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<document id="61465E5C813B9B308CCBD80965915226" ID-DOI="10.1080/00222933.2024.2410028" ID-ISSN="1464-5262" ID-Zenodo-Dep="14243350" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732838534497" checkinUser="plazi" docAuthor="Kamalanathan, Veenakumari, Kolla, Sreedevi, Khan, Farmanur Rahman &amp; Mohanraj, Prashanth" docDate="2024" docId="03FC879BFFBAFFEF9AB7FBDC810804D0" docLanguage="en" docName="JNATHIST.58.45-48.2121-2187.pdf" docOrigin="Journal of Natural History 58 (45 - 48)" docSource="http://dx.doi.org/10.1080/00222933.2024.2410028" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Hadronotus randhra Veenakumari 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="2162" masterDocId="FFC5FFE3FF9CFFC69B20FFB28568056E" masterDocTitle="Hadronotus insulare species group (Hymenoptera: Platygastroidea: Scelionidae) with descriptions of new species from India" masterLastPageNumber="2187" masterPageNumber="2121" pageNumber="2159" updateTime="1732892349642" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="72153B82CA550E62942EC3C5A988B921">Hadronotus insulare species group (Hymenoptera: Platygastroidea: Scelionidae) with descriptions of new species from India</mods:title>
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<mods:namePart id="FE6DE9C659E0AF7AFB9435D603272DF2">Kamalanathan, Veenakumari</mods:namePart>
<mods:affiliation id="05C93D83EBCEB416831FD70E3E28C056">Division of Germplasm Collection and Characterisation, ICAR - National Bureau of Agricultural Insect Resources, Hebbal, India;</mods:affiliation>
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<mods:namePart id="9E09AAB001CB65420329BF1F31DECD2C">Kolla, Sreedevi</mods:namePart>
<mods:affiliation id="7BF500DF37F6D0B1C956F63826BA618E">Division of Germplasm Collection and Characterisation, ICAR - National Bureau of Agricultural Insect Resources, Hebbal, India;</mods:affiliation>
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<mods:namePart id="B2F5B8928C69C93510DD81905A762955">Khan, Farmanur Rahman</mods:namePart>
<mods:affiliation id="06B474F2D7849900C99AB99AA2F82742">Department of Plant Production and Protection, College of Agriculture and Veterinary Medicine, Qassim University, Buraidah, Saudi Arabia</mods:affiliation>
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<mods:namePart id="EABB1B8152BCA1114E37968135CDF9FB">Mohanraj, Prashanth</mods:namePart>
<mods:affiliation id="10F277F2E01A315F8A41ACB7A7F905B6">Division of Germplasm Collection and Characterisation, ICAR - National Bureau of Agricultural Insect Resources, Hebbal, India;</mods:affiliation>
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<mods:title id="653D5D4EF483BF1C392927901F2F0C64">Journal of Natural History</mods:title>
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<mods:date id="C2FF6AA8239D89915BED89420288EC00">2024</mods:date>
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<mods:number id="020550175FAF0D3ED09254808069ACDD">58</mods:number>
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<mods:number id="82A230D6C7119FB0CF96BA2B39D16D59">45 - 48</mods:number>
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<paragraph id="8BEA368DFFBAFFE09AB7FBDC86E501E6" blockId="38.[407,909,1134,1160]" box="[407,909,1134,1160]" pageId="38" pageNumber="2159">
<heading id="D0A281E1FFBAFFE09AB7FBDC86E501E6" bold="true" box="[407,909,1134,1160]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="2159" reason="3">
<emphasis id="B921EA9FFFBAFFE09AB7FBDC86E501E6" bold="true" box="[407,909,1134,1160]" pageId="38" pageNumber="2159">
<taxonomicName id="4C554D0EFFBAFFE09AB7FBDC862901E6" authority="Veenakumari" authorityName="Veenakumari" authorityYear="2024" box="[407,833,1134,1160]" class="Insecta" family="Curculionidae" genus="Hadronotus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="2159" phylum="Arthropoda" rank="species" species="randhra" status="sp. nov.">
<emphasis id="B921EA9FFFBAFFE09AB7FBDC87FA01E6" bold="true" box="[407,658,1134,1160]" italics="true" pageId="38" pageNumber="2159">Hadronotus randhra</emphasis>
Veenakumari
</taxonomicName>
<taxonomicNameLabel id="A21257E4FFBAFFE09869FBDC86E501E6" box="[841,909,1134,1160]" pageId="38" pageNumber="2159" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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http://www.zoobank.org/
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urn:lsid:zoobank.org:pub:
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<subSubSection id="C34F6506FFBAFFE19926FB6384AB00D9" lastPageId="39" lastPageNumber="2160" pageId="38" pageNumber="2159" type="description">
<paragraph id="8BEA368DFFBAFFE09926FB6386750185" blockId="38.[518,797,1233,1259]" box="[518,797,1233,1259]" pageId="38" pageNumber="2159">
(
<figureCitation id="136E2A08FFBAFFE0992EFB6387E20185" box="[526,650,1233,1259]" captionStart="Figure 14" captionStartId="40.[160,225,1544,1567]" captionTargetBox="[163,1153,142,1511]" captionTargetId="figure-3@40.[163,1153,142,1512]" captionTargetPageId="40" captionText="Figure 14. Hadronotus randhra sp. n., female. (a) Habitus (dorsal view); (b) antenna; (c) head and pleuron; (d) vertex; (e) wings; (f) frons; (g) head and mesonotum." figureDoi="http://doi.org/10.5281/zenodo.14243382" httpUri="https://zenodo.org/record/14243382/files/figure.png" pageId="38" pageNumber="2159">Figures 14</figureCitation>
(ag), 26(m))
</paragraph>
<paragraph id="8BEA368DFFBAFFE09B80FAB4816C0071" blockId="38.[160,1028,1286,1311]" box="[160,1028,1286,1311]" pageId="38" pageNumber="2159">
Female body length =
<quantity id="4CAD9B68FFBAFFE09A81FAB4876B0071" box="[417,515,1286,1311]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="38" pageNumber="2159" unit="mm" value="1.53">1.53 mm</quantity>
(n = 1); male body length =
<quantity id="4CAD9B68FFBAFFE0986AFAB486C40071" box="[842,940,1286,1311]" metricMagnitude="-3" metricUnit="m" metricValue="1.82" pageId="38" pageNumber="2159" unit="mm" value="1.82">1.82 mm</quantity>
(n = 1).
</paragraph>
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<emphasis id="B921EA9FFFBAFFE09B80FAF9859D000B" bold="true" box="[160,245,1355,1381]" italics="true" pageId="38" pageNumber="2159">Colour.</emphasis>
Head, mesosoma and metasoma black; all coxae brown, remainder of leg brown-yellow; clypeus yellow-brown; mandible yellow-brown with red-brown teeth; radicle yellow-black; A1 basally orange, remainder brown-black; A2A3 brown-black; A4A6 brown, remaining antennomeres black-brown.
</paragraph>
<paragraph id="8BEA368DFFBAFFE19B80FA4B84450419" blockId="38.[160,1157,1529,1728]" lastBlockId="39.[160,1156,142,375]" lastPageId="39" lastPageNumber="2160" pageId="38" pageNumber="2159">
<emphasis id="B921EA9FFFBAFFE09B80FA4B858D037D" bold="true" box="[160,229,1529,1555]" italics="true" pageId="38" pageNumber="2159">Head.</emphasis>
1.1× as wide as high, 1.6× as high as long. IOS: 0.6× head width, 1.2× eye length. POL: LOL: OOL: 19.2: 11.5: 2.1. OOL: 0.8× MOD. Compound eye: ovate (L: W = 23.6: 16.7). Sculpture of orbital furrow: foveate. Inner orbits in frontal view: converging dorsally. Central keel on frons: present, extending ¼ length of frontal depression. Malar sulcus: with shallow rectangular depressions. Shape of frontal depression: ovoid. L: W of frontal depression: 27.0: 17.5. Width of frontal depression: 0.6× IOS. Sculpture of frontal depression: transversely carinate with smooth area between carinae; polygonal cells present on either side of transverse carinae in basal half. Submedian carina of frontal depression: restricted to dorsal 1/5.
<collectingRegion id="4991F86FFFBBFFE198FDFF66817A0583" box="[989,1042,212,237]" country="Tanzania" name="Pwani" pageId="39" pageNumber="2160">AOD</collectingRegion>
in dorsal view: 6× OOL. Sculpture of frons sans frontal depression: polygonal cells with rugose sculpture, setose. Sculpture of vertex: polygonal cells with rugose sculpture, setose. Hyperoccipital carina: present. Sculpture of gena: polygonal cells with rugose sculpture, setose.
</paragraph>
<paragraph id="8BEA368DFFBBFFE19B80FE0187A20152" blockId="39.[160,1156,435,1084]" pageId="39" pageNumber="2160">
<emphasis id="B921EA9FFFBBFFE19B80FE01844E04A3" bold="true" box="[160,294,435,461]" italics="true" pageId="39" pageNumber="2160">Mesosoma.</emphasis>
L: W of mesoscutum: 27.5: 41.0. Sculpture of mesoscutum: polygonal cells with rugose sculpture, densely setose. Scutoscutellar sulcus: foveate. L: W of mesoscutellum: 17.0: 30.2. Sculpture of mesoscutellum: polygonal cells with rugose sculpture, densely setose laterad. Posterior mesoscutellar sulcus: foveate. Sculpture of metascutellum: foveate. Posterior margin of metascutellum: semicircular. Sculpture of metanotal trough: foveate. Lateral propodeal area: laterally wide and narrowing centrad. Sculpture of lateral propodeal area: with depressions. Posterior propodeal projection: not pointed. Pronotal cervical sulcus: with depressions. Sculpture of lateral pronotal area: dorsally coriaceous reticulate, ventrally transversely carinate. Prespecular sulcus: transversely foveate. Sculpture of speculum: transversely carinate interspersed with foveae. Femoral depression: with irregular foveae. Episternal sulcus: foveate. Postacetabular sulcus: with elongate foveae. Mesepimeral sulcus: foveate. Sculpture of posterior mesepimeral area: narrow and smooth. Mesopleural carina: indicated ventrally. Mesopleural pit: present. Metapleural sulcus: with depressions. Sculpture of dorsal metapleural area: foveate, densely setose. Sculpture of ventral metapleural area: concealed beneath dense setae. Paracoxal sulcus: with depressions. L: W of fore wing: 100.6: 41.9. Ratio of length of m: st: pm: 4.9: 8.2: 5.6. L: W of hind wing: 80.1: 11.4. Length of hind wing marginal cilia: 0.5× width of wing. Shape of submarginalis: straight, distant from anterior margin of wing dipping slightly before reaching marginalis.
</paragraph>
<paragraph id="8BEA368DFFBBFFE19B80FBCA81EC0051" blockId="39.[160,1156,1144,1343]" pageId="39" pageNumber="2160">
<emphasis id="B921EA9FFFBBFFE19B80FBCA844C01FC" bold="true" box="[160,292,1144,1170]" italics="true" pageId="39" pageNumber="2160">Metasoma.</emphasis>
L: W of metasoma: 58.3: 42.5, 0.9 × as long as wide. Anterior margin of T1: with a slight projection medially and laterally. Basal foveae on T1: present. Sculpture of T1: longitudinally costate, space between costae rugose. Sculpture of T2: basally with short longitudinal costae, remainder with depressions except for smooth area on posterior margin; basal foveae absent. Length of T2: 9× length of T3. Sculpture of T3T4: anteriorly areolate, posteriorly smooth. Setation of metasoma: dense laterally and sparse posteriorly.
</paragraph>
<paragraph id="8BEA368DFFBBFFE19B80FAC984AB00D9" blockId="39.[160,1156,1403,1463]" pageId="39" pageNumber="2160">
<emphasis id="B921EA9FFFBBFFE19B80FAC9858900FB" bold="true" box="[160,225,1403,1429]" italics="true" pageId="39" pageNumber="2160">Male.</emphasis>
Similar to female. Length of A1A
<quantity id="4CAD9B68FFBBFFE19940FAC987FF00FA" box="[608,663,1403,1428]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="39" pageNumber="2160" unit="in" value="12.0">12 in</quantity>
ratio of 34.6: 9.3: 9.8: 6.3: 8.8: 7.3: 7.3: 7.3: 7.3: 7.3: 7.8: 13.7, respectively.
</paragraph>
</subSubSection>
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<emphasis id="B921EA9FFFBBFFE19B80FA4184170363" bold="true" box="[160,383,1523,1549]" italics="true" pageId="39" pageNumber="2160">Material examined</emphasis>
</paragraph>
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<typeStatus id="54EE882FFFBBFFE19B80F9A4846D0341" box="[160,261,1558,1583]" pageId="39" pageNumber="2160" type="holotype">Holotype</typeStatus>
, female (
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/
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/P4974),
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<collectingCountry id="F342761DFFBBFFE19960F9A487EC0341" box="[576,644,1558,1583]" name="India" pageId="39" pageNumber="2160">INDIA</collectingCountry>
</emphasis>
:
<collectingRegion id="4991F86FFFBBFFE199AEF9A487930341" box="[654,763,1558,1583]" country="India" name="Karnataka" pageId="39" pageNumber="2160">Karnataka</collectingRegion>
, Shivamogga,
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, Naville,
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,
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, ca
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, YPT, in potato (
<taxonomicName id="4C554D0EFFBBFFE1991AF98B8665033C" box="[570,781,1593,1618]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="39" pageNumber="2160" phylum="Tracheophyta" rank="species" species="tuberosum">
<emphasis id="B921EA9FFFBBFFE1991AF98B8665033C" box="[570,781,1593,1618]" italics="true" pageId="39" pageNumber="2160">Solanum tuberosum</emphasis>
</taxonomicName>
:
<taxonomicName id="4C554D0EFFBBFFE19839F98B86FC033C" box="[793,916,1593,1618]" class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" order="Solanales" pageId="39" pageNumber="2160" phylum="Tracheophyta" rank="family">Solanaceae</taxonomicName>
) field,
<date id="FFEB104DFFBBFFE198FAF98B81E8033C" box="[986,1152,1593,1618]" pageId="39" pageNumber="2160" value="2017-01-08">8 January 2017</date>
.
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:
<specimenCount id="9D53FD04FFBBFFE19A2EF9E9841F031A" box="[270,375,1627,1652]" count="1" pageId="39" pageNumber="2160" type="male">one male</specimenCount>
(
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/
<collectionCode id="ED44AE48FFBBFFE19AE5F9E98765031A" box="[453,525,1627,1652]" pageId="39" pageNumber="2160">NBAIR</collectionCode>
/P4975),
<collectingRegion id="4991F86FFFBBFFE19948F9E987BD031A" box="[616,725,1627,1652]" country="India" name="Karnataka" pageId="39" pageNumber="2160">Karnataka</collectingRegion>
, Shivamogga,
<collectionCode id="ED44AE48FFBBFFE19858F9E986D1031A" box="[888,953,1627,1652]" pageId="39" pageNumber="2160">UAHS</collectionCode>
, Naville,
<geoCoordinate id="EE61504AFFBBFFE19F3DF9E981E8031A" box="[1053,1152,1627,1652]" degrees="13.973" direction="north" orientation="latitude" pageId="39" pageNumber="2160" precision="55" value="13.973">13.973°N</geoCoordinate>
,
<geoCoordinate id="EE61504AFFBBFFE19BBFF9CC859503F9" box="[159,253,1662,1687]" degrees="75.578" direction="east" orientation="longitude" pageId="39" pageNumber="2160" precision="55" value="75.578">75.578°E</geoCoordinate>
, ca
<quantity id="4CAD9B68FFBBFFE19A06F9CC840503F9" box="[294,365,1662,1687]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="39" pageNumber="2160" unit="m" value="610.0">610 m</quantity>
, YPT, in paddy (
<taxonomicName id="4C554D0EFFBBFFE19939F9CC87F203F9" box="[537,666,1662,1687]" class="Liliopsida" family="Poaceae" genus="Oryza" kingdom="Plantae" order="Poales" pageId="39" pageNumber="2160" phylum="Tracheophyta" rank="species" species="sativa">
<emphasis id="B921EA9FFFBBFFE19939F9CC87F203F9" box="[537,666,1662,1687]" italics="true" pageId="39" pageNumber="2160">Oryza sativa</emphasis>
</taxonomicName>
:
<taxonomicName id="4C554D0EFFBBFFE19985F9CC866903F9" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[677,769,1662,1687]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="39" pageNumber="2160" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
) field,
<date id="FFEB104DFFBBFFE19868F9CC868503F9" box="[840,1005,1662,1687]" pageId="39" pageNumber="2160" value="2017-01-09">9 January 2017</date>
.
</paragraph>
</subSubSection>
<caption id="DF2A6605FFB4FFEE9B80F9BA86400352" ID-DOI="http://doi.org/10.5281/zenodo.14243382" ID-Zenodo-Dep="14243382" httpUri="https://zenodo.org/record/14243382/files/figure.png" pageId="40" pageNumber="2161" startId="40.[160,225,1544,1567]" targetBox="[163,1153,142,1511]" targetPageId="40" targetType="figure">
<paragraph id="8BEA368DFFB4FFEE9B80F9BA86400352" blockId="40.[160,1155,1544,1596]" pageId="40" pageNumber="2161">
<emphasis id="B921EA9FFFB4FFEE9B80F9BA84610371" bold="true" box="[160,265,1544,1567]" pageId="40" pageNumber="2161">Figure 14.</emphasis>
<taxonomicName id="4C554D0EFFB4FFEE9A32F9BA84B00371" authorityName="Kamalanathan &amp; Kolla &amp; Khan &amp; Mohanraj" authorityYear="2024" box="[274,472,1544,1567]" class="Insecta" family="Curculionidae" genus="Hadronotus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="2161" phylum="Arthropoda" rank="species" species="randhra" status="sp. nov.">
<emphasis id="B921EA9FFFB4FFEE9A32F9BA84B00371" box="[274,472,1544,1567]" italics="true" pageId="40" pageNumber="2161">Hadronotus randhra</emphasis>
</taxonomicName>
<emphasis id="B921EA9FFFB4FFEE9AC0F9BA87740371" bold="true" box="[480,540,1544,1567]" pageId="40" pageNumber="2161">
<taxonomicNameLabel id="A21257E4FFB4FFEE9AC0F9BA87740371" box="[480,540,1544,1567]" pageId="40" pageNumber="2161" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, female. (a) Habitus (dorsal view); (b) antenna; (c) head and pleuron; (d) vertex; (e) wings; (f) frons; (g) head and mesonotum.
