374 lines
23 KiB
XML
374 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.672.10212" ID-GBIF-Dataset="62682ac8-f1bf-4e19-a4e3-0c24a5764441" ID-PMC="PMC5527339" ID-Pensoft-Pub="1313-2970-672-1" ID-PubMed="28769665" ID-ZBK="48810F7697B940B29A75DB0F98C5311F" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-672-1" ModsDocOrigin="ZooKeys 672" ModsDocTitle="New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae)" checkinTime="1493896456945" checkinUser="pensoft" docAuthor="Chanabun, Ratmanee, Inkavilay, Khamla & Panha, Somsak" docDate="2017" docId="830715AC146E9FADBBD0CED922F826CD" docLanguage="en" docName="ZooKeys 672: 1-34" docOrigin="ZooKeys 672" docSource="http://dx.doi.org/10.3897/zookeys.672.10212" docTitle="Glyphidrilus nanensis Chanabun & Panha, sp. n." docType="treatment" docUuid="E3CF3E43-7045-4C0F-9946-73AAB425425A" docUuidSource="ZooBank" docVersion="5" lastPageNumber="4" masterDocId="C91DFFAFFFFFFFF8F80DFFDC7D30FF88" masterDocTitle="New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae)" masterLastPageNumber="34" masterPageNumber="1" pageNumber="4" updateTime="1668164443210" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chanabun, Ratmanee</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Inkavilay, Khamla</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>672</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>34</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.672.10212</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.672.10212</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-672-1</mods:identifier>
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<mods:identifier type="ZBK">48810F7697B940B29A75DB0F98C5311F</mods:identifier>
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<mods:identifier type="ZooBank">48810F7697B940B29A75DB0F98C5311F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="128835294" LSID="urn:lsid:zoobank.org:act:E3CF3E43-7045-4C0F-9946-73AAB425425A" httpUri="http://treatment.plazi.org/id/830715AC146E9FADBBD0CED922F826CD" lastPageNumber="4" pageId="3" pageNumber="4">
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<subSubSection pageId="3" pageNumber="4" type="nomenclature">
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName LSID="http://zoobank.org/E3CF3E43-7045-4C0F-9946-73AAB425425A" authority="Chanabun & Panha" class="Clitellata" family="Almidae" genus="Glyphidrilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphidrilus nanensis" order="Opisthopora" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="nanensis">
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<pageBreakToken pageId="3" pageNumber="4" start="start">Glyphidrilus</pageBreakToken>
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nanensis Chanabun & Panha
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</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="4">sp. n.</taxonomicNameLabel>
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Figs 1, 3, 4, Table 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="type material">
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<paragraph pageId="3" pageNumber="4">Type material.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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Holotype: One adult (CUMZ 3403) in a rice field near Nan River at Saklek, Phichit, north Thailand (
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.507889">16°30'28.4"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="100.520836">100°31'15.0"E</geoCoordinate>
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), 49 meters elevation on 16 June 2012. Paratypes: 42 adults and 10 juveniles (CUMZ 3404), 2 adults (ZMH 14579), 2 adults (NHMUK), and 2 adults (ZRC), all same collection data as holotype.
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</paragraph>
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<caption pageId="3" pageNumber="4">
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<paragraph pageId="3" pageNumber="4">
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Figure 3. Photographs showing the A
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<taxonomicName class="Clitellata" family="Almidae" genus="Glyphidrilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphidrilus nanensis" order="Haplotaxida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="nanensis">Glyphidrilus nanensis</taxonomicName>
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sp. n. casts B type locality of
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<taxonomicName lsidName="G. nanensis" pageId="3" pageNumber="4" rank="species" species="nanensis">G. nanensis</taxonomicName>
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sp. n. in the rice field near Nan River at Saklek, Phichit, north Thailand C coloration of living paratype (CUMZ 3404).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="diagnosis">
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<paragraph pageId="3" pageNumber="4">Diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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A small sized earthworm with the clitellar wings on the lateral side of the body in XXIV, XXV,
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<normalizedToken originalValue="XXVI–">XXVI-</normalizedToken>
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XXVII, XXVIII, XXIX. Clitellum in XVII,
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<normalizedToken originalValue="XVIII–XXXIII">XVIII-XXXIII</normalizedToken>
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, XXXIV. Female pores, male pores, and spermathecal pores not visible. Genital markings: paired or asymmetrical on aa in X, XI, XII, XIII, XIV and XXVIII, XXIX, XXX; paired or asymmetrical on bc in XV, XVI,
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<normalizedToken originalValue="XVII–XXIII">XVII-XXIII</normalizedToken>
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, XXIV, XXV. Four pairs of seminal vesicles in
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<normalizedToken originalValue="IX–XII">IX-XII</normalizedToken>
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. Intestinal origin in XV. Ovaries in
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<normalizedToken originalValue="XIII–XIV">XIII-XIV</normalizedToken>
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. Spermathecae between 13/14-17/18.
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</paragraph>
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<caption pageId="3" pageNumber="4">
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<paragraph pageId="3" pageNumber="4">
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Figure 4. Morphology of holotype (CUMZ 3403) of
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<taxonomicName class="Clitellata" family="Almidae" genus="Glyphidrilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphidrilus nanensis" order="Haplotaxida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="nanensis">Glyphidrilus nanensis</taxonomicName>
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sp. n. A external ventral view, annular clitellum in
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<normalizedToken originalValue="XVII–XXXIII">XVII-XXXIII</normalizedToken>
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B internal dorsal view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="description">
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<paragraph pageId="3" pageNumber="4">Description of holotype.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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Dimensions: body length 78 mm, diameter 2.6 mm in segment VIII, 4.7 mm before the clitellar wing in segment XXIII, 4.2 mm after wing in segment XXX in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 241 segments. Body color pale brown with variations from red to pink at adjacent tissues of wing portion in different individuals of newly collected specimens. At posterior end dorsal surface considerably broader than the ventral. Clitellar wing on ventro-lateral part of clitellum in
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<normalizedToken originalValue="XXVI–XXIX">XXVI-XXIX</normalizedToken>
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, 3.6 mm in height, and 0.3 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in
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<normalizedToken originalValue="XVII–XXXIII">XVII-XXXIII</normalizedToken>
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. Four pairs of setae per segment from II, setal formula aa:ab:bc:cd:dd = 1.0:0.6:1.3:0.6:1.4 in segment VIII and 1.0:0.6:1.5:0.6:1.5 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: paired on aa in
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<normalizedToken originalValue="XII–XIII">XII-XIII</normalizedToken>
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, laterally paired or asymmetrical on bc in
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<normalizedToken originalValue="XVII–XXIV">XVII-XXIV</normalizedToken>
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.
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</paragraph>
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<paragraph pageId="3" pageNumber="4">
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Septa 5/6-8/9 thicker than between other segments, 9/10-14/15 thick and 15/16 to the last segment thin. Gizzard small, globular in 7/8. Intestine enlarged from XV. Hearts, five pairs in
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<normalizedToken originalValue="VII–XI">VII-XI</normalizedToken>
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, first in VII and last in XI. A pair of holonephridia in each segment from segment XIII onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries, two pairs in
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<normalizedToken originalValue="XIII–XIV">XIII-XIV</normalizedToken>
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. Testes free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, elongated oval or globular between 13/14-17/18, about 0.2-0.3 mm in diameter, two to ten on each side per segment.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="variation">
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<paragraph pageId="3" pageNumber="4">Variations.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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Body lengths of adult (n = 49) and juvenile paratypes (n = 10). Adults ranged from 72-99 mm (85.2
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<normalizedToken originalValue="±">+/-</normalizedToken>
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19.0), with 186-257 segments. Juveniles ranged from 73-93 mm (82.3
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<normalizedToken originalValue="±">+/-</normalizedToken>
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7.5), with 186-251 segments. Wings in XXIV, XXV,
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<normalizedToken originalValue="XXVI–XXVII">XXVI-XXVII</normalizedToken>
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, XXVIII, XXIX, clitellum in XVII, XVIII -XXXIII, XXXIV. Genital markings: paired or asymmetrical on aa in X, XI, XII, XIII, XIV and XXVIII, XXIX, XXX; paired or asymmetrical on bc in XV, XVI,
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<normalizedToken originalValue="XVII–XXIII">XVII-XXIII</normalizedToken>
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, XXIV, XXV.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="distribution">
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<paragraph pageId="3" pageNumber="4">Distribution.</paragraph>
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<paragraph pageId="3" pageNumber="4">The new species was found in the river banks of Nan River at Saklek, Phichit, north Thailand, which was covered with worm casts.</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="etymology">
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<paragraph pageId="3" pageNumber="4">Etymology.</paragraph>
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<paragraph pageId="3" pageNumber="4">The species was named after the locality on the banks of the Nan River. This was the first time that the worm genus was ever recorded from near this river.</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="remarks">
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<paragraph pageId="3" pageNumber="4">Remarks.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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For a summary of the following comparisons please see Table 1. The new species differs from
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<taxonomicName lsidName="G. yunnanensis" pageId="3" pageNumber="4" rank="species" species="yunnanensis">G. yunnanensis</taxonomicName>
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Chen & Xu, 1977 reported from China by the latter having longer wings in
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<normalizedToken originalValue="XXII–XXXII">XXII-XXXII</normalizedToken>
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, the clitellum in
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<normalizedToken originalValue="XVIII–XXXVIII">XVIII-XXXVIII</normalizedToken>
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, paired genital markings on bc in
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<normalizedToken originalValue="XVII–XXI">XVII-XXI</normalizedToken>
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and
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<normalizedToken originalValue="XXXII–XXXIV">XXXII-XXXIV</normalizedToken>
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, lacking a spermathecae. It differs from
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<taxonomicName lsidName="G. vangviengensis" pageId="3" pageNumber="4" rank="species" species="vangviengensis">G. vangviengensis</taxonomicName>
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Chanabun & Panha, 2011 from Laos by
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<taxonomicName lsidName="G. vangviengensis" pageId="3" pageNumber="4" rank="species" species="vangviengensis">G. vangviengensis</taxonomicName>
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having longer wings in XXIV,
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<normalizedToken originalValue="XXV–XXXI">XXV-XXXI</normalizedToken>
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, XXXII, the clitellum in XIX,
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<normalizedToken originalValue="XX–XXXV">XX-XXXV</normalizedToken>
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, XXXVI, XXXVII, and lacking spermathecae. It differs from the species recorded as
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<taxonomicName lsidName="G. mekongensis" pageId="3" pageNumber="4" rank="species" species="mekongensis">G. mekongensis</taxonomicName>
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Panha & Chanabun, 2012 from the Mekong River, Ubon Ratchathani, northeast Thailand by the latter species having longer wings in
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<normalizedToken originalValue="XXIV–½">XXIV-1/2</normalizedToken>
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XXXIII, XXXIII, XXXIV,
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<normalizedToken originalValue="½">1/2</normalizedToken>
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XXXV, the clitellum in
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<normalizedToken originalValue="XIX–XXXVII">XIX-XXXVII</normalizedToken>
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, XXXVIII, and lacking spermathecae. It differs from
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<taxonomicName lsidName="G. borealis" pageId="3" pageNumber="4" rank="species" species="borealis">G. borealis</taxonomicName>
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Chanabun & Panha, 2013 from Chiangmai, north Thailand by the latter having longer wings in XXI,
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<normalizedToken originalValue="XXII–XXVII">XXII-XXVII</normalizedToken>
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, XXVIII, XXIX, a longer clitellum in XIV, XVI,
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<normalizedToken originalValue="XVII–XXXI">XVII-XXXI</normalizedToken>
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, XXXII, XXXIII, XXXIV, XXXV, XXXVI, and the intestine enlarged from XIII. It differs from
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<taxonomicName lsidName="G. vangthongensis" pageId="3" pageNumber="4" rank="species" species="vangthongensis">G. vangthongensis</taxonomicName>
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Chanabun & Panha, 2013 reported from Phitsanulok, north Thailand by
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<taxonomicName lsidName="G. vangthongensis" pageId="3" pageNumber="4" rank="species" species="vangthongensis">G. vangthongensis</taxonomicName>
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having longer wings in XXIV, XXV,
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<normalizedToken originalValue="XXVI–XXXI">XXVI-XXXI</normalizedToken>
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, XXXII, a longer clitellum in XII, XIII, XIV, XV,
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<normalizedToken originalValue="XVI–XL">XVI-XL</normalizedToken>
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, XLI, XLII, an intestine enlarged from XIV, and spermathecae between 12/13-18/19. It differs from
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<taxonomicName lsidName="G. chaophraya" pageId="3" pageNumber="4" rank="species" species="chaophraya">G. chaophraya</taxonomicName>
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Chanabun & Panha, 2013 from Chaophraya River, Nakhonsawan, central Thailand by the latter having longer wings in XXIV,
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<normalizedToken originalValue="XXV–XXXII">XXV-XXXII</normalizedToken>
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, XXXIII, a longer clitellum in
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<normalizedToken originalValue="XX–XLIII">XX-XLIII</normalizedToken>
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, XLIV, XLV, and spermathecae between 16/17-22/23. It differs from
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<taxonomicName lsidName="G. chiensis" pageId="3" pageNumber="4" rank="species" species="chiensis">G. chiensis</taxonomicName>
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Chanabun & Panha, 2013 from Mahasarakham, northeast Thailand by
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<taxonomicName lsidName="G. chiensis" pageId="3" pageNumber="4" rank="species" species="chiensis">G. chiensis</taxonomicName>
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having wings in XXIII, XXIV, XXV,
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<normalizedToken originalValue="XXVI–XXIX">XXVI-XXIX</normalizedToken>
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, XXX, XXXI, XXXII, the clitellum in XVII,
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<normalizedToken originalValue="XVIII–XXXIII">XVIII-XXXIII</normalizedToken>
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, XXXIV, XXXV, XXXVI, XXXVII, XXXVIII, and spermathecae between 12/13-18/19. It differs from
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<taxonomicName lsidName="G. quadratus" pageId="3" pageNumber="4" rank="species" species="quadratus">G. quadratus</taxonomicName>
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Chanabun & Panha, 2013 reported from the Mun River drainage area, northeast Thailand by
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<taxonomicName lsidName="G. quadratus" pageId="3" pageNumber="4" rank="species" species="quadratus">G. quadratus</taxonomicName>
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having wings in XXIII,
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<normalizedToken originalValue="XXIV–XXVIII">XXIV-XXVIII</normalizedToken>
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, XXIX, XXX, XXXI, the clitellum in XV, XVI, XVII,
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<normalizedToken originalValue="XVIII–XXXI">XVIII-XXXI</normalizedToken>
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, XXXII, XXXIII, XXXIV, XXXV, XXXVI, and spermathecae between 12/13-17/18. It differs from
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<taxonomicName lsidName="G. huailuangensis" pageId="3" pageNumber="4" rank="species" species="huailuangensis">G. huailuangensis</taxonomicName>
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Chanabun & Panha, 2013 recorded from Najahlauy National Park, Ubon Ratchathani, northeast Thailand which has wings in XXV,
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<normalizedToken originalValue="XXVI–XXX">XXVI-XXX</normalizedToken>
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, XXXI, a clitellum in XII, XIII,
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<normalizedToken originalValue="XVI–XXXII">XVI-XXXII</normalizedToken>
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, XXXIII, and lacking spermathecae. It differs from
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<taxonomicName lsidName="G. sekongensis" pageId="3" pageNumber="4" rank="species" species="sekongensis">G. sekongensis</taxonomicName>
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sp. n. by
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<taxonomicName lsidName="G. sekongensis" pageId="3" pageNumber="4" rank="species" species="sekongensis">G. sekongensis</taxonomicName>
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sp. n. having longer wings in
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<normalizedToken originalValue="XXV–XXXI">XXV-XXXI</normalizedToken>
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, a longer clitellum in XVI,
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<normalizedToken originalValue="XVII–XXXVI">XVII-XXXVI</normalizedToken>
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, XXXVII, and spermathecae between 12/13-15/16 (see Table 1).
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</paragraph>
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<caption pageId="3" pageNumber="4">
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<paragraph pageId="3" pageNumber="4">
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Table 1. The comparison of characters among
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<taxonomicName class="Clitellata" family="Almidae" genus="Glyphidrilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphidrilus" order="Haplotaxida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">Glyphidrilus</taxonomicName>
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species from Thailand and Laos. The morphological characters of the species described are from type specimens and original description.
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</paragraph>
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</caption>
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<paragraph pageId="3" pageNumber="4">
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<table pageId="3" pageNumber="4">
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<tr pageId="3" pageNumber="4">
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<th colspan="1" pageId="3" pageNumber="4" rowspan="2">Species</th>
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<th colspan="1" pageId="3" pageNumber="4" rowspan="2">Length (mm)</th>
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<th colspan="1" pageId="3" pageNumber="4" rowspan="2">Segments</th>
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<th colspan="1" pageId="3" pageNumber="4" rowspan="2">
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<abbrev id="ABBRID0E1OAE" pageId="3" pageNumber="4">Cli.</abbrev>
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</th>
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<th colspan="1" pageId="3" pageNumber="4" rowspan="2">
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<abbrev id="ABBRID0ECPAE" pageId="3" pageNumber="4">Wi.</abbrev>
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</th>
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<th colspan="3" pageId="3" pageNumber="4" rowspan="1">
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<abbrev id="ABBRID0EKPAE" pageId="3" pageNumber="4">Gm.</abbrev>
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</th>
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<th colspan="1" pageId="3" pageNumber="4" rowspan="2">
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<abbrev id="ABBRID0ESPAE" pageId="3" pageNumber="4">He.</abbrev>
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</th>
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<th colspan="1" pageId="3" pageNumber="4" rowspan="2">
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<abbrev id="ABBRID0E1PAE" pageId="3" pageNumber="4">In.</abbrev>
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</th>
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<th colspan="1" pageId="3" pageNumber="4" rowspan="2">
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<abbrev id="ABBRID0ECQAE" pageId="3" pageNumber="4">Gi.</abbrev>
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</th>
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<th colspan="1" pageId="3" pageNumber="4" rowspan="2">
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<abbrev id="ABBRID0EKQAE" pageId="3" pageNumber="4">Sc.</abbrev>
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</th>
|
||
<th colspan="1" pageId="3" pageNumber="4" rowspan="2">Type locality</th>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<th colspan="1" pageId="3" pageNumber="4" rowspan="1">Paired on bc</th>
|
||
<th colspan="1" pageId="3" pageNumber="4" rowspan="1">Paired on aa</th>
|
||
<th colspan="1" pageId="3" pageNumber="4" rowspan="1">Un paired on aa</th>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. mekongensis" pageId="3" pageNumber="4" rank="species" species="mekongensis">G. mekongensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. vangviengensis" pageId="3" pageNumber="4" rank="species" species="vangviengensis">G. vangviengensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. yunnanensis" pageId="3" pageNumber="4" rank="species" species="yunnanensis">G. yunnanensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. borealis" pageId="3" pageNumber="4" rank="species" species="borealis">G. borealis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. vangthongensis" pageId="3" pageNumber="4" rank="species" species="vangthongensis">G. vangthongensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. chaophraya" pageId="3" pageNumber="4" rank="species" species="chaophraya">G. chaophraya</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. chiensis" pageId="3" pageNumber="4" rank="species" species="chiensis">G. chiensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. quadratus" pageId="3" pageNumber="4" rank="species" species="quadratus">G. quadratus</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. huailuangensis" pageId="3" pageNumber="4" rank="species" species="huailuangensis">G. huailuangensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. trangensis" pageId="3" pageNumber="4" rank="species" species="trangensis">G. trangensis</taxonomicName>
|
||
</td>
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">++</td>
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">++</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. wararamensis" pageId="3" pageNumber="4" rank="species" species="wararamensis">G. wararamensis</taxonomicName>
|
||
</td>
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">+</td>
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">+</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. kratuensis" pageId="3" pageNumber="4" rank="species" species="kratuensis">G. kratuensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. nanensis" pageId="3" pageNumber="4" rank="species" species="nanensis">G. nanensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. satunensis" pageId="3" pageNumber="4" rank="species" species="satunensis">G. satunensis</taxonomicName>
|
||
</td>
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">+</td>
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">+</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. chiangraiensis" pageId="3" pageNumber="4" rank="species" species="chiangraiensis">G. chiangraiensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. namphao" pageId="3" pageNumber="4" rank="species" species="namphao">G. namphao</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. sekongensis" pageId="3" pageNumber="4" rank="species" species="sekongensis">G. sekongensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. namdonensis" pageId="3" pageNumber="4" rank="species" species="namdonensis">G. namdonensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="4">
|
||
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
|
||
<taxonomicName lsidName="G. champasakensis" pageId="3" pageNumber="4" rank="species" species="champasakensis">G. champasakensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |