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<mods:title id="FCB66A15EA5A537504D8D8CA680CC96D">Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records</mods:title>
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<treatment id="F07F612D480164C329AF890E6C1DB8CE" ID-GBIF-Taxon="154127691" LSID="urn:lsid:plazi:treatment:F07F612D480164C329AF890E6C1DB8CE" httpUri="http://treatment.plazi.org/id/F07F612D480164C329AF890E6C1DB8CE" lastPageNumber="68" pageId="67" pageNumber="68">
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<taxonomicName id="522F10321D5CDEA0C9AFDDE91C94E390" ID-CoL="7ZMPC" authority="Arnaud, 1997" authorityName="Arnaud" authorityYear="1997" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophanaeus (Coprophanaeus) edmondsi" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="edmondsi" subGenus="Coprophanaeus">Coprophanaeus (Coprophanaeus) edmondsi Arnaud, 1997</taxonomicName>
Plate 16B
</paragraph>
</subSubSection>
<subSubSection id="DB63532907853B6615C2BAD5E5F37238" pageId="67" pageNumber="68" type="reference_group">
<paragraph id="36231EB5A33A2356A4BF95F06BEFA7EC" pageId="67" pageNumber="68">
<taxonomicName id="7B5DD88BF8C635044796266D5D860495" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophanaeus edmondsi" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="edmondsi">Coprophanaeus edmondsi</taxonomicName>
Arnaud, 1997: 5 (original description. Type locality: Pich. 4600 feet [= 1400 m], 23 km E of
<normalizedToken id="89100467CE355E3F606EA396ECFB0E6C" originalValue="Alluriquín">Alluriquin</normalizedToken>
, Chiriboga Road).
</paragraph>
<paragraph id="D85C7331FCFC9738757D063923D9C87A" pageId="67" pageNumber="68">
<taxonomicName id="AC7021ED56AB7CE2842B680980B19B19" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophanaeus edmondsi" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="edmondsi">Coprophanaeus edmondsi</taxonomicName>
:
<bibRefCitation id="417026646AFBD481F5812572B0F8CF89" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
: 320-321 (cited for Ecuador).
</paragraph>
<paragraph id="12C057FCBFB956D68FE00B0C32A82223" pageId="67" pageNumber="68">
<taxonomicName id="539EAFE77B9C1A515D1926E307154918" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophanaeus (Coprophanaeus) edmondsi" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="edmondsi" subGenus="Coprophanaeus">Coprophanaeus (Coprophanaeus) edmondsi</taxonomicName>
:
<bibRefCitation id="0F6B01EC2A14E73E2028F03F36F9915B" author="Medina, CA" journalOrPublisher="Biota Colombiana" pageId="274" pageNumber="275" pagination="131 - 144" title="Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia." volume="2" year="2001">Medina et al. 2001</bibRefCitation>
: 140 (cited for Colombia);
<bibRefCitation id="0B62320B8D9DB5EC7EF764E0158E9F2F" author="Arnaud, P" journalOrPublisher="Hillside Books, Canterbury, England" pageId="261" pageNumber="262" title="Les Coleopteres du Monde, Vol. 28. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus." year="2002 a">Arnaud 2002a</bibRefCitation>
: 47 (diagnosis);
<bibRefCitation id="436E1FB1CA136F7312456F6BD86B44E5" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
Edmonds and
<normalizedToken id="2F6845F9F6D88B4486B069C92EEE5FAD" originalValue="Zídek">Zidek</normalizedToken>
2010
</bibRefCitation>
: 69 (cited as synonym of
<taxonomicName id="57587FA144FE6C5ACE6F56E3C0421B5C" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophanaeus conocephalus" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="conocephalus">Coprophanaeus conocephalus</taxonomicName>
<normalizedToken id="9F13D4AF86C06F08DC8B6CD35FC45C54" originalValue="dOlsoufieff">d'Olsoufieff</normalizedToken>
, 1924); Arnaud, 2018: 6 (revalidated name), 7 (figure 2b).
</paragraph>
</subSubSection>
<subSubSection id="90AD1FE60925ED4DB46F6B24CBD1684D" pageId="67" pageNumber="68" type="type specimens">
<paragraph id="7B7808F6105D767A482D6168915A4159" pageId="67" pageNumber="68">Type specimens.</paragraph>
<paragraph id="D3741F12227600BCBBA07916DD4D0851" pageId="67" pageNumber="68">
<taxonomicName id="8E2CCBA4A86DB6207C543DA6F0400145" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophanaeus edmondsi" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="edmondsi">Coprophanaeus edmondsi</taxonomicName>
Arnaud, 1997. The holotype (♂) is deposited at the CMNC (formerly in A and H Howden collection, see
<bibRefCitation id="7D37F407AF08A71211606EBA1FB91AAB" author="Arnaud, P" journalOrPublisher="Besoiro" pageId="260" pageNumber="261" pagination="4 - 8" title="Description de nouvelles especes du genre Coprophanaeus Ols." volume="4" year="1997">Arnaud 1997</bibRefCitation>
: 6). Locality: Pich. 4600 feet [= 1400 m], 23 km E of
<normalizedToken id="E47BF9846D266E88C3C97ACFC2EED0B7" originalValue="Alluriquín">Alluriquin</normalizedToken>
, Chiriboga Road, not examined. One paratype is deposited in the MQCAZ. Javin, examined.
</paragraph>
<paragraph id="E89264726D28B338D332C1208DAF017F" pageId="67" pageNumber="68">Paratype (♂): &quot;ECUADOR (CAN) / Javin II. 92 / 850 - 1400m / P. Arnaud leg [p]&quot;, &quot;Coprophanaeus / edmondsi / 94 / P.ARNAUD DET /PARATYPE ♂ [p, pink margin]&quot;.</paragraph>
</subSubSection>
<subSubSection id="2CBA63F93DB115A99DE4C26DBFA21C2E" pageId="67" pageNumber="68" type="distribution">
<paragraph id="9D56D028BD7DC1A39DEA41B0D259E3CD" pageId="67" pageNumber="68">Distribution.</paragraph>
<paragraph id="E2EE8A92A019D557E3F56B541A72E12A" pageId="67" pageNumber="68">Colombia and Ecuador.</paragraph>
</subSubSection>
<subSubSection id="03147E7CDDC8C93F19A4F76FFD8CB45B" pageId="67" pageNumber="68" type="records examined">
<paragraph id="08E4B104B3235D270D10BCFABB8E5C7D" pageId="67" pageNumber="68">Records examined.</paragraph>
<paragraph id="0AD4E0B9FF4517E87024CF07550634AD" pageId="67" pageNumber="68">
<normalizedToken id="6ED7F7E3D63FCCE666D898AF645D0DE8" originalValue="CAÑAR">CANAR</normalizedToken>
: Javin, 850-1400 m (3 specimens CEMT; 1 specimen MQCAZ). CARCHI: Goaltal, Hacienda San Francisco, 1200 m (5 specimens MECN); Maldonado, 1830 m (2 specimens CEMT). PICHINCHA: Curipoglio, Cerro San Cristobal (1 specimen MUTPL); Mindo, 1500 m (1 specimen MUTPL);
<normalizedToken id="1BC9E76388CA3E87737F6CC5CE63B0BC" originalValue="Estación">Estacion</normalizedToken>
<normalizedToken id="6AC286081DDE57B2A2D1C6333A4A23A8" originalValue="Biológica">Biologica</normalizedToken>
la Hesperia (1 specimen MUTPL).
</paragraph>
</subSubSection>
<subSubSection id="8ADF79A4CD926ED968D388E3FB905B30" pageId="67" pageNumber="68" type="literature records">
<paragraph id="6DFCE2C7180EE9F3CF192E30B95D77D5" pageId="67" pageNumber="68">Literature records.</paragraph>
<paragraph id="07F06A140F46D9A935D903F23CC51684" pageId="67" pageNumber="68">PICHINCHA: 4600 feet [1400 m], 23 km E de Alluriquin, Chiriboga Road (Arnaud, 1997: 6); Chiriboga Road 1200-1830, km 12-20 m (Arnaud, 1997: 6).</paragraph>
</subSubSection>
<subSubSection id="F86F53D174D573761B810793F4F40B06" pageId="67" pageNumber="68" type="temporal data">
<paragraph id="D692D272D5726A1BDF94A8D5ECA692FE" pageId="67" pageNumber="68">Temporal data.</paragraph>
<paragraph id="E7A4EBEDAF9E569659F92BBC0FF82B19" pageId="67" pageNumber="68">Collected in February, March, June, September, October, and December.</paragraph>
</subSubSection>
<subSubSection id="5E409179D3DB86C63126976ECA514C6D" pageId="67" pageNumber="68" type="remarks">
<paragraph id="DEC36ECA8B7C06CC4C1D570129588715" pageId="67" pageNumber="68">Remarks.</paragraph>
<paragraph id="461235B5530DFA1AFD3DF140F205AEF6" pageId="67" pageNumber="68">Inhabits coastal foothill evergreen forests from 850-1200 m a.s.l. In the Andean region it has been registered in montane evergreen forests from 1400-1800 m a.s.l. Collected with pitfall traps baited with carrion and human feces.</paragraph>
</subSubSection>
</treatment>
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