</paragraph>
</caption>
<subSubSection id="C34F6506FFB5FFEF9B80FF3C878D0583" pageId="41" pageNumber="2162" type="etymology">
<paragraph id="8BEA368DFFB5FFEF9B80FF3C847505C6" blockId="41.[160,1155,142,237]" box="[160,285,142,168]" pageId="41" pageNumber="2162">
<emphasis id="B921EA9FFFB5FFEF9B80FF3C847505C6" bold="true" box="[160,285,142,168]" italics="true" pageId="41" pageNumber="2162">Etymology</emphasis>
</paragraph>
<paragraph id="8BEA368DFFB5FFEF9B80FF00878D0583" blockId="41.[160,1155,142,237]" pageId="41" pageNumber="2162">The Sanskrit epithet randhra means depressions, referring to the depressions present on the metasoma. The name is treated as an adjective.</paragraph>
</subSubSection>
<subSubSection id="C34F6506FFB5FFEF9B80FEA8810804D0" pageId="41" pageNumber="2162" type="diagnosis">
<paragraph id="8BEA368DFFB5FFEF9B80FEA8847C045A" blockId="41.[160,1156,282,446]" box="[160,276,282,308]" pageId="41" pageNumber="2162">
<emphasis id="B921EA9FFFB5FFEF9B80FEA8847C045A" bold="true" box="[160,276,282,308]" italics="true" pageId="41" pageNumber="2162">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BEA368DFFB5FFEF9B80FE8F810804D0" blockId="41.[160,1156,282,446]" pageId="41" pageNumber="2162">
<taxonomicName id="4C554D0EFFB5FFEF9B80FE8F84150438" authorityName="Kamalanathan &amp; Kolla &amp; Khan &amp; Mohanraj" authorityYear="2024" box="[160,381,317,342]" class="Insecta" family="Curculionidae" genus="Hadronotus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="2162" phylum="Arthropoda" rank="species" species="randhra" status="sp. nov.">
<emphasis id="B921EA9FFFB5FFEF9B80FE8F84150438" box="[160,381,317,342]" italics="true" pageId="41" pageNumber="2162">Hadronotus randhra</emphasis>
</taxonomicName>
<emphasis id="B921EA9FFFB5FFEF9AA3FE8F84AA0438" bold="true" box="[387,450,317,342]" pageId="41" pageNumber="2162">
<taxonomicNameLabel id="A21257E4FFB5FFEF9AA3FE8F84AA0438" box="[387,450,317,342]" pageId="41" pageNumber="2162" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C554D0EFFB5FFEF996DFE8F87DB0438" authorityName="Kamalanathan &amp; Kolla &amp; Khan &amp; Mohanraj" authorityYear="2024" box="[589,691,317,342]" class="Insecta" family="Curculionidae" genus="Hadronotus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="2162" phylum="Arthropoda" rank="species" species="hrdyae" status="sp. n.">
<emphasis id="B921EA9FFFB5FFEF996DFE8F87DB0438" box="[589,691,317,342]" italics="true" pageId="41" pageNumber="2162">H. hrdyae</emphasis>
</taxonomicName>
<emphasis id="B921EA9FFFB5FFEF999AFE8F87910438" bold="true" box="[698,761,317,342]" pageId="41" pageNumber="2162">
<taxonomicNameLabel id="A21257E4FFB5FFEF999AFE8F87910438" box="[698,761,317,342]" pageId="41" pageNumber="2162" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
but differs from it in having a short central keel on frons which extends 1/4 length of frontal depression, and with stigmalis straight on fore wing, whereas in
<taxonomicName id="4C554D0EFFB5FFEF9918FE3187CD04F2" authorityName="Kamalanathan &amp; Kolla &amp; Khan &amp; Mohanraj" authorityYear="2024" box="[568,677,387,412]" class="Insecta" family="Curculionidae" genus="Hadronotus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="2162" phylum="Arthropoda" rank="species" species="hrdyae" status="sp. n.">
<emphasis id="B921EA9FFFB5FFEF9918FE3187CD04F2" box="[568,677,387,412]" italics="true" pageId="41" pageNumber="2162">H. hrdyae</emphasis>
</taxonomicName>
<emphasis id="B921EA9FFFB5FFEF9991FE31879E04F2" bold="true" box="[689,758,387,412]" pageId="41" pageNumber="2162">
<taxonomicNameLabel id="A21257E4FFB5FFEF9991FE31879E04F2" box="[689,758,387,412]" pageId="41" pageNumber="2162" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
central keel on frons is elongate extending 4/5 length of frontal depression, and fore wing with an incurved stigmalis.
</paragraph>
</subSubSection>
</treatment>
</document>