, persistent; lacinia triangular; petals ca. 2.5 ×
<quantity id="4CAD9B51FFADFFE575E0F815FDA0F7E6" box="[428,514,2032,2057]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="114" unit="mm" value="1.5">1.5 mm</quantity>
, white, oval, symmetrical, apex rounded, base truncate, margins not ciliate, glabrous; stamens 10, heteromorphic, smaller anthers ca.
<quantity id="4CAD9B51FFADFFE576D3F7F1FD57F7C2" box="[671,757,2068,2093]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="114" unit="mm" value="2.2">2.2 mm</quantity>
long, larger anthers ca.
<quantity id="4CAD9B51FFADFFE57048F7F1FBE7F7C2" box="[1028,1093,2068,2093]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="114" unit="mm" value="3.0">3 mm</quantity>
long, white, lanceolate, curved, apex attenuate,1 apical pore, connective not prolonged, smaller filaments ca.
<quantity id="4CAD9B51FFAEFFE677B9FF45FBECFF56" box="[1013,1102,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="115" unit="mm" value="2.5">2.5 mm</quantity>
, larger filaments
<quantity id="4CAD9B51FFAEFFE67158FF45FAC9FF56" box="[1300,1387,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="14" pageNumber="115" unit="mm" value="3.7">3.7 mm</quantity>
long, glabrous; ovary ca.
<quantity id="4CAD9B51FFAEFFE6752AFF21FE1EFF32" box="[358,444,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="115" unit="mm" value="1.4">1.4 mm</quantity>
long, half-inferior, placentation axillary, 4-locular, with glandular trichomes; style ca.
<quantity id="4CAD9B51FFAEFFE671C5FF21FF16FEEE" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="14" pageNumber="115" unit="mm" value="6.2">6.2 mm</quantity>
long, curved, glabrous. Berry 3.8 ×
<quantity id="4CAD9B51FFAEFFE67607FF0DFD01FEEE" box="[587,675,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="14" pageNumber="115" unit="mm" value="3.2">3.2 mm</quantity>
, unripe fruit green, ripe fruit orange; seeds numerous, triangular.
</paragraph>
<paragraph id="8BEA36B4FFAEFFE674F1FEE9FE1EFE82" blockId="14.[136,1452,160,545]" pageId="14" pageNumber="115">
<emphasis id="B921EAA6FFAEFFE674F1FEE9FE78FECA" bold="true" box="[189,474,268,293]" pageId="14" pageNumber="115">Specimens examined:—</emphasis>
<materialsCitation id="3B3D3CE9FFAEFFE67596FEE9FEEBFEA6" collectingDate="2012-08-15" collectionCode="A, L, EAFM, HUAM, INPA" country="Brazil" location="Amazonas" municipality="Manaus" pageId="14" pageNumber="115" specimenCount="1">
<collectingCountry id="F3427624FFAEFFE67596FEE9FDE1FECA" box="[474,579,268,293]" name="Brazil" pageId="14" pageNumber="115">BRAZIL</collectingCountry>
.
<location id="8E8A606FFFAEFFE67607FEE9FD66FECA" LSID="urn:lsid:plazi:treatment:03FC87A2FFADFFE674C4F949FE17FDCE:8E8A606FFFAEFFE67607FEE9FD66FECA" box="[587,708,268,293]" country="Brazil" municipality="Manaus" name="Amazonas" pageId="14" pageNumber="115">Amazonas</location>
,
<collectingMunicipality id="6B8EACCEFFAEFFE67682FEE9FC8BFECA" box="[718,809,268,293]" pageId="14" pageNumber="115">Manaus</collectingMunicipality>
, RDS-Tupé,
<date id="FFEB1074FFAEFFE677F6FEE9FBCEFECA" box="[954,1132,268,293]" pageId="14" pageNumber="115" value="2012-08-15">
<collectingDate id="EFAFE99CFFAEFFE677F6FEE9FBCEFECA" box="[954,1132,268,293]" pageId="14" pageNumber="115" value="2012-08-15">15 August 2012</collectingDate>
</date>
, fr.,
<emphasis id="B921EAA6FFAEFFE670D0FEE9FA9CFECA" box="[1180,1342,268,293]" italics="true" pageId="14" pageNumber="115">
<collectionCode id="ED44AE71FFAEFFE670D0FEE9FB0DFECA" box="[1180,1199,268,293]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="14" pageNumber="115" type="Herbarium">A</collectionCode>
.
<collectionCode id="ED44AE71FFAEFFE670FFFEE9FB67FECA" box="[1203,1221,268,293]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="14" pageNumber="115" type="Herbarium">L</collectionCode>
.
<taxonomicName id="4C554D37FFAEFFE67084FEE9FAB9FECA" box="[1224,1307,268,293]" class="Magnoliopsida" family="Rutaceae" genus="Correa" kingdom="Plantae" order="Sapindales" pageId="14" pageNumber="115" phylum="Tracheophyta" rank="genus">Corrêa</taxonomicName>
40
</emphasis>
(
<collectionCode id="ED44AE71FFAEFFE67100FEE9FA01FECA" box="[1356,1443,268,293]" country="Brazil" httpUri="http://grbio.org/cool/fwjk-rkfj" name="Instituto Federal de Educação, Ciência e Tecnologia do Amazonas" pageId="14" pageNumber="115" type="Herbarium">EAFM</collectionCode>
!,
<collectionCode id="ED44AE71FFAEFFE674C4FED5FF44FEA6" box="[136,230,304,329]" pageId="14" pageNumber="115">HUAM</collectionCode>
!,
<collectionCode id="ED44AE71FFAEFFE674B8FED5FE9FFEA6" box="[244,317,304,329]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="14" pageNumber="115">INPA</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="3B3D3CE9FFAEFFE6751FFED5FBA8FEA6" box="[339,1034,304,329]" collectingDate="2012-08-15" collectionCode="A, L, EAFM, HUAM, INPA" country="Brazil" pageId="14" pageNumber="115" specimenCount="1">
<date id="FFEB1074FFAEFFE6751FFED5FDA4FEA6" box="[339,518,304,329]" pageId="14" pageNumber="115" value="2012-08-15">
<collectingDate id="EFAFE99CFFAEFFE6751FFED5FDA4FEA6" box="[339,518,304,329]" pageId="14" pageNumber="115" value="2012-08-15">15 August 2012</collectingDate>
</date>
, fr.,
<emphasis id="B921EAA6FFAEFFE67679FED5FD74FEA6" box="[565,726,304,329]" italics="true" pageId="14" pageNumber="115">
<collectionCode id="ED44AE71FFAEFFE67679FED5FDEAFEA6" box="[565,584,304,329]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="14" pageNumber="115" type="Herbarium">A</collectionCode>
.
<collectionCode id="ED44AE71FFAEFFE67607FED5FDFFFEA6" box="[587,605,304,329]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="14" pageNumber="115" type="Herbarium">L</collectionCode>
.
<taxonomicName id="4C554D37FFAEFFE6762DFED5FD16FEA6" box="[609,692,304,329]" class="Magnoliopsida" family="Rutaceae" genus="Correa" kingdom="Plantae" order="Sapindales" pageId="14" pageNumber="115" phylum="Tracheophyta" rank="genus">Corrêa</taxonomicName>
41
</emphasis>
(
<collectionCode id="ED44AE71FFAEFFE676A8FED5FC99FEA6" box="[740,827,304,329]" country="Brazil" httpUri="http://grbio.org/cool/fwjk-rkfj" name="Instituto Federal de Educação, Ciência e Tecnologia do Amazonas" pageId="14" pageNumber="115" type="Herbarium">EAFM</collectionCode>
!,
<collectionCode id="ED44AE71FFAEFFE67706FED5FC0AFEA6" box="[842,936,304,329]" pageId="14" pageNumber="115">HUAM</collectionCode>
!,
<collectionCode id="ED44AE71FFAEFFE677FAFED5FC5DFEA6" box="[950,1023,304,329]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="14" pageNumber="115">INPA</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="3B3D3CE9FFAEFFE67059FED5FE1AFE82" collectingDate="2013-08-10" collectionCode="A, L, EAFM, HUAM, INPA" country="Brazil" pageId="14" pageNumber="115" specimenCount="1">
<date id="FFEB1074FFAEFFE67059FED5FB65FEA6" box="[1045,1223,304,329]" pageId="14" pageNumber="115" value="2013-08-10">
<collectingDate id="EFAFE99CFFAEFFE67059FED5FB65FEA6" box="[1045,1223,304,329]" pageId="14" pageNumber="115" value="2013-08-10">10 August 2013</collectingDate>
</date>
, fl.,
<emphasis id="B921EAA6FFAEFFE670BAFED5FA09FEA6" box="[1270,1451,304,329]" italics="true" pageId="14" pageNumber="115">
<collectionCode id="ED44AE71FFAEFFE670BAFED5FAA8FEA6" box="[1270,1290,304,329]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="14" pageNumber="115" type="Herbarium">A</collectionCode>
.
<collectionCode id="ED44AE71FFAEFFE67142FED5FA83FEA6" box="[1294,1313,304,329]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="14" pageNumber="115" type="Herbarium">L</collectionCode>
.
<taxonomicName id="4C554D37FFAEFFE67166FED5FAD9FEA6" box="[1322,1403,304,329]" class="Magnoliopsida" family="Rutaceae" genus="Correa" kingdom="Plantae" order="Sapindales" pageId="14" pageNumber="115" phylum="Tracheophyta" rank="genus">Corrêa</taxonomicName>
239
</emphasis>
(
<collectionCode id="ED44AE71FFAEFFE674DCFEB1FF45FE82" box="[144,231,340,365]" country="Brazil" httpUri="http://grbio.org/cool/fwjk-rkfj" name="Instituto Federal de Educação, Ciência e Tecnologia do Amazonas" pageId="14" pageNumber="115" type="Herbarium">EAFM</collectionCode>
!,
<collectionCode id="ED44AE71FFAEFFE674BAFEB1FEF6FE82" box="[246,340,340,365]" pageId="14" pageNumber="115">HUAM</collectionCode>
!,
<collectionCode id="ED44AE71FFAEFFE6752FFEB1FE0EFE82" box="[355,428,340,365]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="14" pageNumber="115">INPA</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F653FFFAEFFE674F1FE9DFCD7FE36" pageId="14" pageNumber="115" type="distribution">
<paragraph id="8BEA36B4FFAEFFE674F1FE9DFCD7FE36" blockId="14.[136,1452,160,545]" pageId="14" pageNumber="115">
<emphasis id="B921EAA6FFAEFFE674F1FE9DFDA4FE7E" bold="true" box="[189,518,376,401]" pageId="14" pageNumber="115">Distribution and habitat:—</emphasis>
<collectingCountry id="F3427624FFAEFFE6764AFE9DFDD5FE7E" box="[518,631,376,401]" name="Colombia" pageId="14" pageNumber="115">Colombia</collectingCountry>
,
<collectingCountry id="F3427624FFAEFFE676CEFE9DFD5AFE7E" box="[642,760,376,401]" name="Venezuela" pageId="14" pageNumber="115">Venezuela</collectingCountry>
,
<collectingCountry id="F3427624FFAEFFE67748FE9DFCFCFE7E" box="[772,862,376,401]" name="Guyana" pageId="14" pageNumber="115">Guyana</collectingCountry>
, French Guiana,
<collectingCountry id="F3427624FFAEFFE67051FE9DFB2AFE7E" box="[1053,1160,376,401]" name="Suriname" pageId="14" pageNumber="115">Suriname</collectingCountry>
(
<bibRefCitation id="EFC44B45FFAEFFE670DBFE9DFA02FE7E" author="Goldenberg, R. &amp; Almeda, F. &amp; Caddah, M. K. &amp; Martins, A. B. &amp; Meirelles, J. &amp; Michelangeli, F. A. &amp; Weiss, M." box="[1175,1440,376,401]" pageId="14" pageNumber="115" pagination="1 - 171" refId="ref30409" refString="Goldenberg, R., Almeda, F., Caddah, M. K., Martins, A. B., Meirelles, J., Michelangeli, F. A. &amp; Weiss, M. (2013) Nomenclator Botanicus for the neotropical genus Miconia (Melastomataceae). Phytotaxa 106: 1 - 171. https: // doi. org / 10.11646 / phytotaxa. 106.1.1" type="journal article" year="2013">
Goldenberg
<emphasis id="B921EAA6FFAEFFE67168FE9DFAFEFE7E" box="[1316,1372,376,401]" italics="true" pageId="14" pageNumber="115">et al.</emphasis>
2013
</bibRefCitation>
). In
<collectingCountry id="F3427624FFAEFFE674EAFE79FF48FE5A" box="[166,234,412,437]" name="Brazil" pageId="14" pageNumber="115">Brazil</collectingCountry>
it occurs in Amazonian forests and Caatinga (
<bibRefCitation id="EFC44B45FFAEFFE676ABFE79FB8AFE5A" author="Goldenberg, R. &amp; Caddah, M. K." box="[743,1064,412,437]" pageId="14" pageNumber="115" refId="ref30549" refString="Goldenberg, R. &amp; Caddah, M. K. (2015) Miconia in Lista de Especies da Flora do Brasil. Jardim Botanico do Rio de Janeiro. Available from: http: // reflora. jbrj. gov. br / jabot / floradobrasil / FB 9666 (accessed 29 May 2017)" type="url" year="2015">Goldenberg &amp; Caddah 2015</bibRefCitation>
). In the reserve, it occurs in second growth plateaus of “terra firme” forest, over well drained clay soil.
</paragraph>
</subSubSection>
<subSubSection id="C34F653FFFAEFFE674F1FE01FE17FDCE" pageId="14" pageNumber="115" type="discussion">
<paragraph id="8BEA36B4FFAEFFE674F1FE01FE17FDCE" blockId="14.[136,1452,160,545]" pageId="14" pageNumber="115">
<emphasis id="B921EAA6FFAEFFE674F1FE01FEC6FE12" bold="true" box="[189,356,484,509]" pageId="14" pageNumber="115">Comments:—</emphasis>
This species is not frequent in the reserve. It can be distinguished from the other species by the isophyllous, sessile leaves.
</paragraph>
</subSubSection>
</treatment>
</document>