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<document id="1E680BC6F6C4828D5565186B0FFBD95F" ID-CLB-Dataset="30423" ID-DOI="10.11646/zootaxa.4399.2.7" ID-GBIF-Dataset="6c5127b3-a849-4ea8-a965-8e915f63a551" ID-ISSN="1175-5326" ID-Zenodo-Dep="1207158" ID-ZooBank="073E1E48-50BE-42DE-B21A-FA9043786E23" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521995446897" checkinUser="plazi" docAuthor="Couri, Márcia S. &amp; Pont, Adrian C." docDate="2018" docId="CD02BC11FFD6E565FF5CD686FBAF51E1" docLanguage="en" docName="zootaxa.4399.2.7.pdf.imf" docOrigin="Zootaxa 4399 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helina lasiopa Stein 1913" docType="treatment" docVersion="7" lastPageNumber="240" masterDocId="313BC469FFD1E562FFCBD567FFB0542C" masterDocTitle="The male terminalia of some African species of Helina Robineau-Desvoidy, 1830 (Diptera, Muscidae)" masterLastPageNumber="247" masterPageNumber="233" pageNumber="240" updateTime="1698533436902" updateUser="ExternalLinkService">
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<mods:title id="F2EBEA574FEB2B7AFBC3B5246687B405">The male terminalia of some African species of Helina Robineau-Desvoidy, 1830 (Diptera, Muscidae)</mods:title>
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<mods:namePart id="8A7BB84D92C3DC5B296E52F1C79533A5">Couri, Márcia S.</mods:namePart>
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<mods:namePart id="4260801EBC0B3C1AF34C94639643AC4B">Pont, Adrian C.</mods:namePart>
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<mods:date id="DBCC08D8D69FDD6702781BA030DC8653">2018</mods:date>
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<mods:number id="D467B7A7CD616FB56C02D7D8129FAF7B">2018-03-21</mods:number>
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<mods:number id="6AF57448883967DE527154DE725C79DB">4399</mods:number>
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<treatment id="CD02BC11FFD6E565FF5CD686FBAF51E1" ID-DOI="http://doi.org/10.5281/zenodo.6489503" ID-GBIF-Taxon="142611013" ID-Zenodo-Dep="6489503" LSID="urn:lsid:plazi:treatment:CD02BC11FFD6E565FF5CD686FBAF51E1" httpUri="http://treatment.plazi.org/id/CD02BC11FFD6E565FF5CD686FBAF51E1" lastPageNumber="240" pageId="7" pageNumber="240">
<subSubSection id="0DB15E8CFFD6E565FF5CD686FDBC57D7" box="[151,524,993,1019]" pageId="7" pageNumber="240" type="nomenclature">
<paragraph id="45140D07FFD6E565FF5CD686FDBC57D7" blockId="7.[151,524,993,1053]" box="[151,524,993,1019]" pageId="7" pageNumber="240">
<heading id="1E5CBA6BFFD6E565FF5CD686FDBC57D7" bold="true" box="[151,524,993,1019]" fontSize="11" level="1" pageId="7" pageNumber="240" reason="1">
<taxonomicName id="82AB7684FFD6E565FF5CD686FDBC57D7" ID-CoL="6LJSD" authority="(Stein, 1913)" authorityName="Stein" authorityYear="1913" box="[151,524,993,1019]" class="Insecta" family="Muscidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="240" phylum="Arthropoda" rank="species" species="lasiopa">
<emphasis id="77DFD115FFD6E565FF5CD686FDBC57D7" bold="true" box="[151,524,993,1019]" pageId="7" pageNumber="240">
<emphasis id="77DFD115FFD6E565FF5CD686FED957D7" bold="true" box="[151,361,993,1019]" italics="true" pageId="7" pageNumber="240">Helina nemoralis</emphasis>
(
<bibRefCitation id="213A70F6FFD6E565FEB2D686FDB357D7" box="[377,515,993,1019]" pageId="7" pageNumber="240" refId="ref5056" refString="Stein, P. (1913) Neue afrikanische Anthomyiden. Annales historico-naturales Musei nationalis Hungarici, 11, 457 - 583." type="journal article">Stein, 1913</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0DB15E8CFFD6E565FF5CD163FE975031" box="[151,295,1028,1053]" pageId="7" pageNumber="240" type="description">
<paragraph id="45140D07FFD6E565FF5CD163FE975031" blockId="7.[151,524,993,1053]" box="[151,295,1028,1053]" pageId="7" pageNumber="240">
(
<figureCitation id="DD901182FFD6E565FF54D163FEAF5031" box="[159,287,1028,1053]" captionStart="FIGURES 35–41" captionStartId="10.[151,264,1916,1939]" captionTargetBox="[199,1366,207,1849]" captionTargetId="figure@10.[189,1392,193,1894]" captionTargetPageId="10" captionText="FIGURES 35–41. Helina nemoralis (Stein, 1913), male: (35) sternite 5, dorsal view; (36) cercal plate and surstylus, dorsal view; (37) cercal plate and surstylus, lateral view; (38) aedeagal complex, lateral view. Helina novarae (Schiner, 1868), male: (39) sternite 5, dorsal view; (40) cercal plate and surstylus, dorsal view; (41) cercal plate, surstylus and aedeagal complex, lateral view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/1207172/files/figure.png" pageId="7" pageNumber="240">Figs 35–38</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0DB15E8CFFD6E565FF5CD12BFE8E50A4" pageId="7" pageNumber="240" type="materials_examined">
<paragraph id="45140D07FFD6E565FF5CD12BFE8E50A4" blockId="7.[151,1436,1100,1485]" pageId="7" pageNumber="240">
<emphasis id="77DFD115FFD6E565FF5CD12BFE735049" bold="true" box="[151,451,1100,1125]" pageId="7" pageNumber="240">
Examined
<typeStatus id="9A10B3A5FFD6E565FED2D12BFEFC5049" box="[281,332,1100,1125]" pageId="7" pageNumber="240">type</typeStatus>
material:
</emphasis>
<typeStatus id="9A10B3A5FFD6E565FE07D12BFD835048" box="[460,563,1100,1124]" pageId="7" pageNumber="240" type="syntype">Syntypes</typeStatus>
destroyed (
<bibRefCitation id="213A70F6FFD6E565FD73D12BFCEF5048" box="[696,863,1100,1124]" pageId="7" pageNumber="240" refId="ref4789" refString="Pont, A. C. (2013) The Fanniidae and Muscidae (Diptera) described by Paul Stein (1852 - 1921). Zoosystematics and Evolution, 89, 31 - 166. https: // doi. org / 10.1002 / zoos. 201300004" type="journal article">Pont 2013: 90</bibRefCitation>
). One male from
<collectingCountry id="3DBC4D97FFD6E565FBE2D12AFBC25049" box="[1065,1138,1101,1125]" name="Kenya" pageId="7" pageNumber="240">Kenya</collectingCountry>
(Mt. Kinangop) dissected and illustrated.
</paragraph>
</subSubSection>
<subSubSection id="0DB15E8CFFD6E565FF0CD1F3FAD1514C" pageId="7" pageNumber="240" type="diagnosis">
<paragraph id="45140D07FFD6E565FF0CD1F3FCD650D9" blockId="7.[151,1436,1100,1485]" pageId="7" pageNumber="240">
<emphasis id="77DFD115FFD6E565FF0CD1F3FEF15081" bold="true" box="[199,321,1172,1197]" pageId="7" pageNumber="240">Diagnosis.</emphasis>
Palpus dark brown; scutum dark brown-grey pollinose; dorsocentrals 1+4; crossvein dm–cu uniformly and sometimes very slightly infuscated; abdominal tergites each with one pair of round dark marks, syntergite 1+2 and tergites 3–4 also with a median brown stripe.
</paragraph>
<paragraph id="45140D07FFD6E565FF0CD198FAD1514C" blockId="7.[151,1436,1100,1485]" pageId="7" pageNumber="240">
<emphasis id="77DFD115FFD6E565FF0CD198FECB5134" box="[199,379,1279,1304]" italics="true" pageId="7" pageNumber="240">Male terminalia</emphasis>
. Sternite 5 longer than wide, with posterior membrane deep, with short setae on disc and lobes (
<figureCitation id="DD901182FFD6E565FF54D043FF455111" box="[159,245,1316,1341]" captionStart="FIGURES 35–41" captionStartId="10.[151,264,1916,1939]" captionTargetBox="[199,1366,207,1849]" captionTargetId="figure@10.[189,1392,193,1894]" captionTargetPageId="10" captionText="FIGURES 35–41. Helina nemoralis (Stein, 1913), male: (35) sternite 5, dorsal view; (36) cercal plate and surstylus, dorsal view; (37) cercal plate and surstylus, lateral view; (38) aedeagal complex, lateral view. Helina novarae (Schiner, 1868), male: (39) sternite 5, dorsal view; (40) cercal plate and surstylus, dorsal view; (41) cercal plate, surstylus and aedeagal complex, lateral view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/1207172/files/figure.png" pageId="7" pageNumber="240">Fig. 35</figureCitation>
); cercal plate and surstylus short (
<figureCitation id="DD901182FFD6E565FD4ED043FD6C5111" box="[645,732,1316,1341]" captionStart="FIGURES 35–41" captionStartId="10.[151,264,1916,1939]" captionTargetBox="[199,1366,207,1849]" captionTargetId="figure@10.[189,1392,193,1894]" captionTargetPageId="10" captionText="FIGURES 35–41. Helina nemoralis (Stein, 1913), male: (35) sternite 5, dorsal view; (36) cercal plate and surstylus, dorsal view; (37) cercal plate and surstylus, lateral view; (38) aedeagal complex, lateral view. Helina novarae (Schiner, 1868), male: (39) sternite 5, dorsal view; (40) cercal plate and surstylus, dorsal view; (41) cercal plate, surstylus and aedeagal complex, lateral view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/1207172/files/figure.png" pageId="7" pageNumber="240">Fig. 36</figureCitation>
); surstylus shorter than cercal plate in lateral view (
<figureCitation id="DD901182FFD6E565FAF3D043FA3E5110" box="[1336,1422,1316,1341]" captionStart="FIGURES 35–41" captionStartId="10.[151,264,1916,1939]" captionTargetBox="[199,1366,207,1849]" captionTargetId="figure@10.[189,1392,193,1894]" captionTargetPageId="10" captionText="FIGURES 35–41. Helina nemoralis (Stein, 1913), male: (35) sternite 5, dorsal view; (36) cercal plate and surstylus, dorsal view; (37) cercal plate and surstylus, lateral view; (38) aedeagal complex, lateral view. Helina novarae (Schiner, 1868), male: (39) sternite 5, dorsal view; (40) cercal plate and surstylus, dorsal view; (41) cercal plate, surstylus and aedeagal complex, lateral view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/1207172/files/figure.png" pageId="7" pageNumber="240">Fig. 37</figureCitation>
); aedeagal complex with phallapodeme short, and epiphallus long and curved, longer than postgonite (
<figureCitation id="DD901182FFD6E565FACBD020FAE4514C" box="[1280,1364,1351,1376]" captionStart="FIGURES 35–41" captionStartId="10.[151,264,1916,1939]" captionTargetBox="[199,1366,207,1849]" captionTargetId="figure@10.[189,1392,193,1894]" captionTargetPageId="10" captionText="FIGURES 35–41. Helina nemoralis (Stein, 1913), male: (35) sternite 5, dorsal view; (36) cercal plate and surstylus, dorsal view; (37) cercal plate and surstylus, lateral view; (38) aedeagal complex, lateral view. Helina novarae (Schiner, 1868), male: (39) sternite 5, dorsal view; (40) cercal plate and surstylus, dorsal view; (41) cercal plate, surstylus and aedeagal complex, lateral view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/1207172/files/figure.png" pageId="7" pageNumber="240">Fig. 38</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0DB15E8CFFD6E565FF0CD00BFBAF51E1" pageId="7" pageNumber="240" type="discussion">
<paragraph id="45140D07FFD6E565FF0CD00BFBAF51E1" blockId="7.[151,1436,1100,1485]" pageId="7" pageNumber="240">
<emphasis id="77DFD115FFD6E565FF0CD00BFEA151A9" bold="true" box="[199,273,1388,1413]" pageId="7" pageNumber="240">Notes.</emphasis>
This species was originally described from
<collectingCountry id="3DBC4D97FFD6E565FCD2D00BFC1E51A9" box="[793,942,1388,1413]" name="South Africa" pageId="7" pageNumber="240">
<collectingRegion id="876FC3E5FFD6E565FCD2D00BFCEA51A9" box="[793,858,1388,1413]" country="Cameroon" name="South" pageId="7" pageNumber="240">South</collectingRegion>
Africa
</collectingCountry>
in the genus
<taxonomicName id="82AB7684FFD6E565FB99D00BFF7F5184" authority="Robineau-Desvoidy, 1830" authorityName="Robineau-Desvoidy" authorityYear="1830" class="Insecta" family="Muscidae" genus="Mydaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="240" phylum="Arthropoda" rank="genus">
<emphasis id="77DFD115FFD6E565FB99D00BFB1C51A9" box="[1106,1196,1388,1413]" italics="true" pageId="7" pageNumber="240">Mydaea</emphasis>
<bibRefCitation id="213A70F6FFD6E565FB7CD00BFF7F5184" pageId="7" pageNumber="240" refId="ref4837" refString="Robineau-Desvoidy, A. J. B. (1830) Essai sur les Myodaires. Memoires Presentes Par divers Savants a l' Academie Royale des Sciences de l'Institut de France, 2, 1 - 813." type="journal article">Robineau-Desvoidy, 1830</bibRefCitation>
</taxonomicName>
and was placed in
<emphasis id="77DFD115FFD6E565FE66D0E8FE495184" box="[429,505,1423,1448]" italics="true" pageId="7" pageNumber="240">Helina</emphasis>
, subgenus
<taxonomicName id="82AB7684FFD6E565FDB2D0E8FC2F5184" authority="Malloch, 1921" authorityName="Malloch" authorityYear="1921" box="[633,927,1423,1448]" class="Insecta" family="Muscidae" genus="Euspilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="240" phylum="Arthropoda" rank="genus">
<emphasis id="77DFD115FFD6E565FDB2D0E8FD405184" box="[633,752,1423,1448]" italics="true" pageId="7" pageNumber="240">Euspilaria</emphasis>
<bibRefCitation id="213A70F6FFD6E565FD33D0E8FC2F5184" box="[760,927,1423,1448]" pageId="7" pageNumber="240" refId="ref4609" refString="Malloch, J. R. (1921) Exotic Muscaridae (Diptera). - III. Annals and Magazine of Natural History, 8 (9), 225 - 239. https: // doi. org / 10.1080 / 00222932108632584" type="journal article">Malloch, 1921</bibRefCitation>
</taxonomicName>
, by
<bibRefCitation id="213A70F6FFD6E565FC19D0E8FBC85184" box="[978,1144,1423,1448]" pageId="7" pageNumber="240" refId="ref4484" refString="Emden, F. I. van (1951) Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoninae. In: Ruwenzori ExPedition 1934 - 35. Vol. 2. British Museum (Natural History), London, pp. 325 - 710." type="book chapter">Emden (1951)</bibRefCitation>
. It is easily recognized in Emden’s key to
<taxonomicName id="82AB7684FFD6E565FE87D0D3FDFD51E1" authorityName="Malloch" authorityYear="1921" box="[332,589,1460,1485]" class="Insecta" family="Muscidae" genus="Euspilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="240" phylum="Arthropoda" rank="genus">
<emphasis id="77DFD115FFD6E565FE87D0D3FE7351E1" box="[332,451,1460,1485]" italics="true" pageId="7" pageNumber="240">Euspilaria</emphasis>
(1951: 614)
</taxonomicName>
by having four postsutural dorsocentrals.
</paragraph>
</subSubSection>
</treatment>
</document>
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