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<mods:title id="C68FBE172BF6751288D88C7141E90F30">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part IX. Lanzatus, Orthochirus, and Somalicharmus (Buthidae), with description of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n.</mods:title>
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<emphasis id="61F5EAAFFFF5E16DEFAAF91FFBB0A3A1" bold="true" box="[911,1099,1685,1709]" italics="true" pageId="11" pageNumber="10">Orthochirus afar</emphasis>
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(
<figureCitation id="CBBA2A38FFF5E16DEFC3F938FB9CA3C5" box="[998,1127,1714,1737]" captionStart-0="Figures 41–44" captionStart-1="Figures 45–48" captionStart-2="Figures 49–65" captionStart-3="Figures 66–71" captionStart-4="Figures 72–73" captionStart-5="Figures 74–75" captionStart-6="Figures 76–77" captionStartId-0="11.[192,278,1602,1625]" captionStartId-1="12.[192,278,1782,1805]" captionStartId-2="13.[192,278,1758,1781]" captionStartId-3="15.[192,278,1806,1829]" captionStartId-4="16.[192,278,1710,1733]" captionStartId-5="17.[192,278,1806,1829]" captionStartId-6="18.[192,278,1857,1880]" captionTargetBox-0="[200,1429,156,1565]" captionTargetBox-1="[191,1440,228,1753]" captionTargetBox-2="[210,1427,222,1724]" captionTargetBox-3="[226,1430,223,1764]" captionTargetBox-4="[191,1440,189,1681]" captionTargetBox-5="[191,1440,214,1777]" captionTargetBox-6="[191,1440,236,1825]" captionTargetId-0="figure-30@11.[192,1439,144,1572]" captionTargetId-1="figure-58@12.[192,1439,229,1752]" captionTargetId-2="figure-95@13.[192,1439,215,1728]" captionTargetId-3="figure-6@15.[192,1439,217,1776]" captionTargetId-4="figure-69@16.[192,1439,190,1680]" captionTargetId-5="figure-18@17.[192,1439,214,1776]" captionTargetId-6="figure-27@18.[192,1439,237,1824]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="15" captionTargetPageId-4="16" captionTargetPageId-5="17" captionTargetPageId-6="18" captionText-0="Figures 41–44: Orthochirus afar sp. n. Figures 41–42. Male holotype in dorsal (41) and ventral (42) views. Figures 43–44. Female paratype in dorsal (43) and ventral (44) views. Scale bar: 10 mm." captionText-1="Figures 45–48: Orthochirus afar sp. n. Figures 45–46. Male holotype, carapace and tergites I–II (45), and coxosternal area and sternites (46). Figures 47–48. Female paratype, carapace and tergites I–III (47), and coxosternal area and sternites (48)." captionText-2="Figures 49–65: Orthochirus afar sp. n. Figures 49, 59, 60. Male holotype, pedipalp chela dorsal (49), movable (59) and fixed (60) finger dentition. Figures 50–58, 61–65. Female paratype, pedipalp chela dorsal (50), external (51) and ventral (52), patella dorsal (53), external (54) and ventral (55), trochanter and femur internal (56) and dorsal (57), movable finger dentition (58), and right legs I–IV, retrolateral aspect (61–64), and right leg IV, externolateral aspect (65). Trichobothrial pattern indicated in Figures 51–54 and 56–57." captionText-3="Figures 66–71: Orthochirus afar sp. n., metasoma and telson. Figures 66–68. Male holotype, lateral (66), ventral (67), and dorsal (68) views. Figures 69–71. Female paratype, lateral (69), ventral (70), and dorsal (71) views. Scale bars: 10 mm." captionText-4="Figures 72–73: Orthochirus afar sp. n., female juvenile paratype in vivo habitus in Ethiopia at locality 12EM (72) and the locality (73)." captionText-5="Figures 74–75: Orthochirus afar sp. n., male holotype (74) and female paratype (75) in vivo habitus in Ethiopia at type locality." captionText-6="Figures 76–77: Orthochirus afar sp. n., the type locality." pageId="11" pageNumber="10">Figs. 41–77</figureCitation>
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<figureCitation id="CBBA2A38FFF5E16DE857F938FB67A3C5" box="[1138,1180,1714,1737]" captionStart="Figure 155" captionStartId="35.[192,268,1422,1445]" captionTargetBox="[196,1434,196,1389]" captionTargetId="figure-260@35.[192,1437,191,1392]" captionTargetPageId="35" captionText="Figure 155: Map showing confirmed distribution of Lanzatus, Orthochirus, and Somalicharmus in Horn of Africa. Most points indicate sites sampled during 2011–2016 expeditions, except for the localities of Lanzatus somalicus in Somalia, and of Orthochirus sp. in Djibouti (the map indicates with arbitrary plotted symbol for two citations: a juvenile cited by Werner (1916: 81) with imprecise coordinates, and of a specimen cited by Lourenço &amp; Leguin (2011: 4, 15, figs. 5, 39); see also comments under Orthochirus in Systematics section." pageId="11" pageNumber="10">155</figureCitation>
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<tableCitation id="1E030306FFF5E16DE882F938FB0DA3C5" box="[1191,1270,1714,1737]" captionStart="Table 1" captionStartId="19.[192,257,836,859]" captionText="Table 1: Comparative measurements of adults of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="11" pageNumber="10">Table 1</tableCitation>
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<emphasis id="61F5EAAFFFF2E16AECE5F97CFE91A201" bold="true" box="[192,362,1782,1805]" pageId="12" pageNumber="11">Figures 45–48:</emphasis>
<taxonomicName id="94814D3EFFF2E16AED51F972FDE2A201" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[372,537,1784,1805]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="afar" status="sp. nov.">
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<emphasis id="61F5EAAFFFF2E16AEE47F972FD0FA201" bold="true" box="[610,756,1784,1805]" pageId="12" pageNumber="11">Figures 45–46</emphasis>
. Male holotype, carapace and tergites I–II (45), and coxosternal area and sternites (46). 
<emphasis id="61F5EAAFFFF2E16AED56F89BFDF9A22A" bold="true" box="[371,514,1809,1830]" pageId="12" pageNumber="11">Figures 47–48</emphasis>
. Female paratype, carapace and tergites I–III (47), and coxosternal area and sternites (48).
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<emphasis id="61F5EAAFFFF3E16BECE5F954FE92A3F9" bold="true" box="[192,361,1758,1781]" pageId="13" pageNumber="12">Figures 49–65:</emphasis>
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<emphasis id="61F5EAAFFFF3E16BEE7DF955FD3EA3F8" bold="true" box="[600,709,1759,1780]" pageId="13" pageNumber="12">Figures 49</emphasis>
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<emphasis id="61F5EAAFFFF3E16BEEF4F955FD12A3F8" bold="true" box="[721,745,1759,1780]" pageId="13" pageNumber="12">59</emphasis>
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<emphasis id="61F5EAAFFFF3E16BEED3F955FCF5A3F8" bold="true" box="[758,782,1759,1780]" pageId="13" pageNumber="12">60</emphasis>
. Male holotype, pedipalp chela dorsal (49), movable (59) and fixed (60) finger dentition. 
<emphasis id="61F5EAAFFFF3E16BEDB6F972FDD8A201" bold="true" box="[403,547,1784,1805]" pageId="13" pageNumber="12">Figures 50–58</emphasis>
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<emphasis id="61F5EAAFFFF3E16BEE14F972FD96A201" bold="true" box="[561,621,1784,1805]" pageId="13" pageNumber="12">61–65</emphasis>
. Female paratype, pedipalp chela dorsal (50), external (51) and ventral (52), patella dorsal (53), external (54) and ventral (55), trochanter and femur internal (56) and dorsal (57), movable finger dentition (58), and right legs I–IV, retrolateral aspect (61–64), and right leg IV, externolateral aspect (65). Trichobothrial pattern indicated in Figures 5 1–54 and 56–57.
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a.s.l.
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(
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<emphasis id="61F5EAAFFFF0E168EEFFFF6CFCE0A5F1" bold="true" box="[730,795,230,253]" pageId="14" pageNumber="13">12EO</emphasis>
, 
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), 
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</date>
, 
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(
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, 
<figureCitation id="CBBA2A38FFF0E168EE9CFE8EFF26A436" captionStart="Figures 41–44" captionStartId="11.[192,278,1602,1625]" captionTargetBox="[200,1429,156,1565]" captionTargetId="figure-30@11.[192,1439,144,1572]" captionTargetPageId="11" captionText="Figures 41–44: Orthochirus afar sp. n. Figures 41–42. Male holotype in dorsal (41) and ventral (42) views. Figures 43–44. Female paratype in dorsal (43) and ventral (44) views. Scale bar: 10 mm." pageId="14" pageNumber="13">Figs. 41– 42</figureCitation>
, 
<figureCitation id="CBBA2A38FFF0E168ECD5FEA9FECDA436" box="[240,310,291,314]" captionStart="Figures 45–48" captionStartId="12.[192,278,1782,1805]" captionTargetBox="[191,1440,228,1753]" captionTargetId="figure-58@12.[192,1439,229,1752]" captionTargetPageId="12" captionText="Figures 45–48: Orthochirus afar sp. n. Figures 45–46. Male holotype, carapace and tergites I–II (45), and coxosternal area and sternites (46). Figures 47–48. Female paratype, carapace and tergites I–III (47), and coxosternal area and sternites (48)." pageId="14" pageNumber="13">45–46</figureCitation>
, 
<figureCitation id="CBBA2A38FFF0E168ED6DFEA9FE45A436" box="[328,446,291,314]" captionStart="Figures 49–65" captionStartId="13.[192,278,1758,1781]" captionTargetBox="[210,1427,222,1724]" captionTargetId="figure-95@13.[192,1439,215,1728]" captionTargetPageId="13" captionText="Figures 49–65: Orthochirus afar sp. n. Figures 49, 59, 60. Male holotype, pedipalp chela dorsal (49), movable (59) and fixed (60) finger dentition. Figures 50–58, 61–65. Female paratype, pedipalp chela dorsal (50), external (51) and ventral (52), patella dorsal (53), external (54) and ventral (55), trochanter and femur internal (56) and dorsal (57), movable finger dentition (58), and right legs I–IV, retrolateral aspect (61–64), and right leg IV, externolateral aspect (65). Trichobothrial pattern indicated in Figures 51–54 and 56–57." pageId="14" pageNumber="13">49, 59–60</figureCitation>
, 
<figureCitation id="CBBA2A38FFF0E168EDF5FEA9FDEDA436" box="[464,534,291,314]" captionStart="Figures 66–71" captionStartId="15.[192,278,1806,1829]" captionTargetBox="[226,1430,223,1764]" captionTargetId="figure-6@15.[192,1439,217,1776]" captionTargetPageId="15" captionText="Figures 66–71: Orthochirus afar sp. n., metasoma and telson. Figures 66–68. Male holotype, lateral (66), ventral (67), and dorsal (68) views. Figures 69–71. Female paratype, lateral (69), ventral (70), and dorsal (71) views. Scale bars: 10 mm." pageId="14" pageNumber="13">66–68</figureCitation>
, 
<figureCitation id="CBBA2A38FFF0E168EE0DFEA9FDBFA436" box="[552,580,291,314]" captionStart="Figures 74–75" captionStartId="17.[192,278,1806,1829]" captionTargetBox="[191,1440,214,1777]" captionTargetId="figure-18@17.[192,1439,214,1776]" captionTargetPageId="17" captionText="Figures 74–75: Orthochirus afar sp. n., male holotype (74) and female paratype (75) in vivo habitus in Ethiopia at type locality." pageId="14" pageNumber="13">74</figureCitation>
) 
<specimenCount id="4587FD34FFF0E168EE7FFEA9FD87A436" box="[602,636,291,314]" pageId="14" pageNumber="13" type="male">1♂</specimenCount>
<specimenCount id="4587FD34FFF0E168EE59FEA9FD66A436" box="[636,669,291,314]" pageId="14" pageNumber="13" type="female">1♀</specimenCount>
(
<typeStatus id="8C3A881FFFF0E168EE96FEA9FCE6A436" box="[691,797,291,314]" pageId="14" pageNumber="13" type="paratype">paratypes</typeStatus>
, 
<figureCitation id="CBBA2A38FFF0E168ECE5FEC8FEBEA455" box="[192,325,322,345]" captionStart="Figures 41–44" captionStartId="11.[192,278,1602,1625]" captionTargetBox="[200,1429,156,1565]" captionTargetId="figure-30@11.[192,1439,144,1572]" captionTargetPageId="11" captionText="Figures 41–44: Orthochirus afar sp. n. Figures 41–42. Male holotype in dorsal (41) and ventral (42) views. Figures 43–44. Female paratype in dorsal (43) and ventral (44) views. Scale bar: 10 mm." pageId="14" pageNumber="13">Figs. 43–44</figureCitation>
, 
<figureCitation id="CBBA2A38FFF0E168ED70FEC8FE61A455" box="[341,410,322,345]" captionStart="Figures 45–48" captionStartId="12.[192,278,1782,1805]" captionTargetBox="[191,1440,228,1753]" captionTargetId="figure-58@12.[192,1439,229,1752]" captionTargetPageId="12" captionText="Figures 45–48: Orthochirus afar sp. n. Figures 45–46. Male holotype, carapace and tergites I–II (45), and coxosternal area and sternites (46). Figures 47–48. Female paratype, carapace and tergites I–III (47), and coxosternal area and sternites (48)." pageId="14" pageNumber="13">47–48</figureCitation>
, 
<figureCitation id="CBBA2A38FFF0E168ED8FFEC8FDBFA455" box="[426,580,322,345]" captionStart="Figures 49–65" captionStartId="13.[192,278,1758,1781]" captionTargetBox="[210,1427,222,1724]" captionTargetId="figure-95@13.[192,1439,215,1728]" captionTargetPageId="13" captionText="Figures 49–65: Orthochirus afar sp. n. Figures 49, 59, 60. Male holotype, pedipalp chela dorsal (49), movable (59) and fixed (60) finger dentition. Figures 50–58, 61–65. Female paratype, pedipalp chela dorsal (50), external (51) and ventral (52), patella dorsal (53), external (54) and ventral (55), trochanter and femur internal (56) and dorsal (57), movable finger dentition (58), and right legs I–IV, retrolateral aspect (61–64), and right leg IV, externolateral aspect (65). Trichobothrial pattern indicated in Figures 51–54 and 56–57." pageId="14" pageNumber="13">50–58, 61–65</figureCitation>
, 
<figureCitation id="CBBA2A38FFF0E168EE76FEC8FD63A455" box="[595,664,322,345]" captionStart="Figures 66–71" captionStartId="15.[192,278,1806,1829]" captionTargetBox="[226,1430,223,1764]" captionTargetId="figure-6@15.[192,1439,217,1776]" captionTargetPageId="15" captionText="Figures 66–71: Orthochirus afar sp. n., metasoma and telson. Figures 66–68. Male holotype, lateral (66), ventral (67), and dorsal (68) views. Figures 69–71. Female paratype, lateral (69), ventral (70), and dorsal (71) views. Scale bars: 10 mm." pageId="14" pageNumber="13">69–71</figureCitation>
, 
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), leg. 
<collectorName id="FE74536BFFF0E168ED56FEEAFE15A47B" box="[371,494,352,375]" pageId="14" pageNumber="13">F. Kovařík</collectorName>
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, 
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<quantity id="94799B58FFF0E168ED2DFEF5FEAAA49A" box="[264,337,383,406]" metricMagnitude="2" metricUnit="m" metricValue="4.57" pageId="14" pageNumber="13" unit="m" value="457.0">457 m</quantity>
a.s.l.
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(
<collectorName id="FE74536BFFF0E168ED87FEF5FDCAA49A" box="[418,561,383,406]" pageId="14" pageNumber="13">Locality No.</collectorName>
<emphasis id="61F5EAAFFFF0E168EE19FEF5FD7AA49A" bold="true" box="[572,641,383,406]" pageId="14" pageNumber="13">12EM</emphasis>
, 
<figureCitation id="CBBA2A38FFF0E168EEB1FEF5FD13A49A" box="[660,744,383,406]" captionStart="Figures 72–73" captionStartId="16.[192,278,1710,1733]" captionTargetBox="[191,1440,189,1681]" captionTargetId="figure-69@16.[192,1439,190,1680]" captionTargetPageId="16" captionText="Figures 72–73: Orthochirus afar sp. n., female juvenile paratype in vivo habitus in Ethiopia at locality 12EM (72) and the locality (73)." pageId="14" pageNumber="13">Fig. 73</figureCitation>
), 
<date id="273F107DFFF0E168EF25FEF5FEE1A4B9" pageId="14" pageNumber="13" value="2012-11-20">
<collectingDate id="377BE995FFF0E168EF25FEF5FEE1A4B9" pageId="14" pageNumber="13" value="2012-11-20">20. XI.2012</collectingDate>
</date>
, 
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juv. (
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, 
<figureCitation id="CBBA2A38FFF0E168EDD8FE14FDA9A4B9" box="[509,594,414,437]" captionStart="Figures 72–73" captionStartId="16.[192,278,1710,1733]" captionTargetBox="[191,1440,189,1681]" captionTargetId="figure-69@16.[192,1439,190,1680]" captionTargetPageId="16" captionText="Figures 72–73: Orthochirus afar sp. n., female juvenile paratype in vivo habitus in Ethiopia at locality 12EM (72) and the locality (73)." pageId="14" pageNumber="13">Fig. 72</figureCitation>
), leg. 
<collectorName id="FE74536BFFF0E168EE84FE14FCE6A4B9" box="[673,797,414,437]" pageId="14" pageNumber="13">F. Kovařík</collectorName>
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, 
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.
</paragraph>
</subSubSection>
<subSubSection id="1B9B6536FFF0E168ECE5FE70FEE4A60A" pageId="14" pageNumber="13" type="etymology">
<paragraph id="533E36BDFFF0E168ECE5FE70FEE4A60A" blockId="14.[192,802,506,774]" pageId="14" pageNumber="13">
ETYMOLOGY. Named after the 
<collectingRegion id="9145F85FFFF0E168EE3CFE70FDB1A71D" box="[537,586,506,529]" country="Ethiopia" name="Afar" pageId="14" pageNumber="13">Afar</collectingRegion>
(
<taxonomicName id="94814D3EFFF0E168EE79FE70FD24A71D" authority=": Qafar" authorityName="Qafar" box="[604,735,506,529]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="afar">
Afar: 
<emphasis id="61F5EAAFFFF0E168EEBAFE70FD24A71D" box="[671,735,506,529]" italics="true" pageId="14" pageNumber="13">Qafár</emphasis>
</taxonomicName>
) people, also known as the Danakil and Adal, an ethnic group inhabiting the Horn of Africa. They reside primarily in the 
<collectingRegion id="9145F85FFFF0E168ED4DFDDCFE0EA761" box="[360,501,598,621]" country="Ethiopia" name="Afar" pageId="14" pageNumber="13">Afar Region</collectingRegion>
of 
<collectingCountry id="2B96762DFFF0E168EE0CFDDCFD78A761" box="[553,643,598,621]" name="Ethiopia" pageId="14" pageNumber="13">Ethiopia</collectingCountry>
and northern 
<collectingCountry id="2B96762DFFF0E168ECE5FDFEFEE7A787" box="[192,284,628,651]" name="Djibouti" pageId="14" pageNumber="13">Djibouti</collectingCountry>
, although some also live in the southern end of 
<collectingCountry id="2B96762DFFF0E168ECE5FD19FEF7A7A6" box="[192,268,659,682]" name="Eritrea" pageId="14" pageNumber="13">Eritrea</collectingCountry>
. The Afars speak the Afar language, which is part of the Cushitic branch of the Afro-Asiatic family. Both recorded localities of this species are within their territory.
</paragraph>
</subSubSection>
<subSubSection id="1B9B6536FFF0E168ECE5FCA6FE21A1DE" pageId="14" pageNumber="13" type="diagnosis">
<paragraph id="533E36BDFFF0E168ECE5FCA6FE21A1DE" blockId="14.[192,802,812,1234]" pageId="14" pageNumber="13">
DIAGNOSIS: Adults from 
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(male) to 
<quantity id="94799B58FFF0E168EEE9FCA6FCD9A64F" box="[716,802,812,835]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="14" pageNumber="13" unit="mm" value="29.0">29 mm</quantity>
(female) long. Base color uniformly black, sternites III– VI brown with yellow zones indicated, pedipalp movable fingers yellow, tarsomeres of legs yellow to white, telson reddish brown. Pectine teeth number 
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males, 
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females. Metasomal segments IV–V ventrally punctate, without median carinae; areas between punctae smooth. Entire metasoma glabrous. Dorsal surface of metasomal segments III–V smooth medially. Pedipalp chela movable finger bear 10 or 11 rows of granules, with internal and external accessory granules. Fixed fingers bear 9 rows of granules, with internal and external accessory granules. Tarsomeres of legs without bristlecombs.
</paragraph>
</subSubSection>
<subSubSection id="1B9B6536FFF0E175ECE5FB72FF07A1FA" lastPageId="19" lastPageNumber="18" pageId="14" pageNumber="13" type="description">
<paragraph id="533E36BDFFF0E168ECE5FB72FE29A309" blockId="14.[192,803,1272,1909]" pageId="14" pageNumber="13">
DESCRIPTION. Adults 
<quantity id="94799B58FFF0E168ED92FB72FDF3A003" box="[439,520,1272,1295]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="14" pageNumber="13" unit="mm" value="23.0">23 mm</quantity>
(male)– 
<quantity id="94799B58FFF0E168EE4CFB72FD41A003" box="[617,698,1272,1295]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="14" pageNumber="13" unit="mm" value="29.0">29 mm</quantity>
(female) long. Habitus as shown in 
<figureCitation id="CBBA2A38FFF0E168EDC7FA9DFD9EA022" box="[482,613,1303,1326]" captionStart="Figures 41–44" captionStartId="11.[192,278,1602,1625]" captionTargetBox="[200,1429,156,1565]" captionTargetId="figure-30@11.[192,1439,144,1572]" captionTargetPageId="11" captionText="Figures 41–44: Orthochirus afar sp. n. Figures 41–42. Male holotype in dorsal (41) and ventral (42) views. Figures 43–44. Female paratype in dorsal (43) and ventral (44) views. Scale bar: 10 mm." pageId="14" pageNumber="13">Figs. 41–44</figureCitation>
. For position and distribution of trichobothria of pedipalps see 
<figureCitation id="CBBA2A38FFF0E168EE86FABCFEC3A067" captionStart="Figures 49–65" captionStartId="13.[192,278,1758,1781]" captionTargetBox="[210,1427,222,1724]" captionTargetId="figure-95@13.[192,1439,215,1728]" captionTargetPageId="13" captionText="Figures 49–65: Orthochirus afar sp. n. Figures 49, 59, 60. Male holotype, pedipalp chela dorsal (49), movable (59) and fixed (60) finger dentition. Figures 50–58, 61–65. Female paratype, pedipalp chela dorsal (50), external (51) and ventral (52), patella dorsal (53), external (54) and ventral (55), trochanter and femur internal (56) and dorsal (57), movable finger dentition (58), and right legs I–IV, retrolateral aspect (61–64), and right leg IV, externolateral aspect (65). Trichobothrial pattern indicated in Figures 51–54 and 56–57." pageId="14" pageNumber="13">Figs. 51–54 and 56–57</figureCitation>
. On femur, trichobothrium 
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missing, distance between 
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and 
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shorter that distance between 
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and 
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, 
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situated between 
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and 
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<subScript id="CF0534F8FFF0E168EE66FA16FDB7A0A7" attach="right" box="[579,588,1436,1451]" fontSize="6" pageId="14" pageNumber="13">4</subScript>
. Only minor sexual dimorphism in pectines (
<figureCitation id="CBBA2A38FFF0E168EDF7FA3AFDD7A0CB" box="[466,556,1456,1479]" captionStart="Figures 45–48" captionStartId="12.[192,278,1782,1805]" captionTargetBox="[191,1440,228,1753]" captionTargetId="figure-58@12.[192,1439,229,1752]" captionTargetPageId="12" captionText="Figures 45–48: Orthochirus afar sp. n. Figures 45–46. Male holotype, carapace and tergites I–II (45), and coxosternal area and sternites (46). Figures 47–48. Female paratype, carapace and tergites I–III (47), and coxosternal area and sternites (48)." pageId="14" pageNumber="13">Figs. 46</figureCitation>
vs. 48), no difference between sexes in relative lengths or widths of pedipalps and metasomal segments.
</paragraph>
<paragraph id="533E36BDFFF0E168ECE5F986FE9AA3B1" blockId="14.[192,803,1272,1909]" pageId="14" pageNumber="13">
<emphasis id="61F5EAAFFFF0E168ECE5F986FEC0A32F" bold="true" box="[192,315,1548,1571]" pageId="14" pageNumber="13">Coloration</emphasis>
(
<figureCitation id="CBBA2A38FFF0E168ED6BF986FE2FA32F" box="[334,468,1548,1571]" captionStart-0="Figures 41–44" captionStart-1="Figures 45–48" captionStartId-0="11.[192,278,1602,1625]" captionStartId-1="12.[192,278,1782,1805]" captionTargetBox-0="[200,1429,156,1565]" captionTargetBox-1="[191,1440,228,1753]" captionTargetId-0="figure-30@11.[192,1439,144,1572]" captionTargetId-1="figure-58@12.[192,1439,229,1752]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="Figures 41–44: Orthochirus afar sp. n. Figures 41–42. Male holotype in dorsal (41) and ventral (42) views. Figures 43–44. Female paratype in dorsal (43) and ventral (44) views. Scale bar: 10 mm." captionText-1="Figures 45–48: Orthochirus afar sp. n. Figures 45–46. Male holotype, carapace and tergites I–II (45), and coxosternal area and sternites (46). Figures 47–48. Female paratype, carapace and tergites I–III (47), and coxosternal area and sternites (48)." pageId="14" pageNumber="13">Figs. 41–48</figureCitation>
). Base color uniformly black, sternites III–VI brown with yellow zones indicated, seventh sternite black. Chelicerae brown to black, reticulate. Pedipalp movable finger yellow, lighter than manus. Tarsomeres of legs yellow to white. Telson reddish brown.
</paragraph>
<paragraph id="533E36BDFFF0E168ECE5F94EFC68A47B" blockId="14.[192,803,1272,1909]" lastBlockId="14.[830,1441,199,1265]" pageId="14" pageNumber="13">
<emphasis id="61F5EAAFFFF0E168ECE5F94EFDF9A3D7" bold="true" box="[192,514,1732,1755]" pageId="14" pageNumber="13">Carapace and mesosoma</emphasis>
(
<figureCitation id="CBBA2A38FFF0E168EE01F94EFD40A3D7" box="[548,699,1732,1755]" captionStart="Figures 45–48" captionStartId="12.[192,278,1782,1805]" captionTargetBox="[191,1440,228,1753]" captionTargetId="figure-58@12.[192,1439,229,1752]" captionTargetPageId="12" captionText="Figures 45–48: Orthochirus afar sp. n. Figures 45–46. Male holotype, carapace and tergites I–II (45), and coxosternal area and sternites (46). Figures 47–48. Female paratype, carapace and tergites I–III (47), and coxosternal area and sternites (48)." pageId="14" pageNumber="13">Figs. 45–48</figureCitation>
). Entire carapace covered by large granules. Carinae absent. Anterior margin of the carapace almost straight, weakly convex medially. Tergites granulated, coarsely so in males. Tergite VII pentacarinate. Pectinal tooth count 18–20 (2x18, 3x19, 1x20) in males, 16–17 (3x16, 1x17) in females. Pectine marginal tips extend to end of fourth sternite in both sexes. The pectines with three marginal lamellae, 6–7 middle lamellae. Lamellae and fulcra bear numerous pale setae. Sternites III–VI smooth without carinae. Sternite VII densely granulated, with one pair of carinae.
</paragraph>
<paragraph id="533E36BDFFF0E168EF1BFEF5FAF1A60A" blockId="14.[830,1441,199,1265]" pageId="14" pageNumber="13">
<emphasis id="61F5EAAFFFF0E168EF1BFEF5FBC2A49A" bold="true" box="[830,1081,383,406]" pageId="14" pageNumber="13">Metasoma and telson</emphasis>
(
<figureCitation id="CBBA2A38FFF0E168E86EFEF5FB28A49A" box="[1099,1235,383,406]" captionStart="Figures 66–71" captionStartId="15.[192,278,1806,1829]" captionTargetBox="[226,1430,223,1764]" captionTargetId="figure-6@15.[192,1439,217,1776]" captionTargetPageId="15" captionText="Figures 66–71: Orthochirus afar sp. n., metasoma and telson. Figures 66–68. Male holotype, lateral (66), ventral (67), and dorsal (68) views. Figures 69–71. Female paratype, lateral (69), ventral (70), and dorsal (71) views. Scale bars: 10 mm." pageId="14" pageNumber="13">Figs. 66–71</figureCitation>
). First and second metasomal segments bear a total of 10 granulated carinae. Segments III–V lacking lateral carinae, segments IV and V with only dorsal and ventrolateral carinae, all smooth, sometimes incomplete. First metasomal segment granulated, second segment partly granulated, other segments smooth. All segments punctate. Punctae weak on first segment, stronger on segments II–V. Areas between punctae smooth. Dorsal surface of all metasomal segments smooth medially except for first segment. Entire metasoma and telson glabrous. Short, thin setae may emerge from some punctae. Telson punctate, lacking granules.
</paragraph>
<paragraph id="533E36BDFFF0E168EF1BFC84FB84A116" blockId="14.[830,1441,199,1265]" pageId="14" pageNumber="13">
<emphasis id="61F5EAAFFFF0E168EF1BFC84FC56A629" bold="true" box="[830,941,782,805]" pageId="14" pageNumber="13">Pedipalps</emphasis>
(
<figureCitation id="CBBA2A38FFF0E168EFEDFC84FBA3A629" box="[968,1112,782,805]" captionStart="Figures 49–65" captionStartId="13.[192,278,1758,1781]" captionTargetBox="[210,1427,222,1724]" captionTargetId="figure-95@13.[192,1439,215,1728]" captionTargetPageId="13" captionText="Figures 49–65: Orthochirus afar sp. n. Figures 49, 59, 60. Male holotype, pedipalp chela dorsal (49), movable (59) and fixed (60) finger dentition. Figures 50–58, 61–65. Female paratype, pedipalp chela dorsal (50), external (51) and ventral (52), patella dorsal (53), external (54) and ventral (55), trochanter and femur internal (56) and dorsal (57), movable finger dentition (58), and right legs I–IV, retrolateral aspect (61–64), and right leg IV, externolateral aspect (65). Trichobothrial pattern indicated in Figures 51–54 and 56–57." pageId="14" pageNumber="13">Figs. 49–60</figureCitation>
). Pedipalp femur granulate, bearing 5 granulate carinae. Patella with 7 smooth, well developed carinae. Chela with smooth carinae which may be discernible throughout length of fixed finger. Movable finger with 10 or 11 rows of granules, flanked by 9 or 10 internal and 8 or 9 external accessory granules, 4 subterminal granules, one terminal granule. Fixed finger with 9 rows of granules, flanked by 8 internal and 8 external accessory granules.
</paragraph>
<paragraph id="533E36BDFFF0E168EF1BFBA8FAB3A1DE" blockId="14.[830,1441,199,1265]" pageId="14" pageNumber="13">
<emphasis id="61F5EAAFFFF0E168EF1BFBA8FC88A135" bold="true" box="[830,883,1058,1081]" pageId="14" pageNumber="13">Legs</emphasis>
(
<figureCitation id="CBBA2A38FFF0E168EFA2FBA8FBF5A135" box="[903,1038,1058,1081]" captionStart="Figures 49–65" captionStartId="13.[192,278,1758,1781]" captionTargetBox="[210,1427,222,1724]" captionTargetId="figure-95@13.[192,1439,215,1728]" captionTargetPageId="13" captionText="Figures 49–65: Orthochirus afar sp. n. Figures 49, 59, 60. Male holotype, pedipalp chela dorsal (49), movable (59) and fixed (60) finger dentition. Figures 50–58, 61–65. Female paratype, pedipalp chela dorsal (50), external (51) and ventral (52), patella dorsal (53), external (54) and ventral (55), trochanter and femur internal (56) and dorsal (57), movable finger dentition (58), and right legs I–IV, retrolateral aspect (61–64), and right leg IV, externolateral aspect (65). Trichobothrial pattern indicated in Figures 51–54 and 56–57." pageId="14" pageNumber="13">Figs. 61–65</figureCitation>
). Legs III and IV with tibial spurs. Retrolateral and prolateral pedal spurs present on all legs. Femur, patella, tibia and partly tarsomere I carinate and smooth. Patella with only a few solitary setae and spines. The tibia with only several spines on the external aspect. Tarsomeres of legs without bristlecombs.
</paragraph>
<paragraph id="533E36BDFFF0E168EF1BFB50FB8EA1FD" blockId="14.[830,1441,199,1265]" box="[830,1141,1242,1265]" pageId="14" pageNumber="13">
<emphasis id="61F5EAAFFFF0E168EF1BFB50FC1EA1FD" bold="true" box="[830,997,1242,1265]" pageId="14" pageNumber="13">Measurements</emphasis>
. See 
<tableCitation id="1E030306FFF0E168E83AFB50FB8EA1FD" box="[1055,1141,1242,1265]" captionStart="Table 1" captionStartId="19.[192,257,836,859]" captionText="Table 1: Comparative measurements of adults of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="14" pageNumber="13">Table 1.</tableCitation>
</paragraph>
<paragraph id="533E36BDFFF0E176EF1BFA9DFEC4A25E" blockId="14.[830,1441,1303,1909]" lastBlockId="16.[192,802,1789,1874]" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="13">
AFFINITIES. The described features distinguish 
<taxonomicName id="94814D3EFFF0E168E96BFA9DFA64A022" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[1358,1439,1303,1326]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="afar" status="sp. nov.">
<emphasis id="61F5EAAFFFF0E168E96BFA9DFA64A022" box="[1358,1439,1303,1326]" italics="true" pageId="14" pageNumber="13">O. afar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AC657D4FFF0E168EF1BFABCFC87A041" box="[830,892,1334,1357]" pageId="14" pageNumber="13" rank="species">
<emphasis id="61F5EAAFFFF0E168EF1BFABCFCACA041" bold="true" box="[830,855,1334,1357]" pageId="14" pageNumber="13">sp</emphasis>
. 
<emphasis id="61F5EAAFFFF0E168EF42FABCFC8DA041" bold="true" box="[871,886,1334,1357]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
from all other species of the genus. 
<taxonomicName id="94814D3EFFF0E168E929FABCFAA1A041" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[1292,1370,1334,1357]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="afar" status="sp. nov.">
<emphasis id="61F5EAAFFFF0E168E929FABCFAA1A041" box="[1292,1370,1334,1357]" italics="true" pageId="14" pageNumber="13">O. afar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AC657D4FFF0E168E946FABCFA5BA041" box="[1379,1440,1334,1357]" pageId="14" pageNumber="13" rank="species">
<emphasis id="61F5EAAFFFF0E168E946FABCFA87A041" bold="true" box="[1379,1404,1334,1357]" pageId="14" pageNumber="13">sp</emphasis>
. 
<emphasis id="61F5EAAFFFF0E168E9AEFABCFA61A041" bold="true" box="[1419,1434,1334,1357]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
is similar to 
<taxonomicName id="94814D3EFFF0E168EFEBFADEFBB1A067" authorityName="" baseAuthorityName="Simon" baseAuthorityYear="1883" box="[974,1098,1364,1387]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="aristidis">
<emphasis id="61F5EAAFFFF0E168EFEBFADEFBB1A067" box="[974,1098,1364,1387]" italics="true" pageId="14" pageNumber="13">O. aristidis</emphasis>
</taxonomicName>
from 
<collectingCountry id="2B96762DFFF0E168E8B7FADEFB29A067" box="[1170,1234,1364,1387]" name="Egypt" pageId="14" pageNumber="13">Egypt</collectingCountry>
and 
<collectingCountry id="2B96762DFFF0E168E928FADEFAABA067" box="[1293,1360,1364,1387]" name="Sudan" pageId="14" pageNumber="13">Sudan</collectingCountry>
. These two can be separated unequivocally by: 
<emphasis id="61F5EAAFFFF0E168E937FAF9FAE4A086" bold="true" box="[1298,1311,1395,1418]" pageId="14" pageNumber="13">1</emphasis>
) fixed and movable fingers of pedipalp with 9 and 10–11 rows of granules in 
<taxonomicName id="94814D3EFFF0E168EF9EFA3AFBF2A0CB" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[955,1033,1456,1479]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="afar" status="sp. nov.">
<emphasis id="61F5EAAFFFF0E168EF9EFA3AFBF2A0CB" box="[955,1033,1456,1479]" italics="true" pageId="14" pageNumber="13">O. afar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AC657D4FFF0E168E82AFA3AFBB1A0CB" box="[1039,1098,1456,1479]" pageId="14" pageNumber="13" rank="species">
<emphasis id="61F5EAAFFFF0E168E82AFA3AFBD3A0CB" bold="true" box="[1039,1064,1456,1479]" pageId="14" pageNumber="13">sp</emphasis>
. 
<emphasis id="61F5EAAFFFF0E168E810FA3AFBBFA0CB" bold="true" box="[1077,1092,1456,1479]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
, vs. 8 and 9 rows of granules in 
<taxonomicName id="94814D3EFFF0E168EF1BFA45FC43A0EA" authorityName="" baseAuthorityName="Simon" baseAuthorityYear="1883" box="[830,952,1487,1510]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="aristidis">
<emphasis id="61F5EAAFFFF0E168EF1BFA45FC43A0EA" box="[830,952,1487,1510]" italics="true" pageId="14" pageNumber="13">O. aristidis</emphasis>
</taxonomicName>
; 
<emphasis id="61F5EAAFFFF0E168EFE3FA45FC28A0EA" bold="true" box="[966,979,1487,1510]" pageId="14" pageNumber="13">2</emphasis>
) sternites III–VI brown with yellow zones indicated in 
<taxonomicName id="94814D3EFFF0E168EFECFA64FBE2A309" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[969,1049,1518,1541]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="afar" status="sp. nov.">
<emphasis id="61F5EAAFFFF0E168EFECFA64FBE2A309" box="[969,1049,1518,1541]" italics="true" pageId="14" pageNumber="13">O. afar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AC657D4FFF0E168E801FA64FB98A309" box="[1060,1123,1518,1541]" pageId="14" pageNumber="13" rank="species">
<emphasis id="61F5EAAFFFF0E168E801FA64FBC6A309" bold="true" box="[1060,1085,1518,1541]" pageId="14" pageNumber="13">sp</emphasis>
. 
<emphasis id="61F5EAAFFFF0E168E868FA64FBA7A309" bold="true" box="[1101,1116,1518,1541]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="CBBA2A38FFF0E168E850FA64FAD3A309" box="[1141,1320,1518,1541]" captionStart="Figures 45–48" captionStartId="12.[192,278,1782,1805]" captionTargetBox="[191,1440,228,1753]" captionTargetId="figure-58@12.[192,1439,229,1752]" captionTargetPageId="12" captionText="Figures 45–48: Orthochirus afar sp. n. Figures 45–46. Male holotype, carapace and tergites I–II (45), and coxosternal area and sternites (46). Figures 47–48. Female paratype, carapace and tergites I–III (47), and coxosternal area and sternites (48)." pageId="14" pageNumber="13">Figs. 46 and 48</figureCitation>
), vs. black without yellow zones in 
<taxonomicName id="94814D3EFFF0E168E868F986FB3CA32F" authorityName="" baseAuthorityName="Simon" baseAuthorityYear="1883" box="[1101,1223,1548,1571]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="aristidis">
<emphasis id="61F5EAAFFFF0E168E868F986FB3CA32F" box="[1101,1223,1548,1571]" italics="true" pageId="14" pageNumber="13">O. aristidis</emphasis>
</taxonomicName>
(fig. 
<quantity id="94799B58FFF0E168E923F986FACAA32F" box="[1286,1329,1548,1571]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="14" pageNumber="13" unit="in" value="4.0">4 in</quantity>
Lourenço &amp; Leguin, 2011: 3); 
<emphasis id="61F5EAAFFFF0E168E802F9A1FBCFA34E" bold="true" box="[1063,1076,1579,1602]" pageId="14" pageNumber="13">3</emphasis>
) sternite VII densely granulated, bearing one pair of carinae in 
<taxonomicName id="94814D3EFFF0E168E8B2F9C0FB1CA36D" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[1175,1255,1610,1633]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="afar" status="sp. nov.">
<emphasis id="61F5EAAFFFF0E168E8B2F9C0FB1CA36D" box="[1175,1255,1610,1633]" italics="true" pageId="14" pageNumber="13">O. afar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AC657D4FFF0E168E8D7F9C0FAC8A36D" box="[1266,1331,1610,1633]" pageId="14" pageNumber="13" rank="species">
<emphasis id="61F5EAAFFFF0E168E8D7F9C0FAF0A36D" bold="true" box="[1266,1291,1610,1633]" pageId="14" pageNumber="13">sp</emphasis>
. 
<emphasis id="61F5EAAFFFF0E168E938F9C0FAD7A36D" bold="true" box="[1309,1324,1610,1633]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="CBBA2A38FFF0E168E960F9C0FC74A373" captionStart="Figures 45–48" captionStartId="12.[192,278,1782,1805]" captionTargetBox="[191,1440,228,1753]" captionTargetId="figure-58@12.[192,1439,229,1752]" captionTargetPageId="12" captionText="Figures 45–48: Orthochirus afar sp. n. Figures 45–46. Male holotype, carapace and tergites I–II (45), and coxosternal area and sternites (46). Figures 47–48. Female paratype, carapace and tergites I–III (47), and coxosternal area and sternites (48)." pageId="14" pageNumber="13">Figs. 46 and 48</figureCitation>
), vs. two pairs of moderate carinae with some minute granulations in 
<taxonomicName id="94814D3EFFF0E168E81CF90DFC67A3B1" authority="(Lourenco &amp; Leguin, 2011: 2)" baseAuthorityName="Lourenco &amp; Leguin" baseAuthorityPageNumber="2" baseAuthorityYear="2011" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="aristidis">
<emphasis id="61F5EAAFFFF0E168E81CF90DFB49A392" box="[1081,1202,1671,1694]" italics="true" pageId="14" pageNumber="13">O. aristidis</emphasis>
(Lourenço &amp; Leguin, 2011: 2)
</taxonomicName>
; 
<emphasis id="61F5EAAFFFF0E168EF8DF92CFC4EA3B1" bold="true" box="[936,949,1702,1725]" pageId="14" pageNumber="13">4</emphasis>
) on pedipalp femur, distance between trichobothria 
<emphasis id="61F5EAAFFFF0E168EFB1F94EFC5AA3D7" box="[916,929,1732,1755]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<subScript id="CF0534F8FFF0E168EF87F944FC50A3D1" attach="left" box="[930,939,1742,1757]" fontSize="6" pageId="14" pageNumber="13">1</subScript>
and 
<emphasis id="61F5EAAFFFF0E168EFC3F94EFC08A3D7" box="[998,1011,1732,1755]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<subScript id="CF0534F8FFF0E168EFD1F944FC06A3D1" attach="left" box="[1012,1021,1742,1757]" fontSize="6" pageId="14" pageNumber="13">3</subScript>
shorter than distance between 
<emphasis id="61F5EAAFFFF0E168E97CF94EFA9DA3D7" box="[1369,1382,1732,1755]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<subScript id="CF0534F8FFF0E168E942F944FA8BA3D1" attach="left" box="[1383,1392,1742,1757]" fontSize="6" pageId="14" pageNumber="13">3</subScript>
and 
<emphasis id="61F5EAAFFFF0E168EF1BF969FCB0A3F6" box="[830,843,1763,1786]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<subScript id="CF0534F8FFF0E168EF69F967FCAEA3F0" attach="right" box="[844,853,1773,1788]" fontSize="6" pageId="14" pageNumber="13">4</subScript>
, trichobothrium 
<emphasis id="61F5EAAFFFF0E168E834F969FBE6A3F6" box="[1041,1053,1763,1786]" italics="true" pageId="14" pageNumber="13">e</emphasis>
<subScript id="CF0534F8FFF0E168E838F967FBDDA3F0" attach="left" box="[1053,1062,1773,1788]" fontSize="6" pageId="14" pageNumber="13">1</subScript>
situated between 
<emphasis id="61F5EAAFFFF0E168E8D6F969FAFBA3F6" box="[1267,1280,1763,1786]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<subScript id="CF0534F8FFF0E168E925F967FAF2A3F0" attach="left" box="[1280,1289,1773,1788]" fontSize="6" pageId="14" pageNumber="13">3</subScript>
and 
<emphasis id="61F5EAAFFFF0E168E960F969FAA9A3F6" box="[1349,1362,1763,1786]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<quantity id="94799B58FFF0E168E976F967FA80A3F6" box="[1363,1403,1763,1788]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="14" pageNumber="13" unit="in" value="4.0">
<subScript id="CF0534F8FFF0E168E976F967FAA7A3F0" attach="left" box="[1363,1372,1773,1788]" fontSize="6" pageId="14" pageNumber="13">4</subScript>
in
</quantity>
<taxonomicName id="94814D3EFFF0E168E9A3F969FC90A215" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="afar" status="sp. nov.">
<emphasis id="61F5EAAFFFF0E168E9A3F969FC90A215" italics="true" pageId="14" pageNumber="13">O. afar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AC657D4FFF0E168EF5DF888FC40A215" box="[888,955,1794,1817]" pageId="14" pageNumber="13" rank="species">
<emphasis id="61F5EAAFFFF0E168EF5DF888FC6AA215" bold="true" box="[888,913,1794,1817]" pageId="14" pageNumber="13">sp</emphasis>
. 
<emphasis id="61F5EAAFFFF0E168EF80F888FC4FA215" bold="true" box="[933,948,1794,1817]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="CBBA2A38FFF0E168EFEAF888FBDEA215" box="[975,1061,1794,1817]" captionStart="Figures 49–65" captionStartId="13.[192,278,1758,1781]" captionTargetBox="[210,1427,222,1724]" captionTargetId="figure-95@13.[192,1439,215,1728]" captionTargetPageId="13" captionText="Figures 49–65: Orthochirus afar sp. n. Figures 49, 59, 60. Male holotype, pedipalp chela dorsal (49), movable (59) and fixed (60) finger dentition. Figures 50–58, 61–65. Female paratype, pedipalp chela dorsal (50), external (51) and ventral (52), patella dorsal (53), external (54) and ventral (55), trochanter and femur internal (56) and dorsal (57), movable finger dentition (58), and right legs I–IV, retrolateral aspect (61–64), and right leg IV, externolateral aspect (65). Trichobothrial pattern indicated in Figures 51–54 and 56–57." pageId="14" pageNumber="13">Fig. 57</figureCitation>
), vs. distance between 
<emphasis id="61F5EAAFFFF0E168E911F888FABAA215" box="[1332,1345,1794,1817]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<subScript id="CF0534F8FFF0E168E964F886FAB1A217" attach="left" box="[1345,1354,1804,1819]" fontSize="6" pageId="14" pageNumber="13">1</subScript>
and 
<emphasis id="61F5EAAFFFF0E168E9AFF888FA6CA215" box="[1418,1431,1794,1817]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<subScript id="CF0534F8FFF0E168E9B2F886FA5BA217" attach="left" box="[1431,1440,1804,1819]" fontSize="6" pageId="14" pageNumber="13">3</subScript>
approximately equal to that between 
<emphasis id="61F5EAAFFFF0E168E8D2F8AAFAFFA23B" box="[1271,1284,1824,1847]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<subScript id="CF0534F8FFF0E168E921F8A0FAF6A235" attach="left" box="[1284,1293,1834,1849]" fontSize="6" pageId="14" pageNumber="13">3</subScript>
and 
<emphasis id="61F5EAAFFFF0E168E976F8AAFA9BA23B" box="[1363,1376,1824,1847]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<subScript id="CF0534F8FFF0E168E944F8A0FA92A235" attach="right" box="[1377,1385,1834,1849]" fontSize="6" pageId="14" pageNumber="13">4</subScript>
, trichobothrium 
<emphasis id="61F5EAAFFFF0E168EFF0F8B5FC11A254" box="[981,1002,1855,1880]" italics="true" pageId="14" pageNumber="13">
e 
<subScript id="CF0534F8FFF0E168EFC4F8C3FC11A254" attach="left" box="[993,1002,1865,1880]" fontSize="6" pageId="14" pageNumber="13">1</subScript>
</emphasis>
is situated between 
<emphasis id="61F5EAAFFFF0E168E8C6F8B5FB0BA25A" box="[1251,1264,1855,1878]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<subScript id="CF0534F8FFF0E168E8D4F8C3FB01A254" attach="left" box="[1265,1274,1865,1880]" fontSize="6" pageId="14" pageNumber="13">2</subScript>
and 
<emphasis id="61F5EAAFFFF0E168E91BF8B5FAB0A25A" box="[1342,1355,1855,1878]" italics="true" pageId="14" pageNumber="13">d</emphasis>
<quantity id="94799B58FFF0E168E96EF8C3FA83A25A" box="[1355,1400,1855,1880]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="14" pageNumber="13" unit="in" value="3.0">
<subScript id="CF0534F8FFF0E168E96EF8C3FAAFA254" attach="left" box="[1355,1364,1865,1880]" fontSize="6" pageId="14" pageNumber="13">3</subScript>
in
</quantity>
<taxonomicName id="94814D3EFFF0E168E9A3F8B5FC6DA279" baseAuthorityName="Lourenco &amp; Leguin" baseAuthorityYear="2011" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="aristidis">
<emphasis id="61F5EAAFFFF0E168E9A3F8B5FC6DA279" italics="true" pageId="14" pageNumber="13">O. aristidis</emphasis>
</taxonomicName>
(fig. 
<quantity id="94799B58FFF0E168EFF2F8D4FBF6A279" box="[983,1037,1886,1909]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" pageId="14" pageNumber="13" unit="in" value="27.0">27 in</quantity>
Lourenço &amp; Leguin, 2011: 14); 
<emphasis id="61F5EAAFFFF0E168E9A0F8D4FA69A279" bold="true" box="[1413,1426,1886,1909]" pageId="14" pageNumber="13">5</emphasis>
) pectinal tooth count 
<quantity id="94799B58FFEEE176EDB9F977FE01A218" box="[412,506,1789,1812]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.08" metricValueMin="4.572" pageId="16" pageNumber="15" unit="in" value="19.0" valueMax="20.0" valueMin="18.0">18–20 in</quantity>
males, 
<quantity id="94799B58FFEEE176EE69F977FD50A218" box="[588,683,1789,1812]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.318" metricValueMin="4.064" pageId="16" pageNumber="15" unit="in" value="16.5" valueMax="17.0" valueMin="16.0">16–17 in</quantity>
females of 
<taxonomicName id="94814D3EFFEEE176ECE5F896FEF4A23F" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[192,271,1820,1843]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="afar" status="sp. nov.">
<emphasis id="61F5EAAFFFEEE176ECE5F896FEF4A23F" box="[192,271,1820,1843]" italics="true" pageId="16" pageNumber="15">O. afar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AC657D4FFEEE176ED3CF896FEADA23F" box="[281,342,1820,1843]" pageId="16" pageNumber="15" rank="species">
<emphasis id="61F5EAAFFFEEE176ED3CF896FEC9A23F" bold="true" box="[281,306,1820,1843]" pageId="16" pageNumber="15">sp</emphasis>
. 
<emphasis id="61F5EAAFFFEEE176ED67F896FEAAA23F" bold="true" box="[322,337,1820,1843]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
, vs. 
<quantity id="94799B58FFEEE176EDABF896FE14A23F" box="[398,495,1820,1843]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="16" pageNumber="15" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16–18 in</quantity>
males, 
<quantity id="94799B58FFEEE176EE63F896FD5CA23F" box="[582,679,1820,1843]" metricMagnitude="-1" metricUnit="m" metricValue="3.683" metricValueMax="4.064" metricValueMin="3.302" pageId="16" pageNumber="15" unit="in" value="14.5" valueMax="16.0" valueMin="13.0">13–16 in</quantity>
females of 
<taxonomicName id="94814D3EFFEEE176ECE5F8B1FEC3A25E" baseAuthorityName="Lourenco &amp; Leguin" baseAuthorityYear="2011" box="[192,312,1851,1874]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="aristidis">
<emphasis id="61F5EAAFFFEEE176ECE5F8B1FEC3A25E" box="[192,312,1851,1874]" italics="true" pageId="16" pageNumber="15">O. aristidis</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="07FE6635FFF1E169ECE5F884FABBA231" pageId="15" pageNumber="14" startId="15.[192,278,1806,1829]" targetBox="[226,1430,223,1764]" targetPageId="15">
<paragraph id="533E36BDFFF1E169ECE5F884FABBA231" blockId="15.[192,1440,1806,1853]" pageId="15" pageNumber="14">
<emphasis id="61F5EAAFFFF1E169ECE5F884FE91A229" bold="true" box="[192,362,1806,1829]" pageId="15" pageNumber="14">Figures 66–71:</emphasis>
<taxonomicName id="94814D3EFFF1E169ED56F885FDE2A228" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[371,537,1807,1828]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="afar" status="sp. nov.">
<emphasis id="61F5EAAFFFF1E169ED56F885FDE2A228" box="[371,537,1807,1828]" italics="true" pageId="15" pageNumber="14">Orthochirus afar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AC657D4FFF1E169EE04F885FDA2A228" box="[545,601,1807,1828]" pageId="15" pageNumber="14" rank="species">
<emphasis id="61F5EAAFFFF1E169EE04F885FDC3A228" bold="true" box="[545,568,1807,1828]" pageId="15" pageNumber="14">sp</emphasis>
. 
<emphasis id="61F5EAAFFFF1E169EE63F885FDA8A228" bold="true" box="[582,595,1807,1828]" pageId="15" pageNumber="14">n</emphasis>
.
</taxonomicNameLabel>
, metasoma and telson. 
<emphasis id="61F5EAAFFFF1E169EF60F885FC2CA228" bold="true" box="[837,983,1807,1828]" pageId="15" pageNumber="14">Figures 66–68</emphasis>
. Male holotype, lateral (66), ventral (67), and dorsal (68) views. 
<emphasis id="61F5EAAFFFF1E169ED56F8A2FDF9A231" bold="true" box="[371,514,1832,1853]" pageId="15" pageNumber="14">Figures 69–71</emphasis>
. Female paratype, lateral (69), ventral (70), and dorsal (71) views. Scale bars: 10 mm.
</paragraph>
</caption>
<caption id="07FE6635FFEEE176ECE5F924FEC6A3D2" pageId="16" pageNumber="15" startId="16.[192,278,1710,1733]" targetBox="[191,1440,189,1681]" targetPageId="16">
<paragraph id="533E36BDFFEEE176ECE5F924FEC6A3D2" blockId="16.[192,1440,1710,1758]" pageId="16" pageNumber="15">
<emphasis id="61F5EAAFFFEEE176ECE5F924FE90A3C9" bold="true" box="[192,363,1710,1733]" pageId="16" pageNumber="15">Figures 72–73:</emphasis>
<taxonomicName id="94814D3EFFEEE176ED51F93AFDE2A3C9" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[372,537,1712,1733]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="afar" status="sp. nov.">
<emphasis id="61F5EAAFFFEEE176ED51F93AFDE2A3C9" box="[372,537,1712,1733]" italics="true" pageId="16" pageNumber="15">Orthochirus afar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AC657D4FFEEE176EE07F93AFDA1A3C9" box="[546,602,1712,1733]" pageId="16" pageNumber="15" rank="species">
<emphasis id="61F5EAAFFFEEE176EE07F93AFDC2A3C9" bold="true" box="[546,569,1712,1733]" pageId="16" pageNumber="15">sp</emphasis>
. 
<emphasis id="61F5EAAFFFEEE176EE6DF93AFDAEA3C9" bold="true" box="[584,597,1712,1733]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
, female juvenile paratype in vivo habitus in Ethiopia at locality 12EM (72) and the locality (73).
</paragraph>
</caption>
<paragraph id="533E36BDFFEEE175EF1BF977FF07A1FA" blockId="16.[830,1440,1789,1874]" lastBlockId="19.[192,802,940,1270]" lastPageId="19" lastPageNumber="18" pageId="16" pageNumber="15">
COMMENTS ON LOCALITIES AND LIFE STRATEGY. The first author visited the 
<typeStatus id="8C3A881FFFEEE176E813F896FB9FA23F" box="[1078,1124,1820,1843]" pageId="16" pageNumber="15">type</typeStatus>
locality 
<emphasis id="61F5EAAFFFEEE176E8E0F896FAFCA23F" bold="true" box="[1221,1287,1820,1843]" pageId="16" pageNumber="15">12EO</emphasis>
(
<figureCitation id="CBBA2A38FFEEE176E932F896FA62A23F" box="[1303,1433,1820,1843]" captionStart="Figures 76–77" captionStartId="18.[192,278,1857,1880]" captionTargetBox="[191,1440,236,1825]" captionTargetId="figure-27@18.[192,1439,237,1824]" captionTargetPageId="18" captionText="Figures 76–77: Orthochirus afar sp. n., the type locality." pageId="16" pageNumber="15">Figs. 76–77</figureCitation>
) on 
<date id="273F107DFFEEE176EF45F8B1FBA6A25E" box="[864,1117,1851,1874]" pageId="16" pageNumber="15" value="2012-11-23" valueMax="2012-11-24" valueMin="2012-11-23">23–24 November 2012</date>
and collected 
<specimenCount id="4587FD34FFEEE176E8DBF8B1FAA4A25E" box="[1278,1375,1851,1874]" pageId="16" pageNumber="15" type="generic" typeStatus="type">four type</specimenCount>
specimens in a narrow area on the margin of a hillside (
<figureCitation id="CBBA2A38FFEDE175EED2FC26FF27A6EE" captionStart="Figures 76–77" captionStartId="18.[192,278,1857,1880]" captionTargetBox="[191,1440,236,1825]" captionTargetId="figure-27@18.[192,1439,237,1824]" captionTargetPageId="18" captionText="Figures 76–77: Orthochirus afar sp. n., the type locality." pageId="19" pageNumber="18">Fig. 76</figureCitation>
) by UV detection at night. In addition to 
<taxonomicName id="94814D3EFFEDE175EE8FFC41FD02A6EE" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2016" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2016" box="[682,761,971,994]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="afar" status="sp. nov.">
<emphasis id="61F5EAAFFFEDE175EE8FFC41FD02A6EE" box="[682,761,971,994]" italics="true" pageId="19" pageNumber="18">O. afar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AC657D4FFEDE175EF27FC41FF2DA10D" pageId="19" pageNumber="18" rank="species">
<emphasis id="61F5EAAFFFEDE175EF27FC41FCE0A6EE" bold="true" box="[770,795,971,994]" pageId="19" pageNumber="18">sp</emphasis>
. 
<emphasis id="61F5EAAFFFEDE175ECE5FC60FF34A10D" bold="true" box="[192,207,1002,1025]" pageId="19" pageNumber="18">n</emphasis>
.
</taxonomicNameLabel>
which was very rare there, the first author also recorded at this locality 
<taxonomicName id="94814D3EFFEDE175ED8AFB82FCE5A113" authority="Kovarik, 2011" authorityName="Kovarik" authorityYear="2011" box="[431,798,1032,1055]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="awashensis">
<emphasis id="61F5EAAFFFEDE175ED8AFB82FD81A113" box="[431,634,1032,1055]" italics="true" pageId="19" pageNumber="18">Buthus awashensis</emphasis>
<bibRefCitation id="37104B4CFFEDE175EEA5FB82FCE5A113" author="KOVARIK" box="[640,798,1032,1055]" pageId="19" pageNumber="18" pagination="1 - 9" refId="ref13305" refString="KOVARIK, F. &amp; T. MAZUCH. 2011. Hemiscorpius novaki sp. n. from Somaliland (Scorpiones: Hemiscorpiidae). Euscorpius, 126: 1 - 9." type="journal article" year="2011">Kovařík, 2011</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="94814D3EFFEDE175ECE5FBADFD67A132" authority="(Birula, 1903)" baseAuthorityName="Birula" baseAuthorityYear="1903" box="[192,668,1063,1086]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="abyssinicus">
<emphasis id="61F5EAAFFFEDE175ECE5FBADFE12A132" box="[192,489,1063,1086]" italics="true" pageId="19" pageNumber="18">Compsobuthus abyssinicus</emphasis>
(Birula, 1903)
</taxonomicName>
, 
<taxonomicName id="94814D3EFFEDE175EE95FBADFDA7A151" authority="Kovarik et Lowe, 2012" authorityName="Kovarik et Lowe" authorityYear="2012" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="awashensis">
<emphasis id="61F5EAAFFFEDE175EE95FBADFEC1A151" italics="true" pageId="19" pageNumber="18">Neobuthus awashensis</emphasis>
Kovařík et Lowe, 2012
</taxonomicName>
, and 
<taxonomicName id="94814D3EFFEDE175EE80FBCCFE1AA177" authority="Pocock, 1901" authorityName="Pocock" authorityYear="1901" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="abyssinicus">
<emphasis id="61F5EAAFFFEDE175EE80FBCCFEC0A177" italics="true" pageId="19" pageNumber="18">Parabuthus abyssinicus</emphasis>
Pocock, 1901
</taxonomicName>
. The first author visited the second locality 
<emphasis id="61F5EAAFFFEDE175ED4BFB09FE4FA196" bold="true" box="[366,436,1155,1178]" pageId="19" pageNumber="18">12EM</emphasis>
(
<figureCitation id="CBBA2A38FFEDE175EDE3FB09FDECA196" box="[454,535,1155,1178]" captionStart="Figures 72–73" captionStartId="16.[192,278,1710,1733]" captionTargetBox="[191,1440,189,1681]" captionTargetId="figure-69@16.[192,1439,190,1680]" captionTargetPageId="16" captionText="Figures 72–73: Orthochirus afar sp. n., female juvenile paratype in vivo habitus in Ethiopia at locality 12EM (72) and the locality (73)." pageId="19" pageNumber="18">Fig. 73</figureCitation>
) on 
<date id="273F107DFFEDE175EE68FB09FCDAA196" box="[589,801,1155,1178]" pageId="19" pageNumber="18" value="2012-11-20">20 November 2012</date>
shortly after sunset and also recorded at this locality (UV detection) 
<taxonomicName id="94814D3EFFEDE175ED1FFB4AFDF6A1DB" authorityName="Kovarik" authorityYear="2011" box="[314,525,1216,1239]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="awashensis">
<emphasis id="61F5EAAFFFEDE175ED1FFB4AFDF6A1DB" box="[314,525,1216,1239]" italics="true" pageId="19" pageNumber="18">Buthus awashensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="94814D3EFFEDE175EE76FB4AFF0EA1FA" authorityName="Kovarik et Lowe" authorityYear="2012" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="awashensis">
<emphasis id="61F5EAAFFFEDE175EE76FB4AFF0EA1FA" italics="true" pageId="19" pageNumber="18">Neobuthus awashensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>