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<document id="8FC7910C1EDBD74DCA62C3D2BC1B7B50" ID-CLB-Dataset="293416" ID-DOI="10.11646/zootaxa.5433.3.1" ID-GBIF-Dataset="68f4f3c7-6a6e-41a5-9c7a-6ffd996ce89c" ID-ISSN="1175-5326" ID-Zenodo-Dep="10954352" ID-ZooBank="17F2DD04-A68B-47F9-8B9C-792370B35072" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1712752477619" checkinUser="plazi" docAuthor="Qi, Zhi-Hao, Zhou, Zheng, Su, Rong-Xiang, Duan, Lei, Zhang, Yi-Feng &amp; Song, Hai-Tian" docDate="2024" docId="F16FB810A03AFFE5FF0CF9E276A0F982" docLanguage="en" docName="zootaxa.5433.3.1.pdf" docOrigin="Zootaxa 5433 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5433.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Capreolucanus Didier 1928" docType="key" docVersion="1" lastPageNumber="318" masterDocId="0D56C068A035FFF4FF9BFFD4733DFF95" masterDocTitle="Capreolucanus yanxui Qi &amp; Zhou, new species, the third species of the genus from Yunnan (China) based on morphological and molecular characters (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="320" masterPageNumber="301" pageNumber="316" updateTime="1712846505441" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="49EB93A01EEC3647884DB1EC59F15D43">Capreolucanus yanxui Qi &amp; Zhou, new species, the third species of the genus from Yunnan (China) based on morphological and molecular characters (Coleoptera: Lucanidae: Lucaninae)</mods:title>
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<mods:namePart id="43FFF843BF22FCB79C65B9D9184C2608">Qi, Zhi-Hao</mods:namePart>
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<mods:affiliation id="87042DA8CF5D44FDB3A8D559170FD1CC">College of Plant Protection, Fujian Agriculture and Forestry University, Fuzhou, Fujian, 350002, China &amp; Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China</mods:affiliation>
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<mods:namePart id="7FC740E2B2678BBF60D38A871B8A103A">Zhou, Zheng</mods:namePart>
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<mods:affiliation id="52B3DDD90C18D6E7334711950F693BC9">School of Ecology and Environment, Northwestern Polytechnical University, Xi’an, Shaanxi, 710072, China</mods:affiliation>
<mods:nameIdentifier id="688C2126F0D6266296B0927323E23CC9" type="email">zhouzheng_insect@yeah.net</mods:nameIdentifier>
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<mods:namePart id="48CFFF453C09F28F942D8AD19113F569">Su, Rong-Xiang</mods:namePart>
<mods:nameIdentifier id="4D178C38FEA6816B04D2ED60BFA93F58" type="ORCID">0000-0002-6575-5776</mods:nameIdentifier>
<mods:affiliation id="B8EE0EAEFE547737A4839C6D97B3967B">Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China</mods:affiliation>
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<mods:namePart id="F98B8D660889E43AF8C12BAEF8F1FAE3">Duan, Lei</mods:namePart>
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<mods:affiliation id="667F564E40450F43BAAE8360FC83B10E">School of Ecology and Environment, Northwestern Polytechnical University, Xi’an, Shaanxi, 710072, China</mods:affiliation>
<mods:nameIdentifier id="C5A89D82E4B5662815318AE7AFCA8C18" type="email">duanlei@mail.nwpu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="6E196E9440AC04E192B0466CA32C1C4B">Zhang, Yi-Feng</mods:namePart>
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<mods:affiliation id="93B7827CDE4112BE87D0784BFF7CB33D">Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China</mods:affiliation>
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<mods:namePart id="A3F1C4A2EDB3370452253E16170E971E">Song, Hai-Tian</mods:namePart>
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<mods:affiliation id="8A9E7FBCCF0C9897A4FC6FF822D127EF">Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China</mods:affiliation>
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<treatment id="F16FB810A03AFFE5FF0CF9E276A0F982" ID-DOI="http://doi.org/10.5281/zenodo.10960039" ID-GBIF-Taxon="225315123" ID-Zenodo-Dep="10960039" LSID="urn:lsid:plazi:treatment:F16FB810A03AFFE5FF0CF9E276A0F982" httpUri="http://treatment.plazi.org/id/F16FB810A03AFFE5FF0CF9E276A0F982" lastPageId="17" lastPageNumber="318" pageId="15" pageNumber="316">
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Key to males of
<taxonomicName id="BEC67285A03AFFFBFEFBF9E17125F9C5" ID-CoL="3H5Q" authorityName="Didier" authorityYear="1928" box="[352,536,1589,1616]" class="Insecta" family="Lucanidae" genus="Capreolucanus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="4BB2D514A03AFFFBFEFBF9E17125F9C5" bold="true" box="[352,536,1589,1616]" italics="true" pageId="15" pageNumber="316">Capreolucanus</emphasis>
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1. Bicolor species, pronotum black to brown with two distinct cross-like orange red stripes flank midline, all femurs orange red in about apical third (
<figureCitation id="E1FD1583A03AFFFBFE0CF94C72DEF938" box="[407,483,1688,1710]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[223,1360,183,1885]" captionTargetId="figure-17@3.[221,1365,181,1885]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Capreolucanus species. A–C, C. yanxui Qi &amp; Zhou, new species (holotype, 18.6 mm, Pingbian, Yunnan, China); D–F, C. sicardi Didier, 1928 (18.9 mm, Lushui, Yunnan, China); G–I, C. zhuchuangi Wang, 2020 (15.0 mm, Bo Kluea, Nan, Thailand, Photo by Wuttipon Pathomwattananurak). A, D, G, dorsal view; B, E, H, lateral view; C, F, I, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10954354" httpUri="https://zenodo.org/record/10954354/files/figure.png" pageId="15" pageNumber="316">Fig. 1G</figureCitation>
); mentum subtriangular (
<figureCitation id="E1FD1583A03AFFFBFD4DF94C702AF938" box="[726,791,1688,1710]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[223,1360,183,1885]" captionTargetId="figure-17@3.[221,1365,181,1885]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Capreolucanus species. A–C, C. yanxui Qi &amp; Zhou, new species (holotype, 18.6 mm, Pingbian, Yunnan, China); D–F, C. sicardi Didier, 1928 (18.9 mm, Lushui, Yunnan, China); G–I, C. zhuchuangi Wang, 2020 (15.0 mm, Bo Kluea, Nan, Thailand, Photo by Wuttipon Pathomwattananurak). A, D, G, dorsal view; B, E, H, lateral view; C, F, I, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10954354" httpUri="https://zenodo.org/record/10954354/files/figure.png" pageId="15" pageNumber="316">Fig. 1I</figureCitation>
)...............................
<taxonomicName id="BEC67285A03AFFFBFB02F94C76A1F938" ID-CoL="7TBMT" authority="Wang, 2020" authorityName="Wang" authorityYear="2020" box="[1177,1436,1688,1709]" class="Insecta" family="Lucanidae" genus="Capreolucanus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="zhuchuangi">
<emphasis id="4BB2D514A03AFFFBFB02F94C7794F938" box="[1177,1193,1688,1709]" italics="true" pageId="15" pageNumber="316">C</emphasis>
.
<emphasis id="4BB2D514A03AFFFBFB2EF94C7619F938" box="[1205,1316,1688,1709]" italics="true" pageId="15" pageNumber="316">zhuchuangi</emphasis>
<bibRefCitation id="1D5774F7A03AFFFBFAB2F94C76A1F938" author="Wang, C. B." box="[1321,1436,1688,1709]" pageId="15" pageNumber="316" pagination="2 - 7" refId="ref8939" refString="Wang, C. B. (2020) The second species of Capreolucanus Didier, 1928 (Coleoptera: Lucanidae: Lucaninae) from Yunnan, China. Beetles world, 22, 2 - 7." type="journal article" year="2020">Wang, 2020</bibRefCitation>
</taxonomicName>
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<paragraph id="79790906A03AFFFBFF0CF96276A1F97D" blockId="15.[151,1437,1659,1945]" pageId="15" pageNumber="316">
<keyLead id="C237AA33A03AFFFBFF0CF96276A1F97D" pageId="15" pageNumber="316">
- Monochromatic species, pronotum black to brown without stripes, all femurs black (
<figureCitation id="E1FD1583A03AFFFBFBBBF96177B0F95F" box="[1056,1165,1717,1739]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[223,1360,183,1885]" captionTargetId="figure-17@3.[221,1365,181,1885]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Capreolucanus species. A–C, C. yanxui Qi &amp; Zhou, new species (holotype, 18.6 mm, Pingbian, Yunnan, China); D–F, C. sicardi Didier, 1928 (18.9 mm, Lushui, Yunnan, China); G–I, C. zhuchuangi Wang, 2020 (15.0 mm, Bo Kluea, Nan, Thailand, Photo by Wuttipon Pathomwattananurak). A, D, G, dorsal view; B, E, H, lateral view; C, F, I, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10954354" httpUri="https://zenodo.org/record/10954354/files/figure.png" pageId="15" pageNumber="316">Fig. 1A, D</figureCitation>
); mentum not subtriangular (
<figureCitation id="E1FD1583A03AFFFBFF72F9077268F97D" box="[233,341,1747,1769]" captionStart="FIGURE 3" captionStartId="5.[151,250,991,1015]" captionTargetBox="[151,1435,181,967]" captionTargetId="figure-17@5.[151,1435,181,967]" captionTargetPageId="5" captionText="FIGURE 3. Male canthus (A–F), clypeolabrum (G–L) and mentum (M–R) of Capreolucanus species. A–C, G–I, M–O, C. yanxui Qi &amp; Zhou, new species; A, G, M, holotype; B–C, H–I, N–O, paratypes; D–F, J–L, P–R, C. sicardi Didier, 1928. A–L, dorsal view; M–R, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10954358" httpUri="https://zenodo.org/record/10954358/files/figure.png" pageId="15" pageNumber="316">Fig. 3M–R</figureCitation>
)......................................................................................... 2
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<keyStep id="C23211A3A03AFFFBFF0CF92476A0F80D" pageId="15" pageNumber="316">
<paragraph id="79790906A03AFFFBFF0CF92476A1F8D5" blockId="15.[151,1437,1659,1945]" pageId="15" pageNumber="316">
<keyLead id="C237AA33A03AFFFBFF0CF92476A1F8D5" pageId="15" pageNumber="316">
2. Head with a pair of rounded anterolateral angles; when dorsal tooth present, mandible with two dorsal teeth, smaller one subapical and larger one near dorsal base (
<figureCitation id="E1FD1583A03AFFFBFDEDF8D971FFF8B7" box="[630,706,1805,1827]" captionStart="FIGURE 2" captionStartId="4.[151,250,1176,1200]" captionTargetBox="[158,1436,188,1146]" captionTargetId="figure-21@4.[151,1436,181,1152]" captionTargetPageId="4" captionText="FIGURE 2. Male head details of Capreolucanus species in dorsolateral view. A–B, C. yanxui Qi &amp; Zhou, new species; A, holotype; B, paratype; C–D, C. sicardi Didier, 1928; A, C, large sized; B, D, small sized." figureDoi="http://doi.org/10.5281/zenodo.10954356" httpUri="https://zenodo.org/record/10954356/files/figure.png" pageId="15" pageNumber="316">Fig. 2C</figureCitation>
); mentum without notched at anterior margin (
<figureCitation id="E1FD1583A03AFFFBFBE4F8D977D8F8B7" box="[1151,1253,1805,1827]" captionStart="FIGURE 3" captionStartId="5.[151,250,991,1015]" captionTargetBox="[151,1435,181,967]" captionTargetId="figure-17@5.[151,1435,181,967]" captionTargetPageId="5" captionText="FIGURE 3. Male canthus (A–F), clypeolabrum (G–L) and mentum (M–R) of Capreolucanus species. A–C, G–I, M–O, C. yanxui Qi &amp; Zhou, new species; A, G, M, holotype; B–C, H–I, N–O, paratypes; D–F, J–L, P–R, C. sicardi Didier, 1928. A–L, dorsal view; M–R, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10954358" httpUri="https://zenodo.org/record/10954358/files/figure.png" pageId="15" pageNumber="316">Fig. 3P–R</figureCitation>
); flagellum usually about 6.0 times as long as aedeagus (
<figureCitation id="E1FD1583A03AFFFBFDDEF8FF7197F8D4" box="[581,682,1835,1857]" captionStart="FIGURE 5" captionStartId="7.[151,250,1747,1771]" captionTargetBox="[157,1431,187,1716]" captionTargetId="figure-17@7.[151,1436,181,1723]" captionTargetPageId="7" captionText="FIGURE 5. Aedeagus of Capreolucanus species in ventral view. A–C, C. yanxui Qi &amp; Zhou, new species. A, holotype; B–C, paratypes; D–F, C. sicardi Didier, 1928. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10954360" httpUri="https://zenodo.org/record/10954360/files/figure.png" pageId="15" pageNumber="316">Fig. 5D–F</figureCitation>
)............................................
<taxonomicName id="BEC67285A03AFFFBFB5BF8FF76A1F8D5" ID-CoL="QSF2" authority="Didier, 1928" authorityName="Didier" authorityYear="1928" box="[1216,1436,1835,1857]" class="Insecta" family="Lucanidae" genus="Capreolucanus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="sicardi">
<emphasis id="4BB2D514A03AFFFBFB5BF8FF77EDF8D5" box="[1216,1232,1835,1856]" italics="true" pageId="15" pageNumber="316">C</emphasis>
.
<emphasis id="4BB2D514A03AFFFBFB47F8FF7623F8D5" box="[1244,1310,1835,1856]" italics="true" pageId="15" pageNumber="316">sicardi</emphasis>
<bibRefCitation id="1D5774F7A03AFFFBFABFF8FF76A1F8D5" author="Didier, R." box="[1316,1436,1835,1857]" pageId="15" pageNumber="316" pagination="65 - 67" refId="ref8340" refString="Didier, R. (1928) Etudes sur les Coleopteres Lucanides du globe. V. Descriptions de Lucanides nouveaux ou peu connus. 1. Description d'un genre nouveau et d'une espece nouvelle. Capreolucanus sicardi. Librairie speciale Agricole, Paris, Fascicule, 3, 65 - 67." type="journal article" year="1928">Didier, 1928</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="79790906A03AFFFBFF0CF89C76A0F80D" blockId="15.[151,1437,1659,1945]" pageId="15" pageNumber="316">
<keyLead id="C237AA33A03AFFFBFF0CF89C76A0F80D" pageId="15" pageNumber="316">
- Head with a pair of triangular anterolateral angles; when dorsal tooth present, mandible only with one dorsal tooth behind the apex (
<figureCitation id="E1FD1583A03AFFFBFE86F8B172B5F8EF" box="[285,392,1893,1915]" captionStart="FIGURE 2" captionStartId="4.[151,250,1176,1200]" captionTargetBox="[158,1436,188,1146]" captionTargetId="figure-21@4.[151,1436,181,1152]" captionTargetPageId="4" captionText="FIGURE 2. Male head details of Capreolucanus species in dorsolateral view. A–B, C. yanxui Qi &amp; Zhou, new species; A, holotype; B, paratype; C–D, C. sicardi Didier, 1928; A, C, large sized; B, D, small sized." figureDoi="http://doi.org/10.5281/zenodo.10954356" httpUri="https://zenodo.org/record/10954356/files/figure.png" pageId="15" pageNumber="316">Fig. 2A–B</figureCitation>
); mentum with anterior margin distinct notched (
<figureCitation id="E1FD1583A03AFFFBFCFFF8B170E9F8EF" box="[868,980,1893,1915]" captionStart="FIGURE 3" captionStartId="5.[151,250,991,1015]" captionTargetBox="[151,1435,181,967]" captionTargetId="figure-17@5.[151,1435,181,967]" captionTargetPageId="5" captionText="FIGURE 3. Male canthus (A–F), clypeolabrum (G–L) and mentum (M–R) of Capreolucanus species. A–C, G–I, M–O, C. yanxui Qi &amp; Zhou, new species; A, G, M, holotype; B–C, H–I, N–O, paratypes; D–F, J–L, P–R, C. sicardi Didier, 1928. A–L, dorsal view; M–R, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10954358" httpUri="https://zenodo.org/record/10954358/files/figure.png" pageId="15" pageNumber="316">Fig. 3M–O</figureCitation>
); flagellum usually about 10.0 times as long as aedeagus (
<figureCitation id="E1FD1583A03AFFFBFEDCF8577293F80C" box="[327,430,1923,1945]" captionStart="FIGURE 5" captionStartId="7.[151,250,1747,1771]" captionTargetBox="[157,1431,187,1716]" captionTargetId="figure-17@7.[151,1436,181,1723]" captionTargetPageId="7" captionText="FIGURE 5. Aedeagus of Capreolucanus species in ventral view. A–C, C. yanxui Qi &amp; Zhou, new species. A, holotype; B–C, paratypes; D–F, C. sicardi Didier, 1928. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10954360" httpUri="https://zenodo.org/record/10954360/files/figure.png" pageId="15" pageNumber="316">Fig. 5A–C</figureCitation>
).......................................................
<taxonomicName id="BEC67285A03AFFFBFBD4F8577620F80C" authority="Qi &amp; Zhou" authorityName="Qi &amp; Zhou" authorityYear="2024" box="[1103,1309,1923,1945]" class="Insecta" family="Lucanidae" genus="Capreolucanus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="yanxui" status="new species">
<emphasis id="4BB2D514A03AFFFBFBD4F8577762F80D" box="[1103,1119,1923,1944]" italics="true" pageId="15" pageNumber="316">C</emphasis>
.
<emphasis id="4BB2D514A03AFFFBFBF0F8577796F80D" box="[1131,1195,1923,1944]" italics="true" pageId="15" pageNumber="316">yanxui</emphasis>
Qi &amp; Zhou
</taxonomicName>
,
<emphasis id="4BB2D514A03AFFFBFABDF85776A0F80D" bold="true" box="[1318,1437,1922,1944]" pageId="15" pageNumber="316">
<taxonomicNameLabel id="5081686FA03AFFFBFABDF85776A0F80D" box="[1318,1437,1922,1944]" pageId="15" pageNumber="316" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<caption id="2DB9598EA025FFE4FF0CF8437254F862" ID-DOI="http://doi.org/10.5281/zenodo.10954386" ID-Zenodo-Dep="10954386" httpUri="https://zenodo.org/record/10954386/files/figure.png" pageId="16" pageNumber="317" startId="16.[151,250,1943,1967]" targetBox="[203,1385,181,1919]" targetPageId="16" targetType="figure">
<paragraph id="79790906A025FFE4FF0CF8437254F862" blockId="16.[151,1436,1943,2039]" pageId="16" pageNumber="317">
<emphasis id="4BB2D514A025FFE4FF0CF8437222F83A" bold="true" box="[151,287,1943,1967]" pageId="16" pageNumber="317">FIGURE 14.</emphasis>
Natural habitats of
<taxonomicName id="BEC67285A025FFE4FE7FF84371D1F83A" box="[484,748,1943,1967]" class="Insecta" family="Lucanidae" genus="Capreolucanus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="4BB2D514A025FFE4FE7FF8437143F83A" box="[484,638,1943,1967]" italics="true" pageId="16" pageNumber="317">Capreolucanus</emphasis>
species.
<emphasis id="4BB2D514A025FFE4FD42F84371D1F83A" bold="true" box="[729,748,1943,1967]" pageId="16" pageNumber="317">A</emphasis>
</taxonomicName>
<emphasis id="4BB2D514A025FFE4FD77F84371CCF83A" bold="true" box="[748,753,1943,1967]" pageId="16" pageNumber="317">,</emphasis>
Daweishan National Nature Reserve, Pingbian County, Yunnan, the natural habitat of
<taxonomicName id="BEC67285A025FFE4FED1F868711EF846" authority="Qi &amp; Zhou" authorityName="Qi &amp; Zhou" authorityYear="2024" box="[330,547,1979,2003]" class="Insecta" family="Lucanidae" genus="Capreolucanus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="yanxui" status="new species">
<emphasis id="4BB2D514A025FFE4FED1F8687266F846" box="[330,347,1980,2003]" italics="true" pageId="16" pageNumber="317">C</emphasis>
.
<emphasis id="4BB2D514A025FFE4FEF3F8687291F846" box="[360,428,1980,2003]" italics="true" pageId="16" pageNumber="317">yanxui</emphasis>
Qi &amp; Zhou
</taxonomicName>
,
<emphasis id="4BB2D514A025FFE4FDB5F86F7196F846" bold="true" box="[558,683,1979,2003]" pageId="16" pageNumber="317">
<taxonomicNameLabel id="5081686FA025FFE4FDB5F86F7196F846" box="[558,683,1979,2003]" pageId="16" pageNumber="317" rank="species">new species</taxonomicNameLabel>
</emphasis>
;
<emphasis id="4BB2D514A025FFE4FD2CF86F71F3F846" bold="true" box="[695,718,1979,2003]" pageId="16" pageNumber="317">B,</emphasis>
the mountains of Longchuan County, Yunnan, the natural habitat of
<taxonomicName id="BEC67285A025FFE4FA1EF8687258F862" authority="Didier, 1928" authorityName="Didier" authorityYear="1928" class="Insecta" family="Lucanidae" genus="Capreolucanus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="sicardi">
<emphasis id="4BB2D514A025FFE4FA1EF86876ABF846" box="[1413,1430,1980,2003]" italics="true" pageId="16" pageNumber="317">C</emphasis>
.
<emphasis id="4BB2D514A025FFE4FF0CF80B73E1F862" box="[151,220,2015,2039]" italics="true" pageId="16" pageNumber="317">sicardi</emphasis>
<bibRefCitation id="1D5774F7A025FFE4FF78F80B7258F862" author="Didier, R." box="[227,357,2015,2039]" pageId="16" pageNumber="317" pagination="65 - 67" refId="ref8340" refString="Didier, R. (1928) Etudes sur les Coleopteres Lucanides du globe. V. Descriptions de Lucanides nouveaux ou peu connus. 1. Description d'un genre nouveau et d'une espece nouvelle. Capreolucanus sicardi. Librairie speciale Agricole, Paris, Fascicule, 3, 65 - 67." type="journal article" year="1928">Didier, 1928</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="2DB9598EA024FFE5FF0CFBF17655FBF7" ID-DOI="http://doi.org/10.5281/zenodo.10954388" ID-Zenodo-Dep="10954388" httpUri="https://zenodo.org/record/10954388/files/figure.png" pageId="17" pageNumber="318" startId="17.[151,250,1061,1085]" targetBox="[151,1436,181,1037]" targetPageId="17" targetType="figure">
<paragraph id="79790906A024FFE5FF0CFBF17655FBF7" blockId="17.[151,1436,1061,1122]" pageId="17" pageNumber="318">
<emphasis id="4BB2D514A024FFE5FF0CFBF17222FBA8" bold="true" box="[151,287,1061,1085]" pageId="17" pageNumber="318">FIGURE 15.</emphasis>
Adults of
<taxonomicName id="BEC67285A024FFE5FE12FBF271CFFBAB" authority="Didier, 1928" authorityName="Didier" authorityYear="1928" box="[393,754,1062,1086]" class="Insecta" family="Lucanidae" genus="Capreolucanus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="sicardi">
<emphasis id="4BB2D514A024FFE5FE12FBF27150FBAB" box="[393,621,1062,1086]" italics="true" pageId="17" pageNumber="318">Capreolucanus sicardi</emphasis>
<bibRefCitation id="1D5774F7A024FFE5FDE8FBF271CFFBAB" author="Didier, R." box="[627,754,1062,1086]" pageId="17" pageNumber="318" pagination="65 - 67" refId="ref8340" refString="Didier, R. (1928) Etudes sur les Coleopteres Lucanides du globe. V. Descriptions de Lucanides nouveaux ou peu connus. 1. Description d'un genre nouveau et d'une espece nouvelle. Capreolucanus sicardi. Librairie speciale Agricole, Paris, Fascicule, 3, 65 - 67." type="journal article" year="1928">Didier, 1928</bibRefCitation>
</taxonomicName>
mating on standing decaying wood while the female biting the bark in broad-leaved forests of Gaoligongshan, Yunnan at an altitude of 1800 m at night in August (photo by Yan-Dong Chen).
</paragraph>
</caption>
<subSubSection id="31DC5A8DA024FFE5FF0CFB607113FB5B" box="[151,558,1203,1230]" pageId="17" pageNumber="318" type="multiple">
<paragraph id="79790906A024FFE5FF0CFB607113FB5B" blockId="17.[151,558,1203,1230]" box="[151,558,1203,1230]" pageId="17" pageNumber="318">
<emphasis id="4BB2D514A024FFE5FF0CFB607113FB5B" bold="true" box="[151,558,1203,1230]" pageId="17" pageNumber="318">
Key to females of
<taxonomicName id="BEC67285A024FFE5FEEDFB677113FB5B" ID-CoL="3H5Q" authorityName="Didier" authorityYear="1928" box="[374,558,1203,1230]" class="Insecta" family="Lucanidae" genus="Capreolucanus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="4BB2D514A024FFE5FEEDFB677113FB5B" bold="true" box="[374,558,1203,1230]" italics="true" pageId="17" pageNumber="318">Capreolucanus</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="31DC5A8DA024FFE5FF0CFB2D76A0F982" pageId="17" pageNumber="318" type="key">
<keyStep id="C23211A3A024FFE5FF0CFB2D76A1FAF3" pageId="17" pageNumber="318">
<paragraph id="79790906A024FFE5FF0CFB2D76A1FABE" blockId="17.[151,1437,1273,1559]" pageId="17" pageNumber="318">
<keyLead id="C237AA33A024FFE5FF0CFB2D76A1FABE" pageId="17" pageNumber="318">
1. Bicolor species, pronotum black to brown with two cross-like orange red stripes flank midline, all femurs orange red in about apical third (
<figureCitation id="E1FD1583A024FFE5FEC7FAC27294FABE" box="[348,425,1302,1324]" captionStart="FIGURE 7" captionStartId="9.[151,250,1908,1932]" captionTargetBox="[229,1359,184,1884]" captionTargetId="figure-17@9.[223,1365,181,1885]" captionTargetPageId="9" captionText="FIGURE 7. Female habitus of Capreolucanus species. A–C, C. yanxui Qi &amp; Zhou, new species (paratype, 19.4 mm, Pingbian, Yunnan, China); D–F, C. sicardi Didier, 1928 (19.0 mm, Yingjiang, Yunnan, China); G–I, C. zhuchuangi Wang, 2020 (16.8 mm, Bo Kluea, Nan, Thailand, photo by Wuttipon Pathomwattananurak). A, D, G, dorsal view; B, E, H, lateral view; C, F, I, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10954364" httpUri="https://zenodo.org/record/10954364/files/figure.png" pageId="17" pageNumber="318">Fig. 7G</figureCitation>
); mentum subtriangular (
<figureCitation id="E1FD1583A024FFE5FD00FAC271E1FABE" box="[667,732,1302,1324]" captionStart="FIGURE 7" captionStartId="9.[151,250,1908,1932]" captionTargetBox="[229,1359,184,1884]" captionTargetId="figure-17@9.[223,1365,181,1885]" captionTargetPageId="9" captionText="FIGURE 7. Female habitus of Capreolucanus species. A–C, C. yanxui Qi &amp; Zhou, new species (paratype, 19.4 mm, Pingbian, Yunnan, China); D–F, C. sicardi Didier, 1928 (19.0 mm, Yingjiang, Yunnan, China); G–I, C. zhuchuangi Wang, 2020 (16.8 mm, Bo Kluea, Nan, Thailand, photo by Wuttipon Pathomwattananurak). A, D, G, dorsal view; B, E, H, lateral view; C, F, I, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10954364" httpUri="https://zenodo.org/record/10954364/files/figure.png" pageId="17" pageNumber="318">Fig. 7I</figureCitation>
)...................................
<taxonomicName id="BEC67285A024FFE5FB02FAC276A1FABE" ID-CoL="7TBMT" authority="Wang, 2020" authorityName="Wang" authorityYear="2020" box="[1177,1436,1302,1324]" class="Insecta" family="Lucanidae" genus="Capreolucanus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="zhuchuangi">
<emphasis id="4BB2D514A024FFE5FB02FAC27794FABE" box="[1177,1193,1302,1323]" italics="true" pageId="17" pageNumber="318">C</emphasis>
.
<emphasis id="4BB2D514A024FFE5FB2EFAC27619FABE" box="[1205,1316,1302,1323]" italics="true" pageId="17" pageNumber="318">zhuchuangi</emphasis>
<bibRefCitation id="1D5774F7A024FFE5FAB2FAC376A1FABE" author="Wang, C. B." box="[1321,1436,1302,1324]" pageId="17" pageNumber="318" pagination="2 - 7" refId="ref8939" refString="Wang, C. B. (2020) The second species of Capreolucanus Didier, 1928 (Coleoptera: Lucanidae: Lucaninae) from Yunnan, China. Beetles world, 22, 2 - 7." type="journal article" year="2020">Wang, 2020</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="79790906A024FFE5FF0CFAE076A1FAF3" blockId="17.[151,1437,1273,1559]" pageId="17" pageNumber="318">
<keyLead id="C237AA33A024FFE5FF0CFAE076A1FAF3" pageId="17" pageNumber="318">
- Monochromatic species, pronotum black to brown without stripes, all femurs black (
<figureCitation id="E1FD1583A024FFE5FBBBFAE777B0FADC" box="[1056,1165,1331,1353]" captionStart="FIGURE 7" captionStartId="9.[151,250,1908,1932]" captionTargetBox="[229,1359,184,1884]" captionTargetId="figure-17@9.[223,1365,181,1885]" captionTargetPageId="9" captionText="FIGURE 7. Female habitus of Capreolucanus species. A–C, C. yanxui Qi &amp; Zhou, new species (paratype, 19.4 mm, Pingbian, Yunnan, China); D–F, C. sicardi Didier, 1928 (19.0 mm, Yingjiang, Yunnan, China); G–I, C. zhuchuangi Wang, 2020 (16.8 mm, Bo Kluea, Nan, Thailand, photo by Wuttipon Pathomwattananurak). A, D, G, dorsal view; B, E, H, lateral view; C, F, I, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10954364" httpUri="https://zenodo.org/record/10954364/files/figure.png" pageId="17" pageNumber="318">Fig. 7A, D</figureCitation>
); mentum not subtriangular (
<figureCitation id="E1FD1583A024FFE5FF72FA857268FAF3" box="[233,341,1361,1383]" captionStart="FIGURE 8" captionStartId="10.[151,250,927,951]" captionTargetBox="[151,1435,181,904]" captionTargetId="figure-21@10.[151,1435,181,904]" captionTargetPageId="10" captionText="FIGURE 8. Female canthus (A–F), clypeolabrum (G–L), and mentum (M–R) of Capreolucanus species. A–B, G–H, M–N, C. yanxui Qi &amp; Zhou, new species (paratypes); C–F, I–L, O–R, C. sicardi Didier, 1928. A–L, dorsal view; M–R, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10954370" httpUri="https://zenodo.org/record/10954370/files/figure.png" pageId="17" pageNumber="318">Fig. 8M–R</figureCitation>
)......................................................................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C23211A3A024FFE5FF0CFABA76A0F982" pageId="17" pageNumber="318">
<paragraph id="79790906A024FFE5FF0CFABA76A1FA2B" blockId="17.[151,1437,1273,1559]" pageId="17" pageNumber="318">
<keyLead id="C237AA33A024FFE5FF0CFABA76A1FA2B" pageId="17" pageNumber="318">
2. Canthus less developed, dividing about 1/6 of the eye (
<figureCitation id="E1FD1583A024FFE5FD62FABA7062FA16" box="[761,863,1390,1412]" captionStart="FIGURE 8" captionStartId="10.[151,250,927,951]" captionTargetBox="[151,1435,181,904]" captionTargetId="figure-21@10.[151,1435,181,904]" captionTargetPageId="10" captionText="FIGURE 8. Female canthus (A–F), clypeolabrum (G–L), and mentum (M–R) of Capreolucanus species. A–B, G–H, M–N, C. yanxui Qi &amp; Zhou, new species (paratypes); C–F, I–L, O–R, C. sicardi Didier, 1928. A–L, dorsal view; M–R, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10954370" httpUri="https://zenodo.org/record/10954370/files/figure.png" pageId="17" pageNumber="318">Fig. 8C–F</figureCitation>
); mentum with rounded apex (
<figureCitation id="E1FD1583A024FFE5FB12FABA77C9FA16" box="[1161,1268,1390,1412]" captionStart="FIGURE 8" captionStartId="10.[151,250,927,951]" captionTargetBox="[151,1435,181,904]" captionTargetId="figure-21@10.[151,1435,181,904]" captionTargetPageId="10" captionText="FIGURE 8. Female canthus (A–F), clypeolabrum (G–L), and mentum (M–R) of Capreolucanus species. A–B, G–H, M–N, C. yanxui Qi &amp; Zhou, new species (paratypes); C–F, I–L, O–R, C. sicardi Didier, 1928. A–L, dorsal view; M–R, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10954370" httpUri="https://zenodo.org/record/10954370/files/figure.png" pageId="17" pageNumber="318">Fig. 8O–R</figureCitation>
); outer margin of hemisternite membranous at middle, base of the outer margin thickening of sclerotization and extension backward obviously (
<figureCitation id="E1FD1583A024FFE5FF72FA7D7270FA2B" box="[233,333,1449,1471]" captionStart="FIGURE 9" captionStartId="11.[151,250,1401,1425]" captionTargetBox="[151,1432,188,1377]" captionTargetId="figure-17@11.[151,1435,181,1377]" captionTargetPageId="11" captionText="FIGURE 9. Female abdominal tergite VIII (A, F, K, P), abdominal ventrite VIII (B, G, L, Q) and genitalia (C, H, M, R) including details of hemisternite (D, I, N, S), spermatheca and spermathecal gland (E, J, O, T) of Capreolucanus species. A–J, C. yanxui Qi &amp; Zhou, new species (paratypes); K–T, C. sicardi Didier, 1928. A, F, K, P, dorsal view; B–E, G–J, L–O, Q–T, ventral view. Scale bar for E, J, O, T = 0.2 mm, for the others = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10960034" httpUri="https://zenodo.org/record/10960034/files/figure.png" pageId="17" pageNumber="318">Fig. 9N, S</figureCitation>
).........................................................................
<taxonomicName id="BEC67285A024FFE5FB5BFA7D76A1FA2B" ID-CoL="QSF2" authority="Didier, 1928" authorityName="Didier" authorityYear="1928" box="[1216,1436,1449,1471]" class="Insecta" family="Lucanidae" genus="Capreolucanus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="sicardi">
<emphasis id="4BB2D514A024FFE5FB5BFA7D77EDFA2B" box="[1216,1232,1449,1470]" italics="true" pageId="17" pageNumber="318">C</emphasis>
.
<emphasis id="4BB2D514A024FFE5FB47FA7D7623FA2B" box="[1244,1310,1449,1470]" italics="true" pageId="17" pageNumber="318">sicardi</emphasis>
<bibRefCitation id="1D5774F7A024FFE5FABFFA7D76A1FA2B" author="Didier, R." box="[1316,1436,1449,1471]" pageId="17" pageNumber="318" pagination="65 - 67" refId="ref8340" refString="Didier, R. (1928) Etudes sur les Coleopteres Lucanides du globe. V. Descriptions de Lucanides nouveaux ou peu connus. 1. Description d'un genre nouveau et d'une espece nouvelle. Capreolucanus sicardi. Librairie speciale Agricole, Paris, Fascicule, 3, 65 - 67." type="journal article" year="1928">Didier, 1928</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="79790906A024FFE5FF0CFA1376A0F982" blockId="17.[151,1437,1273,1559]" pageId="17" pageNumber="318">
<keyLead id="C237AA33A024FFE5FF0CFA1376A0F982" pageId="17" pageNumber="318">
- Canthus more developed, dividing about 1/4 of the eye (
<figureCitation id="E1FD1583A024FFE5FC94FA127046FA4E" box="[783,891,1478,1500]" captionStart="FIGURE 8" captionStartId="10.[151,250,927,951]" captionTargetBox="[151,1435,181,904]" captionTargetId="figure-21@10.[151,1435,181,904]" captionTargetPageId="10" captionText="FIGURE 8. Female canthus (A–F), clypeolabrum (G–L), and mentum (M–R) of Capreolucanus species. A–B, G–H, M–N, C. yanxui Qi &amp; Zhou, new species (paratypes); C–F, I–L, O–R, C. sicardi Didier, 1928. A–L, dorsal view; M–R, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10954370" httpUri="https://zenodo.org/record/10954370/files/figure.png" pageId="17" pageNumber="318">Fig. 8A–B</figureCitation>
); mentum subtrapezoidal (
<figureCitation id="E1FD1583A024FFE5FB1BFA1277CCFA4E" box="[1152,1265,1478,1500]" captionStart="FIGURE 8" captionStartId="10.[151,250,927,951]" captionTargetBox="[151,1435,181,904]" captionTargetId="figure-21@10.[151,1435,181,904]" captionTargetPageId="10" captionText="FIGURE 8. Female canthus (A–F), clypeolabrum (G–L), and mentum (M–R) of Capreolucanus species. A–B, G–H, M–N, C. yanxui Qi &amp; Zhou, new species (paratypes); C–F, I–L, O–R, C. sicardi Didier, 1928. A–L, dorsal view; M–R, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10954370" httpUri="https://zenodo.org/record/10954370/files/figure.png" pageId="17" pageNumber="318">Fig. 8M–N</figureCitation>
); outer margin of hemisternite sclerotized at middle, base of the outer margin not thickening and extension obviously (
<figureCitation id="E1FD1583A024FFE5FB33FA37763AFA6C" box="[1192,1287,1507,1529]" captionStart="FIGURE 9" captionStartId="11.[151,250,1401,1425]" captionTargetBox="[151,1432,188,1377]" captionTargetId="figure-17@11.[151,1435,181,1377]" captionTargetPageId="11" captionText="FIGURE 9. Female abdominal tergite VIII (A, F, K, P), abdominal ventrite VIII (B, G, L, Q) and genitalia (C, H, M, R) including details of hemisternite (D, I, N, S), spermatheca and spermathecal gland (E, J, O, T) of Capreolucanus species. A–J, C. yanxui Qi &amp; Zhou, new species (paratypes); K–T, C. sicardi Didier, 1928. A, F, K, P, dorsal view; B–E, G–J, L–O, Q–T, ventral view. Scale bar for E, J, O, T = 0.2 mm, for the others = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10960034" httpUri="https://zenodo.org/record/10960034/files/figure.png" pageId="17" pageNumber="318">Fig. 9D, I</figureCitation>
)...................................................................................
<taxonomicName id="BEC67285A024FFE5FBD4F9D57620F982" authority="Qi &amp; Zhou" authorityName="Qi &amp; Zhou" authorityYear="2024" box="[1103,1309,1537,1559]" class="Insecta" family="Lucanidae" genus="Capreolucanus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="yanxui" status="new species">
<emphasis id="4BB2D514A024FFE5FBD4F9D57762F983" box="[1103,1119,1537,1558]" italics="true" pageId="17" pageNumber="318">C</emphasis>
.
<emphasis id="4BB2D514A024FFE5FBF0F9D57796F983" box="[1131,1195,1537,1558]" italics="true" pageId="17" pageNumber="318">yanxui</emphasis>
Qi &amp; Zhou
</taxonomicName>
,
<emphasis id="4BB2D514A024FFE5FABDF9D576A0F982" bold="true" box="[1318,1437,1537,1559]" pageId="17" pageNumber="318">
<taxonomicNameLabel id="5081686FA024FFE5FABDF9D576A0F982" box="[1318,1437,1537,1559]" pageId="17" pageNumber="318" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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