343 lines
57 KiB
XML
343 lines
57 KiB
XML
<document id="4D0A738EA86C1ED3A35441F7F505094F" ID-DOI="10.5281/zenodo.195647" ID-GBIF-Dataset="5dbc17e2-43e6-4ca4-9921-56eb43a9aec3" ID-ISSN="1175-5326" ID-Zenodo-Dep="195647" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460231312424" checkinUser="plazi" docAuthor="Zaragoza, Juan A." docDate="2010" docId="0386879EFFA8B868FF5F72519F65FDF4" docLanguage="en" docName="zt02491p060.pdf" docOrigin="Zootaxa 2491" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Arcanobisium comasi Zaragoza, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="48" masterDocId="FFBFFFE6FFABB86FFFC8743C9C0FFA23" masterDocTitle="Arcanobisium, a remarkable new genus, representing a new subfamily with a relictual distribution from eastern Spain (Arachnida: Pseudoscorpiones: Syarinidae)" masterLastPageNumber="60" masterPageNumber="41" pageNumber="44" updateTime="1698239000673" updateUser="plazi">
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<mods:title id="F16239007044EEC8252694123F850986">Arcanobisium, a remarkable new genus, representing a new subfamily with a relictual distribution from eastern Spain (Arachnida: Pseudoscorpiones: Syarinidae)</mods:title>
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<mods:name id="D2729B82A2EA7EC4D9E19C3A398C5EA7" type="personal">
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<mods:namePart id="C99548BCE83F28D79A5E65ED7BC82494">Zaragoza, Juan A.</mods:namePart>
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<mods:title id="D0A32446869809577E2440B84FC6AC34">Zootaxa</mods:title>
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<mods:part id="C5D6096CAA5BF64147373BFBB2DBF102">
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<mods:date id="A92A7EEFCE72D74310BC03820AB07473">2010</mods:date>
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<mods:number id="A65C821A3AC7A9154427351FB51956AD">2491</mods:number>
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<mods:identifier id="A4E925522CBA2A6C1177D8B5D18698E9" type="DOI">10.5281/zenodo.195647</mods:identifier>
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<treatment id="0386879EFFA8B868FF5F72519F65FDF4" ID-DOI="http://doi.org/10.5281/zenodo.6208433" ID-GBIF-Taxon="119387336" ID-Zenodo-Dep="6208433" LSID="urn:lsid:plazi:treatment:0386879EFFA8B868FF5F72519F65FDF4" httpUri="http://treatment.plazi.org/id/0386879EFFA8B868FF5F72519F65FDF4" lastPageId="7" lastPageNumber="48" pageId="3" pageNumber="44">
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<subSubSection id="C3356503FFA8B86CFF5F72519D11FC89" pageId="3" pageNumber="44" type="nomenclature">
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<paragraph id="8B903688FFA8B86CFF5F72519E0DFCA4" blockId="3.[151,514,1645,1706]" box="[151,514,1645,1671]" pageId="3" pageNumber="44">
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<heading id="D0D881E4FFA8B86CFF5F72519E0DFCA4" bold="true" box="[151,514,1645,1671]" fontSize="11" level="1" pageId="3" pageNumber="44" reason="1">
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<emphasis id="B95BEA9AFFA8B86CFF5F72519E0DFCA4" bold="true" box="[151,514,1645,1671]" pageId="3" pageNumber="44">
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<taxonomicName id="4C2F4D0BFFA8B86CFF5F72519D91FCA4" ID-CoL="G7CN" box="[151,414,1645,1671]" class="Arachnida" family="Syarinidae" genus="Arcanobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="comasi" status="sp. nov.">
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<emphasis id="B95BEA9AFFA8B86CFF5F72519D91FCA4" bold="true" box="[151,414,1645,1671]" italics="true" pageId="3" pageNumber="44">Arcanobisium comasi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26857E1FFA8B86CFE6D72519E0DFCA4" box="[421,514,1645,1671]" pageId="3" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B903688FFA8B86CFF5F72AC9D11FC89" blockId="3.[151,514,1645,1706]" box="[151,286,1680,1706]" pageId="3" pageNumber="44">
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(
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<figureCitation id="13142A0DFFA8B86CFF5772AC9D19FC89" box="[159,278,1680,1706]" captionStart-0="FIGURES 2 – 3" captionStart-1="FIGURES 4 – 12" captionStart-2="FIGURES 13 – 18" captionStart-3="FIGURES 19 – 25" captionStartId-0="4.[151,269,1024,1048]" captionStartId-1="5.[151,271,1882,1906]" captionStartId-2="6.[151,269,1871,1895]" captionStartId-3="8.[151,270,1883,1907]" captionTargetBox-0="[210,1376,200,995]" captionTargetBox-1="[220,1300,190,1843]" captionTargetBox-2="[274,1377,205,1833]" captionTargetBox-3="[152,1397,186,1882]" captionTargetId-0="figure@4.[207,1379,197,1000]" captionTargetId-1="figure@5.[185,1400,179,1853]" captionTargetId-2="figure@6.[186,1410,194,1857]" captionTargetId-3="figure@8.[150,1435,178,1902]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionText-0="FIGURES 2 – 3. Arcanobisium comasi sp. nov., male holotype. 2. habitus, dorsal view; 3. habitus, ventral view." captionText-1="FIGURES 4 – 12. Arcanobisium comasi sp. nov., male holotype. 4. carapace; 5. anterior margin of carapace, showing epistome; 6. right chelicera; 7. partial view of fingers of right chelicera; 8. cheliceral rallum; 9. right leg I, lateral view; 10. right leg IV, lateral view; 11. distal end of tarsus and apotele of right leg IV, lateral view; 12. subterminal seta of tarsus right leg IV, partial view. Abbreviations: P glandular pore; TS tactile seta. Scale bars (mm): 0.01 (Fig. 12), 0.05 (Fig. 5, 7, 8, 11), 0.10 (Fig. 6), 0.20 (Fig. 4, 9, 10)." captionText-2="FIGURES 13 – 18. Arcanobisium comasi sp. nov., male holotype. 13. right pedipalp, without chela, dorsal view; 14. right chela, dorsal view; 15. right chela, antiaxial view; 16. tips of fixed and movable finger of right chela, antiaxial view; 17. tips of fixed and movable finger of right chela, dorsal view; 18. trichobothrium t. Abbreviations: af? pore or sensillum; pc doubled pore sensillum (other abbreviations correspond to trichobothria). Scale bars (mm): 0.05 (Fig. 18), 0.10 (Figs 16 – 17), 0.20 (Figs 13 – 15)." captionText-3="FIGURES 19 – 25. Arcanobisium comasi sp. nov., male holotype. 19. coxae of pedipalp and leg I; 20. genital sternites, ventral; 21. genitalia, ventral; 22. stigmata of sternites III and IV; 23. tergites-sternites VIII – XII, posterior view, drawing in perspective, entire setae only illustrated in sternites X – XI; 24. tergites-sternites VIII – XII, lateral view; 25. habitus, lateral view; pedipalps, legs and one chelicera omitted. Abbreviations: I – XII segments I – XII of opisthosoma. Scale bars (mm): 0.05 (Fig. 21), 0.10 (Figs 20, 22, 24, 25), 0.20 (Figs 19, 23)." httpUri-0="https://zenodo.org/record/195649/files/figure.png" httpUri-1="https://zenodo.org/record/195650/files/figure.png" httpUri-2="https://zenodo.org/record/195651/files/figure.png" httpUri-3="https://zenodo.org/record/195652/files/figure.png" pageId="3" pageNumber="44">Figs 2–25</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3356503FFA8B86CFF5F72E69F99FD1C" pageId="3" pageNumber="44" type="materials_examined">
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<paragraph id="8B903688FFA8B86CFF5F72E69F99FD1C" blockId="3.[151,1436,1754,1967]" pageId="3" pageNumber="44">
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<emphasis id="B95BEA9AFFA8B86CFF5F72E69D40FCD7" bold="true" box="[151,335,1754,1780]" pageId="3" pageNumber="44">
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<typeStatus id="5494882AFFA8B86CFF5F72E69CDCFCD7" box="[151,211,1754,1780]" pageId="3" pageNumber="44">Type</typeStatus>
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material.
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</emphasis>
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Male
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<typeStatus id="5494882AFFA8B86CFE5772E69E03FCD7" box="[415,524,1754,1780]" pageId="3" pageNumber="44" type="holotype">holotype</typeStatus>
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:
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<collectingCountry id="F3387618FFA8B86CFDD072E69E50FCD7" box="[536,607,1754,1780]" name="Spain" pageId="3" pageNumber="44">Spain</collectingCountry>
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, Valencian Community, Castellón province, La Plana Alta, Cabanes, Avenc d’en Serenge, UTM coordinates: 31TBE501469, altitude
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<quantity id="4CD79B6DFFA8B86CFC69733C9FFFFD39" box="[929,1008,1792,1818]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="3" pageNumber="44" unit="m" value="305.0">305 m</quantity>
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,
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<date id="FF911048FFA8B86CFC35733C98B7FD39" box="[1021,1208,1792,1818]" pageId="3" pageNumber="44" value="1982-03-14">14 March 1982</date>
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, collected by Jordi Comas, deposited in
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<collectionCode id="ED3EAE4DFFA8B86CFE4473199DE8FD1C" box="[396,487,1829,1855]" httpUri="http://grbio.org/cool/fs2t-231f" name="Museu de Ciències Naturals de Barcelona" pageId="3" pageNumber="44">MCNB</collectionCode>
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: register number: 2009 - 4118
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<collectionCode id="ED3EAE4DFFA8B86CFC8673199F9EFD1C" box="[846,913,1829,1855]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="44">MZB</collectionCode>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3356503FFA8B86CFF0C73769CD3FD8C" pageId="3" pageNumber="44" type="etymology">
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<paragraph id="8B903688FFA8B86CFF0C73769CD3FD8C" blockId="3.[151,1436,1754,1967]" pageId="3" pageNumber="44">
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<emphasis id="B95BEA9AFFA8B86CFF0C73769D5DFD47" bold="true" box="[196,338,1866,1892]" pageId="3" pageNumber="44">Etymology.</emphasis>
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The species is named after the biospeleologist Jordi Comas, from Barcelona, who collected the unique specimen and to honour his work collecting and studying cave-dwelling arthropods for the past 40 years.
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</paragraph>
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</subSubSection>
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<caption id="DF506600FFAFB86BFF5F703C9933FE34" box="[151,1340,1024,1048]" httpUri="https://zenodo.org/record/195649/files/figure.png" pageId="4" pageNumber="45" targetBox="[210,1376,200,995]" targetPageId="4">
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<paragraph id="8B903688FFAFB86BFF5F703C9933FE34" blockId="4.[151,1340,1024,1048]" box="[151,1340,1024,1048]" pageId="4" pageNumber="45">
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<emphasis id="B95BEA9AFFAFB86BFF5F703C9D4CFE3B" bold="true" box="[151,323,1024,1048]" pageId="4" pageNumber="45">FIGURES 2–3.</emphasis>
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<taxonomicName id="4C2F4D0BFFAFB86BFE81703C9E3FFE34" box="[329,560,1024,1047]" class="Arachnida" family="Syarinidae" genus="Arcanobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="comasi" status="sp. nov.">
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<emphasis id="B95BEA9AFFAFB86BFE81703C9E3FFE34" box="[329,560,1024,1047]" italics="true" pageId="4" pageNumber="45">Arcanobisium comasi</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B95BEA9AFFAFB86BFDFF703C9E83FE3B" bold="true" box="[567,652,1024,1048]" pageId="4" pageNumber="45">
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<taxonomicNameLabel id="A26857E1FFAFB86BFDFF703C9E83FE3B" box="[567,652,1024,1048]" pageId="4" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, male holotype. 2. habitus, dorsal view; 3. habitus, ventral view.
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</paragraph>
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</caption>
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<subSubSection id="C3356503FFAFB868FF0C707E9D60FECF" lastPageId="7" lastPageNumber="48" pageId="4" pageNumber="45" type="description">
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<paragraph id="8B903688FFAFB86BFF0C707E9916FEA1" blockId="4.[151,1437,1090,1976]" pageId="4" pageNumber="45">
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<emphasis id="B95BEA9AFFAFB86BFF0C707E9D53FE7F" bold="true" box="[196,348,1090,1116]" pageId="4" pageNumber="45">Description.</emphasis>
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A small pseudoscorpion (body length
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<quantity id="4CD79B6DFFAFB86BFCE8707F9F81FE7E" box="[800,910,1091,1117]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" pageId="4" pageNumber="45" unit="mm" value="1.64">1.64 mm</quantity>
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); legs pale, carapace, opisthosomal segments and pedipalps pale brown in colour and showing in general a weak polygonal or scaly reticulation.
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</paragraph>
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<paragraph id="8B903688FFAFB86BFF0E70B29F45FED1" blockId="4.[151,1437,1090,1976]" pageId="4" pageNumber="45">
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Carapace (
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<figureCitation id="13142A0DFFAFB86BFE8C70B29D80FE8B" box="[324,399,1166,1192]" captionStart="FIGURES 4 – 12" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[220,1300,190,1843]" captionTargetId="figure@5.[185,1400,179,1853]" captionTargetPageId="5" captionText="FIGURES 4 – 12. Arcanobisium comasi sp. nov., male holotype. 4. carapace; 5. anterior margin of carapace, showing epistome; 6. right chelicera; 7. partial view of fingers of right chelicera; 8. cheliceral rallum; 9. right leg I, lateral view; 10. right leg IV, lateral view; 11. distal end of tarsus and apotele of right leg IV, lateral view; 12. subterminal seta of tarsus right leg IV, partial view. Abbreviations: P glandular pore; TS tactile seta. Scale bars (mm): 0.01 (Fig. 12), 0.05 (Fig. 5, 7, 8, 11), 0.10 (Fig. 6), 0.20 (Fig. 4, 9, 10)." httpUri="https://zenodo.org/record/195650/files/figure.png" pageId="4" pageNumber="45">Fig. 4</figureCitation>
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). Elongated, maximum width at posterior margin. Without eyes or eye-spots. Epistome small and rounded (
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<figureCitation id="13142A0DFFAFB86BFE48708F9DC7FEEE" box="[384,456,1203,1229]" captionStart="FIGURES 4 – 12" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[220,1300,190,1843]" captionTargetId="figure@5.[185,1400,179,1853]" captionTargetPageId="5" captionText="FIGURES 4 – 12. Arcanobisium comasi sp. nov., male holotype. 4. carapace; 5. anterior margin of carapace, showing epistome; 6. right chelicera; 7. partial view of fingers of right chelicera; 8. cheliceral rallum; 9. right leg I, lateral view; 10. right leg IV, lateral view; 11. distal end of tarsus and apotele of right leg IV, lateral view; 12. subterminal seta of tarsus right leg IV, partial view. Abbreviations: P glandular pore; TS tactile seta. Scale bars (mm): 0.01 (Fig. 12), 0.05 (Fig. 5, 7, 8, 11), 0.10 (Fig. 6), 0.20 (Fig. 4, 9, 10)." httpUri="https://zenodo.org/record/195650/files/figure.png" pageId="4" pageNumber="45">Fig. 5</figureCitation>
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). Chaetotaxy: 21 setae, formula: 4:4:4:3:6. Two glandular pores on each side of the anterior third. Three microlyrifissures on the anterior zone.
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</paragraph>
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<paragraph id="8B903688FFAFB86BFF0E70C29FAFFF41" blockId="4.[151,1437,1090,1976]" pageId="4" pageNumber="45">
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Coxal area (
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<figureCitation id="13142A0DFFAFB86BFE9070C29DBEFF3B" box="[344,433,1278,1304]" captionStart="FIGURES 19 – 25" captionStartId="8.[151,270,1883,1907]" captionTargetBox="[152,1397,186,1882]" captionTargetId="figure@8.[150,1435,178,1902]" captionTargetPageId="8" captionText="FIGURES 19 – 25. Arcanobisium comasi sp. nov., male holotype. 19. coxae of pedipalp and leg I; 20. genital sternites, ventral; 21. genitalia, ventral; 22. stigmata of sternites III and IV; 23. tergites-sternites VIII – XII, posterior view, drawing in perspective, entire setae only illustrated in sternites X – XI; 24. tergites-sternites VIII – XII, lateral view; 25. habitus, lateral view; pedipalps, legs and one chelicera omitted. Abbreviations: I – XII segments I – XII of opisthosoma. Scale bars (mm): 0.05 (Fig. 21), 0.10 (Figs 20, 22, 24, 25), 0.20 (Figs 19, 23)." httpUri="https://zenodo.org/record/195652/files/figure.png" pageId="4" pageNumber="45">Fig. 19</figureCitation>
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). Apex of pedipalpal coxa triangular, long and acute, with 2 distal setae. Pedipalpal coxa with 7–8 setae, pedal coxa I: 5, II: 4, III: 3, IV: 4. Anterolateral process of coxa I rounded and moderately prominent, with some granules at the apex; medial process absent.
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</paragraph>
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<paragraph id="8B903688FFAFB86BFF0E71529E86FCDE" blockId="4.[151,1437,1090,1976]" pageId="4" pageNumber="45">
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Opisthosoma. Ovate (
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<figureCitation id="13142A0DFFAFB86BFE0071529E3CFFAB" box="[456,563,1390,1416]" captionStart="FIGURES 2 – 3" captionStartId="4.[151,269,1024,1048]" captionTargetBox="[210,1376,200,995]" captionTargetId="figure@4.[207,1379,197,1000]" captionTargetPageId="4" captionText="FIGURES 2 – 3. Arcanobisium comasi sp. nov., male holotype. 2. habitus, dorsal view; 3. habitus, ventral view." httpUri="https://zenodo.org/record/195649/files/figure.png" pageId="4" pageNumber="45">Figs 2–3</figureCitation>
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), pleural membrane laterally striate and medially weakly granulated (
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<figureCitation id="13142A0DFFAFB86BFAA471529CB5FF8E" captionStart="FIGURES 19 – 25" captionStartId="8.[151,270,1883,1907]" captionTargetBox="[152,1397,186,1882]" captionTargetId="figure@8.[150,1435,178,1902]" captionTargetPageId="8" captionText="FIGURES 19 – 25. Arcanobisium comasi sp. nov., male holotype. 19. coxae of pedipalp and leg I; 20. genital sternites, ventral; 21. genitalia, ventral; 22. stigmata of sternites III and IV; 23. tergites-sternites VIII – XII, posterior view, drawing in perspective, entire setae only illustrated in sternites X – XI; 24. tergites-sternites VIII – XII, lateral view; 25. habitus, lateral view; pedipalps, legs and one chelicera omitted. Abbreviations: I – XII segments I – XII of opisthosoma. Scale bars (mm): 0.05 (Fig. 21), 0.10 (Figs 20, 22, 24, 25), 0.20 (Figs 19, 23)." httpUri="https://zenodo.org/record/195652/files/figure.png" pageId="4" pageNumber="45">Fig. 25</figureCitation>
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). Tergite I longer than other tergites. Only tergites and sternites I–IX visible dorsally and ventrally, respectively (
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<figureCitation id="13142A0DFFAFB86BFEFB71849D92FFF1" box="[307,413,1464,1490]" captionStart="FIGURES 2 – 3" captionStartId="4.[151,269,1024,1048]" captionTargetBox="[210,1376,200,995]" captionTargetId="figure@4.[207,1379,197,1000]" captionTargetPageId="4" captionText="FIGURES 2 – 3. Arcanobisium comasi sp. nov., male holotype. 2. habitus, dorsal view; 3. habitus, ventral view." httpUri="https://zenodo.org/record/195649/files/figure.png" pageId="4" pageNumber="45">Figs 2–3</figureCitation>
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,
|
||
<figureCitation id="13142A0DFFAFB86BFE6171849DF8FFF1" box="[425,503,1464,1490]" captionStart="FIGURES 19 – 25" captionStartId="8.[151,270,1883,1907]" captionTargetBox="[152,1397,186,1882]" captionTargetId="figure@8.[150,1435,178,1902]" captionTargetPageId="8" captionText="FIGURES 19 – 25. Arcanobisium comasi sp. nov., male holotype. 19. coxae of pedipalp and leg I; 20. genital sternites, ventral; 21. genitalia, ventral; 22. stigmata of sternites III and IV; 23. tergites-sternites VIII – XII, posterior view, drawing in perspective, entire setae only illustrated in sternites X – XI; 24. tergites-sternites VIII – XII, lateral view; 25. habitus, lateral view; pedipalps, legs and one chelicera omitted. Abbreviations: I – XII segments I – XII of opisthosoma. Scale bars (mm): 0.05 (Fig. 21), 0.10 (Figs 20, 22, 24, 25), 0.20 (Figs 19, 23)." httpUri="https://zenodo.org/record/195652/files/figure.png" pageId="4" pageNumber="45">23–25</figureCitation>
|
||
): tergal and sternal plates X–XI extremely reduced in size, difficult to recognize and only distinctly visible laterally or ventrally (
|
||
<figureCitation id="13142A0DFFAFB86BFD2F71E29F79FFDB" box="[743,886,1502,1528]" captionStart="FIGURES 19 – 25" captionStartId="8.[151,270,1883,1907]" captionTargetBox="[152,1397,186,1882]" captionTargetId="figure@8.[150,1435,178,1902]" captionTargetPageId="8" captionText="FIGURES 19 – 25. Arcanobisium comasi sp. nov., male holotype. 19. coxae of pedipalp and leg I; 20. genital sternites, ventral; 21. genitalia, ventral; 22. stigmata of sternites III and IV; 23. tergites-sternites VIII – XII, posterior view, drawing in perspective, entire setae only illustrated in sternites X – XI; 24. tergites-sternites VIII – XII, lateral view; 25. habitus, lateral view; pedipalps, legs and one chelicera omitted. Abbreviations: I – XII segments I – XII of opisthosoma. Scale bars (mm): 0.05 (Fig. 21), 0.10 (Figs 20, 22, 24, 25), 0.20 (Figs 19, 23)." httpUri="https://zenodo.org/record/195652/files/figure.png" pageId="4" pageNumber="45">Figs 23–25</figureCitation>
|
||
); tergite XI limited to a narrow band without setae, sternite XI divided into two lateral isolated diminutive plates, each bearing one seta. Sternites IV and V distinctly wider than the others. Tergal chaetotaxy I–XI: 8: 8: 10: 8: 8: 8: 7: 6: 6: 4: 0. Anal cone with 2 ventral and 2 dorsal setae. Genital area with 11 setae on sternite II plus 13 small setae on the anterior genital opening; sternite III with 11 setae, both medial setae thickened, 3 small setae on anterior margin (
|
||
<figureCitation id="13142A0DFFAFB86BFB0B724F9911FCAE" box="[1219,1310,1651,1677]" captionStart="FIGURES 19 – 25" captionStartId="8.[151,270,1883,1907]" captionTargetBox="[152,1397,186,1882]" captionTargetId="figure@8.[150,1435,178,1902]" captionTargetPageId="8" captionText="FIGURES 19 – 25. Arcanobisium comasi sp. nov., male holotype. 19. coxae of pedipalp and leg I; 20. genital sternites, ventral; 21. genitalia, ventral; 22. stigmata of sternites III and IV; 23. tergites-sternites VIII – XII, posterior view, drawing in perspective, entire setae only illustrated in sternites X – XI; 24. tergites-sternites VIII – XII, lateral view; 25. habitus, lateral view; pedipalps, legs and one chelicera omitted. Abbreviations: I – XII segments I – XII of opisthosoma. Scale bars (mm): 0.05 (Fig. 21), 0.10 (Figs 20, 22, 24, 25), 0.20 (Figs 19, 23)." httpUri="https://zenodo.org/record/195652/files/figure.png" pageId="4" pageNumber="45">Fig. 20</figureCitation>
|
||
); genitalia with 3+3 internal setae (
|
||
<figureCitation id="13142A0DFFAFB86BFE7472A49E19FC91" box="[444,534,1688,1714]" captionStart="FIGURES 19 – 25" captionStartId="8.[151,270,1883,1907]" captionTargetBox="[152,1397,186,1882]" captionTargetId="figure@8.[150,1435,178,1902]" captionTargetPageId="8" captionText="FIGURES 19 – 25. Arcanobisium comasi sp. nov., male holotype. 19. coxae of pedipalp and leg I; 20. genital sternites, ventral; 21. genitalia, ventral; 22. stigmata of sternites III and IV; 23. tergites-sternites VIII – XII, posterior view, drawing in perspective, entire setae only illustrated in sternites X – XI; 24. tergites-sternites VIII – XII, lateral view; 25. habitus, lateral view; pedipalps, legs and one chelicera omitted. Abbreviations: I – XII segments I – XII of opisthosoma. Scale bars (mm): 0.05 (Fig. 21), 0.10 (Figs 20, 22, 24, 25), 0.20 (Figs 19, 23)." httpUri="https://zenodo.org/record/195652/files/figure.png" pageId="4" pageNumber="45">Fig. 21</figureCitation>
|
||
). Chaetotaxy of sternites IV–XI: 8: 9: 11: 11: 11: 6: 5: 2. One microseta in front of each stigma of sternite III, no setae in front of stigmata of sternite IV (
|
||
<figureCitation id="13142A0DFFAFB86BFBAD728298CDFCFB" box="[1125,1218,1726,1752]" captionStart="FIGURES 19 – 25" captionStartId="8.[151,270,1883,1907]" captionTargetBox="[152,1397,186,1882]" captionTargetId="figure@8.[150,1435,178,1902]" captionTargetPageId="8" captionText="FIGURES 19 – 25. Arcanobisium comasi sp. nov., male holotype. 19. coxae of pedipalp and leg I; 20. genital sternites, ventral; 21. genitalia, ventral; 22. stigmata of sternites III and IV; 23. tergites-sternites VIII – XII, posterior view, drawing in perspective, entire setae only illustrated in sternites X – XI; 24. tergites-sternites VIII – XII, lateral view; 25. habitus, lateral view; pedipalps, legs and one chelicera omitted. Abbreviations: I – XII segments I – XII of opisthosoma. Scale bars (mm): 0.05 (Fig. 21), 0.10 (Figs 20, 22, 24, 25), 0.20 (Figs 19, 23)." httpUri="https://zenodo.org/record/195652/files/figure.png" pageId="4" pageNumber="45">Fig. 22</figureCitation>
|
||
). Glandular pores present on sternites IV–IX: 6: 9: 7: 7: 7: 7.
|
||
</paragraph>
|
||
<paragraph id="8B903688FFAFB86BFF0E73349DF1FD9B" blockId="4.[151,1437,1090,1976]" pageId="4" pageNumber="45">
|
||
Chelicera (
|
||
<figureCitation id="13142A0DFFAFB86BFE8073349D80FD01" box="[328,399,1800,1826]" captionStart="FIGURES 4 – 12" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[220,1300,190,1843]" captionTargetId="figure@5.[185,1400,179,1853]" captionTargetPageId="5" captionText="FIGURES 4 – 12. Arcanobisium comasi sp. nov., male holotype. 4. carapace; 5. anterior margin of carapace, showing epistome; 6. right chelicera; 7. partial view of fingers of right chelicera; 8. cheliceral rallum; 9. right leg I, lateral view; 10. right leg IV, lateral view; 11. distal end of tarsus and apotele of right leg IV, lateral view; 12. subterminal seta of tarsus right leg IV, partial view. Abbreviations: P glandular pore; TS tactile seta. Scale bars (mm): 0.01 (Fig. 12), 0.05 (Fig. 5, 7, 8, 11), 0.10 (Fig. 6), 0.20 (Fig. 4, 9, 10)." httpUri="https://zenodo.org/record/195650/files/figure.png" pageId="4" pageNumber="45">Fig. 6</figureCitation>
|
||
) with 5 setae on hand and one seta on movable finger, 0.64 from base. Galea long and simple. Fixed finger with 16 small to medium teeth; movable finger with 5 apical protuberances and 6 teeth, two medial teeth larger than others (
|
||
<figureCitation id="13142A0DFFAFB86BFD8A736F9E85FD4E" box="[578,650,1875,1901]" captionStart="FIGURES 4 – 12" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[220,1300,190,1843]" captionTargetId="figure@5.[185,1400,179,1853]" captionTargetPageId="5" captionText="FIGURES 4 – 12. Arcanobisium comasi sp. nov., male holotype. 4. carapace; 5. anterior margin of carapace, showing epistome; 6. right chelicera; 7. partial view of fingers of right chelicera; 8. cheliceral rallum; 9. right leg I, lateral view; 10. right leg IV, lateral view; 11. distal end of tarsus and apotele of right leg IV, lateral view; 12. subterminal seta of tarsus right leg IV, partial view. Abbreviations: P glandular pore; TS tactile seta. Scale bars (mm): 0.01 (Fig. 12), 0.05 (Fig. 5, 7, 8, 11), 0.10 (Fig. 6), 0.20 (Fig. 4, 9, 10)." httpUri="https://zenodo.org/record/195650/files/figure.png" pageId="4" pageNumber="45">Fig. 7</figureCitation>
|
||
); light scaly reticulation on hand and movable finger. Rallum with 5 finely serrate blades, at least the basal one shorter than others (
|
||
<figureCitation id="13142A0DFFAFB86BFC7D7344980FFDB1" box="[949,1024,1912,1938]" captionStart="FIGURES 4 – 12" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[220,1300,190,1843]" captionTargetId="figure@5.[185,1400,179,1853]" captionTargetPageId="5" captionText="FIGURES 4 – 12. Arcanobisium comasi sp. nov., male holotype. 4. carapace; 5. anterior margin of carapace, showing epistome; 6. right chelicera; 7. partial view of fingers of right chelicera; 8. cheliceral rallum; 9. right leg I, lateral view; 10. right leg IV, lateral view; 11. distal end of tarsus and apotele of right leg IV, lateral view; 12. subterminal seta of tarsus right leg IV, partial view. Abbreviations: P glandular pore; TS tactile seta. Scale bars (mm): 0.01 (Fig. 12), 0.05 (Fig. 5, 7, 8, 11), 0.10 (Fig. 6), 0.20 (Fig. 4, 9, 10)." httpUri="https://zenodo.org/record/195650/files/figure.png" pageId="4" pageNumber="45">Fig. 8</figureCitation>
|
||
). Serrula exterior with 23 blades, serrula interior with 14 blades.
|
||
</paragraph>
|
||
<caption id="DF506600FFAEB86AFF5F73669EBDFDD1" httpUri="https://zenodo.org/record/195650/files/figure.png" pageId="5" pageNumber="46" targetBox="[220,1300,190,1843]" targetPageId="5">
|
||
<paragraph id="8B903688FFAEB86AFF5F73669EBDFDD1" blockId="5.[151,1437,1882,2034]" pageId="5" pageNumber="46">
|
||
<emphasis id="B95BEA9AFFAEB86AFF5F73669D5CFD51" bold="true" box="[151,339,1882,1906]" pageId="5" pageNumber="46">FIGURES 4–12.</emphasis>
|
||
<taxonomicName id="4C2F4D0BFFAEB86AFE9373679E48FD51" box="[347,583,1883,1906]" class="Arachnida" family="Syarinidae" genus="Arcanobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="comasi" status="sp. nov.">
|
||
<emphasis id="B95BEA9AFFAEB86AFE9373679E48FD51" box="[347,583,1883,1906]" italics="true" pageId="5" pageNumber="46">Arcanobisium comasi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26857E1FFAEB86AFD8673669EA9FD51" box="[590,678,1882,1906]" pageId="5" pageNumber="46" rank="species">
|
||
<emphasis id="B95BEA9AFFAEB86AFD8673669EA9FD51" bold="true" box="[590,678,1882,1906]" pageId="5" pageNumber="46">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male holotype. 4. carapace; 5. anterior margin of carapace, showing epistome; 6. right chelicera; 7. partial view of fingers of right chelicera; 8. cheliceral rallum; 9. right leg I, lateral view; 10. right leg IV, lateral view; 11. distal end of tarsus and apotele of right leg IV, lateral view; 12. subterminal seta of tarsus right leg IV, partial view. Abbreviations:
|
||
<emphasis id="B95BEA9AFFAEB86AFD5573879EA2FDF1" box="[669,685,1979,2002]" italics="true" pageId="5" pageNumber="46">P</emphasis>
|
||
glandular pore;
|
||
<emphasis id="B95BEA9AFFAEB86AFCA973879F72FDF1" box="[865,893,1979,2002]" italics="true" pageId="5" pageNumber="46">TS</emphasis>
|
||
tactile seta. Scale bars (mm): 0.01 (Fig. 12), 0.05 (Fig. 5, 7, 8, 11), 0.10 (Fig. 6), 0.20 (Fig. 4, 9, 10).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF506600FFADB869FF5F73739DBAFDC5" httpUri="https://zenodo.org/record/195651/files/figure.png" pageId="6" pageNumber="47" targetBox="[274,1377,205,1833]" targetPageId="6">
|
||
<paragraph id="8B903688FFADB869FF5F73739DBAFDC5" blockId="6.[151,1436,1871,2022]" pageId="6" pageNumber="47">
|
||
<emphasis id="B95BEA9AFFADB869FF5F73739D52FD44" bold="true" box="[151,349,1871,1895]" pageId="6" pageNumber="47">FIGURES 13–18.</emphasis>
|
||
<taxonomicName id="4C2F4D0BFFADB869FEAB736C9E46FD44" box="[355,585,1872,1895]" class="Arachnida" family="Syarinidae" genus="Arcanobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="comasi" status="sp. nov.">
|
||
<emphasis id="B95BEA9AFFADB869FEAB736C9E46FD44" box="[355,585,1872,1895]" italics="true" pageId="6" pageNumber="47">Arcanobisium comasi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26857E1FFADB869FD8773739EABFD44" box="[591,676,1871,1895]" pageId="6" pageNumber="47" rank="species">
|
||
<emphasis id="B95BEA9AFFADB869FD8773739EABFD44" bold="true" box="[591,676,1871,1895]" pageId="6" pageNumber="47">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male holotype. 13. right pedipalp, without chela, dorsal view; 14. right chela, dorsal view; 15. right chela, antiaxial view; 16. tips of fixed and movable finger of right chela, antiaxial view; 17. tips of fixed and movable finger of right chela, dorsal view; 18. trichobothrium
|
||
<emphasis id="B95BEA9AFFADB869FC3C73AC9FF4FD84" box="[1012,1019,1936,1959]" italics="true" pageId="6" pageNumber="47">t</emphasis>
|
||
. Abbreviations:
|
||
<emphasis id="B95BEA9AFFADB869FB7873AC98DDFD84" box="[1200,1234,1936,1959]" italics="true" pageId="6" pageNumber="47">af?</emphasis>
|
||
pore or sensillum;
|
||
<emphasis id="B95BEA9AFFADB869FF5F738C9CBFFDE4" box="[151,176,1968,1991]" italics="true" pageId="6" pageNumber="47">pc</emphasis>
|
||
doubled pore sensillum (other abbreviations correspond to trichobothria). Scale bars (mm): 0.05 (Fig. 18), 0.10 (Figs 16–17), 0.20 (Figs 13–15).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B903688FFACB868FF0E74A4994AF89F" blockId="7.[151,1438,152,2007]" pageId="7" pageNumber="48">
|
||
Pedipalps (
|
||
<figureCitation id="13142A0DFFACB868FE8374A49DD9FA91" box="[331,470,152,178]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,269,1871,1895]" captionTargetBox="[274,1377,205,1833]" captionTargetId="figure@6.[186,1410,194,1857]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Arcanobisium comasi sp. nov., male holotype. 13. right pedipalp, without chela, dorsal view; 14. right chela, dorsal view; 15. right chela, antiaxial view; 16. tips of fixed and movable finger of right chela, antiaxial view; 17. tips of fixed and movable finger of right chela, dorsal view; 18. trichobothrium t. Abbreviations: af? pore or sensillum; pc doubled pore sensillum (other abbreviations correspond to trichobothria). Scale bars (mm): 0.05 (Fig. 18), 0.10 (Figs 16 – 17), 0.20 (Figs 13 – 15)." httpUri="https://zenodo.org/record/195651/files/figure.png" pageId="7" pageNumber="48">Figs 13–15</figureCitation>
|
||
). Trochanter and basal half of femur granulate, paraxial face of patella partially granulate. Femur with one glandular pore mediodistally, patella with one pore at base of pedicel and 5 more pores along medial line of article (
|
||
<figureCitation id="13142A0DFFACB868FDFA74DE9E84FADF" box="[562,651,226,252]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,269,1871,1895]" captionTargetBox="[274,1377,205,1833]" captionTargetId="figure@6.[186,1410,194,1857]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Arcanobisium comasi sp. nov., male holotype. 13. right pedipalp, without chela, dorsal view; 14. right chela, dorsal view; 15. right chela, antiaxial view; 16. tips of fixed and movable finger of right chela, antiaxial view; 17. tips of fixed and movable finger of right chela, dorsal view; 18. trichobothrium t. Abbreviations: af? pore or sensillum; pc doubled pore sensillum (other abbreviations correspond to trichobothria). Scale bars (mm): 0.05 (Fig. 18), 0.10 (Figs 16 – 17), 0.20 (Figs 13 – 15)." httpUri="https://zenodo.org/record/195651/files/figure.png" pageId="7" pageNumber="48">Fig. 13</figureCitation>
|
||
). Chela (
|
||
<figureCitation id="13142A0DFFACB868FD3F74DE9F71FADF" box="[759,894,226,252]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,269,1871,1895]" captionTargetBox="[274,1377,205,1833]" captionTargetId="figure@6.[186,1410,194,1857]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Arcanobisium comasi sp. nov., male holotype. 13. right pedipalp, without chela, dorsal view; 14. right chela, dorsal view; 15. right chela, antiaxial view; 16. tips of fixed and movable finger of right chela, antiaxial view; 17. tips of fixed and movable finger of right chela, dorsal view; 18. trichobothrium t. Abbreviations: af? pore or sensillum; pc doubled pore sensillum (other abbreviations correspond to trichobothria). Scale bars (mm): 0.05 (Fig. 18), 0.10 (Figs 16 – 17), 0.20 (Figs 13 – 15)." httpUri="https://zenodo.org/record/195651/files/figure.png" pageId="7" pageNumber="48">Figs 14–15</figureCitation>
|
||
) smooth, hand and fixed finger with peculiar and irregular outline, hand with a marked wide depression mid-way on antiaxial face, trichobothria
|
||
<emphasis id="B95BEA9AFFACB868FAEC75349994FB02" box="[1316,1435,264,290]" italics="true" pageId="7" pageNumber="48">isb -esb -eb</emphasis>
|
||
on a wide and low bump, fixed finger with an angular kink from trichobothrium
|
||
<emphasis id="B95BEA9AFFACB868FBAE75119873FB65" box="[1126,1148,301,326]" italics="true" pageId="7" pageNumber="48">et</emphasis>
|
||
to the tip (
|
||
<figureCitation id="13142A0DFFACB868FAC975119982FB64" box="[1281,1421,301,327]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,269,1871,1895]" captionTargetBox="[274,1377,205,1833]" captionTargetId="figure@6.[186,1410,194,1857]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Arcanobisium comasi sp. nov., male holotype. 13. right pedipalp, without chela, dorsal view; 14. right chela, dorsal view; 15. right chela, antiaxial view; 16. tips of fixed and movable finger of right chela, antiaxial view; 17. tips of fixed and movable finger of right chela, dorsal view; 18. trichobothrium t. Abbreviations: af? pore or sensillum; pc doubled pore sensillum (other abbreviations correspond to trichobothria). Scale bars (mm): 0.05 (Fig. 18), 0.10 (Figs 16 – 17), 0.20 (Figs 13 – 15)." httpUri="https://zenodo.org/record/195651/files/figure.png" pageId="7" pageNumber="48">Figs 14, 17</figureCitation>
|
||
), movable finger markedly curved and partly overlapping fixed finger distally. Hand with a distal group of glandular pores on antiaxial and paraxial faces, another pore on pedicel, trichobothrium
|
||
<emphasis id="B95BEA9AFFACB868FB0D754498D3FBB2" box="[1221,1244,376,401]" italics="true" pageId="7" pageNumber="48">ib</emphasis>
|
||
on a prominent tubercle and located in basal half of hand, trichobothria
|
||
<emphasis id="B95BEA9AFFACB868FCE875A19F97FB95" box="[800,920,413,439]" italics="true" pageId="7" pageNumber="48">isb -esb -eb</emphasis>
|
||
forming a triangular group on distal third of hand. Fixed finger with only four trichobothria,
|
||
<emphasis id="B95BEA9AFFACB868FD0475FE9EE7FBF8" box="[716,744,450,475]" italics="true" pageId="7" pageNumber="48">ist</emphasis>
|
||
isolated in basal half,
|
||
<emphasis id="B95BEA9AFFACB868FC3975FE981DFBF8" box="[1009,1042,450,475]" italics="true" pageId="7" pageNumber="48">est</emphasis>
|
||
relatively close and basal to
|
||
<emphasis id="B95BEA9AFFACB868FAAF75FE9973FBF8" box="[1383,1404,450,475]" italics="true" pageId="7" pageNumber="48">et</emphasis>
|
||
,
|
||
<emphasis id="B95BEA9AFFACB868FA4475FE9993FBF8" box="[1420,1436,450,475]" italics="true" pageId="7" pageNumber="48">it</emphasis>
|
||
far from the latter and situated close to tip; with 68 contiguous teeth ending at level of trichobothrium
|
||
<emphasis id="B95BEA9AFFACB868FABF75D4999CF822" box="[1399,1427,488,513]" italics="true" pageId="7" pageNumber="48">ist</emphasis>
|
||
; venom duct short,
|
||
<emphasis id="B95BEA9AFFACB868FEB876319E2EF805" box="[368,545,525,550]" italics="true" pageId="7" pageNumber="48">nodus ramosus</emphasis>
|
||
slightly distal of trichobothrium
|
||
<emphasis id="B95BEA9AFFACB868FC6C76319FBBF805" box="[932,948,525,550]" italics="true" pageId="7" pageNumber="48">it</emphasis>
|
||
; one pore or sensillum (
|
||
<emphasis id="B95BEA9AFFACB868FB18763198E8F805" box="[1232,1255,525,550]" italics="true" pageId="7" pageNumber="48">af</emphasis>
|
||
?) on a tubercle distal of
|
||
<emphasis id="B95BEA9AFFACB868FF36760E9D01F868" box="[254,270,562,587]" italics="true" pageId="7" pageNumber="48">it</emphasis>
|
||
(
|
||
<figureCitation id="13142A0DFFACB868FED6760E9DA8F86F" box="[286,423,562,588]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,269,1871,1895]" captionTargetBox="[274,1377,205,1833]" captionTargetId="figure@6.[186,1410,194,1857]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Arcanobisium comasi sp. nov., male holotype. 13. right pedipalp, without chela, dorsal view; 14. right chela, dorsal view; 15. right chela, antiaxial view; 16. tips of fixed and movable finger of right chela, antiaxial view; 17. tips of fixed and movable finger of right chela, dorsal view; 18. trichobothrium t. Abbreviations: af? pore or sensillum; pc doubled pore sensillum (other abbreviations correspond to trichobothria). Scale bars (mm): 0.05 (Fig. 18), 0.10 (Figs 16 – 17), 0.20 (Figs 13 – 15)." httpUri="https://zenodo.org/record/195651/files/figure.png" pageId="7" pageNumber="48">Figs 16–17</figureCitation>
|
||
). Movable finger with 72 contiguous teeth, long and apically rounded, ending basal to trichobothrium
|
||
<emphasis id="B95BEA9AFFACB868FE8676649D52F852" box="[334,349,600,625]" italics="true" pageId="7" pageNumber="48">b</emphasis>
|
||
; trichobothrium
|
||
<emphasis id="B95BEA9AFFACB868FDED76649E36F852" box="[549,569,600,625]" italics="true" pageId="7" pageNumber="48">st</emphasis>
|
||
close to
|
||
<emphasis id="B95BEA9AFFACB868FD6C76649EA3F852" box="[676,684,600,625]" italics="true" pageId="7" pageNumber="48">t</emphasis>
|
||
and both near middle of finger,
|
||
<emphasis id="B95BEA9AFFACB868FBE27664984BF852" box="[1066,1092,600,625]" italics="true" pageId="7" pageNumber="48">sb</emphasis>
|
||
slightly closer to
|
||
<emphasis id="B95BEA9AFFACB868FAD176649922F852" box="[1305,1325,600,625]" italics="true" pageId="7" pageNumber="48">st</emphasis>
|
||
than to
|
||
<emphasis id="B95BEA9AFFACB868FA4576649993F852" box="[1421,1436,600,625]" italics="true" pageId="7" pageNumber="48">b</emphasis>
|
||
but not forming a group with
|
||
<emphasis id="B95BEA9AFFACB868FE3F76419E13F8B5" box="[503,540,637,663]" italics="true" pageId="7" pageNumber="48">st -t</emphasis>
|
||
; trichobothrium
|
||
<emphasis id="B95BEA9AFFACB868FD2D76419EE2F8B5" box="[741,749,637,662]" italics="true" pageId="7" pageNumber="48">t</emphasis>
|
||
slightly lanceolate toward tip (
|
||
<figureCitation id="13142A0DFFACB868FB97764198B8F8B4" box="[1119,1207,637,663]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,269,1871,1895]" captionTargetBox="[274,1377,205,1833]" captionTargetId="figure@6.[186,1410,194,1857]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Arcanobisium comasi sp. nov., male holotype. 13. right pedipalp, without chela, dorsal view; 14. right chela, dorsal view; 15. right chela, antiaxial view; 16. tips of fixed and movable finger of right chela, antiaxial view; 17. tips of fixed and movable finger of right chela, dorsal view; 18. trichobothrium t. Abbreviations: af? pore or sensillum; pc doubled pore sensillum (other abbreviations correspond to trichobothria). Scale bars (mm): 0.05 (Fig. 18), 0.10 (Figs 16 – 17), 0.20 (Figs 13 – 15)." httpUri="https://zenodo.org/record/195651/files/figure.png" pageId="7" pageNumber="48">Fig. 18</figureCitation>
|
||
); one doubled pore sensillum (
|
||
<emphasis id="B95BEA9AFFACB868FED0769E9D3CF898" box="[280,307,674,699]" italics="true" pageId="7" pageNumber="48">pc</emphasis>
|
||
) close to dental margin, basal to trichobothrium
|
||
<emphasis id="B95BEA9AFFACB868FCA7769E9F86F898" box="[879,905,674,699]" italics="true" pageId="7" pageNumber="48">sb</emphasis>
|
||
. Lyrifissures as shown in
|
||
<figureCitation id="13142A0DFFACB868FB73769E994EF89F" box="[1211,1345,674,700]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,269,1871,1895]" captionTargetBox="[274,1377,205,1833]" captionTargetId="figure@6.[186,1410,194,1857]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Arcanobisium comasi sp. nov., male holotype. 13. right pedipalp, without chela, dorsal view; 14. right chela, dorsal view; 15. right chela, antiaxial view; 16. tips of fixed and movable finger of right chela, antiaxial view; 17. tips of fixed and movable finger of right chela, dorsal view; 18. trichobothrium t. Abbreviations: af? pore or sensillum; pc doubled pore sensillum (other abbreviations correspond to trichobothria). Scale bars (mm): 0.05 (Fig. 18), 0.10 (Figs 16 – 17), 0.20 (Figs 13 – 15)." httpUri="https://zenodo.org/record/195651/files/figure.png" pageId="7" pageNumber="48">Figs 14–15</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B903688FFACB868FF0E76F498B2F954" blockId="7.[151,1438,152,2007]" pageId="7" pageNumber="48">
|
||
Leg I (
|
||
<figureCitation id="13142A0DFFACB868FED076F49D6EF8C1" box="[280,353,712,738]" captionStart="FIGURES 4 – 12" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[220,1300,190,1843]" captionTargetId="figure@5.[185,1400,179,1853]" captionTargetPageId="5" captionText="FIGURES 4 – 12. Arcanobisium comasi sp. nov., male holotype. 4. carapace; 5. anterior margin of carapace, showing epistome; 6. right chelicera; 7. partial view of fingers of right chelicera; 8. cheliceral rallum; 9. right leg I, lateral view; 10. right leg IV, lateral view; 11. distal end of tarsus and apotele of right leg IV, lateral view; 12. subterminal seta of tarsus right leg IV, partial view. Abbreviations: P glandular pore; TS tactile seta. Scale bars (mm): 0.01 (Fig. 12), 0.05 (Fig. 5, 7, 8, 11), 0.10 (Fig. 6), 0.20 (Fig. 4, 9, 10)." httpUri="https://zenodo.org/record/195650/files/figure.png" pageId="7" pageNumber="48">Fig. 9</figureCitation>
|
||
). Femur without basi-dorsal mound; femur and patella granulate, tibia and metatarsus with weak scales, tarsus smooth; claws smooth; subterminal setae with three apical rami. Leg IV (
|
||
<figureCitation id="13142A0DFFACB868FB1876D19928F924" box="[1232,1319,749,775]" captionStart="FIGURES 4 – 12" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[220,1300,190,1843]" captionTargetId="figure@5.[185,1400,179,1853]" captionTargetPageId="5" captionText="FIGURES 4 – 12. Arcanobisium comasi sp. nov., male holotype. 4. carapace; 5. anterior margin of carapace, showing epistome; 6. right chelicera; 7. partial view of fingers of right chelicera; 8. cheliceral rallum; 9. right leg I, lateral view; 10. right leg IV, lateral view; 11. distal end of tarsus and apotele of right leg IV, lateral view; 12. subterminal seta of tarsus right leg IV, partial view. Abbreviations: P glandular pore; TS tactile seta. Scale bars (mm): 0.01 (Fig. 12), 0.05 (Fig. 5, 7, 8, 11), 0.10 (Fig. 6), 0.20 (Fig. 4, 9, 10)." httpUri="https://zenodo.org/record/195650/files/figure.png" pageId="7" pageNumber="48">Fig. 10</figureCitation>
|
||
): junction between femur and patella perpendicular; femur slightly longer than patella, both granulate, the other segments generally smooth; claws smooth; subterminal setae with three apical rami (
|
||
<figureCitation id="13142A0DFFACB868FBBF770498F1F971" box="[1143,1278,824,850]" captionStart="FIGURES 4 – 12" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[220,1300,190,1843]" captionTargetId="figure@5.[185,1400,179,1853]" captionTargetPageId="5" captionText="FIGURES 4 – 12. Arcanobisium comasi sp. nov., male holotype. 4. carapace; 5. anterior margin of carapace, showing epistome; 6. right chelicera; 7. partial view of fingers of right chelicera; 8. cheliceral rallum; 9. right leg I, lateral view; 10. right leg IV, lateral view; 11. distal end of tarsus and apotele of right leg IV, lateral view; 12. subterminal seta of tarsus right leg IV, partial view. Abbreviations: P glandular pore; TS tactile seta. Scale bars (mm): 0.01 (Fig. 12), 0.05 (Fig. 5, 7, 8, 11), 0.10 (Fig. 6), 0.20 (Fig. 4, 9, 10)." httpUri="https://zenodo.org/record/195650/files/figure.png" pageId="7" pageNumber="48">Figs 11–12</figureCitation>
|
||
); tactile setae (TS) ratios: tibia 0.54, metatarsus 0.27, tarsus 0.31. Arolium shorter than claws on all legs.
|
||
</paragraph>
|
||
<paragraph id="8B903688FFACB868FF0E77BE9D60FECF" blockId="7.[151,1438,152,2007]" pageId="7" pageNumber="48">Ratios and measurements. Body: 1.64. Carapace: 0.60/0.44 (1.35×). Chelicera: hand: 0.34/0.18 (1.91×); finger: 0.21. Pedipalp: trochanter: 0.32/0.12 (2.79×); femur: 0.67/0.14 (4.81×); patella: 0.64/0.17 (3.76×), pedicel: 0.14, club: 0.50/0.17 (2.92×), club/pedicel: 3.44×; hand with pedicel: 0.40/0.22 (1.85×), hand without pedicel: 0.34/0.22 (1.55×); finger: 0.56; chela with pedicel: 0.96/0.22 (4.42×), chela without pedicel: 0.90/ 0.22 (4.09×); chela/carapace: 1.59×; femur/carapace: 1.12×; femur/finger: 1.20×; femur/patella: 1.05×; patella/hand: 1.60×; finger/hand: 1.40×. Leg I: femur: 0.33/0.09 (3.82×); patella: 0.28/0.09 (3.29×); tibia 0.31/ 0.06 (5.06×); metatarsus 0.20/0.05 (4.06×); tarsus 0.25/0.04 (5.56×); femur/patella: 1.16×; tarsus/metatarsus: 1.23×. Leg IV: femur: 0.31; patella: 0.30; femur+patella: 0.61/0.09 (6.47×); tibia 0.62/0.06 (9.97×); metatarsus: 0.26/0.05 (4.95×); tarsus 0.27/0.05 (5.51×); femur/patella: 1.05×; tibia/femur: 1.01×; tarsus/ metatarsus: 1.04×.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3356503FFACB868FF0C70CB9E5BFF14" pageId="7" pageNumber="48" type="distribution">
|
||
<paragraph id="8B903688FFACB868FF0C70CB9E5BFF14" blockId="7.[151,1438,152,2007]" pageId="7" pageNumber="48">
|
||
<emphasis id="B95BEA9AFFACB868FF0C70CB9D6AFF32" bold="true" box="[196,357,1271,1297]" pageId="7" pageNumber="48">Distribution.</emphasis>
|
||
The new species has only been found in the cave Avenc d’en Serenge, Castellón province, Valencian Community,
|
||
<collectingCountry id="F3387618FFACB868FE6271219DE2FF14" box="[426,493,1309,1335]" name="Spain" pageId="7" pageNumber="48">Spain</collectingCountry>
|
||
(
|
||
<figureCitation id="13142A0DFFACB868FE3571219E4DFF14" box="[509,578,1309,1335]" captionStart="FIGURE 1" captionStartId="1.[151,255,1129,1153]" captionTargetBox="[292,1292,495,1100]" captionTargetId="figure@1.[289,1299,495,1103]" captionTargetPageId="1" captionText="FIGURE 1. Map showing location of the type locality of Arcanobisium comasi sp. nov., the cave Avenc d’en Serenge in Castellón province, Valencian Community, Spain." httpUri="https://zenodo.org/record/195648/files/figure.png" pageId="7" pageNumber="48">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3356503FFACB868FF0C717E9F65FDF4" pageId="7" pageNumber="48" type="biology_ecology">
|
||
<paragraph id="8B903688FFACB868FF0C717E9979FF84" blockId="7.[151,1438,152,2007]" pageId="7" pageNumber="48">
|
||
<emphasis id="B95BEA9AFFACB868FF0C717E9D22FF7F" bold="true" box="[196,301,1346,1372]" pageId="7" pageNumber="48">Biology.</emphasis>
|
||
The species shows troglomorphic characteristics such as the absence of eyes or eye-spots, elongated carapace, pedipalps and legs, and unpigmented cuticle, but the size of the body is more like that of epigean taxa. The new species seems to be troglobitic, but an edaphic existence should also be considered.
|
||
</paragraph>
|
||
<paragraph id="8B903688FFACB868FF0C718E9E1EFD3F" blockId="7.[151,1438,152,2007]" pageId="7" pageNumber="48">
|
||
<emphasis id="B95BEA9AFFACB868FF0C718E9D8AFFEF" bold="true" box="[196,389,1458,1484]" pageId="7" pageNumber="48">Biogeography.</emphasis>
|
||
The relictual condition of
|
||
<taxonomicName id="4C2F4D0BFFACB868FD29718F9FFBFFEF" box="[737,1012,1459,1484]" class="Arachnida" family="Syarinidae" genus="Arcanobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="comasi" status="sp. nov.">
|
||
<emphasis id="B95BEA9AFFACB868FD29718F9FFBFFEF" box="[737,1012,1459,1484]" italics="true" pageId="7" pageNumber="48">Arcanobisium comasi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26857E1FFACB868FC35718E986BFFEF" box="[1021,1124,1458,1484]" pageId="7" pageNumber="48" rank="species">
|
||
<emphasis id="B95BEA9AFFACB868FC35718E986BFFEF" bold="true" box="[1021,1124,1458,1484]" pageId="7" pageNumber="48">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is implied by its unique morphological features and this is reinforced by its isolated location in a cave, the “Avenc d’en Serenge”, which belongs to an independent biogeographic region in
|
||
<collectingCountry id="F3387618FFACB868FC4C71C19FDFFC34" box="[900,976,1533,1559]" name="Spain" pageId="7" pageNumber="48">Spain</collectingCountry>
|
||
, considered as refuge for Laurasian arthropods (
|
||
<bibRefCitation id="EFBE4B79FFACB868FEEE721E9E06FC1F" author="Ortuno" box="[294,521,1570,1596]" pageId="7" pageNumber="59" refString="Ortuno, V. M., Sendra, A., Montagud, S. & Teruel, S. (2005) Systematique et biologie d'une espece paleoendemique hypogee de la peninsule Iberique: Ildobates neboti Espanol, 1966 (Coleoptera: Carabidae: Dryptinae). Annales de la Societe Entomologique de France (Nouvelle Serie), 40, 459 - 475." type="journal article" year="2005">
|
||
Ortuño
|
||
<emphasis id="B95BEA9AFFACB868FE4C721F9DB6FC1F" box="[388,441,1571,1596]" italics="true" pageId="7" pageNumber="48">et al</emphasis>
|
||
. 2005
|
||
</bibRefCitation>
|
||
). Important relictual troglobites have been found in Avenc d’en Serenge and nearby caves, most of them monotypic genera and endemic to this area. These include the pseudoscorpion
|
||
<taxonomicName id="4C2F4D0BFFACB868FF5F72519E9BFCA4" authority="Ellingsen, 1912" authorityName="Ellingsen" authorityYear="1912" box="[151,660,1645,1671]" class="Arachnida" family="Bochicidae" genus="Troglobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="racovitzai">
|
||
<emphasis id="B95BEA9AFFACB868FF5F72519DB8FCA5" box="[151,439,1645,1670]" italics="true" pageId="7" pageNumber="48">Troglobisium racovitzai</emphasis>
|
||
(Ellingsen, 1912)
|
||
</taxonomicName>
|
||
, the only representative of the family
|
||
<taxonomicName id="4C2F4D0BFFACB868FBAA725198E6FCA4" box="[1122,1257,1645,1671]" class="Arachnida" family="Bochicidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="48" phylum="Arthropoda" rank="family">Bochicidae</taxonomicName>
|
||
in Europe, the remarkable beetle
|
||
<taxonomicName id="4C2F4D0BFFACB868FEBB72AF9EE9FC8F" authority="Espanol, 1966" authorityName="Espanol" authorityYear="1966" box="[371,742,1682,1708]" class="Insecta" family="Carabidae" genus="Ildobates" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="neboti">
|
||
<emphasis id="B95BEA9AFFACB868FEBB72AF9E3BFC8F" box="[371,564,1683,1708]" italics="true" pageId="7" pageNumber="48">Ildobates neboti</emphasis>
|
||
Español, 1966
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C2F4D0BFFACB868FD3E72AE9F71FC8F" box="[758,894,1682,1708]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C2F4D0BFFACB868FC4272AE9808FC8F" box="[906,1031,1682,1708]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
|
||
), the campodeid
|
||
<taxonomicName id="4C2F4D0BFFACB868FB1A72AF9E93FCF1" authority="Bareth & Conde, 1981" authorityName="Bareth & Conde" authorityYear="1981" class="Entognatha" family="Campodeidae" genus="Paratachycampa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="hispanica">
|
||
<emphasis id="B95BEA9AFFACB868FB1A72AF9D04FCF2" italics="true" pageId="7" pageNumber="48">Paratachycampa hispanica</emphasis>
|
||
Bareth & Condé, 1981 (Diplura)
|
||
</taxonomicName>
|
||
[this genus is also known as relicts in caves in
|
||
<collectingCountry id="F3387618FFACB868FB1472849938FCF1" box="[1244,1335,1720,1746]" name="Mexico" pageId="7" pageNumber="48">Mexico</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFBE4B79FFACB868FA8F72849CDAFCD4" author="Sendra" pageId="7" pageNumber="59" refString="Sendra, A. (2003) Distribucion y colonizacion de los Campodeidos cavernicolas en la Peninsula Iberica e Islas Baleares. Boletin de la Sociedad Espanola de Espeleologia y Ciencias del Karst, 4, 12 - 20." type="journal article" year="2003">Sendra 2003</bibRefCitation>
|
||
, 2006)] and the enigmatic japygid
|
||
<taxonomicName id="4C2F4D0BFFACB868FDA272E198F8FCD4" authority="Sendra & Ortuno, 2006" authorityName="Sendra & Ortuno" authorityYear="2006" box="[618,1271,1757,1783]" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="smeagol">
|
||
<emphasis id="B95BEA9AFFACB868FDA272E19F65FCD5" box="[618,874,1757,1782]" italics="true" pageId="7" pageNumber="48">Gollumjapyx smeagol</emphasis>
|
||
Sendra & Ortuño, 2006 (Diplura)
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFBE4B79FFACB868FACE72E19CDBFD3F" author="Ortuno" pageId="7" pageNumber="59" refString="Ortuno, V. M., Sendra, A., Montagud, S. & Teruel, S. (2005) Systematique et biologie d'une espece paleoendemique hypogee de la peninsule Iberique: Ildobates neboti Espanol, 1966 (Coleoptera: Carabidae: Dryptinae). Annales de la Societe Entomologique de France (Nouvelle Serie), 40, 459 - 475." type="journal article" year="2005">
|
||
Ortuño
|
||
<emphasis id="B95BEA9AFFACB868FAAA72E1999AFCD5" box="[1378,1429,1757,1782]" italics="true" pageId="7" pageNumber="48">et al</emphasis>
|
||
. 2005
|
||
</bibRefCitation>
|
||
, 2006; Sendra
|
||
<emphasis id="B95BEA9AFFACB868FE4B733F9DBAFD3F" box="[387,437,1795,1820]" italics="true" pageId="7" pageNumber="48">et al</emphasis>
|
||
. 2006).
|
||
</paragraph>
|
||
<paragraph id="8B903688FFACB868FF0C73149F65FDF4" blockId="7.[151,1438,152,2007]" pageId="7" pageNumber="48">
|
||
The relictual condition of European syarinid genera has been discussed in literature.
|
||
<taxonomicName id="4C2F4D0BFFACB868FB6373149962FD62" box="[1195,1389,1832,1857]" class="Arachnida" family="Syarinidae" genus="Syarinus" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B95BEA9AFFACB868FB6373149962FD62" box="[1195,1389,1832,1857]" italics="true" pageId="7" pageNumber="48">Syarinus strandi</emphasis>
|
||
</taxonomicName>
|
||
has been linked to a pre-glacial fauna by
|
||
<bibRefCitation id="EFBE4B79FFACB868FD9373719F34FD44" author="Schawaller" box="[603,827,1869,1895]" pageId="7" pageNumber="59" refString="Schawaller, W. (1987) Erstnachweis der Familie Syarinidae in Deutschland: ein Reliktvorkommen von Syarinus strandi im Oberen Donautal (Arachnida: Pseudoscorpiones). Jahreshefte, Gesellschaft fur Naturkunde in Wurttemberg, 142, 287 - 292." type="journal article" year="1987">Schawaller (1987)</bibRefCitation>
|
||
, based on the fact that other representatives of the genus are distributed in North
|
||
<collectingCountry id="F3387618FFACB868FDE5734E9EAEFDAF" box="[557,673,1906,1932]" name="United States of America" pageId="7" pageNumber="48">America</collectingCountry>
|
||
. The European genera formerly placed by
|
||
<bibRefCitation id="EFBE4B79FFACB868FB1B734E9974FDAF" author="Beier" box="[1235,1403,1906,1932]" pageId="7" pageNumber="58" refString="Beier, M. (1963) Ordnung Pseudoscorpionidea (Afterskorpione). Bestimmungsbucher zur Bodenfauna Europas. Vol. 1. Akademie-Verlag. Berlin, 313 pp." type="book" year="1963">Beier (1963)</bibRefCitation>
|
||
in Microcreagrellinae also show relict distribution (
|
||
<bibRefCitation id="EFBE4B79FFACB868FD1573A49F69FD91" author="Beier" box="[733,870,1944,1970]" pageId="7" pageNumber="58" refString="Beier, M. (1969) Reliktformen in der Pseudoscorpioniden-Fauna Europas. Memorie della Societa Entomologica Italiana, 48, 317 - 323." type="journal article" year="1969">Beier 1969</bibRefCitation>
|
||
), with
|
||
<taxonomicName id="4C2F4D0BFFACB868FC7173A49852FD92" box="[953,1117,1944,1969]" class="Arachnida" family="Syarinidae" genus="Hadoblothrus" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95BEA9AFFACB868FC7173A49852FD92" box="[953,1117,1944,1969]" italics="true" pageId="7" pageNumber="48">Hadoblothrus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2F4D0BFFACB868FB5073A4995DFD92" box="[1176,1362,1944,1969]" class="Arachnida" family="Syarinidae" genus="Pseudoblothrus" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95BEA9AFFACB868FB5073A4995DFD92" box="[1176,1362,1944,1969]" italics="true" pageId="7" pageNumber="48">Pseudoblothrus</emphasis>
|
||
</taxonomicName>
|
||
being palaeo-endemic troglobites and survivors of an archaic fauna.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |