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<document id="FF8D71C430BFABF7499C648B0C24E062" ID-DOI="10.5281/zenodo.5176618" ID-ISSN="0374-1036" ID-Zenodo-Dep="5176618" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628601973277" checkinUser="carolina" docAuthor="Hájek, Jiří &amp; Šťastný, Jaroslav" docDate="2005" docId="FE0987CFFFFEE93FFE67E99AFEBE17F0" docLanguage="en" docName="ActaEntMusNatPra.45.51-58.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 45" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Laccophilus boukali Hájek &amp; Šťastný 2005, sp. nov." docType="treatment" docVersion="4" lastPageNumber="53" masterDocId="0230FFB7FFFFE93DFFDCEB58FFFA1347" masterDocTitle="Two new species of Laccophilus Leach, 1815 (Coleoptera: Dytiscidae) from India and Borneo with notes on other species of the genus" masterLastPageNumber="58" masterPageNumber="51" pageNumber="52" updateTime="1699047269640" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="36833482D68727D056528D7371F5306A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="ED034EDBDFFDE8C5E53E61866791BE83">
<mods:title id="70E7602EC1D7F4A189AC56C43621562F">Two new species of Laccophilus Leach, 1815 (Coleoptera: Dytiscidae) from India and Borneo with notes on other species of the genus</mods:title>
</mods:titleInfo>
<mods:name id="EB6EA86A3FDDABEF0665930905641D80" type="personal">
<mods:role id="D621C6404CF8D23D48B8BD4853CFC663">
<mods:roleTerm id="A56485EC9A726F6A171504705F72CA2E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="853CA35CAEFF1F884E4F68F507F7CC2C">Hájek, Jiří</mods:namePart>
<mods:affiliation id="4C1338B5176BC72704065004BF27640D">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: jiri _ hajek @ nm. cz</mods:affiliation>
</mods:name>
<mods:name id="B686F076E401CA1A87CA69D692AFFD77" type="personal">
<mods:role id="9687F8C32F0ED30907FCFEBF37202E1D">
<mods:roleTerm id="DF56238FDA12F26B927AC8DA416DE024">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5F2795AF087F5B9E449B9A72694DF2AA">Šťastný, Jaroslav</mods:namePart>
<mods:affiliation id="8678E51A201A4F5AC1103390C4557EBF">Kosmonautů 359, CZ- 460 05 Liberec, Czech Republic; e-mail: stastnyj @ jergym. hiedu. cz</mods:affiliation>
</mods:name>
<mods:typeOfResource id="D45F8CC9405FD2A06DA6D34B66ADC329">text</mods:typeOfResource>
<mods:relatedItem id="5E5E2B0D337CA9E88DFD43BEBBA4D5B8" type="host">
<mods:titleInfo id="71851551D4C0B0350BB959C535A29D10">
<mods:title id="79F36A88C9960A3F9FBE23E3AA254220">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="34EFF3D7336BE5F81B724DF6CBD03CB8">
<mods:date id="B136D129B2106EFAC4279196B9C05AEC">2005</mods:date>
<mods:detail id="C83C25C2A77D60C20E1565478899A73F" type="pubDate">
<mods:number id="8A74B7042F0227B087997ED5FF1DDDBB">2005-12-31</mods:number>
</mods:detail>
<mods:detail id="8FD5B5F6A351D2397097278C7D263473" type="volume">
<mods:number id="3D47BA3DDFE92BC64D30D325295B981D">45</mods:number>
</mods:detail>
<mods:extent id="D69C4B9FA3C83B22BE2FE4810F0721B7" unit="page">
<mods:start id="A44035419C7A06830CE50DABCAD57664">51</mods:start>
<mods:end id="8AD44424A9DEEF7B234B42ABC268F6FA">58</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="324800B0DC3E316C1CA3C91BA80620F1">journal article</mods:classification>
<mods:identifier id="413C693987920D4F35FF889C38D846A4" type="DOI">10.5281/zenodo.5176618</mods:identifier>
<mods:identifier id="40B3B1DC8808F5D9A94F628CB974D110" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="735D94177D2A9F5AE8ED019EB1A69192" type="Zenodo-Dep">5176618</mods:identifier>
</mods:mods>
<treatment id="FE0987CFFFFEE93FFE67E99AFEBE17F0" ID-DOI="http://doi.org/10.5281/zenodo.5188234" ID-Zenodo-Dep="5188234" LSID="urn:lsid:plazi:treatment:FE0987CFFFFEE93FFE67E99AFEBE17F0" httpUri="http://treatment.plazi.org/id/FE0987CFFFFEE93FFE67E99AFEBE17F0" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
<subSubSection id="3EBA6552FFFEE93CFE67E99AFD0D119D" box="[443,759,706,730]" pageId="1" pageNumber="52" type="nomenclature">
<paragraph id="761F36D9FFFEE93CFE67E99AFD0D119D" blockId="1.[443,759,706,730]" box="[443,759,706,730]" pageId="1" pageNumber="52">
<heading id="2D5781B5FFFEE93CFE67E99AFD0D119D" bold="true" box="[443,759,706,730]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="52" reason="3">
<emphasis id="44D4EACBFFFEE93CFE67E99AFD0D119D" bold="true" box="[443,759,706,730]" pageId="1" pageNumber="52">
<taxonomicName id="B1A04D5AFFFEE93CFE67E99AFD60119D" ID-CoL="3RM5M" authority="Hájek &amp; Šťastný, 2005" authorityName="Hájek &amp; Šťastný" authorityYear="2005" box="[443,666,706,730]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="boukali" status="sp. nov.">
<emphasis id="44D4EACBFFFEE93CFE67E99AFD60119D" bold="true" box="[443,666,706,730]" italics="true" pageId="1" pageNumber="52">Laccophilus boukali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FE757B0FFFEE93CFD7EE99AFD0D119D" box="[674,759,706,730]" pageId="1" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFEE93CFDF6E9BAFD7211B2" box="[554,648,738,758]" pageId="1" pageNumber="52" type="description">
<paragraph id="761F36D9FFFEE93CFDF6E9BAFD7211B2" blockId="1.[554,648,738,758]" box="[554,648,738,758]" pageId="1" pageNumber="52">
(
<figureCitation id="EE9B2A5CFFFEE93CFDECE9BAFD7811B2" box="[560,642,738,758]" captionStart="Figs 1-4. 1, 3" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[167,1112,215,1451]" captionTargetId="figure-9@4.[167,1112,215,1451]" captionTargetPageId="4" captionText="Figs 1-4. 1, 3 – Laccophilus boukali sp. nov. 1 – habitus; 3 – median lobe in lateral view. 2, 4 – L. uncletan sp. nov. 2 – habitus; 4 – aedeagus (a – median lobe in lateral view; b – apex of median lobe in dorsal view; c – left paramere; d – right paramere). Scale bars: 2 mm (Figs 1-2); 0.5 mm (Figs 3-4)." figureDoi="http://doi.org/10.5281/zenodo.5176620" httpUri="https://zenodo.org/record/5176620/files/figure.png" pageId="1" pageNumber="52">Figs. 1, 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFEE93CFFADE840FC6C1757" pageId="1" pageNumber="52" type="materials_examined">
<paragraph id="761F36D9FFFEE93CFFADE840FD571068" blockId="1.[113,1089,791,1040]" box="[113,685,791,816]" pageId="1" pageNumber="52">
<emphasis id="44D4EACBFFFEE93CFFADE840FEF21068" bold="true" box="[113,264,791,816]" pageId="1" pageNumber="52">
<typeStatus id="A91B887BFFFEE93CFFADE840FF521077" box="[113,168,792,816]" pageId="1" pageNumber="52">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="0EB77649FFFEE93CFED2E840FEBD1077" box="[270,327,792,816]" name="India" pageId="1" pageNumber="52">India</collectingCountry>
,
<collectingRegion id="B464F83BFFFEE93CFE8EE840FE381077" box="[338,450,792,816]" country="India" name="Karnataka" pageId="1" pageNumber="52">Karnataka</collectingRegion>
, Coorg, Kakkabe env.
</paragraph>
<paragraph id="761F36D9FFFEE93CFFADE860FC6C1757" blockId="1.[113,1089,791,1040]" pageId="1" pageNumber="52">
<emphasis id="44D4EACBFFFEE93CFFADE860FEE81008" bold="true" box="[113,274,823,848]" pageId="1" pageNumber="52">Type material.</emphasis>
<materialsCitation id="C6C83C84FFFEE93CFECAE860FD361037" collectingDate="1998-12-21" collectorName="D. Boukal" country="India" elevation="1050" latitude="12.25" location="Kakkabe" longLatPrecision="1293" longitude="75.583336" municipality="Coorg" pageId="1" pageNumber="52" specimenCount="1" stateProvince="Karnataka" typeStatus="holotype">
<typeStatus id="A91B887BFFFEE93CFECAE860FE811009" box="[278,379,824,847]" pageId="1" pageNumber="52" type="holotype">HOLOTYPE</typeStatus>
: J, ‘
<collectingCountry id="0EB77649FFFEE93CFE75E860FE061017" box="[425,508,824,848]" name="India" pageId="1" pageNumber="52">INDIA</collectingCountry>
:
<collectingRegion id="B464F83BFFFEE93CFDD8E860FD881017" box="[516,626,824,848]" country="India" name="Karnataka" pageId="1" pageNumber="52">Karnataka</collectingRegion>
,
<collectingMunicipality id="967BACA3FFFEE93CFDA6E860FD411017" box="[634,699,824,848]" pageId="1" pageNumber="52">Coorg</collectingMunicipality>
/
<location id="737F6002FFFEE93CFD17E860FCD21017" LSID="urn:lsid:plazi:treatment:FE0987CFFFFEE93FFE67E99AFEBE17F0:737F6002FFFEE93CFD17E860FCD21017" box="[715,808,824,848]" country="India" latitude="12.25" longLatPrecision="1293" longitude="75.583336" municipality="Coorg" name="Kakkabe" pageId="1" pageNumber="52" stateProvince="Karnataka">Kakkabe</location>
env.,
<date id="021E1019FFFEE93CFCBFE860FC221017" box="[867,984,824,848]" pageId="1" pageNumber="52" value="1998-12-21">
<collectingDate id="125AE9F1FFFEE93CFCBFE860FC221017" box="[867,984,824,848]" pageId="1" pageNumber="52" value="1998-12-21">21.12.1998</collectingDate>
</date>
/
<geoCoordinate id="1394501EFFFEE93CFC35E860FBBB1008" box="[1001,1089,822,848]" degrees="12" direction="north" minutes="15" orientation="latitude" pageId="1" pageNumber="52" precision="925" value="12.25">12°15′N</geoCoordinate>
<geoCoordinate id="1394501EFFFEE93CFFADE800FF331028" box="[113,201,854,880]" degrees="75" direction="east" minutes="35" orientation="longitude" pageId="1" pageNumber="52" precision="925" value="75.583336">75°35′E</geoCoordinate>
,
<quantity id="B1589B3CFFFEE93CFF0BE800FE981037" box="[215,354,856,880]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="1" pageNumber="52" unit="m" value="1050.0" valueMax="1200.0" valueMin="900.0">
<elevation id="FD8DD1EAFFFEE93CFF0BE800FE981037" box="[215,354,856,880]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="1" pageNumber="52" unit="m" value="1050.0" valueMax="1200.0" valueMin="900.0">900 - 1200m</elevation>
</quantity>
/ leg.
<collectorName id="DB55530FFFFEE93CFE77E800FDE11037" box="[427,539,856,880]" pageId="1" pageNumber="52">D. Boukal</collectorName>
(12) [printed] //
</materialsCitation>
<materialsCitation id="C6C83C84FFFEE93CFD09E800FC5510D7" collectionCode="NMW" country="India" determinerName="J. Hajek &amp; J. Stastny" pageId="1" pageNumber="52" specimenCount="1" stateProvince="Karnataka" typeStatus="holotype">
<typeStatus id="A91B887BFFFEE93CFD09E800FC9B1037" box="[725,865,856,880]" pageId="1" pageNumber="52" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="B1A04D5AFFFEE93CFCA6E800FBCB1037" authorityName="Leach" authorityYear="1815" box="[890,1073,856,880]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">LACCOPHILUS</taxonomicName>
/ boukali
<taxonomicNameLabel id="5FE757B0FFFEE93CFF14E820FEE010C8" box="[200,282,888,911]" pageId="1" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
/
<determinerName id="83C3DA1DFFFEE93CFEF3E820FE7F10D7" box="[303,389,888,912]" pageId="1" pageNumber="52">J. Hájek</determinerName>
&amp;
<determinerName id="83C3DA1DFFFEE93CFE74E820FDE810D7" box="[424,530,882,912]" pageId="1" pageNumber="52">J. Šťastný</determinerName>
det. 2005’ [printed, red label] (
<collectionCode id="10B1AE1CFFFEE93CFC82E820FC5210D7" box="[862,936,888,912]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="52">NMW</collectionCode>
)
</materialsCitation>
.
<typeStatus id="A91B887BFFFEE93CFC65E820FBDF10C9" box="[953,1061,888,911]" pageId="1" pageNumber="52" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="60A6FD50FFFEE93CFBEFE820FF7410F6" pageId="1" pageNumber="52" type="female">2 ♀♀</specimenCount>
, same label data as
<typeStatus id="A91B887BFFFEE93CFEACE8C0FE3710F7" box="[368,461,920,944]" pageId="1" pageNumber="52" type="holotype">holotype</typeStatus>
(
<collectionCode id="10B1AE1CFFFEE93CFE00E8C0FDDC10F7" box="[476,550,920,944]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="52">NMW</collectionCode>
). 1 J, ‘
<collectingCountry id="0EB77649FFFEE93CFDAFE8C0FD3D10F7" box="[627,711,920,944]" name="India" pageId="1" pageNumber="52">INDIA</collectingCountry>
:
<collectingRegion id="B464F83BFFFEE93CFD0EE8C0FCB810F7" box="[722,834,920,944]" country="India" name="Karnataka" pageId="1" pageNumber="52">Karnataka</collectingRegion>
, Coorg / Madikeri env.,
<date id="021E1019FFFEE93CFFADE8E0FF131097" box="[113,233,952,976]" pageId="1" pageNumber="52" value="1998-12-19">19.12.1998</date>
/
<quantity id="B1589B3CFFFEE93CFED9E8E0FEAB1097" box="[261,337,952,976]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="52" unit="m" value="1300.0">1300m</quantity>
, Talakaveri /
<geoCoordinate id="1394501EFFFEE93CFE37E8E0FDBE1088" box="[491,580,950,976]" degrees="12" direction="north" minutes="23" orientation="latitude" pageId="1" pageNumber="52" precision="925" value="12.383333">12°23′N</geoCoordinate>
<geoCoordinate id="1394501EFFFEE93CFD92E8E0FD5F1088" box="[590,677,950,976]" degrees="75" direction="east" minutes="30" orientation="longitude" pageId="1" pageNumber="52" precision="925" value="75.5">75°30′E</geoCoordinate>
/ leg. D. Boukal (5)’ (
<collectionCode id="10B1AE1CFFFEE93CFC79E8E0FC0D1097" box="[933,1015,952,976]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="52">NMPC</collectionCode>
). Each
<typeStatus id="A91B887BFFFEE93CFFADE880FF3610B7" box="[113,204,984,1008]" pageId="1" pageNumber="52" type="paratype">paratype</typeStatus>
is provided with a red label similar to that of the
<typeStatus id="A91B887BFFFEE93CFD2EE880FCAB10B7" box="[754,849,984,1008]" pageId="1" pageNumber="52" type="holotype">holotype</typeStatus>
, except ‘PARATYPE’ instead of ‘HOLOTYPE’, the respective sex symbol and collection number.
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFEE93FFFADEF7FFC4B1170" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52" type="description">
<paragraph id="761F36D9FFFEE93CFFADEF7FFBFB1719" blockId="1.[113,1090,1063,1599]" pageId="1" pageNumber="52">
<emphasis id="44D4EACBFFFEE93CFFADEF7FFF011778" bold="true" box="[113,251,1063,1087]" pageId="1" pageNumber="52">Description.</emphasis>
Oval and oblong; head and pronotum testaceous, larger part of elytra brown with testaceous pattern as depicted in
<figureCitation id="EE9B2A5CFFFEE93CFDD9EF1FFDB91718" box="[517,579,1095,1119]" captionStart="Figs 1-4. 1, 3" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[167,1112,215,1451]" captionTargetId="figure-9@4.[167,1112,215,1451]" captionTargetPageId="4" captionText="Figs 1-4. 1, 3 – Laccophilus boukali sp. nov. 1 – habitus; 3 – median lobe in lateral view. 2, 4 – L. uncletan sp. nov. 2 – habitus; 4 – aedeagus (a – median lobe in lateral view; b – apex of median lobe in dorsal view; c – left paramere; d – right paramere). Scale bars: 2 mm (Figs 1-2); 0.5 mm (Figs 3-4)." figureDoi="http://doi.org/10.5281/zenodo.5176620" httpUri="https://zenodo.org/record/5176620/files/figure.png" pageId="1" pageNumber="52">Fig. 1</figureCitation>
. Ventral parts and appendages testaceous.
</paragraph>
<paragraph id="761F36D9FFFEE93CFF4CEF3FFCD617D8" blockId="1.[113,1090,1063,1599]" pageId="1" pageNumber="52">
Measurements. Body length
<quantity id="B1589B3CFFFEE93CFE1FEF3FFDA21739" box="[451,600,1127,1151]" metricMagnitude="-3" metricUnit="m" metricValue="3.275" metricValueMax="3.35" metricValueMin="3.2" pageId="1" pageNumber="52" unit="mm" value="3.275" valueMax="3.35" valueMin="3.2">3.20-3.35 mm</quantity>
(mean value =
<quantity id="B1589B3CFFFEE93CFCDDEF3FFC991739" box="[769,867,1127,1151]" metricMagnitude="-3" metricUnit="m" metricValue="3.26" pageId="1" pageNumber="52" unit="mm" value="3.26">3.26 mm</quantity>
,
<typeStatus id="A91B887BFFFEE93CFCB2EF3FFC311738" box="[878,971,1127,1151]" pageId="1" pageNumber="52" type="holotype">holotype</typeStatus>
<quantity id="B1589B3CFFFEE93CFC0EEF3FFBCF1738" box="[978,1077,1127,1151]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="52" unit="mm" value="3.2">3.20 mm</quantity>
), width
<quantity id="B1589B3CFFFEE93CFF68EFDFFEB317D9" box="[180,329,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="1.825" metricValueMax="1.85" metricValueMin="1.8" pageId="1" pageNumber="52" unit="mm" value="1.825" valueMax="1.85" valueMin="1.8">1.80-1.85 mm</quantity>
(mean value =
<quantity id="B1589B3CFFFEE93CFE33EFDFFDAA17D9" box="[495,592,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="1.81" pageId="1" pageNumber="52" unit="mm" value="1.81">1.81 mm</quantity>
,
<typeStatus id="A91B887BFFFEE93CFD86EFDFFD4D17D8" box="[602,695,1159,1183]" pageId="1" pageNumber="52" type="holotype">holotype</typeStatus>
<quantity id="B1589B3CFFFEE93CFD61EFDFFCDA17D8" box="[701,800,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="52" unit="mm" value="1.8">1.80 mm</quantity>
).
</paragraph>
<paragraph id="761F36D9FFFEE93CFF4CEFFFFDFD1659" blockId="1.[113,1090,1063,1599]" pageId="1" pageNumber="52">Head. Coloration testaceous, in some specimens with indistinct light brown markings. Reticulation double, composed of moderately deeply impressed transverse polygonal meshes on anterior part and irregular meshes on posterior part, and microreticulation of very fine round meshes. Mouthparts testaceous.</paragraph>
<paragraph id="761F36D9FFFEE93CFF4CEE7FFCA31639" blockId="1.[113,1090,1063,1599]" pageId="1" pageNumber="52">Pronotum. Coloration testaceous, with two small brown spots at basal margin, and with anterior margin slightly darkened. Reticulation double, similar to that of head, but meshes larger. Anterior margin and posterior angles with numerous punctures.</paragraph>
<paragraph id="761F36D9FFFEE93FFF4CEEDFFE421270" blockId="1.[113,1090,1063,1599]" lastBlockId="2.[151,1127,223,1207]" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">Elytra. Elytral disc brown with testaceous markings, lateral parts testaceous. Subbasal testaceous band with anterior border almost straight, posterior border with distinct elongation near elytral suture and in most specimens also with smaller discal elongation. Subapical band divided into five longitudinal stripes. Remaining elytral surface with sinuous stripes, often indistinct in middle third of elytra, but widening apically, with elytral apex nearly testaceous in some specimens. Reticulation on whole elytra distinctly double, composed of irregular polygonal meshes and microreticulation, which is moderately deeply impressed and more distinct than on head or pronotum. Each elytron with sutural, discal and lateral row of large serial setigerous punctures.</paragraph>
<paragraph id="761F36D9FFFDE93FFF6AEA67FDB812DF" blockId="2.[151,1127,223,1207]" pageId="2" pageNumber="53">Ventral side. Testaceous and microreticulated. Prosternal process long and acute. Abdominal sternites laterally with oblique grooves. Anal sternite tectiform, its surface with large punctures, setae and transverse grooves.</paragraph>
<paragraph id="761F36D9FFFDE93FFF6AEAF8FF3612B0" blockId="2.[151,1127,223,1207]" pageId="2" pageNumber="53">Legs. Completely testaceous. Male tarsomeres 1-3 of fore and middle legs weakly dilated with microtrichia on ventral side. Claws of fore tarsi simple and equal, as well as those of mid tarsi.</paragraph>
<paragraph id="761F36D9FFFDE93FFF6AEAA7FD3F1150" blockId="2.[151,1127,223,1207]" box="[182,709,511,535]" pageId="2" pageNumber="53">
Male genitalia. Shape of median lobe as in
<figureCitation id="EE9B2A5CFFFDE93FFD5DEAA7FD3B1150" box="[641,705,511,535]" captionStart="Figs 1-4. 1, 3" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[167,1112,215,1451]" captionTargetId="figure-9@4.[167,1112,215,1451]" captionTargetPageId="4" captionText="Figs 1-4. 1, 3 – Laccophilus boukali sp. nov. 1 – habitus; 3 – median lobe in lateral view. 2, 4 – L. uncletan sp. nov. 2 – habitus; 4 – aedeagus (a – median lobe in lateral view; b – apex of median lobe in dorsal view; c – left paramere; d – right paramere). Scale bars: 2 mm (Figs 1-2); 0.5 mm (Figs 3-4)." figureDoi="http://doi.org/10.5281/zenodo.5176620" httpUri="https://zenodo.org/record/5176620/files/figure.png" pageId="2" pageNumber="53">Fig. 3</figureCitation>
.
</paragraph>
<paragraph id="761F36D9FFFDE93FFF6AE947FC4B1170" blockId="2.[151,1127,223,1207]" box="[182,945,543,567]" pageId="2" pageNumber="53">Female. Similar to male; tarsomeres of fore and middle legs not dilated.</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFDE93FFF4BE967FDF01030" pageId="2" pageNumber="53" type="diagnosis">
<paragraph id="761F36D9FFFDE93FFF4BE967FDF01030" blockId="2.[151,1127,223,1207]" pageId="2" pageNumber="53">
<emphasis id="44D4EACBFFFDE93FFF4BE967FE751110" bold="true" box="[151,399,575,599]" pageId="2" pageNumber="53">Differential diagnosis.</emphasis>
Based on the double reticulation of pronotum and elytra and the elytral pattern (brown sinuous stripes), the new species belongs to the
<taxonomicName id="B1A04D5AFFFDE93FFCAEE907FBA71130" authorityName="Boheman" authorityYear="1858" box="[882,1117,607,631]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="44D4EACBFFFDE93FFCAEE907FBA71130" box="[882,1117,607,631]" italics="true" pageId="2" pageNumber="53">Laccophilus chinensis</emphasis>
</taxonomicName>
- group as defined by
<bibRefCitation id="12314B28FFFDE93FFEABE9D8FDC311D0" author="BRANCUCCI M." box="[375,569,639,663]" pageId="2" pageNumber="53" pagination="241 - 426" refId="ref3760" refString="BRANCUCCI M. 1983: Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey 31 / 32: 241 - 426." type="journal article" year="1983">BRANCUCCI (1983)</bibRefCitation>
. Among the
<collectingRegion id="B464F83BFFFDE93FFD18E927FCE111D0" box="[708,795,639,663]" country="Democratic Republic of the Congo" name="Orientale" pageId="2" pageNumber="53">Oriental</collectingRegion>
species of this group, the new species is similar to
<taxonomicName id="B1A04D5AFFFDE93FFEAAE9F8FD5511F0" authority="Regimbart, 1880" authorityName="Regimbart" authorityYear="1880" box="[374,687,671,695]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="ritsemae">
<emphasis id="44D4EACBFFFDE93FFEAAE9F8FE1511F0" box="[374,495,672,695]" italics="true" pageId="2" pageNumber="53">L. ritsemae</emphasis>
Régimbart, 1880
</taxonomicName>
, which it resembles in the habitus, body length and coloration. Both species can only be distinguished by the male genitalia. The median lobe of
<taxonomicName id="B1A04D5AFFFDE93FFEE4E9B8FE5B11B0" authority="Hájek &amp; Šťastný, 2005" authorityName="Hájek &amp; Šťastný" authorityYear="2005" box="[312,417,735,759]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="boukali" status="sp. nov.">
<emphasis id="44D4EACBFFFDE93FFEE4E9B8FE5B11B0" box="[312,417,735,759]" italics="true" pageId="2" pageNumber="53">L. boukali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FE757B0FFFDE93FFE7BE9B8FE0211B0" box="[423,504,736,759]" pageId="2" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
is short and broad with a distinct ventral angle in the middle third of its length, while the median lobe of
<taxonomicName id="B1A04D5AFFFDE93FFDAFE858FD111050" authorityName="Regimbart" authorityYear="1880" box="[627,747,768,791]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="ritsemae">
<emphasis id="44D4EACBFFFDE93FFDAFE858FD111050" box="[627,747,768,791]" italics="true" pageId="2" pageNumber="53">L. ritsemae</emphasis>
</taxonomicName>
is long and slender with its ventral edge sinuous. From other sympatrically occurring species,
<taxonomicName id="B1A04D5AFFFDE93FFCD7E878FBC01070" authority="(Walker, 1859)" baseAuthorityName="Walker" baseAuthorityYear="1859" box="[779,1082,799,823]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="inefficiens">
<emphasis id="44D4EACBFFFDE93FFCD7E878FC6E1070" box="[779,916,799,823]" italics="true" pageId="2" pageNumber="53">L. inefficiens</emphasis>
(Walker, 1859)
</taxonomicName>
and
<taxonomicName id="B1A04D5AFFFDE93FFF4BE818FE3B1010" authority="Brancucci, 1983" authorityName="Brancucci" authorityYear="1983" box="[151,449,831,855]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="44D4EACBFFFDE93FFF4BE818FEFF1010" box="[151,261,831,855]" italics="true" pageId="2" pageNumber="53">L. kaszabi</emphasis>
<bibRefCitation id="12314B28FFFDE93FFED3E867FE3B1010" author="BRANCUCCI M." box="[271,449,831,855]" pageId="2" pageNumber="53" pagination="241 - 426" refId="ref3760" refString="BRANCUCCI M. 1983: Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey 31 / 32: 241 - 426." type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
</taxonomicName>
, the new species differs by the generally smaller size and the different shape of the median lobe.
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFDE93FFF4BE827FEA11710" pageId="2" pageNumber="53" type="distribution">
<paragraph id="761F36D9FFFDE93FFF4BE827FF2F10F0" blockId="2.[151,1127,223,1207]" pageId="2" pageNumber="53">
<emphasis id="44D4EACBFFFDE93FFF4BE827FEDD10D0" bold="true" box="[151,295,895,919]" pageId="2" pageNumber="53">Distribution.</emphasis>
So far known only from two close localities in
<collectingRegion id="B464F83BFFFDE93FFCCDE827FC4E10D0" box="[785,948,895,919]" country="India" name="Karnataka" pageId="2" pageNumber="53">Karnataka state</collectingRegion>
in south-western
<collectingCountry id="0EB77649FFFDE93FFF4BE8C7FF2B10F0" box="[151,209,927,951]" name="India" pageId="2" pageNumber="53">India</collectingCountry>
.
</paragraph>
<paragraph id="761F36D9FFFDE93FFF4BE8E7FEA11710" blockId="2.[151,1127,223,1207]" pageId="2" pageNumber="53">
<emphasis id="44D4EACBFFFDE93FFF4BE8E7FEF81090" bold="true" box="[151,258,959,983]" pageId="2" pageNumber="53">Bionomy.</emphasis>
The specimens from Kakkabe were collected in a larger stream with the main channel ca
<quantity id="B1589B3CFFFDE93FFED2E887FECD10B0" box="[270,311,991,1015]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="53" unit="m" value="5.0">5 m</quantity>
wide, with large boulders, a rocky bed with some stones and gravel in riffles and silt in pools. The specimen from Madikeri was collected in an artificial reservoir at a hilltop; the examined site had a dense shore vegetation, muddy bed, and was little shaded (D. S. BOUKAL in litt.).
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFDE93FFF4BEF07FEBE17F0" pageId="2" pageNumber="53" type="etymology">
<paragraph id="761F36D9FFFDE93FFF4BEF07FEBE17F0" blockId="2.[151,1127,223,1207]" pageId="2" pageNumber="53">
<emphasis id="44D4EACBFFFDE93FFF4BEF07FEED1730" bold="true" box="[151,279,1119,1143]" pageId="2" pageNumber="53">Etymology.</emphasis>
The new species is dedicated to its collector David S. Boukal (České Budějovice,
<collectingCountry id="0EB77649FFFDE93FFF0CEF27FE8617D0" box="[208,380,1151,1175]" name="Czech Republic" pageId="2" pageNumber="53">Czech Republic</collectingCountry>
), our friend, theoretical mathematician, photographer, and specialist on the Dryopoidea.
</paragraph>
</subSubSection>
</treatment>
</document>
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