597 lines
36 KiB
XML
597 lines
36 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.845.33566" ID-GBIF-Dataset="93d412c8-defe-4417-a5a4-f0b978d55e3f" ID-PMC="PMC6535459" ID-Pensoft-Pub="1313-2970-845-99" ID-PubMed="31156329" ID-ZBK="2C4523C4409749048797D03F55DD47D9" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-845-99" ModsDocOrigin="ZooKeys 845" ModsDocTitle="Two new species of the genus Symphylella (Symphyla, Scolopendrellidae) from Tibet, China" checkinTime="1557997603692" checkinUser="pensoft" docAuthor="Jin, Ya-Li, Bu, Yun & Jiang, Yue" docDate="2019" docId="F8E6F69223EF4DC8D49CBF8460C34AE4" docLanguage="en" docName="ZooKeys 845: 99-117" docOrigin="ZooKeys 845" docSource="http://dx.doi.org/10.3897/zookeys.845.33566" docTitle="Symphylella zhongi Jin & Bu, sp. n." docType="treatment" docUuid="159AD15C-EB70-409D-8E5F-3A1E25B62C22" docUuidSource="ZooBank" docVersion="6" lastPageNumber="113" masterDocId="FFBECB349B6FFFC11442FF9DFFDCFFBE" masterDocTitle="Two new species of the genus Symphylella (Symphyla, Scolopendrellidae) from Tibet, China" masterLastPageNumber="117" masterPageNumber="99" pageNumber="106" updateTime="1668167255525" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of the genus Symphylella (Symphyla, Scolopendrellidae) from Tibet, China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jin, Ya-Li</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bu, Yun</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jiang, Yue</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>845</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>99</mods:start>
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<mods:end>117</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.845.33566</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.845.33566</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-845-99</mods:identifier>
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<mods:identifier type="ZBK">2C4523C4409749048797D03F55DD47D9</mods:identifier>
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<mods:identifier type="ZooBank">2C4523C4409749048797D03F55DD47D9</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="157167780" LSID="urn:lsid:zoobank.org:act:159AD15C-EB70-409D-8E5F-3A1E25B62C22" httpUri="http://treatment.plazi.org/id/F8E6F69223EF4DC8D49CBF8460C34AE4" lastPageId="14" lastPageNumber="113" pageId="7" pageNumber="106">
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<subSubSection pageId="7" pageNumber="106" type="nomenclature">
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<paragraph pageId="7" pageNumber="106">
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<taxonomicName LSID="http://zoobank.org/159AD15C-EB70-409D-8E5F-3A1E25B62C22" authority="Jin & Bu" class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella zhongi" order="Symphyla" pageId="7" pageNumber="106" phylum="Arthropoda" rank="species" species="zhongi">Symphylella zhongi Jin & Bu</taxonomicName>
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<taxonomicNameLabel pageId="7" pageNumber="106">sp. n.</taxonomicNameLabel>
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Figures 3, 4, Tables 4, 5, 6, 7
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="107" type="diagnosis">
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<paragraph pageId="8" pageNumber="107">
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<pageBreakToken pageId="8" pageNumber="107" start="start">Diagnosis</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="8" pageNumber="107">
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<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella zhongi" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="zhongi">Symphylella zhongi</taxonomicName>
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sp. n. is characterized by a globular
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<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
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organ with small and roundish opening, processes on tergites mostly longer than their basal width, ovoid swollen ends of processes, and mushroom-shaped sensory organs present on apical antennal segments.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="107" type="materials_examined">
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<paragraph pageId="8" pageNumber="107">Material Examined.</paragraph>
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<paragraph pageId="8" pageNumber="107">
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Holotype, female (slide no. XZ-SY2015049) (SNHM), China, Tibet, Linzhi City, Bomi county, Songzong town, extracted from soil samples from broadleaf forest, alt. 3000 m,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="30.266666">29°76'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="96.6">95°96'E</geoCoordinate>
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, 7-XI-2015, coll. Y. Bu & G. Yang. Paratypes, 2 females (slides nos. XZ-SY2015047-XZ-SY2015048) (SNHM), same date as holotype. Other materials: 3 juvenile with 8 or 9 pairs of legs (slides nos. XZ-SY2015050-XZ-SY2015052) (SNHM), same date as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="13" lastPageNumber="112" pageId="8" pageNumber="107" type="description">
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<paragraph pageId="8" pageNumber="107">Description.</paragraph>
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<paragraph pageId="8" pageNumber="107">Adult body 2.48 mm long on average (2.22-2.93 mm, n = 3), holotype 2.93 mm (Fig. 3A).</paragraph>
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<caption pageId="8" pageNumber="107">
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<paragraph pageId="8" pageNumber="107">
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Figure 3.
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<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella zhongi" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="zhongi">Symphylella zhongi</taxonomicName>
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sp. n. A habitus B right antenna, 8
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<normalizedToken originalValue="th–">th-</normalizedToken>
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11th segments, dorsal view (bo-bladder-shaped organ, co-cavity-shaped organ) C right
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<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
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organ D head, dorsal view E 1
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<normalizedToken originalValue="st–">st-</normalizedToken>
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2nd tergites F 3rd tergite G female gonopore, styli, and coxal sacs on base of leg 4 (arrows indicate styli) H 4
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<normalizedToken originalValue="th–">th-</normalizedToken>
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5th tergite I 8th tergite, right side J 14
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<normalizedToken originalValue="th–">th-</normalizedToken>
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15th tergites K leg 1 (arrow indicates the leg). Scale bars: 200
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(A); 20
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<normalizedToken originalValue="B–K">B-K</normalizedToken>
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).
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</paragraph>
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</caption>
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<paragraph pageId="8" pageNumber="107">
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Head length 250-262
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, width 262-287
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, with widest part somewhat behind the middle on a level with the points of articulation of mandibles. Central rod distinct in both anterior (65-70
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<normalizedToken originalValue="μm">μm</normalizedToken>
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) and posterior (75-90
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<normalizedToken originalValue="μm">μm</normalizedToken>
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) portions, with a middle knot. Anterior branches well developed, median branches vestigial. Dorsal side of head moderately covered with setae of different length, longest setae (27-35
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<normalizedToken originalValue="μm">μm</normalizedToken>
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) located most anterior on head, at least 3.0 times as long as central ones (8-11
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<normalizedToken originalValue="μm">μm</normalizedToken>
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). Cuticle at anterolateral part of head with rather coarse granulation.
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<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
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organ surrounded by fine granulation, other area with faint dense granulation (Fig. 3D).
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</paragraph>
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<paragraph pageId="8" pageNumber="107">
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<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
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organ globular, length19-24
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, width 16-22
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, about half of greatest diameter of 3rd antennal segments (40-42
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<normalizedToken originalValue="μm">μm</normalizedToken>
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), opening small and roundish (length 8
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, width 3-6
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<normalizedToken originalValue="μm">μm</normalizedToken>
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) (Figs 3C, D).
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</paragraph>
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<paragraph pageId="8" pageNumber="107">Mouthparts. Mandible with two fused lamellae and 11 teeth in total (Fig. 4A). First maxilla has two lobes, inner lobe with four hook-shaped teeth, palp bud-like with one distal point close to outer lobe (Fig. 4B). Anterior part of second maxilla with many small protuberances which carry one seta each, distal setae more thick and hard; posterior part with sparse setae. Cuticle of maxilla and labium covered with pubescence.</paragraph>
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<caption pageId="8" pageNumber="107">
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<paragraph pageId="8" pageNumber="107">
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Figure 4.
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<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella zhongi" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="zhongi">Symphylella zhongi</taxonomicName>
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sp. n. A mandible B first maxilla
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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18
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<normalizedToken originalValue="th–">th-</normalizedToken>
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21th segments of right antenna C dorsal view (bo-bladder-shaped organ, co-cavity-shaped organ, mo-mushroom-shaped organ, rso-rudimentary spined sensory organ, so-spined sensory organ) D ventral view E 1
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<normalizedToken originalValue="st–">st-</normalizedToken>
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3rd segments of right antenna, dorsal view F stylus on base of 5th leg G 12th leg H right cercus, dorsal view. Scale bars: 20
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<normalizedToken originalValue="A–E">A-E</normalizedToken>
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, G, H); 5
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(F).
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</paragraph>
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</caption>
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<paragraph lastPageId="11" lastPageNumber="110" pageId="9" pageNumber="108">
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<pageBreakToken pageId="9" pageNumber="108" start="start">Antennae</pageBreakToken>
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with 18-23 segments (left antenna with 19, right antenna with 21 in holotype), length 513-663
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(663
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<normalizedToken originalValue="μm">μm</normalizedToken>
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in holotype), about one fourth of body length. 1st segment cylindrical, greatest diameter somewhat wider than long (40-42
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, 23-40
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<normalizedToken originalValue="μm">μm</normalizedToken>
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), with 1 or 2 microsetae and 6 or 7 mesosetae in one whorl, longest
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<pageBreakToken pageId="10" pageNumber="109" start="start">seta</pageBreakToken>
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(20-22
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<normalizedToken originalValue="μm">μm</normalizedToken>
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) inserted at inner side and distinctly longer than outer ones (14-15
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<normalizedToken originalValue="μm">μm</normalizedToken>
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). 2nd segment wider (25-28
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<normalizedToken originalValue="μm">μm</normalizedToken>
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) than long (40
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<normalizedToken originalValue="μm">μm</normalizedToken>
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), with 7 or 8 setae evenly inserted around the antennal wall with interior setae (20-21
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<normalizedToken originalValue="μm">μm</normalizedToken>
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) slightly longer than exterior ones (14-15
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<normalizedToken originalValue="μm">μm</normalizedToken>
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). Chaetotaxy of 3rd segment similar to preceding ones. Setae
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<pageBreakToken pageId="11" pageNumber="110" start="start">on</pageBreakToken>
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basal segments longer and on distal segments shorter. Basal antennae segments with only primary whorl of setae, in middle and subapical segments several setae in secondary whorl. Four kinds of sensory organs on antenna: rso on dorsal side of most segments (Figs 4C, E); co on 7
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<normalizedToken originalValue="th–">th-</normalizedToken>
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19th segments; bo on 9-11 segments next to apical one increasing in number on subdistal segments to a maximum of 21 (Figs 3B, 4C); mo on distal 2 segments (Fig. 4C). Apical segment subspherical, somewhat longer than wide (width 24-27
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, length 32-35
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<normalizedToken originalValue="μm">μm</normalizedToken>
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), with 15-18 setae on distal half; 3-5 spined sensory organs consisting of 3 or 4 curved spines around a central pillar in depressions in distal surface; 2 baculiform organs on apex of segment (Figs 4C, D). All segment covered with short pubescence. Chaetotaxy and sensory organs of antennae are given in Table 4.
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</paragraph>
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<caption pageId="11" pageNumber="110">
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<paragraph pageId="11" pageNumber="110">Table 4. Numbers of setae and sensory organs of antennae (holotype).</paragraph>
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</caption>
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<paragraph pageId="11" pageNumber="110">
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<table pageId="11" pageNumber="110">
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">Segments</td>
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">No. of primary whorl setae</td>
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">No. of secondary whorl setae</td>
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">Rudimentary spined sensory organs (rso)</td>
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">Cavity-shaped organs on dorsal side (co)</td>
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<td colspan="2" pageId="11" pageNumber="110" rowspan="1">Bladder-shaped organs (bo)</td>
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">Mushroom-shaped organs (mo)</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">Dorsal</td>
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">Ventral</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">st</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">nd</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">rd</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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<tr pageId="11" pageNumber="110">
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<td colspan="1" pageId="11" pageNumber="110" rowspan="1">th</td>
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</tr>
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</table>
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</paragraph>
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<paragraph lastPageId="12" lastPageNumber="111" pageId="11" pageNumber="110">
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Trunk with 17 dorsal tergites. Most tergites overlap, with intertergal zones present between adjacent tergites, except for borders between 14th and 15th, as well as 16th and 17th tergite. Tergites 2-13 and 15 each with one pair of triangular processes. Length from base to tip of processes somewhat longer than its basal width except for the 4th, 7th, 10th and 13th tergites, where processes are broader than long; basal distance between processes of 4
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<normalizedToken originalValue="th–">th-</normalizedToken>
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13th and 15th tergite longer than their length from base to tip (Table 5). Triangular processes with ovoid swollen ends (Fig. 3I). Anterolateral setae of 2nd, 3rd, 4th, 6th, 7th, 9th and10th tergite dis
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<pageBreakToken pageId="12" pageNumber="111" start="start">tinctly</pageBreakToken>
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longer than other lateromarginal setae, that of 5th, 8th, 11th -13th and 15th subequal or slightly longer than other lateromarginal. One to two inserted setae. All tergites pubescent.
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</paragraph>
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<caption pageId="12" pageNumber="111">
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<paragraph pageId="12" pageNumber="111">
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Table 5. Measurements of tergites and processes (in
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<normalizedToken originalValue="μm">μm</normalizedToken>
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, n = 3) (holotype in brackets).
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</paragraph>
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</caption>
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<paragraph pageId="12" pageNumber="111">
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<table pageId="12" pageNumber="111">
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<tr pageId="12" pageNumber="111">
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<td colspan="1" pageId="12" pageNumber="111" rowspan="1">No. of tergites</td>
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<td colspan="1" pageId="12" pageNumber="111" rowspan="1">Length</td>
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<td colspan="1" pageId="12" pageNumber="111" rowspan="1">Width</td>
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<td colspan="1" pageId="12" pageNumber="111" rowspan="1">Length of processes</td>
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<td colspan="1" pageId="12" pageNumber="111" rowspan="1">Basal width of processes</td>
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<td colspan="1" pageId="12" pageNumber="111" rowspan="1">Basal distance between processes</td>
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</tr>
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<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">st</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">nd</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">rd</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph pageId="12" pageNumber="111">
|
||
Tergites. 1st tergite reduced and build up of two short plates separated in the median axis of the body, with 8 short setae in a row. 2nd tergite complete, with two slender posterior processes, 6-8 lateromarginal setae, 1 or 2 inserted setae, 2 central setae, with anterolateral setae distinctly longer than other lateromarginal ones, processes approximately 1.1 times as long as broad, basal distance between processes approximately 0.8 times as long as their length (Fig. 3E). 3rd tergite complete, broader and longer than preceding one with ratios of 1.1 and 0.9 respectively, 8 or 9 lateromarginal setae (Fig. 3F). 4th tergite broader than 3rd tergite, with ratios approximately 0.8 and 1.5 respectively, 6 or 7 lateromarginal setae (Fig. 3H). Chaetotaxy of 5
|
||
<normalizedToken originalValue="th–">th-</normalizedToken>
|
||
7th, 8
|
||
<normalizedToken originalValue="th–">th-</normalizedToken>
|
||
10th, and 11
|
||
<normalizedToken originalValue="th–">th-</normalizedToken>
|
||
13th tergites similar to 2
|
||
<normalizedToken originalValue="nd–">nd-</normalizedToken>
|
||
4th tergites. Pattern of alternating tergite lengths of two short tergites followed by a long tergite only disrupted at the caudal end (Table 5). Within short tergites (1, 2, 4, 5, 7, 8, 10, 11, 13, 14, 16) the length augments toward caudal. Same is generally true for long tergites (3, 6, 9, 12, 15, 17), but for the 15th tergite being shorter than the others, with likewise smaller processes. 14th and 16th tergite without processes with 13-18 and 10-15 marginal setae respectively (Fig. 3J). 17th tergite with 32-44 setae. Chaetotaxy and measurements of tergites are given in Tables 5 and 6.
|
||
</paragraph>
|
||
<caption pageId="12" pageNumber="111">
|
||
<paragraph pageId="12" pageNumber="111">Table 6. Chaetotaxy of tergites (holotype in brackets).</paragraph>
|
||
</caption>
|
||
<paragraph pageId="12" pageNumber="111">
|
||
<table pageId="12" pageNumber="111">
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">No. of tergites</td>
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">lateromarginal</td>
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">Inserted seta</td>
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">Central setae</td>
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">Other setae</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">st</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">nd</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">rd</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
<tr pageId="12" pageNumber="111">
|
||
<td colspan="1" pageId="12" pageNumber="111" rowspan="1">th</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph lastPageId="13" lastPageNumber="112" pageId="12" pageNumber="111">
|
||
Legs. 1st pair of legs reduced to two small hairy cupules, each with two long setae (Fig. 3G). Basal areas of legs 2-12 each with 5-7 setae. Leg 12 about 0.1 times the length of the body, about same length like head. Trochanter distinctly longer than wide (70-83
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
, 40-50
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
), with 7 or 8 subequal setae. Femur as long as wide (38-43
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
,
|
||
<pageBreakToken pageId="13" pageNumber="112" start="start">35</pageBreakToken>
|
||
-40
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
), with 5 setae and one of them (22
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
) distinctly longer than others (12-18
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
); trochanter and femur pubescent dorsally, ventrally with cuticular thickenings in pattern of large scales. Tibia nearly 2 times longer than wide (50-60
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
, 27-30
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
), with 6 dorsal setae and longest one nearly the same length as greatest diameter of tibia (21-30
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
). Tarsus subcylindrical, about 3.5 times as long as wide (68-75
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
, 20
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
) with 5 dorsal setae: 3 straight and protruding, 2 curved and depressed; longest setae (20-22
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
) about same length of greatest width of podomere; 2 ventral setae inserted near claw distinctly shorter than dorsal ones. Claws curved, anterior one somewhat longer and broader than posterior one, the latter more curved than the former (Fig. 4G). All legs covered with dense pubescences except areas with cuticular thickenings.
|
||
</paragraph>
|
||
<paragraph pageId="13" pageNumber="112">
|
||
Coxal sacs present at bases of 3
|
||
<normalizedToken originalValue="rd–">rd-</normalizedToken>
|
||
9th pairs of legs, fully developed, each with 4 setae on surface (Fig. 3G).
|
||
</paragraph>
|
||
<paragraph pageId="13" pageNumber="112">
|
||
Styli present at base of 3
|
||
<normalizedToken originalValue="rd–">rd-</normalizedToken>
|
||
12th pairs of legs, subconical (length 5-8
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
, width 3-4
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
), basal part with straight hairs; distal quarter hairless and blunt (3-4
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
) (Figs 3G, 4F).
|
||
</paragraph>
|
||
<paragraph pageId="13" pageNumber="112">
|
||
<taxonomicName genus="Sense" lsidName="Sense calicles" pageId="13" pageNumber="112" rank="species" species="calicles">Sense calicles</taxonomicName>
|
||
with smooth margin to pit, about same length as outer diameter (28-30
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
, 26-30
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
). Sensory seta inserted in cup center, extremely long, length 170-180
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
, at least 5.5 times longer than other two lateral setae (20-24
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
, 13-20
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
respectively) inserted at edge of cup.
|
||
</paragraph>
|
||
<paragraph pageId="13" pageNumber="112">
|
||
Cerci subuliform, about 0.7 of head length and leg 12, length at least three times as long as its greatest width (150-188
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
, 45-58
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
), moderately covered with subequal length and slightly curved setae, with longest (25-28
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
) at least half of greatest width of cerci, terminal area (24-28
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
) short, circled by 6-8 layers of curved ridges. Terminal setae length 25
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(Fig. 4H).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="113" type="etymology">
|
||
<paragraph pageId="14" pageNumber="113">
|
||
<pageBreakToken pageId="14" pageNumber="113" start="start">Etymology</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="14" pageNumber="113">We dedicate this new species in honor of the late Professor Zhong Yang (1964-2017) who was an eminent botanist from Fudan University, for his great contribution to the knowledge of flora and biodiversity of Tibet.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="113" type="distribution">
|
||
<paragraph pageId="14" pageNumber="113">Distribution.</paragraph>
|
||
<paragraph pageId="14" pageNumber="113">Known only from the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="113" type="remarks">
|
||
<paragraph pageId="14" pageNumber="113">Remarks.</paragraph>
|
||
<paragraph pageId="14" pageNumber="113">
|
||
<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella zhongi" pageId="14" pageNumber="113" phylum="Arthropoda" rank="species" species="zhongi">Symphylella zhongi</taxonomicName>
|
||
sp. n. is most similar to
|
||
<taxonomicName lsidName="S. multisetosa" pageId="14" pageNumber="113" rank="species" species="multisetosa">S. multisetosa</taxonomicName>
|
||
Scheller, 1971 in the shape of the
|
||
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
|
||
organ, as well as the shape and chaetotaxy of the tergites 1-4, 1st leg, but it deviates distinctly in the shape of the ends of the processes (with ovoid swollen ends in
|
||
<taxonomicName lsidName="S. zhongi" pageId="14" pageNumber="113" rank="species" species="zhongi">S. zhongi</taxonomicName>
|
||
sp. n. vs without ovoid swollen ends in
|
||
<taxonomicName lsidName="S. multisetosa" pageId="14" pageNumber="113" rank="species" species="multisetosa">S. multisetosa</taxonomicName>
|
||
), chaetotaxy of the cerci (most setae subequal length and slightly curved in
|
||
<taxonomicName lsidName="S. zhongi" pageId="14" pageNumber="113" rank="species" species="zhongi">S. zhongi</taxonomicName>
|
||
sp. n. vs long, straight, erect setae on dorsal, ventral, and outer sides of cerci arranged in one or two rows in
|
||
<taxonomicName lsidName="S. multisetosa" pageId="14" pageNumber="113" rank="species" species="multisetosa">S. multisetosa</taxonomicName>
|
||
). It is also similar to
|
||
<taxonomicName lsidName="S. simplex" pageId="14" pageNumber="113" rank="species" species="simplex">S. simplex</taxonomicName>
|
||
(Hansen, 1903) in the shape and chaetotaxy of first two tergites, sensory organs of antennal segments and 1st leg, but differs in the shape of 1stantennal segment (moderate in
|
||
<taxonomicName lsidName="S. zhongi" pageId="14" pageNumber="113" rank="species" species="zhongi">S. zhongi</taxonomicName>
|
||
sp. n. vs very short in
|
||
<taxonomicName lsidName="S. simplex" pageId="14" pageNumber="113" rank="species" species="simplex">S. simplex</taxonomicName>
|
||
), chaetatoxy of 3rd tergite (8 or 9 lateromarginal setae in
|
||
<taxonomicName lsidName="S. zhongi" pageId="14" pageNumber="113" rank="species" species="zhongi">S. zhongi</taxonomicName>
|
||
sp. n. vs 11 or 12 in
|
||
<taxonomicName lsidName="S. simplex" pageId="14" pageNumber="113" rank="species" species="simplex">S. simplex</taxonomicName>
|
||
) and 4th tergite (6 or 7 lateromarginal setae in
|
||
<taxonomicName lsidName="S. zhongi" pageId="14" pageNumber="113" rank="species" species="zhongi">S. zhongi</taxonomicName>
|
||
sp. n. vs 8 in
|
||
<taxonomicName lsidName="S. simplex" pageId="14" pageNumber="113" rank="species" species="simplex">S. simplex</taxonomicName>
|
||
), chaetotaxy of cerci (8-10 longish setae protruding, others short and depressed in
|
||
<taxonomicName lsidName="S. simplex" pageId="14" pageNumber="113" rank="species" species="simplex">S. simplex</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="14" pageNumber="113">The new species is compared with similar species in Table 7.</paragraph>
|
||
<caption ID-Table-UUID="712A2424BDAE8B52A8C2DD4BDB66E316" httpUri="http://table.plazi.org/id/712A2424BDAE8B52A8C2DD4BDB66E316" pageId="14" pageNumber="113">
|
||
<paragraph pageId="14" pageNumber="113">
|
||
Table 7. Comparison of
|
||
<taxonomicName lsidName="S. macropora" pageId="14" pageNumber="113" rank="species" species="macropora">S. macropora</taxonomicName>
|
||
sp. n.,
|
||
<taxonomicName lsidName="S. zhongi" pageId="14" pageNumber="113" rank="species" species="zhongi">S. zhongi</taxonomicName>
|
||
sp. n. and the similar species.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="14" pageNumber="113">
|
||
<table pageId="14" pageNumber="113">
|
||
<tr pageId="14" pageNumber="113">
|
||
<td colspan="1" pageId="14" pageNumber="113" rowspan="1">Characters</td>
|
||
<td colspan="1" pageId="14" pageNumber="113" rowspan="1">
|
||
<taxonomicName lsidName="S. macropora" pageId="14" pageNumber="113" rank="species" species="macropora">S. macropora</taxonomicName>
|
||
sp. n.
|
||
</td>
|
||
<td colspan="1" pageId="14" pageNumber="113" rowspan="1">
|
||
<taxonomicName lsidName="S. javanens" pageId="14" pageNumber="113" rank="species" species="javanens">S. javanens</taxonomicName>
|
||
</td>
|
||
<td colspan="1" pageId="14" pageNumber="113" rowspan="1">
|
||
<taxonomicName lsidName="S. asiatica" pageId="14" pageNumber="113" rank="species" species="asiatica">S. asiatica</taxonomicName>
|
||
</td>
|
||
<td colspan="1" pageId="14" pageNumber="113" rowspan="1">
|
||
<taxonomicName lsidName="S. zhongi" pageId="14" pageNumber="113" rank="species" species="zhongi">S. zhongi</taxonomicName>
|
||
sp. n.
|
||
</td>
|
||
<td colspan="1" pageId="14" pageNumber="113" rowspan="1">
|
||
<taxonomicName lsidName="S. multisetosa" pageId="14" pageNumber="113" rank="species" species="multisetosa">S. multisetosa</taxonomicName>
|
||
</td>
|
||
<td colspan="1" pageId="14" pageNumber="113" rowspan="1">
|
||
<taxonomicName lsidName="S. simplex" pageId="14" pageNumber="113" rank="species" species="simplex">S. simplex</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph pageId="14" pageNumber="113">
|
||
The 43 species of the genus
|
||
<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella" pageId="14" pageNumber="113" phylum="Arthropoda" rank="genus">Symphylella</taxonomicName>
|
||
can be distinguished by the following key, but six species (
|
||
<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella natala" pageId="14" pageNumber="113" phylum="Arthropoda" rank="species" species="natala">Symphylella natala</taxonomicName>
|
||
Hilton, 1938,
|
||
<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella vaca" pageId="14" pageNumber="113" phylum="Arthropoda" rank="species" species="vaca">Symphylella vaca</taxonomicName>
|
||
Hilton, 1938,
|
||
<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella elongata" pageId="14" pageNumber="113" phylum="Arthropoda" rank="species" species="elongata">Symphylella elongata</taxonomicName>
|
||
Scheller, 1952,
|
||
<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella foucquei" pageId="14" pageNumber="113" phylum="Arthropoda" rank="species" species="foucquei">Symphylella foucquei</taxonomicName>
|
||
Jupeau, 1954,
|
||
<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella maorica" pageId="14" pageNumber="113" phylum="Arthropoda" rank="species" species="maorica">Symphylella maorica</taxonomicName>
|
||
Adam & Burtel, 1956, and
|
||
<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella" pageId="14" pageNumber="113" phylum="Arthropoda" rank="genus">Symphylella</taxonomicName>
|
||
sp. Rochaix, 1956), which have very brief original descriptions, are not included.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |