192 lines
21 KiB
XML
192 lines
21 KiB
XML
<document id="7857AA03DCA7738AA19A39CC0A60F06F" ID-DOI="10.11646/zootaxa.3949.3.3" ID-GBIF-Dataset="25ece91c-6566-4122-a406-236f147b58b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="238900" ID-ZooBank="CBEF804D-5FBF-48D2-8CCD-036B70FE6ECD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461046281391" checkinUser="plazi" docAuthor="Arias, Andrés & Paxton, Hannelore" docDate="2015" docId="03BA8797FFB9FFA2FF2BF895FBD4C67E" docLanguage="en" docName="zt03949p369.pdf" docOrigin="Zootaxa 3949 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onuphis pancerii Claparede 1868" docType="treatment" docVersion="14" lastPageNumber="356" masterDocId="FF83FFEFFFB3FFA9FFBCFFE0FFA7C254" masterDocTitle="Onuphis and Aponuphis (Annelida: Onuphidae) from southwestern Europe, with the description of a new species" masterLastPageNumber="369" masterPageNumber="345" pageNumber="355" updateTime="1698577207772" updateUser="plazi">
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<mods:title id="04884CE1DC3376E42EFEEBC9ED50A4A9">Onuphis and Aponuphis (Annelida: Onuphidae) from southwestern Europe, with the description of a new species</mods:title>
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<mods:namePart id="4136C9F259A1A360999DE47EA748CA37">Arias, Andrés</mods:namePart>
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<mods:namePart id="4953C37F5CAE467DE17D8C470C18AE7E">Paxton, Hannelore</mods:namePart>
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<mods:typeOfResource id="E802A82BC5A157D074EA02EAC0759BF6">text</mods:typeOfResource>
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<mods:title id="901A2DE83553E0C2718EA693F33C9CF3">Zootaxa</mods:title>
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<mods:date id="FD595B526D27E2EE52DD141B8B20F95E">2015</mods:date>
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<mods:number id="CA651A35C111BAAE567C144B67010C16">3949</mods:number>
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<treatment id="03BA8797FFB9FFA2FF2BF895FBD4C67E" ID-DOI="http://doi.org/10.5281/zenodo.3513280" ID-GBIF-Taxon="119610573" ID-Zenodo-Dep="3513280" LSID="urn:lsid:plazi:treatment:03BA8797FFB9FFA2FF2BF895FBD4C67E" httpUri="http://treatment.plazi.org/id/03BA8797FFB9FFA2FF2BF895FBD4C67E" lastPageId="11" lastPageNumber="356" pageId="10" pageNumber="355">
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<subSubSection id="C309650AFFB9FFA3FF2BF895FD9AC5DB" box="[151,573,1909,1935]" pageId="10" pageNumber="355" type="nomenclature">
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<paragraph id="8BAC3681FFB9FFA3FF2BF895FD9AC5DB" blockId="10.[151,573,1909,1935]" box="[151,573,1909,1935]" pageId="10" pageNumber="355">
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<heading id="D0E481EDFFB9FFA3FF2BF895FD9AC5DB" bold="true" box="[151,573,1909,1935]" fontSize="11" level="1" pageId="10" pageNumber="355" reason="1">
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<taxonomicName id="4C134D02FFB9FFA3FF2BF895FD9AC5DB" ID-CoL="49QKV" authority="Claparede, 1868" authorityName="Claparede" authorityYear="1868" box="[151,573,1909,1935]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="10" pageNumber="355" phylum="Annelida" rank="species" species="pancerii">
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<emphasis id="B967EA93FFB9FFA3FF2BF895FD9AC5DB" bold="true" box="[151,573,1909,1935]" pageId="10" pageNumber="355">
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<emphasis id="B967EA93FFB9FFA3FF2BF895FECEC5DB" bold="true" box="[151,361,1909,1935]" italics="true" pageId="10" pageNumber="355">Onuphis pancerii</emphasis>
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<bibRefCitation id="EF824B70FFB9FFA3FECDF895FD9AC5DB" author="Claparede" box="[369,573,1909,1935]" pageId="10" pageNumber="355" refString="Claparede, E. (1868) Les Annelides Chetopodes du Golfe de Naples. Memoires de la Societe de physique et d'histoire naturelle de Geneve, 19, 313 - 584." type="journal article" year="1868">Claparède, 1868</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C309650AFFB9FFA2FF2BF85CFD4BC298" lastPageId="11" lastPageNumber="356" pageId="10" pageNumber="355" type="reference_group">
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<paragraph id="8BAC3681FFB9FFA3FF2BF85CFDDBC5A4" blockId="10.[151,1436,1980,2032]" pageId="10" pageNumber="355">
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<treatmentCitationGroup id="AB0311AFFFB9FFA3FF2BF85CFDDBC5A4" pageId="10" pageNumber="355">
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<taxonomicName id="4C134D02FFB9FFA3FF2BF85CFD80C586" ID-CoL="49QKV" authority="Claparede, 1868: 438" authorityName="Claparede" authorityPageNumber="438" authorityYear="1868" box="[151,551,1980,2002]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="10" pageNumber="355" phylum="Annelida" rank="species" species="pancerii">
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<emphasis id="B967EA93FFB9FFA3FF2BF85CFEE0C586" box="[151,327,1980,2002]" italics="true" pageId="10" pageNumber="355">Onuphis pancerii</emphasis>
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<treatmentCitation id="0AB21090FFB9FFA3FEF1F85CFD80C586" author="Claparede" box="[333,551,1980,2002]" page="438" pageId="10" pageNumber="355" year="1868">
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<bibRefCitation id="EF824B70FFB9FFA3FEF1F85CFE50C586" author="Claparede" box="[333,503,1980,2002]" pageId="10" pageNumber="355" refString="Claparede, E. (1868) Les Annelides Chetopodes du Golfe de Naples. Memoires de la Societe de physique et d'histoire naturelle de Geneve, 19, 313 - 584." type="journal article" year="1868">Claparède, 1868</bibRefCitation>
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: 438
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</treatmentCitation>
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||
</taxonomicName>
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–440, pl. VIII fig. 1; 1870: 387, pl. V fig. 5a;
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<treatmentCitation id="0AB21090FFB9FFA3FC48F85CFB0EC586" author="Rioja" box="[1012,1193,1980,2002]" page="39" pageId="10" pageNumber="355" year="1918">
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<bibRefCitation id="EF824B70FFB9FFA3FC48F85CFBDEC586" author="Rioja" box="[1012,1145,1980,2002]" pageId="10" pageNumber="355" refString="Rioja, E. (1918) Datos para el conocimiento de la fauna de anelidos poliquetos del Cantabrico (2 ª Parte). Trabajos del Museo Nacional de Ciencias Naturales, Madrid, Serie Zoologica, 37, 1 - 99." type="journal article" year="1918">Rioja, 1918</bibRefCitation>
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: 39
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</treatmentCitation>
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, fig. 10 (Santander);
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<treatmentCitation id="0AB21090FFB9FFA3FF7BF83AFE70C5A4" ID-DOI="http://doi.org/10.5281/zenodo.3507259" ID-Zenodo-Dep="3507259" author="Arias" box="[199,471,2010,2032]" httpUri="http://treatment.plazi.org/id/03CA87FEFF919C17FF020418FF34FB50" page="158" pageId="10" pageNumber="355" year="2014">
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<bibRefCitation id="EF824B70FFB9FFA3FF7BF83AFE0FC5A4" author="Arias" box="[199,424,2010,2032]" pageId="10" pageNumber="355" refString="Arias, A. & Paxton, H. (2014) Hidden diversity within the polychaete Onuphis eremita sensu lato (Annelida: Onuphidae) - redescription of O. eremita Audouin & Milne-Edwards, 1833 and reinstatement of Onuphis pancerii Claparede, 1868. Zootaxa, 3861 (2), 145 - 169." type="journal article" year="2014">Arias & Paxton, 2014</bibRefCitation>
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: 158
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</treatmentCitation>
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–164, figs 9–12.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BAC3681FFB8FFA2FF2BFF78FD4BC298" blockId="11.[151,1436,152,204]" pageId="11" pageNumber="356">
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<emphasis id="B967EA93FFB8FFA2FF2BFF78FEE1C2FA" box="[151,326,152,174]" italics="true" pageId="11" pageNumber="356">
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<taxonomicName id="4C134D02FFB8FFA2FF2BFF78FEE5C2FA" ID-CoL="49QJ7" box="[151,322,152,174]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="356" phylum="Annelida" rank="species" species="eremita">Onuphis eremita</taxonomicName>
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.
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</emphasis>
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—
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<bibRefCitation id="EF824B70FFB8FFA2FEDCFF78FE44C2FA" author="Fauvel" box="[352,483,152,174]" pageId="11" pageNumber="356" refString="Fauvel, P. (1923) Faune de France: Polychetes errantes. Paul Lechevalier, Paris, 488 pp." type="book" year="1923">Fauvel, 1923</bibRefCitation>
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(in part): 413–415, fig. 163. Not
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<bibRefCitation id="EF824B70FFB8FFA2FC8AFF78FB35C2FA" author="Audouin" box="[822,1170,152,174]" pageId="11" pageNumber="356" refString="Audouin, J. V. & Milne Edwards, H. (1833) Classification des Annelides et description de celles qui habitent les cotes de la France. Annales des sciences naturelles, 29, 195 - 269." type="journal article" year="1833">Audouin & Milne Edwards, 1833</bibRefCitation>
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.—
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<bibRefCitation id="EF824B70FFB8FFA2FB0CFF78FAE3C2FA" author="Paxton" box="[1200,1348,152,174]" pageId="11" pageNumber="356" refString="Paxton, H. (1986) Generic revision and relationships of the family Onuphidae (Annelida: Polychaeta). Records of the Australian Museum, 38, 1 - 74. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.38.1986.175" type="journal article" year="1986">Paxton, 1986</bibRefCitation>
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: 56–58, figs 3, 6–8, 10. Not
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<bibRefCitation id="EF824B70FFB8FFA2FE2DFF56FD4FC298" author="Audouin" box="[401,744,182,204]" pageId="11" pageNumber="356" refString="Audouin, J. V. & Milne Edwards, H. (1833) Classification des Annelides et description de celles qui habitent les cotes de la France. Annales des sciences naturelles, 29, 195 - 269." type="journal article" year="1833">Audouin & Milne Edwards, 1833</bibRefCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C309650AFFB8FFA2FF2BFF19FCF9C392" pageId="11" pageNumber="356" type="materials_examined">
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||
<paragraph id="8BAC3681FFB8FFA2FF2BFF19FE5BC362" blockId="11.[151,1437,249,1066]" pageId="11" pageNumber="356">
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||
<emphasis id="B967EA93FFB8FFA2FF2BFF19FED9C346" bold="true" box="[151,382,249,274]" pageId="11" pageNumber="356">Material examined</emphasis>
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||
.
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<typeStatus id="54A88823FFB8FFA2FE2DFF1AFE6FC346" box="[401,456,250,274]" pageId="11" pageNumber="356">Type</typeStatus>
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material.
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||
<typeStatus id="54A88823FFB8FFA2FDFFFF1AFD05C346" box="[579,674,250,274]" pageId="11" pageNumber="356" type="neotype">Neotype</typeStatus>
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(
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<collectionCode id="ED02AE44FFB8FFA2FD09FF19FCACC345" LSID="urn:lsid:biocol.org:col:35275" box="[693,779,249,273]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="356">MNCN</collectionCode>
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16.01/311) Naples,
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<collectingCountry id="F3047611FFB8FFA2FC40FF19FB97C346" box="[1020,1072,249,274]" name="Italy" pageId="11" pageNumber="356">Italy</collectingCountry>
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, specimen Num. 173 from the Collection of Rioja,
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<date id="FFAD1041FFB8FFA2FEC0FEFEFE5FC362" box="[380,504,286,310]" pageId="11" pageNumber="356" value="1894" valueMax="1917">1894-1917</date>
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.
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</paragraph>
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<paragraph id="8BAC3681FFB8FFA2FF7BFEA2FDBBC3F5" blockId="11.[151,1437,249,1066]" pageId="11" pageNumber="356">
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Non-type material. Nine specimens (
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<collectionCode id="ED02AE44FFB8FFA2FDDEFEA1FD1FC30D" LSID="urn:lsid:biocol.org:col:35275" box="[610,696,321,345]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="356">MNCN</collectionCode>
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16.01/310) Enmedio sandy flat, Santander Bay,
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<collectingCountry id="F3047611FFB8FFA2FB66FEA1FABAC30E" box="[1242,1309,321,346]" name="Spain" pageId="11" pageNumber="356">Spain</collectingCountry>
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, specimens Num. 686 from the Rioja Collection
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<date id="FFAD1041FFB8FFA2FD85FE86FD11C32A" box="[569,694,358,382]" pageId="11" pageNumber="356" value="1894" valueMax="1917">1894-1917</date>
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||
. For a detailed listing of additional material from
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<collectingCountry id="F3047611FFB8FFA2FB50FE85FA87C32A" box="[1260,1312,357,382]" name="Italy" pageId="11" pageNumber="356">Italy</collectingCountry>
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,
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<collectingCountry id="F3047611FFB8FFA2FA90FE85FACBC32A" box="[1324,1388,357,382]" name="Spain" pageId="11" pageNumber="356">Spain</collectingCountry>
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and
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<collectingCountry id="F3047611FFB8FFA2FF2BFE6AFF44C3F5" box="[151,227,394,417]" name="France" pageId="11" pageNumber="356">France</collectingCountry>
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see
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<bibRefCitation id="EF824B70FFB8FFA2FEA9FE69FDBFC3F5" author="Arias" box="[277,536,393,418]" pageId="11" pageNumber="356" refString="Arias, A. & Paxton, H. (2014) Hidden diversity within the polychaete Onuphis eremita sensu lato (Annelida: Onuphidae) - redescription of O. eremita Audouin & Milne-Edwards, 1833 and reinstatement of Onuphis pancerii Claparede, 1868. Zootaxa, 3861 (2), 145 - 169." type="journal article" year="2014">Arias & Paxton (2014)</bibRefCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8BAC3681FFB8FFA2FF7BFE4DFCF9C392" blockId="11.[151,1437,249,1066]" box="[199,862,429,454]" pageId="11" pageNumber="356">
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||
<emphasis id="B967EA93FFB8FFA2FF7BFE4DFEC2C392" bold="true" box="[199,357,429,454]" pageId="11" pageNumber="356">
|
||
<typeStatus id="54A88823FFB8FFA2FF7BFE4DFEA5C392" box="[199,258,429,454]" pageId="11" pageNumber="356">Type</typeStatus>
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locality.
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||
</emphasis>
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||
Gulf of Naples,
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<collectingCountry id="F3047611FFB8FFA2FD9CFE4DFDF3C392" box="[544,596,429,454]" name="Italy" pageId="11" pageNumber="356">Italy</collectingCountry>
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||
, central Mediterranean.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C309650AFFB8FFA2FF7BFE31FD64C15E" pageId="11" pageNumber="356" type="diagnosis">
|
||
<paragraph id="8BAC3681FFB8FFA2FF7BFE31FD64C15E" blockId="11.[151,1437,249,1066]" pageId="11" pageNumber="356">
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||
<emphasis id="B967EA93FFB8FFA2FF7BFE31FEE6C3BE" bold="true" box="[199,321,465,490]" pageId="11" pageNumber="356">Diagnosis.</emphasis>
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Prostomium anteriorly extended. Eye spots absent. Palps reaching chaetiger 1–2, lateral antennae reaching chaetiger 5–6 and median antenna chaetiger 2–4. Ceratophores long and strongly ringed, palpal ceratophores with 22–26 rings, lateral antennae with 21–29 and median antenna with 15–20 rings. Subulate ventral cirri in first five to six chaetigers; distinct subulate postchaetal lobes in first 10–19 chaetigers. Small interramal papilla present between chaetigers 4–5 to 9. Bi- and tridentate (or even multidentate) pseudocompound hooks in first three chaetigers. All hooks of equal thickness, slender long-appendaged ones absent. Subacicular hooks from chaetiger 10. Flat, distally oblique pectinate chaetae with 13–17 teeth. Single branchial filaments from chaetiger 1 to 19–24, thereafter number increasing to maximum of six to eight. Tube with soft inner layer and outer layer of vegetal material, shell-fragments and sand grains.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C309650AFFB8FFA2FF7BFCF5FA92C651" pageId="11" pageNumber="356" type="discussion">
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||
<paragraph id="8BAC3681FFB8FFA2FF7BFCF5FA92C651" blockId="11.[151,1437,249,1066]" pageId="11" pageNumber="356">
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||
<emphasis id="B967EA93FFB8FFA2FF7BFCF5FE9CC17A" bold="true" box="[199,315,789,814]" pageId="11" pageNumber="356">Remarks.</emphasis>
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This species was synonymised with
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<taxonomicName id="4C134D02FFB8FFA2FD66FCF6FCF5C179" box="[730,850,790,813]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="356" phylum="Annelida" rank="species" species="eremita">
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||
<emphasis id="B967EA93FFB8FFA2FD66FCF6FCF5C179" box="[730,850,790,813]" italics="true" pageId="11" pageNumber="356">O. eremita</emphasis>
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</taxonomicName>
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||
by
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<bibRefCitation id="EF824B70FFB8FFA2FCC0FCF5FBBDC17A" author="Fauvel" box="[892,1050,789,814]" pageId="11" pageNumber="356" refString="Fauvel, P. (1923) Faune de France: Polychetes errantes. Paul Lechevalier, Paris, 488 pp." type="book" year="1923">Fauvel (1923)</bibRefCitation>
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||
and remained as such for nearly a century. However,
|
||
<bibRefCitation id="EF824B70FFB8FFA2FE38FCD9FD0DC106" author="Arias" box="[388,682,825,850]" pageId="11" pageNumber="356" refString="Arias, A. & Paxton, H. (2014) Hidden diversity within the polychaete Onuphis eremita sensu lato (Annelida: Onuphidae) - redescription of O. eremita Audouin & Milne-Edwards, 1833 and reinstatement of Onuphis pancerii Claparede, 1868. Zootaxa, 3861 (2), 145 - 169." type="journal article" year="2014">Arias & Paxton (2014)</bibRefCitation>
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||
demonstrated that
|
||
<taxonomicName id="4C134D02FFB8FFA2FC19FCDAFB96C105" box="[933,1073,826,849]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="356" phylum="Annelida" rank="species" species="pancerii">
|
||
<emphasis id="B967EA93FFB8FFA2FC19FCDAFB96C105" box="[933,1073,826,849]" italics="true" pageId="11" pageNumber="356">O. pancerii</emphasis>
|
||
</taxonomicName>
|
||
is actually a valid species, morphologically different from
|
||
<taxonomicName id="4C134D02FFB8FFA2FE46FCBEFDD7C121" box="[506,624,862,885]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="356" phylum="Annelida" rank="species" species="eremita">
|
||
<emphasis id="B967EA93FFB8FFA2FE46FCBEFDD7C121" box="[506,624,862,885]" italics="true" pageId="11" pageNumber="356">O. eremita</emphasis>
|
||
</taxonomicName>
|
||
. Although the two species are closely related and thus share a number of parapodial and chaetal characteristics, they can easily be differentiated in that
|
||
<taxonomicName id="4C134D02FFB8FFA2FBB7FC62FB2DC1CD" box="[1035,1162,898,921]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="356" phylum="Annelida" rank="species" species="pancerii">
|
||
<emphasis id="B967EA93FFB8FFA2FBB7FC62FB2DC1CD" box="[1035,1162,898,921]" italics="true" pageId="11" pageNumber="356">O. pancerii</emphasis>
|
||
</taxonomicName>
|
||
is a much larger, robust species than
|
||
<emphasis id="B967EA93FFB8FFA2FE93FC46FE14C1E9" box="[303,435,934,957]" italics="true" pageId="11" pageNumber="356">
|
||
<taxonomicName id="4C134D02FFB8FFA2FE93FC46FE09C1E9" box="[303,430,934,957]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="356" phylum="Annelida" rank="species" species="eremita">O. eremita</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The anterior parapodia of
|
||
<taxonomicName id="4C134D02FFB8FFA2FD48FC46FCDFC1E9" box="[756,888,934,957]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="356" phylum="Annelida" rank="species" species="pancerii">
|
||
<emphasis id="B967EA93FFB8FFA2FD48FC46FCDFC1E9" box="[756,888,934,957]" italics="true" pageId="11" pageNumber="356">O. pancerii</emphasis>
|
||
</taxonomicName>
|
||
have bi- and tridentate (or even multidentate) pseudocompound hooks of similar thickness and length of appendages. In contrast,
|
||
<taxonomicName id="4C134D02FFB8FFA2FBF7FC2AFB64C1B5" box="[1099,1219,970,993]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="356" phylum="Annelida" rank="species" species="eremita">
|
||
<emphasis id="B967EA93FFB8FFA2FBF7FC2AFB64C1B5" box="[1099,1219,970,993]" italics="true" pageId="11" pageNumber="356">O. eremita</emphasis>
|
||
</taxonomicName>
|
||
has only tridentate pseudocompound hooks of two
|
||
<typeStatus id="54A88823FFB8FFA2FE41FC0EFD9CC652" box="[509,571,1006,1030]" pageId="11" pageNumber="356">types</typeStatus>
|
||
, robust short-appendaged hooks and slender long-appendaged ones.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309650AFFB8FFA2FF7BFBF1FBD4C67E" box="[199,1139,1041,1066]" pageId="11" pageNumber="356" type="distribution">
|
||
<paragraph id="8BAC3681FFB8FFA2FF7BFBF1FBD4C67E" blockId="11.[151,1437,249,1066]" box="[199,1139,1041,1066]" pageId="11" pageNumber="356">
|
||
<emphasis id="B967EA93FFB8FFA2FF7BFBF1FEC7C67E" bold="true" box="[199,352,1041,1066]" pageId="11" pageNumber="356">Distribution.</emphasis>
|
||
East Atlantic (Bay of Biscay) and western and central Mediterranean.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |