198 lines
41 KiB
XML
198 lines
41 KiB
XML
<document id="9830AB76AC44689F02BB3EC35E06C902" ID-ISSN="1536-9307" ID-ZooBank="4553AE99-6098-4764-8227-AAD8427739D6" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638251357920" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme & Šťáhlavský, František" docDate="2016" docId="DB2887ABFFEDE172ED6FFA91FE63A279" docLanguage="en" docName="Euscorpius.232.1-38.pdf" docOrigin="Euscorpius 232" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Somalicharmus Kovarik 1998" docType="treatment" docVersion="3" lastPageNumber="19" masterDocId="2711FFD3FFFEE166EC25FF8AFFFBA50C" masterDocTitle="Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part IX. Lanzatus, Orthochirus, and Somalicharmus (Buthidae), with description of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n." masterLastPageNumber="38" masterPageNumber="1" pageNumber="18" updateTime="1699435416033" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="8BE5D33F1984E7A0857DDB3B8B1621C1">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part IX. Lanzatus, Orthochirus, and Somalicharmus (Buthidae), with description of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n.</mods:title>
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<mods:namePart id="D240596DB2D3D8AE4A940F883827E6DD">Kovařík, František</mods:namePart>
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<mods:namePart id="AC087C4ECC1F28500CECAB290BC5293C">Lowe, Graeme</mods:namePart>
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<mods:namePart id="27E0C1BA53CACB68A9FBE90E382E82EA">Šťáhlavský, František</mods:namePart>
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<mods:title id="8AFA61A7B8205D253461B370E352D04C">Euscorpius</mods:title>
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<mods:date id="D402B8925E2DE32E42621A2AFA134A85">2016</mods:date>
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<treatment id="DB2887ABFFEDE172ED6FFA91FE63A279" ID-DOI="http://doi.org/10.5281/zenodo.5739616" ID-Zenodo-Dep="5739616" LSID="urn:lsid:plazi:treatment:DB2887ABFFEDE172ED6FFA91FE63A279" httpUri="http://treatment.plazi.org/id/DB2887ABFFEDE172ED6FFA91FE63A279" lastPageId="20" lastPageNumber="19" pageId="19" pageNumber="18">
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<taxonomicName id="94814D3EFFEDE175ED6FFA91FD63A03F" ID-CoL="7JKT" authority="Kovarik, 1998" authorityName="Kovarik" authorityYear="1998" box="[330,664,1307,1331]" class="Arachnida" family="Buthidae" genus="Somalicharmus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="61F5EAAFFFEDE175ED6FFA91FE0CA03F" bold="true" box="[330,503,1307,1331]" italics="true" pageId="19" pageNumber="18">Somalicharmus</emphasis>
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Kovařík, 1998
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1B9B6536FFEDE175ED1BFAB1FD5FA05E" box="[318,676,1339,1362]" pageId="19" pageNumber="18" type="description">
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<paragraph id="533E36BDFFEDE175ED1BFAB1FD5FA05E" blockId="19.[318,676,1307,1362]" box="[318,676,1339,1362]" pageId="19" pageNumber="18">
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(
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<figureCitation id="CBBA2A38FFEDE175ED63FAB1FE2FA05E" box="[326,468,1339,1362]" captionStart-0="Figures 78–81" captionStart-1="Figures 82–85" captionStart-2="Figures 86–89" captionStart-3="Figures 90–92" captionStart-4="Figures 93–109" captionStart-5="Figures 110–112" captionStart-6="Figures 113–120" captionStart-7="Figures 121–122" captionStartId-0="20.[192,278,1642,1665]" captionStartId-1="21.[192,278,1419,1442]" captionStartId-2="22.[192,278,1404,1427]" captionStartId-3="23.[192,278,1838,1861]" captionStartId-4="24.[192,278,1289,1312]" captionStartId-5="25.[192,278,1730,1753]" captionStartId-6="26.[192,278,1643,1666]" captionStartId-7="27.[192,278,1359,1382]" captionTargetBox-0="[208,1429,175,1600]" captionTargetBox-1="[199,1430,192,1380]" captionTargetBox-2="[204,1437,192,1359]" captionTargetBox-3="[191,1440,191,1808]" captionTargetBox-4="[217,1433,194,1253]" captionTargetBox-5="[191,1440,191,1701]" captionTargetBox-6="[191,1440,191,1614]" captionTargetBox-7="[191,1437,191,1329]" captionTargetId-0="figure-17@20.[192,1439,168,1612]" captionTargetId-1="figure-6@21.[192,1437,168,1389]" captionTargetId-2="figure-16@22.[192,1437,168,1374]" captionTargetId-3="figure-35@23.[192,1439,192,1807]" captionTargetId-4="figure-16@24.[192,1437,192,1260]" captionTargetId-5="figure-41@25.[192,1439,192,1700]" captionTargetId-6="figure-118@26.[192,1439,192,1613]" captionTargetId-7="figure-167@27.[192,1437,192,1329]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionTargetPageId-3="23" captionTargetPageId-4="24" captionTargetPageId-5="25" captionTargetPageId-6="26" captionTargetPageId-7="27" captionText-0="Figures 78–81: Somalicharmus whitmanae. Figures 78–79. Male from locality 16EA in dorsal (78) and ventral (79) views. Figures 80–81. Female from locality 16EF in dorsal (80) and ventral (81) views. Scale bar: 10 mm." captionText-1="Figures 82–85: Somalicharmus whitmanae, male from locality 16EA. Figure 82. Metasomal segments IV–V and telson lateral. Figures 83–85. Metasoma and telson lateral (83), ventral (84), and dorsal (85) views. Scale bar: 5 mm (83–85)." captionText-2="Figures 86–89: Somalicharmus whitmanae, female from locality 16EF. Figure 86. Metasomal segments IV–V and telson lateral. Figures 87–89. Metasoma and telson lateral (87), ventral (88), and dorsal (89) views. Scale bar: 5 mm (87–89)." captionText-3="Figures 90–92: Somalicharmus whitmanae, posterior metasoma and telson of male from locality 16EA. Dorsal (90), lateral (91) and ventral (92) views. UV fluorescence images. Scale bar: 2 mm." captionText-4="Figures 93–109: Somalicharmus whitmanae, pedipalp segments of male from locality 16EA (93–101) and female from locality 16EF (102–109). Pedipalp chela dorsal (93, 102), external (94, 103) and ventral (95, 104), patella dorsal (96, 105), external (97, 106) and ventral (98, 107), trochanter and femur internal/internodorsal (99, 108) and dorsal (100, 109), and movable finger dentition (101). Trichobothrial pattern indicated in Figures 93–97 and 99–100." captionText-5="Figures 110–112: Somalicharmus whitmanae, male from locality 16EA. Right pedipalp chela, external (110) and ventrointernal (111) views, and left pectine, ventral view (112). UV fluorescence images. Scale bars: 1 mm (110–111) and 500 µm (112)." captionText-6="Figures 113–120: Somalicharmus whitmanae. Figures 113, 115, 117–120. Male from locality 16EA, carapace and tergites I– III (113), coxosternal area and sternites III–V (115) and left legs I–IV, retrolateral aspect (117–120). Figures 114, 116. Female from locality 16EF, carapace and tergites I–III (114), and coxosternal area and sternites III–V (116)." captionText-7="Figures 121–122: Somalicharmus whitmanae, male from locality 16EA. Figure 121. Dorsal view of body showing carapace, tergites, pedipalp trochanter and femur. Figure 122. Ventral view of body showing coxosternal area, pectines and sternites. Composite white light/ UV fluorescence images. Scale bar: 1 mm." pageId="19" pageNumber="18">Figs. 78–122</figureCitation>
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,
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<figureCitation id="CBBA2A38FFEDE175EDFAFAB1FDC4A05E" box="[479,575,1339,1362]" captionStart-0="Figures 123–128" captionStart-1="Figures 129–133" captionStart-2="Figures 149–150" captionStart-3="Figures 151–152" captionStart-4="Figures 153–154" captionStart-5="Figure 155" captionStartId-0="28.[192,278,1758,1781]" captionStartId-1="29.[192,278,1270,1293]" captionStartId-2="32.[192,278,1821,1844]" captionStartId-3="33.[192,278,1820,1843]" captionStartId-4="34.[192,278,1840,1863]" captionStartId-5="35.[192,268,1422,1445]" captionTargetBox-0="[209,1417,187,1729]" captionTargetBox-1="[199,1428,177,1225]" captionTargetBox-2="[191,1440,215,1791]" captionTargetBox-3="[191,1440,191,1790]" captionTargetBox-4="[191,1440,191,1811]" captionTargetBox-5="[196,1434,196,1389]" captionTargetId-0="figure-115@28.[210,1416,188,1728]" captionTargetId-1="figure-6@29.[192,1437,168,1241]" captionTargetId-2="figure-24@32.[192,1439,216,1790]" captionTargetId-3="figure-6@33.[192,1439,192,1789]" captionTargetId-4="figure-19@34.[192,1439,192,1810]" captionTargetId-5="figure-260@35.[192,1437,191,1392]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionTargetPageId-2="32" captionTargetPageId-3="33" captionTargetPageId-4="34" captionTargetPageId-5="35" captionText-0="Figures 123–128: Figures 123–126: Diagrams of cheliceral finger dentition. Figures 123–124. Lanzatus somalilandus sp. n., female paratype, dorsal (123) and ventral (124) views, showing typical pattern of buthid dentition. Reconstructed from specimen with worn teeth (Figs. 13–14). Figures 125–125. Somalicharmus whitmanae, male from locality 16EA, dorsal (125) and ventral (126) views, showing pattern of dentition highly aberrant for a buthid. Figures 127–128. Somalicharmus whitmanae, left chelicera of male from locality 16EA, dorsal (127) and ventral (128) views. Composite white light/ UV fluorescence images. Scale bar: 500 µm." captionText-1="Figures 129–133: Somalicharmus whitmanae, right hemispermatophore of male from locality 16EA. Whole hemispermatophore in ventroexternal view (129), and magnified views showing capsule region, flagellum and distal trunk in internal (130), dorsal (131), external (132) and ventral (133) views. Abbreviations: bl, basal lobe; il, internal lobe; ml, median lobe. Scale bars: 1 mm (129) and 500 µm (130–133)." captionText-2="Figures 149–150: Somalicharmus whitmanae in vivo habitus. Male from locality 16EA (149), and female from locality 16EF (150)." captionText-3="Figures 151–152: Somalicharmus whitmanae, female from locality 16EF with 13 newborns (151) and the same female with juveniles after first ecdysis (152)." captionText-4="Figures 153–154: Somalicharmus whitmanae, localities 16EA (153), Ethiopia, Turmi, and 16EF (154), Ethiopia, 20 km SE Konzo." captionText-5="Figure 155: Map showing confirmed distribution of Lanzatus, Orthochirus, and Somalicharmus in Horn of Africa. Most points indicate sites sampled during 2011–2016 expeditions, except for the localities of Lanzatus somalicus in Somalia, and of Orthochirus sp. in Djibouti (the map indicates with arbitrary plotted symbol for two citations: a juvenile cited by Werner (1916: 81) with imprecise coordinates, and of a specimen cited by Lourenço & Leguin (2011: 4, 15, figs. 5, 39); see also comments under Orthochirus in Systematics section." pageId="19" pageNumber="18">125–155</figureCitation>
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,
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<tableCitation id="1E030306FFEDE175EE6FFAB1FD60A05E" box="[586,667,1339,1362]" captionStart="Table 2" captionStartId="31.[192,257,627,650]" captionText="Table 2: The diploid numbers, the relative lengths of chromosomes of the diploid set, postpachytene (Ppt) configuration (after Mattos et al., 2013) and the localities of analyzed Somalicharmus whitmanae from Ethiopia." pageId="19" pageNumber="18">Table 2</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="1B9B6536FFEDE172ECE5FAF2FE5CA21F" lastPageId="20" lastPageNumber="19" pageId="19" pageNumber="18" type="reference_group">
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<paragraph id="533E36BDFFEDE175ECE5FAF2FE03A34B" blockId="19.[192,802,1400,1607]" pageId="19" pageNumber="18">
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<taxonomicName id="94814D3EFFEDE175ECE5FAF2FDB6A083" ID-CoL="7JKT" authority="Kovarik, 1998: 116" authorityName="Kovarik" authorityPageNumber="116" authorityYear="1998" box="[192,589,1400,1423]" class="Arachnida" family="Buthidae" genus="Somalicharmus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="61F5EAAFFFEDE175ECE5FAF2FE9DA083" box="[192,358,1400,1423]" italics="true" pageId="19" pageNumber="18">Somalicharmus</emphasis>
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Kovařík, 1998: 116
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</taxonomicName>
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, 119–122, figs. 9– 15; Fet & Lowe, 2000: 224; Soleglad & Fet, 2003b: 88, Tab. 9; Fet & Soleglad, 2005: 11; Fet et al., 2005: 3, 14, 22,
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<tableCitation id="1E030306FFEDE175ED82FA5EFE17A0E7" box="[423,492,1492,1515]" captionStart="Table 1" captionStartId="19.[192,257,836,859]" captionText="Table 1: Comparative measurements of adults of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="19" pageNumber="18">Tab. 1</tableCitation>
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,
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<figureCitation id="CBBA2A38FFEDE175EDD9FA5EFDB6A0E7" box="[508,589,1492,1515]" captionStart="Figures 15–29" captionStartId="7.[192,278,1567,1590]" captionTargetBox="[192,1439,175,1532]" captionTargetId="figure-6@7.[192,1439,168,1538]" captionTargetPageId="7" captionText="Figures 15–29: Lanzatus. Figures 15–28. L. somalilandus sp. n. Figures 15–27. Female paratype. Figures 15–22. Pedipalp segments, movable finger dentition (15), chela dorsal (16), external (17) and ventral (18), patella dorsal (19), external (20) and ventral (21), trochanter and femur dorsal (22). Trichobothrial pattern indicated in Figures 17–20 and 22. Figures 23–26. Left legs I–IV, retrolateral aspect. Figures 27–29. Telson lateral. Figures 27–28. L. somalilandus sp. n., female paratype (27) and male holotype (28). Figure 29. L. somalicus, male paratype." pageId="19" pageNumber="18">Fig. 23</figureCitation>
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; Prendini & Wheeler, 2005: 481; Dupré, 2007: 10, 13, 16;
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<bibRefCitation id="37104B4CFFEDE175EE8AFA79FE7CA325" author="KOVARIK" pageId="19" pageNumber="18" pagination="201 - 209" refId="ref13435" refString="KOVARIK, F., M. E. SOLEGLAD & V. FET. 2007. A new species of scorpion in the " Charmus " group from India (Scorpiones: Buthidae). Boletin Sociedad Entomologica Aragonesa, 40: 201 - 209." type="journal article" year="2007">Kovařík et al., 2007: 201</bibRefCitation>
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, 205–208, fig. 15;
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<bibRefCitation id="37104B4CFFEDE175EE73F998FCE6A325" author="KOVARIK" box="[598,797,1554,1577]" pageId="19" pageNumber="18" pagination="1 - 12" refId="ref13482" refString="KOVARIK, F., F. STAHLAVSKY, T. KORIN- KOVA, J. KRAL & T. VAN DER ENDE. 2009. Tityus ythieri Lourenco, 2007 is a synonym of Tityus magnimanus Pocock, 1897 (Scorpiones: Buthidae): a combined approach using morphology, hybridization experiments, chromosomes, and mitochondrial DNA. Euscorpius, 77: 1 - 12." type="journal article" year="2009">Kovařík, 2009: 21</bibRefCitation>
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, 31;
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<bibRefCitation id="37104B4CFFEDE175ED3CF9BAFE09A34B" author="MIRZA" box="[281,498,1584,1607]" pageId="19" pageNumber="18" pagination="1 - 11" refId="ref14166" refString="MIRZA, Z. A., R. V. SANAP & K. KUNTE. 2016. A new species of the genus Thaicharmus Kovarik, 1995 (Scorpiones: Buthidae) from northeast India. Euscorpius, 215: 1 - 11." type="journal article" year="2016">Mirza et al., 2016: 1</bibRefCitation>
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.
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</paragraph>
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<paragraph id="533E36BDFFEDE172ECE5F9E4FE5CA21F" blockId="19.[192,802,1645,1916]" lastBlockId="20.[192,802,1727,1811]" lastPageId="20" lastPageNumber="19" pageId="19" pageNumber="18">
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EMENDED DIAGNOSIS. Total length 18.5–
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<quantity id="94799B58FFEDE175EEA8F9E7FD1AA388" box="[653,737,1645,1668]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="18" unit="mm" value="25.0">25 mm</quantity>
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. Pedipalps orthobothriotaxic
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<typeStatus id="8C3A881FFFEDE175EDEBF906FE07A3AF" box="[462,508,1676,1699]" pageId="19" pageNumber="18">type</typeStatus>
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A (Vachon, 1974); dorsal trichobothria of femur arranged in
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<emphasis id="61F5EAAFFFEDE175EE5EF921FD72A3CE" box="[635,649,1707,1730]" italics="true" pageId="19" pageNumber="18">α</emphasis>
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-configuration (Vachon, 1975) (
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<figureCitation id="CBBA2A38FFEDE175ED5BF943FE24A3EC" box="[382,479,1737,1760]" captionStart="Figures 93–109" captionStartId="24.[192,278,1289,1312]" captionTargetBox="[217,1433,194,1253]" captionTargetId="figure-16@24.[192,1437,192,1260]" captionTargetPageId="24" captionText="Figures 93–109: Somalicharmus whitmanae, pedipalp segments of male from locality 16EA (93–101) and female from locality 16EF (102–109). Pedipalp chela dorsal (93, 102), external (94, 103) and ventral (95, 104), patella dorsal (96, 105), external (97, 106) and ventral (98, 107), trochanter and femur internal/internodorsal (99, 108) and dorsal (100, 109), and movable finger dentition (101). Trichobothrial pattern indicated in Figures 93–97 and 99–100." pageId="19" pageNumber="18">Fig. 100</figureCitation>
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); trichobothrium
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<emphasis id="61F5EAAFFFEDE175EE84F943FD55A3EC" box="[673,686,1737,1760]" italics="true" pageId="19" pageNumber="18">d</emphasis>
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<subScript id="CF0534F8FFEDE175EE8BF95EFD4CA3EF" attach="left" box="[686,695,1748,1763]" fontSize="6" pageId="19" pageNumber="18">2</subScript>
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of femur fenced by granules associated with dorsointernal carina, position may appear either dorsal or internal to carina; patella with dorsomedian carina absent, but trichobothrium
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<emphasis id="61F5EAAFFFEDE175ED0FF8CEFECCA257" box="[298,311,1860,1883]" italics="true" pageId="19" pageNumber="18">d</emphasis>
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<subScript id="CF0534F8FFEDE175ED1DF8C4FEBAA251" attach="left" box="[312,321,1870,1885]" fontSize="6" pageId="19" pageNumber="18">3</subScript>
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located external to
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<emphasis id="61F5EAAFFFEDE175EE3EF8CEFDD3A257" box="[539,552,1860,1883]" italics="true" pageId="19" pageNumber="18">d</emphasis>
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<subScript id="CF0534F8FFEDE175EE0DF8C4FDCAA251" attach="left" box="[552,561,1870,1885]" fontSize="6" pageId="19" pageNumber="18">4</subScript>
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; chela manus with
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<emphasis id="61F5EAAFFFEDE175EF2CF8CEFCE2A257" box="[777,793,1860,1883]" italics="true" pageId="19" pageNumber="18">V</emphasis>
|
||
<subScript id="CF0534F8FFEDE175EF3CF8C4FCD9A251" attach="left" box="[793,802,1870,1885]" fontSize="6" pageId="19" pageNumber="18">2</subScript>
|
||
displaced externally relative to
|
||
<emphasis id="61F5EAAFFFEDE175EE2AF8E9FDE4A276" box="[527,543,1891,1914]" italics="true" pageId="19" pageNumber="18">V</emphasis>
|
||
<subScript id="CF0534F8FFEDE175EE05F8E7FDD2A270" attach="right" box="[544,553,1901,1916]" fontSize="6" pageId="19" pageNumber="18">1</subScript>
|
||
;
|
||
<emphasis id="61F5EAAFFFEDE175EE13F8E9FDAAA276" box="[566,593,1891,1914]" italics="true" pageId="19" pageNumber="18">db</emphasis>
|
||
on distal manus,
|
||
<emphasis id="61F5EAAFFFEDE175EF28F8E9FCD9A276" box="[781,802,1891,1914]" italics="true" pageId="19" pageNumber="18">dt</emphasis>
|
||
on base of fixed finger (
|
||
<figureCitation id="CBBA2A38FFEDE175E87EFC26FB4BA6CF" box="[1115,1200,940,963]" captionStart="Figures 93–109" captionStartId="24.[192,278,1289,1312]" captionTargetBox="[217,1433,194,1253]" captionTargetId="figure-16@24.[192,1437,192,1260]" captionTargetPageId="24" captionText="Figures 93–109: Somalicharmus whitmanae, pedipalp segments of male from locality 16EA (93–101) and female from locality 16EF (102–109). Pedipalp chela dorsal (93, 102), external (94, 103) and ventral (95, 104), patella dorsal (96, 105), external (97, 106) and ventral (98, 107), trochanter and femur internal/internodorsal (99, 108) and dorsal (100, 109), and movable finger dentition (101). Trichobothrial pattern indicated in Figures 93–97 and 99–100." pageId="19" pageNumber="18">Fig. 93</figureCitation>
|
||
);
|
||
<emphasis id="61F5EAAFFFEDE175E8ECFC26FB23A6CF" box="[1225,1240,940,963]" italics="true" pageId="19" pageNumber="18">it</emphasis>
|
||
on base of fixed finger (
|
||
<figureCitation id="CBBA2A38FFEDE175EFABFC41FC12A6EE" box="[910,1001,971,994]" captionStart="Figures 93–109" captionStartId="24.[192,278,1289,1312]" captionTargetBox="[217,1433,194,1253]" captionTargetId="figure-16@24.[192,1437,192,1260]" captionTargetPageId="24" captionText="Figures 93–109: Somalicharmus whitmanae, pedipalp segments of male from locality 16EA (93–101) and female from locality 16EF (102–109). Pedipalp chela dorsal (93, 102), external (94, 103) and ventral (95, 104), patella dorsal (96, 105), external (97, 106) and ventral (98, 107), trochanter and femur internal/internodorsal (99, 108) and dorsal (100, 109), and movable finger dentition (101). Trichobothrial pattern indicated in Figures 93–97 and 99–100." pageId="19" pageNumber="18">Figs. 95</figureCitation>
|
||
,
|
||
<figureCitation id="CBBA2A38FFEDE175EFD3FC41FBDAA6EE" box="[1014,1057,971,994]" captionStart="Figures 110–112" captionStartId="25.[192,278,1730,1753]" captionTargetBox="[191,1440,191,1701]" captionTargetId="figure-41@25.[192,1439,192,1700]" captionTargetPageId="25" captionText="Figures 110–112: Somalicharmus whitmanae, male from locality 16EA. Right pedipalp chela, external (110) and ventrointernal (111) views, and left pectine, ventral view (112). UV fluorescence images. Scale bars: 1 mm (110–111) and 500 µm (112)." pageId="19" pageNumber="18">111</figureCitation>
|
||
). Pectines with fulcra, hirsute (
|
||
<figureCitation id="CBBA2A38FFEDE175E950FC41FC91A10D" captionStart="Figures 110–112" captionStartId="25.[192,278,1730,1753]" captionTargetBox="[191,1440,191,1701]" captionTargetId="figure-41@25.[192,1439,192,1700]" captionTargetPageId="25" captionText="Figures 110–112: Somalicharmus whitmanae, male from locality 16EA. Right pedipalp chela, external (110) and ventrointernal (111) views, and left pectine, ventral view (112). UV fluorescence images. Scale bars: 1 mm (110–111) and 500 µm (112)." pageId="19" pageNumber="18">Fig. 112</figureCitation>
|
||
). Pectine teeth number 11–14. Basal middle lamella of pectines extremely dilated and rounded in females (
|
||
<figureCitation id="CBBA2A38FFEDE175EF63FBADFC55A132" box="[838,942,1063,1086]" captionStart="Figures 113–120" captionStartId="26.[192,278,1643,1666]" captionTargetBox="[191,1440,191,1614]" captionTargetId="figure-118@26.[192,1439,192,1613]" captionTargetPageId="26" captionText="Figures 113–120: Somalicharmus whitmanae. Figures 113, 115, 117–120. Male from locality 16EA, carapace and tergites I– III (113), coxosternal area and sternites III–V (115) and left legs I–IV, retrolateral aspect (117–120). Figures 114, 116. Female from locality 16EF, carapace and tergites I–III (114), and coxosternal area and sternites III–V (116)." pageId="19" pageNumber="18">Figs. 116</figureCitation>
|
||
vs. 115). Males with genital papillae. Hemispermatophore flagelliform with lobes of capsule in 2+1 configuration, basal lobe enlarged, elongate, median lobe not fused to flagellum. Sternum subpentagonal (
|
||
<figureCitation id="CBBA2A38FFEDE175EF63FB28FC1CA1B5" box="[838,999,1186,1209]" captionStart="Figures 113–120" captionStartId="26.[192,278,1643,1666]" captionTargetBox="[191,1440,191,1614]" captionTargetId="figure-118@26.[192,1439,192,1613]" captionTargetPageId="26" captionText="Figures 113–120: Somalicharmus whitmanae. Figures 113, 115, 117–120. Male from locality 16EA, carapace and tergites I– III (113), coxosternal area and sternites III–V (115) and left legs I–IV, retrolateral aspect (117–120). Figures 114, 116. Female from locality 16EF, carapace and tergites I–III (114), and coxosternal area and sternites III–V (116)." pageId="19" pageNumber="18">Figs. 115, 116</figureCitation>
|
||
,
|
||
<figureCitation id="CBBA2A38FFEDE175EFD6FB28FBE4A1B5" box="[1011,1055,1186,1209]" captionStart="Figures 121–122" captionStartId="27.[192,278,1359,1382]" captionTargetBox="[191,1437,191,1329]" captionTargetId="figure-167@27.[192,1437,192,1329]" captionTargetPageId="27" captionText="Figures 121–122: Somalicharmus whitmanae, male from locality 16EA. Figure 121. Dorsal view of body showing carapace, tergites, pedipalp trochanter and femur. Figure 122. Ventral view of body showing coxosternal area, pectines and sternites. Composite white light/ UV fluorescence images. Scale bar: 1 mm." pageId="19" pageNumber="18">122</figureCitation>
|
||
). Long tibial spurs present on third and fourth legs (
|
||
<figureCitation id="CBBA2A38FFEDE175EFD7FB4AFB6AA1DB" box="[1010,1169,1216,1239]" captionStart="Figures 113–120" captionStartId="26.[192,278,1643,1666]" captionTargetBox="[191,1440,191,1614]" captionTargetId="figure-118@26.[192,1439,192,1613]" captionTargetPageId="26" captionText="Figures 113–120: Somalicharmus whitmanae. Figures 113, 115, 117–120. Male from locality 16EA, carapace and tergites I– III (113), coxosternal area and sternites III–V (115) and left legs I–IV, retrolateral aspect (117–120). Figures 114, 116. Female from locality 16EF, carapace and tergites I–III (114), and coxosternal area and sternites III–V (116)." pageId="19" pageNumber="18">Figs. 119–120</figureCitation>
|
||
). Chelicera with reduced dentition, movable finger ventral margin smooth, fixed finger margin with 2 non-distal denticles, lacking bicusp, ventral surface of fixed finger without denticles (
|
||
<figureCitation id="CBBA2A38FFEDE175E94FFA96FC5BA05E" captionStart="Figures 123–128" captionStartId="28.[192,278,1758,1781]" captionTargetBox="[209,1417,187,1729]" captionTargetId="figure-115@28.[210,1416,188,1728]" captionTargetPageId="28" captionText="Figures 123–128: Figures 123–126: Diagrams of cheliceral finger dentition. Figures 123–124. Lanzatus somalilandus sp. n., female paratype, dorsal (123) and ventral (124) views, showing typical pattern of buthid dentition. Reconstructed from specimen with worn teeth (Figs. 13–14). Figures 125–125. Somalicharmus whitmanae, male from locality 16EA, dorsal (125) and ventral (126) views, showing pattern of dentition highly aberrant for a buthid. Figures 127–128. Somalicharmus whitmanae, left chelicera of male from locality 16EA, dorsal (127) and ventral (128) views. Composite white light/ UV fluorescence images. Scale bar: 500 µm." pageId="19" pageNumber="18">Figs. 125–128</figureCitation>
|
||
). Carapace granulated without carinae (
|
||
<figureCitation id="CBBA2A38FFEDE175E94FFAB1FC65A07D" captionStart="Figures 113–120" captionStartId="26.[192,278,1643,1666]" captionTargetBox="[191,1440,191,1614]" captionTargetId="figure-118@26.[192,1439,192,1613]" captionTargetPageId="26" captionText="Figures 113–120: Somalicharmus whitmanae. Figures 113, 115, 117–120. Male from locality 16EA, carapace and tergites I– III (113), coxosternal area and sternites III–V (115) and left legs I–IV, retrolateral aspect (117–120). Figures 114, 116. Female from locality 16EF, carapace and tergites I–III (114), and coxosternal area and sternites III–V (116)." pageId="19" pageNumber="18">Figs. 113–114</figureCitation>
|
||
,
|
||
<figureCitation id="CBBA2A38FFEDE175EF8EFAD0FC2DA07D" box="[939,982,1370,1393]" captionStart="Figures 121–122" captionStartId="27.[192,278,1359,1382]" captionTargetBox="[191,1437,191,1329]" captionTargetId="figure-167@27.[192,1437,192,1329]" captionTargetPageId="27" captionText="Figures 121–122: Somalicharmus whitmanae, male from locality 16EA. Figure 121. Dorsal view of body showing carapace, tergites, pedipalp trochanter and femur. Figure 122. Ventral view of body showing coxosternal area, pectines and sternites. Composite white light/ UV fluorescence images. Scale bar: 1 mm." pageId="19" pageNumber="18">121</figureCitation>
|
||
), in lateral view with entire dorsal surface horizontal or nearly so. Lateral eyes number 2 pairs (
|
||
<typeStatus id="8C3A881FFFEDE175EF63FA1DFC8DA0A2" box="[838,886,1431,1454]" pageId="19" pageNumber="18">type</typeStatus>
|
||
2A layout,
|
||
<bibRefCitation id="37104B4CFFEDE175E82BFA1DFADCA0A2" author="LORIA & PRENDINI" box="[1038,1319,1431,1454]" pageId="19" pageNumber="18" pagination="112913" refId="ref13807" refString="LORIA, S. & L. PRENDINI. 2014. Homology of the lateral eyes of scorpiones: a six-ocellus model. PLoS ONE 9 (12): e 112913. doi: 10. 1371 / journal. pone. 0112913." type="journal article" year="2014">Loria & Prendini, 2014</bibRefCitation>
|
||
). Movable finger of pedipalp with 8–9 rows and fixed finger with 7–8 rows of granules forming imbricated diagonal rows, with external and internal accessory granules (
|
||
<figureCitation id="CBBA2A38FFEDE175E911FA79FA60A306" box="[1332,1435,1523,1546]" captionStart="Figures 93–109" captionStartId="24.[192,278,1289,1312]" captionTargetBox="[217,1433,194,1253]" captionTargetId="figure-16@24.[192,1437,192,1260]" captionTargetPageId="24" captionText="Figures 93–109: Somalicharmus whitmanae, pedipalp segments of male from locality 16EA (93–101) and female from locality 16EF (102–109). Pedipalp chela dorsal (93, 102), external (94, 103) and ventral (95, 104), patella dorsal (96, 105), external (97, 106) and ventral (98, 107), trochanter and femur internal/internodorsal (99, 108) and dorsal (100, 109), and movable finger dentition (101). Trichobothrial pattern indicated in Figures 93–97 and 99–100." pageId="19" pageNumber="18">Figs. 101</figureCitation>
|
||
,
|
||
<figureCitation id="CBBA2A38FFEDE175EF1BF998FC5BA325" box="[830,928,1554,1577]" captionStart="Figures 110–112" captionStartId="25.[192,278,1730,1753]" captionTargetBox="[191,1440,191,1701]" captionTargetId="figure-41@25.[192,1439,192,1700]" captionTargetPageId="25" captionText="Figures 110–112: Somalicharmus whitmanae, male from locality 16EA. Right pedipalp chela, external (110) and ventrointernal (111) views, and left pectine, ventral view (112). UV fluorescence images. Scale bars: 1 mm (110–111) and 500 µm (112)." pageId="19" pageNumber="18">110–111</figureCitation>
|
||
). Tergites densely granulate in males (
|
||
<figureCitation id="CBBA2A38FFEDE175E91AF998FA60A325" box="[1343,1435,1554,1577]" captionStart="Figures 113–120" captionStartId="26.[192,278,1643,1666]" captionTargetBox="[191,1440,191,1614]" captionTargetId="figure-118@26.[192,1439,192,1613]" captionTargetPageId="26" captionText="Figures 113–120: Somalicharmus whitmanae. Figures 113, 115, 117–120. Male from locality 16EA, carapace and tergites I– III (113), coxosternal area and sternites III–V (115) and left legs I–IV, retrolateral aspect (117–120). Figures 114, 116. Female from locality 16EF, carapace and tergites I–III (114), and coxosternal area and sternites III–V (116)." pageId="19" pageNumber="18">Fig. 113</figureCitation>
|
||
,
|
||
<figureCitation id="CBBA2A38FFEDE175EF1BF9BAFC93A34B" box="[830,872,1584,1607]" captionStart="Figures 121–122" captionStartId="27.[192,278,1359,1382]" captionTargetBox="[191,1437,191,1329]" captionTargetId="figure-167@27.[192,1437,192,1329]" captionTargetPageId="27" captionText="Figures 121–122: Somalicharmus whitmanae, male from locality 16EA. Figure 121. Dorsal view of body showing carapace, tergites, pedipalp trochanter and femur. Figure 122. Ventral view of body showing coxosternal area, pectines and sternites. Composite white light/ UV fluorescence images. Scale bar: 1 mm." pageId="19" pageNumber="18">121</figureCitation>
|
||
), more so than in females (
|
||
<figureCitation id="CBBA2A38FFEDE175E8B6F9BAFB09A34B" box="[1171,1266,1584,1607]" captionStart="Figures 113–120" captionStartId="26.[192,278,1643,1666]" captionTargetBox="[191,1440,191,1614]" captionTargetId="figure-118@26.[192,1439,192,1613]" captionTargetPageId="26" captionText="Figures 113–120: Somalicharmus whitmanae. Figures 113, 115, 117–120. Male from locality 16EA, carapace and tergites I– III (113), coxosternal area and sternites III–V (115) and left legs I–IV, retrolateral aspect (117–120). Figures 114, 116. Female from locality 16EF, carapace and tergites I–III (114), and coxosternal area and sternites III–V (116)." pageId="19" pageNumber="18">Fig. 114</figureCitation>
|
||
); carinae absent or barely indicated by series of more closely spaced granules. Stigmata small, subovoid or hemielliptic slits. Metasoma stout, segments I with 10 carinae, II–III with 8 carinae; segments I–III with ventrosubmedian and ventrolateral carinae strong, the latter posteriorly converging and connecting to form a U-shaped margin carina (
|
||
<figureCitation id="CBBA2A38FFEDE175EFAAF88DFC11A212" box="[911,1002,1799,1822]" captionStart="Figures 82–85" captionStartId="21.[192,278,1419,1442]" captionTargetBox="[199,1430,192,1380]" captionTargetId="figure-6@21.[192,1437,168,1389]" captionTargetPageId="21" captionText="Figures 82–85: Somalicharmus whitmanae, male from locality 16EA. Figure 82. Metasomal segments IV–V and telson lateral. Figures 83–85. Metasoma and telson lateral (83), ventral (84), and dorsal (85) views. Scale bar: 5 mm (83–85)." pageId="19" pageNumber="18">Figs. 84</figureCitation>
|
||
,
|
||
<figureCitation id="CBBA2A38FFEDE175EFD3F88DFBEFA212" box="[1014,1044,1799,1822]" captionStart="Figures 86–89" captionStartId="22.[192,278,1404,1427]" captionTargetBox="[204,1437,192,1359]" captionTargetId="figure-16@22.[192,1437,168,1374]" captionTargetPageId="22" captionText="Figures 86–89: Somalicharmus whitmanae, female from locality 16EF. Figure 86. Metasomal segments IV–V and telson lateral. Figures 87–89. Metasoma and telson lateral (87), ventral (88), and dorsal (89) views. Scale bar: 5 mm (87–89)." pageId="19" pageNumber="18">88</figureCitation>
|
||
); ventral surfaces of segments IV–V coarsely granulated without carinae (
|
||
<figureCitation id="CBBA2A38FFEDE175E8F9F8ACFAC2A231" box="[1244,1337,1830,1853]" captionStart="Figures 82–85" captionStartId="21.[192,278,1419,1442]" captionTargetBox="[199,1430,192,1380]" captionTargetId="figure-6@21.[192,1437,168,1389]" captionTargetPageId="21" captionText="Figures 82–85: Somalicharmus whitmanae, male from locality 16EA. Figure 82. Metasomal segments IV–V and telson lateral. Figures 83–85. Metasoma and telson lateral (83), ventral (84), and dorsal (85) views. Scale bar: 5 mm (83–85)." pageId="19" pageNumber="18">Figs. 84</figureCitation>
|
||
,
|
||
<figureCitation id="CBBA2A38FFEDE175E96CF8ACFA9DA231" box="[1353,1382,1830,1853]" captionStart="Figures 86–89" captionStartId="22.[192,278,1404,1427]" captionTargetBox="[204,1437,192,1359]" captionTargetId="figure-16@22.[192,1437,168,1374]" captionTargetPageId="22" captionText="Figures 86–89: Somalicharmus whitmanae, female from locality 16EF. Figure 86. Metasomal segments IV–V and telson lateral. Figures 87–89. Metasoma and telson lateral (87), ventral (88), and dorsal (89) views. Scale bar: 5 mm (87–89)." pageId="19" pageNumber="18">88</figureCitation>
|
||
,
|
||
<figureCitation id="CBBA2A38FFEDE175E953F8ACFA6FA231" box="[1398,1428,1830,1853]" captionStart="Figures 90–92" captionStartId="23.[192,278,1838,1861]" captionTargetBox="[191,1440,191,1808]" captionTargetId="figure-35@23.[192,1439,192,1807]" captionTargetPageId="23" captionText="Figures 90–92: Somalicharmus whitmanae, posterior metasoma and telson of male from locality 16EA. Dorsal (90), lateral (91) and ventral (92) views. UV fluorescence images. Scale bar: 2 mm." pageId="19" pageNumber="18">92</figureCitation>
|
||
). Telson strongly swollen, nearly spherical, with a minute subaculear tubercle, its ventral surface densely hirsute with long setae, covered with prominent, pointed, darkbrown to black granules, aculeus shorter than vesicle. (
|
||
<figureCitation id="CBBA2A38FFEAE172ECEDF976FEDAA21F" box="[200,289,1788,1811]" captionStart="Figures 82–85" captionStartId="21.[192,278,1419,1442]" captionTargetBox="[199,1430,192,1380]" captionTargetId="figure-6@21.[192,1437,168,1389]" captionTargetPageId="21" captionText="Figures 82–85: Somalicharmus whitmanae, male from locality 16EA. Figure 82. Metasomal segments IV–V and telson lateral. Figures 83–85. Metasoma and telson lateral (83), ventral (84), and dorsal (85) views. Scale bar: 5 mm (83–85)." pageId="20" pageNumber="19">Figs. 82</figureCitation>
|
||
,
|
||
<figureCitation id="CBBA2A38FFEAE172ED09F976FEB2A21F" box="[300,329,1788,1811]" captionStart="Figures 86–89" captionStartId="22.[192,278,1404,1427]" captionTargetBox="[204,1437,192,1359]" captionTargetId="figure-16@22.[192,1437,168,1374]" captionTargetPageId="22" captionText="Figures 86–89: Somalicharmus whitmanae, female from locality 16EF. Figure 86. Metasomal segments IV–V and telson lateral. Figures 87–89. Metasoma and telson lateral (87), ventral (88), and dorsal (89) views. Scale bar: 5 mm (87–89)." pageId="20" pageNumber="19">86</figureCitation>
|
||
,
|
||
<figureCitation id="CBBA2A38FFEAE172ED71F976FE60A21F" box="[340,411,1788,1811]" captionStart="Figures 90–92" captionStartId="23.[192,278,1838,1861]" captionTargetBox="[191,1440,191,1808]" captionTargetId="figure-35@23.[192,1439,192,1807]" captionTargetPageId="23" captionText="Figures 90–92: Somalicharmus whitmanae, posterior metasoma and telson of male from locality 16EA. Dorsal (90), lateral (91) and ventral (92) views. UV fluorescence images. Scale bar: 2 mm." pageId="20" pageNumber="19">90–92</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="07FE6635FFEAE172ECE5F9E0FB84A395" pageId="20" pageNumber="19" startId="20.[192,278,1642,1665]" targetBox="[208,1429,175,1600]" targetPageId="20">
|
||
<paragraph id="533E36BDFFEAE172ECE5F9E0FB84A395" blockId="20.[192,1441,1642,1689]" pageId="20" pageNumber="19">
|
||
<emphasis id="61F5EAAFFFEAE172ECE5F9E0FE90A38D" bold="true" box="[192,363,1642,1665]" pageId="20" pageNumber="19">Figures 78–81:</emphasis>
|
||
<taxonomicName id="94814D3EFFEAE172ED56F9E1FD81A38C" authorityName="Kovarik" authorityYear="1998" box="[371,634,1643,1664]" class="Arachnida" family="Buthidae" genus="Somalicharmus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="whitmanae">
|
||
<emphasis id="61F5EAAFFFEAE172ED56F9E1FD81A38C" box="[371,634,1643,1664]" italics="true" pageId="20" pageNumber="19">Somalicharmus whitmanae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="61F5EAAFFFEAE172EEACF9E1FCE0A38C" bold="true" box="[649,795,1643,1664]" pageId="20" pageNumber="19">Figures 78–79</emphasis>
|
||
. Male from locality 16EA in dorsal (78) and ventral (79) views.
|
||
<emphasis id="61F5EAAFFFEAE172ECE5F90EFEB4A395" bold="true" box="[192,335,1668,1689]" pageId="20" pageNumber="19">Figures 80–81</emphasis>
|
||
. Female from locality 16EF in dorsal (80) and ventral (81) views. Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="1B9B6536FFEAE172ECE5F8CAFE63A279" pageId="20" pageNumber="19" type="materials_examined">
|
||
<paragraph id="533E36BDFFEAE172ECE5F8CAFE63A279" blockId="20.[192,741,1856,1909]" pageId="20" pageNumber="19">
|
||
<materialsCitation id="E3E93CE0FFEAE172ECE5F8CAFE68A279" collectionCode="SPECIES" pageId="20" pageNumber="19" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="8C3A881FFFEAE172ECE5F8CAFF03A25A" box="[192,248,1856,1879]" pageId="20" pageNumber="19">TYPE</typeStatus>
|
||
<collectionCode id="3590AE78FFEAE172ECDBF8C9FEAAA25A" box="[254,337,1859,1878]" pageId="20" pageNumber="19">SPECIES</collectionCode>
|
||
.
|
||
<taxonomicName id="94814D3EFFEAE172ED7BF8CAFF03A279" authority="Kovarik, 1998" authorityName="Kovarik" authorityYear="1998" class="Arachnida" family="Buthidae" genus="Somalicharmus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="whitmanae">
|
||
<emphasis id="61F5EAAFFFEAE172ED7BF8CAFD7BA25B" box="[350,640,1856,1879]" italics="true" pageId="20" pageNumber="19">Somalicharmus whitmanae</emphasis>
|
||
Kovařík, 1998
|
||
</taxonomicName>
|
||
, by monotypy
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |