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<mods:title id="AED1850F71E948258BE72CD63913650A">A Revision of Mesopodagrion McLachlan, 1896 (Odonata: Zygoptera: Megapodagrionidae)</mods:title>
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<taxonomicName id="7BA3D8B6FFA9D97DFF6BFDB1E45DFDDE" ID-CoL="42586" authority="McLachlan, 1896" authorityName="McLachlan" authorityYear="1896" box="[151,821,536,562]" class="Insecta" family="Megapodagrionidae" genus="Mesopodagrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="64" phylum="Arthropoda" rank="subSpecies" species="tibetanum" subSpecies="tibetanum">
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<emphasis id="8ED77F27FFA9D97DFF6BFDB1E53EFDDE" bold="true" box="[151,598,536,562]" italics="true" pageId="5" pageNumber="64">Mesopodagrion tibetanum tibetanum</emphasis>
<bibRefCitation id="D832DEC4FFA9D97DFDA1FDB1E45DFDDE" author="McLachlan" box="[605,821,536,562]" pageId="5" pageNumber="68" refString="McLachlan, R. (1896) On Odonata from the Province of Szechuen in Western China, and from Moupin in eastern Thibet. The Annals and magazine of natural history: zoology, botany, and geology, (6) 17, 372 - 373." type="journal article" year="1896">McLachlan, 1896</bibRefCitation>
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<emphasis id="8ED77F27FFA9D97DFF6BFDCBE6DDFD95" box="[151,437,610,633]" italics="true" pageId="5" pageNumber="64">Mesopodagrion tibetanum</emphasis>
:
<treatmentCitation id="3D028524FFA9D97DFE39FDCBE5A8FD95" author="McLachlan" box="[453,704,610,633]" page="372" pageId="5" pageNumber="64" year="1896">
<bibRefCitation id="D832DEC4FFA9D97DFE39FDCBE5E1FD95" author="McLachlan" box="[453,649,610,633]" pageId="5" pageNumber="68" refString="McLachlan, R. (1896) On Odonata from the Province of Szechuen in Western China, and from Moupin in eastern Thibet. The Annals and magazine of natural history: zoology, botany, and geology, (6) 17, 372 - 373." type="journal article" year="1896">McLachlan, 1896</bibRefCitation>
: 372
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273, “Moupin (now Baoxing, Sichuan,
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)”;
<treatmentCitation id="3D028524FFA9D97DFB14FDCBE798FD75" author="Needham" page="239" pageId="5" pageNumber="64" year="1930">
<bibRefCitation id="D832DEC4FFA9D97DFB14FDCBE2F1FD95" author="Needham" box="[1256,1433,610,633]" pageId="5" pageNumber="68" refString="Needham, J. G. (1930) A manual of the dragonflies of China. Peiping: The Fan Memorial Institute of Biology, 239 - 240." type="book chapter" year="1930">Needham, 1930</bibRefCitation>
: 239
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240, “Tibet”;
<treatmentCitation id="3D028524FFA9D97DFE6CFD2BE528FD75" author="Fraser" box="[400,576,642,665]" page="96" pageId="5" pageNumber="64" year="1933">
<bibRefCitation id="D832DEC4FFA9D97DFE6CFD2BE571FD75" author="Fraser" box="[400,537,642,665]" pageId="5" pageNumber="68" refString="Fraser, F. C. (1933) The Fauna of British India-Odonata. VI. London: Taylor and Francis, 96 - 98." type="book chapter" year="1933">Fraser, 1933</bibRefCitation>
: 96
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98, figs. 44, 45, 46 (c, d), “Tibet, Sichuan”;
<treatmentCitation id="3D028524FFA9D97DFBD5FD2BE385FD75" author="Lieftinck" box="[1065,1261,642,665]" page="7" pageId="5" pageNumber="64" year="1948">
<bibRefCitation id="D832DEC4FFA9D97DFBD5FD2BE3BBFD75" author="Lieftinck" box="[1065,1235,642,665]" pageId="5" pageNumber="68" refString="Lieftinck, M. A. (1948) Entomological results from the Swedish Expedition 1934 to Burma and British India. Odonata. Arkiv For Zoologi, 41 (A), 7." type="journal article" year="1948">Lieftinck, 1948</bibRefCitation>
: 7
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, “
<collectingCountry id="C4B4E3A5FFA9D97DFAFFFD2BE238FD75" box="[1283,1360,642,665]" name="Myanmar" pageId="5" pageNumber="64">Burma</collectingCountry>
,
<collectingCountry id="C4B4E3A5FFA9D97DFAA0FD2BE2F4FD75" box="[1372,1436,642,665]" name="China" pageId="5" pageNumber="64">China</collectingCountry>
(Shaanxi)” (in part);
<treatmentCitation id="3D028524FFA9D97DFE54FD0BE513FD55" author="Asahina" box="[424,635,674,697]" page="130" pageId="5" pageNumber="64" year="1955">
<bibRefCitation id="D832DEC4FFA9D97DFE54FD0BE52FFD55" author="Asahina" box="[424,583,674,697]" pageId="5" pageNumber="68" refString="Asahina, S. (1955) Odonata from South Shensi (North China) in the collection of the Zoological Museum, Copenhagen. Entomologiske Meddelielser, 27, 130 - 133." type="journal article" year="1955">Asahina, 1955</bibRefCitation>
: 130
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133, figs. 16, “
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,
<collectingCountry id="C4B4E3A5FFA9D97DFC70FD0BE4A4FD55" box="[908,972,674,697]" name="China" pageId="5" pageNumber="64">China</collectingCountry>
(Shaanxi, Sichuan, Zhejiang, Yunnan)” (in part);
<treatmentCitation id="3D028524FFA9D97DFEF6FD6BE567FD35" author="SUI" box="[266,527,706,729]" page="294" pageId="5" pageNumber="64" year="1986">
<bibRefCitation id="D832DEC4FFA9D97DFEF6FD6BE6B1FD35" author="SUI" box="[266,473,706,729]" pageId="5" pageNumber="68" refString="SUI, J-Z. &amp; SUN, H-G .. (1986) Common species of dragonflies from China. Agriculture Publishing House, Beijing, 294 - 295." type="book chapter" year="1986">SUI &amp; SUN, 1986</bibRefCitation>
: 294
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295, Fig. 201, “
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,
<collectingCountry id="C4B4E3A5FFA9D97DFCDEFD6BE40AFD35" box="[802,866,706,729]" name="China" pageId="5" pageNumber="64">China</collectingCountry>
(Xizang, Yunnan, Zhejiang)” (in part);
<treatmentCitation id="3D028524FFA9D97DFAEBFD6BE799FD15" author="Chao" page="112" pageId="5" pageNumber="64" year="1987">
<bibRefCitation id="D832DEC4FFA9D97DFAEBFD6BE2F0FD35" author="Chao" box="[1303,1432,706,729]" pageId="5" pageNumber="68" refString="Chao, H. f. (1987) On the synonymy of a species of Mesopodagrion (Odonata: Megapodagrionidae). Entomotaxonomia, 9 (2), 112, 120." type="journal article" year="1987">Chao, 1987</bibRefCitation>
: 112
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, 120; “
<collectingCountry id="C4B4E3A5FFA9D97DFEBFFD4BE6F8FD15" box="[323,400,738,761]" name="Myanmar" pageId="5" pageNumber="64">Burma</collectingCountry>
,
<collectingCountry id="C4B4E3A5FFA9D97DFE63FD4BE6B7FD15" box="[415,479,738,761]" name="China" pageId="5" pageNumber="64">China</collectingCountry>
(Shaanxi, Sichuan, Yunnan, Zhejiang)” (in part);
<treatmentCitation id="3D028524FFA9D97DFBECFD4BE39DFD15" author="Kalkman" box="[1040,1269,738,761]" page="187" pageId="5" pageNumber="64" year="2008">
<bibRefCitation id="D832DEC4FFA9D97DFBECFD4BE3D5FD15" author="Kalkman" box="[1040,1213,738,761]" pageId="5" pageNumber="68" refString="Kalkman, V. J., (2008) Taxonomy, behaviour, and habitat of Mesopodagrion and Sinocnemis. Notes on old world Megapodagrionidae 3 (Odonata). International Journal of Odonatology 11 (2): 185 - 193." type="journal article" year="2008">Kalkman, 2008</bibRefCitation>
: 187
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, habits, key to species.
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<emphasis id="8ED77F27FFA9D97DFF6BFCE1E67AFC8E" bold="true" box="[151,274,840,866]" pageId="5" pageNumber="64">Material.</emphasis>
1 3,
<collectingCountry id="C4B4E3A5FFA9D97DFEA7FCE0E6C0FC8F" box="[347,424,841,867]" name="China" pageId="5" pageNumber="64">China</collectingCountry>
, Moupin (now Baoxing, Sichuan), 189?,. (
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) [
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]; 2 3, 1 Ƥ,
<collectingCountry id="C4B4E3A5FFA9D97DFAB7FCE0E2F0FC8F" box="[1355,1432,841,867]" name="China" pageId="5" pageNumber="64">China</collectingCountry>
, Sichuan, Wolong nature reserve,
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,
<date id="C81D85F5FFA9D97DFD87FCC6E47AFC65" box="[635,786,879,905]" pageId="5" pageNumber="64" value="2005-07-24">24-VII-2005</date>
, Xin Yu leg. (
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).
</paragraph>
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<subSubSection id="F4B9F0BEFFA9D97DFF39FC3CE6E8F9F7" pageId="5" pageNumber="64" type="description">
<paragraph id="BC1CA335FFA9D97DFF39FC3CE62DFA4A" blockId="5.[151,1437,840,1678]" pageId="5" pageNumber="64">
<emphasis id="8ED77F27FFA9D97DFF39FC3CE635FC43" bold="true" box="[197,349,917,943]" pageId="5" pageNumber="64">Description.</emphasis>
Male. Labium black; labrum, bases of mandibles, genae, anteclypeus and sides of frons pale; postclypeus, middle of frons, and top of head, including antennae, black, except for a pair of short pale stripes between lateral ocelli and bases of antennae; postocular spots absent, pale occipital bar present (
<figureCitation id="2498BFB0FFA9D97DFACDFC4AE214FC11" box="[1329,1404,995,1021]" captionStart="FIGURE 4" captionStartId="4.[151,256,1119,1143]" captionTargetBox="[357,1232,526,1088]" captionTargetId="figure@4.[350,1237,520,1095]" captionTargetPageId="4" captionText="FIGURE 4. Pale occipital bar and antehumeral stripe. a, Mesopodagrion yachowensis (China, Henan, Songxian, Mt. Baiyun); b, Mesopodagrion tibetanum tibetanum (China, Sichuan, Wolong Nature Reserve); c, Mesopodagrion tibetanum australe ssp. Nov. (China, Yunnan, Lijiang, Mt. Yulong); d, Mesopodagrion tibetanum australe ssp. Nov. (China, Yunnan, Yuanjiang, Nanxi)." httpUri="https://zenodo.org/record/189649/files/figure.png" pageId="5" pageNumber="64">Fig. 4</figureCitation>
b); upper border of Rear of head pale, remainder black (
<figureCitation id="2498BFB0FFA9D97DFCFDFBA3E42FFBC8" box="[769,839,1034,1060]" captionStart="FIGURE 3" captionStartId="4.[151,256,373,397]" captionTargetBox="[388,1200,204,342]" captionTargetId="figure@4.[381,1205,194,350]" captionTargetPageId="4" captionText="FIGURE 3. Color pattern of rear of head. a, Mesopodagrion yachowensis (China, Shaanxi, Ningshan, Huoditang); b, Mesopodagrion tibetanum tibetanum (China, Sichuan, Wolong Nature Reserve); c, Mesopodagrion tibetanum australe ssp. Nov. (China, Yunnan, Lijiang, Mt. Yulong)." httpUri="https://zenodo.org/record/189648/files/figure.png" pageId="5" pageNumber="64">Fig. 3</figureCitation>
b). Prothorax black except for a pair of pale stripes on propleura and lower-most margin; synthorax black dorsally with wide pale antehumeral stripe about 1/2 width of mespisternum, sides of synthorax pale except for ventral border (
<figureCitation id="2498BFB0FFA9D97DFBF3FBFEE330FB9D" box="[1039,1112,1111,1137]" captionStart="FIGURE 4" captionStartId="4.[151,256,1119,1143]" captionTargetBox="[357,1232,526,1088]" captionTargetId="figure@4.[350,1237,520,1095]" captionTargetPageId="4" captionText="FIGURE 4. Pale occipital bar and antehumeral stripe. a, Mesopodagrion yachowensis (China, Henan, Songxian, Mt. Baiyun); b, Mesopodagrion tibetanum tibetanum (China, Sichuan, Wolong Nature Reserve); c, Mesopodagrion tibetanum australe ssp. Nov. (China, Yunnan, Lijiang, Mt. Yulong); d, Mesopodagrion tibetanum australe ssp. Nov. (China, Yunnan, Yuanjiang, Nanxi)." httpUri="https://zenodo.org/record/189649/files/figure.png" pageId="5" pageNumber="64">Fig. 4</figureCitation>
b); legs black, pale at base; wings hyaline; pterostigmata brown, braced, covering three cells. S1-2 black dorsally, pale laterally; S2 with black stripe laterally; S3-6 black with pale basal spot and lateral stripe gradually narrowing distally; S7-8 black except for pale basal spot and a pair of ventral narrow stripes (figs. 2a, 2b); S9 black except for a pale latero-ventral spot; S10 black except for pale postero-lateral spot (
<figureCitation id="2498BFB0FFA9D97DFC54FB5BE487FAE0" box="[936,1007,1266,1292]" captionStart="FIGURE 5" captionStartId="4.[151,255,1761,1785]" captionTargetBox="[151,1424,1327,1702]" captionTargetId="figure@4.[151,1436,1301,1737]" captionTargetPageId="4" captionText="FIGURE 5. Color pattern of 3 S 10 and appendages, dorsal and lateral. a, Mesopodagrion yachowensis (China, Henan, Songxian, Mt. Baiyun); b, Mesopodagrion tibetanum tibetanum (China, Sichuan, Wolong Nature Reserve); c, Mesopodagrion tibetanum australe ssp. Nov. (China, Yunnan, Lijiang, Mt. Yulong); d, Mesopodagrion tibetanum australe ssp. Nov. (China, Yunnan, Yuanjiang, Nanxi)." httpUri="https://zenodo.org/record/189650/files/figure.png" pageId="5" pageNumber="64">Fig. 5</figureCitation>
d); bifurcate process longer than 1/2 of S10, apex acute, outer margin inflectional, not smooth, hemline of bifurcate process about 1/2 width of S10 (
<figureCitation id="2498BFB0FFA9D97DFF5CFA96E79CFAB5" box="[160,244,1343,1369]" captionStart="FIGURE 5" captionStartId="4.[151,255,1761,1785]" captionTargetBox="[151,1424,1327,1702]" captionTargetId="figure@4.[151,1436,1301,1737]" captionTargetPageId="4" captionText="FIGURE 5. Color pattern of 3 S 10 and appendages, dorsal and lateral. a, Mesopodagrion yachowensis (China, Henan, Songxian, Mt. Baiyun); b, Mesopodagrion tibetanum tibetanum (China, Sichuan, Wolong Nature Reserve); c, Mesopodagrion tibetanum australe ssp. Nov. (China, Yunnan, Lijiang, Mt. Yulong); d, Mesopodagrion tibetanum australe ssp. Nov. (China, Yunnan, Yuanjiang, Nanxi)." httpUri="https://zenodo.org/record/189650/files/figure.png" pageId="5" pageNumber="64">Figs. 5</figureCitation>
c, 11), foot-like from latero-ventral view (
<figureCitation id="2498BFB0FFA9D97DFD0DFA96E423FAB5" box="[753,843,1343,1369]" captionStart="FIGURES 8 20" captionStartId="7.[151,272,1843,1867]" captionTargetBox="[213,1403,185,1800]" captionTargetId="figure@7.[199,1405,177,1816]" captionTargetPageId="7" captionText="FIGURES 8 20. SEM photos for cerci and bifurcate process of 3 Mesopodagrion, all specimens from China (8 15 dorsal, 16 19 ventro-lateral, and 20 Ƥ pronotum, dorsal): 8) Gansu, Kangxian; 9) Henan, Songxian; 10) Shaanxi, Ningshan; 11) Sichuan, Wolong nature reserve; 12) Yunnan, Mt. Ailao; 13) Yunnan, Dali; 14) Yunnan, Lijiang; 15) Yunnan, Yuanjiang; 16) Henan, Songxian; 17) Shaanxi, Ningshan; 18) Sichuan, Wolong; 19) Yunnan, Yuanjiang; 20) Sichuan, Wolong. FJ = false joint, EP = epiproct. (Scale bar = 1 mm)." httpUri="https://zenodo.org/record/189653/files/figure.png" pageId="5" pageNumber="64">Fig. 18</figureCitation>
); caudal appendages black, cerci almost twice as long as S10, paraprocts vestigial (
<figureCitation id="2498BFB0FFA9D97DFDD6FACFE515FA6C" box="[554,637,1382,1408]" captionStart="FIGURE 5" captionStartId="4.[151,255,1761,1785]" captionTargetBox="[151,1424,1327,1702]" captionTargetId="figure@4.[151,1436,1301,1737]" captionTargetPageId="4" captionText="FIGURE 5. Color pattern of 3 S 10 and appendages, dorsal and lateral. a, Mesopodagrion yachowensis (China, Henan, Songxian, Mt. Baiyun); b, Mesopodagrion tibetanum tibetanum (China, Sichuan, Wolong Nature Reserve); c, Mesopodagrion tibetanum australe ssp. Nov. (China, Yunnan, Lijiang, Mt. Yulong); d, Mesopodagrion tibetanum australe ssp. Nov. (China, Yunnan, Yuanjiang, Nanxi)." httpUri="https://zenodo.org/record/189650/files/figure.png" pageId="5" pageNumber="64">Figs. 5</figureCitation>
d, 11); genital ligula simple, apical lobs bifurcated beyond the base (
<figureCitation id="2498BFB0FFA9D97DFF63FA25E78FFA4A" box="[159,231,1420,1446]" captionStart="FIGURE 2" captionStartId="3.[151,255,1798,1822]" captionTargetBox="[413,1170,837,1762]" captionTargetId="figure@3.[409,1178,822,1774]" captionTargetPageId="3" captionText="FIGURE 2. Mesopodagrion tibetanum (China, Sichuan, Wolong Nature Reserve): (a) body and wings, dorsal view, 3; (b) abdomen, lateral view, 3; (c) abdomen, lateral view, Ƥ; (d) genital ligula, ventral view, 3; (e) genital ligula, lateral view, 3." httpUri="https://zenodo.org/record/189647/files/figure.png" pageId="5" pageNumber="64">Fig. 2</figureCitation>
, d &amp; e).
</paragraph>
<paragraph id="BC1CA335FFA9D97DFF39FA1AE6E8F9F7" blockId="5.[151,1437,840,1678]" pageId="5" pageNumber="64">
Female. Similar as male by color pattern but slightly shorter in body length and more robust (
<figureCitation id="2498BFB0FFA9D97DFAF8FA1AE223FA21" box="[1284,1355,1459,1485]" captionStart="FIGURE 2" captionStartId="3.[151,255,1798,1822]" captionTargetBox="[413,1170,837,1762]" captionTargetId="figure@3.[409,1178,822,1774]" captionTargetPageId="3" captionText="FIGURE 2. Mesopodagrion tibetanum (China, Sichuan, Wolong Nature Reserve): (a) body and wings, dorsal view, 3; (b) abdomen, lateral view, 3; (c) abdomen, lateral view, Ƥ; (d) genital ligula, ventral view, 3; (e) genital ligula, lateral view, 3." httpUri="https://zenodo.org/record/189647/files/figure.png" pageId="5" pageNumber="64">Fig. 2</figureCitation>
c). S10 wholly black; ending before level of cerci, black except for pale tip (
<figureCitation id="2498BFB0FFA9D97DFC48FA73E492FA18" box="[948,1018,1498,1524]" captionStart="FIGURE 6" captionStartId="5.[151,259,389,413]" captionTargetBox="[332,1240,201,331]" captionTargetId="figure@5.[332,1254,194,366]" captionTargetPageId="5" captionText="FIGURE 6. FAE. a, (China, Henan, Huixian, Baligou); b, Mesopodagrion tibetanum tibetanum (China, Sichuan, Wolong Nature Reserve); c, Mesopodagrion tibetanum australe ssp. Nov. (China, Yunnan, Dali)." httpUri="https://zenodo.org/record/189651/files/figure.png" pageId="5" pageNumber="64">Fig. 6</figureCitation>
a). Pale body color pattern on living female apple-green.
</paragraph>
</subSubSection>
<subSubSection id="F4B9F0BEFFA9D97DFF39F98EE587F962" pageId="5" pageNumber="64" type="distribution">
<paragraph id="BC1CA335FFA9D97DFF39F98EE587F962" blockId="5.[151,1437,840,1678]" pageId="5" pageNumber="64">
<emphasis id="8ED77F27FFA9D97DFF39F98EE60DF9AD" bold="true" box="[197,357,1575,1601]" pageId="5" pageNumber="64">Distribution.</emphasis>
Known from northwestern
<collectingCountry id="C4B4E3A5FFA9D97DFD57F98EE593F9AD" box="[683,763,1575,1601]" name="Myanmar" pageId="5" pageNumber="64">Burma</collectingCountry>
and central Sichuan,
<collectingCountry id="C4B4E3A5FFA9D97DFC05F98EE32BF9AD" box="[1017,1091,1575,1601]" name="China" pageId="5" pageNumber="64">China</collectingCountry>
. The nominotypic subspecies occurs principally north of
<taxonomicName id="7BA3D8B6FFA9D97DFE2AF9E7E51CF98B" box="[470,628,1614,1639]" class="Insecta" family="Megapodagrionidae" genus="Mesopodagrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="64" phylum="Arthropoda" rank="subSpecies" species="tibetanum" subSpecies="australe">
<emphasis id="8ED77F27FFA9D97DFE2AF9E7E51CF98B" box="[470,628,1614,1639]" italics="true" pageId="5" pageNumber="64">M. t. australe</emphasis>
</taxonomicName>
but south of
<taxonomicName id="7BA3D8B6FFA9D97DFCF3F9E7E4A3F98B" box="[783,971,1614,1639]" class="Insecta" family="Megapodagrionidae" genus="Mesopodagrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="yachowensis">
<emphasis id="8ED77F27FFA9D97DFCF3F9E7E4A3F98B" box="[783,971,1614,1639]" italics="true" pageId="5" pageNumber="64">M. yachowensis</emphasis>
</taxonomicName>
(
<figureCitation id="2498BFB0FFA9D97DFC27F9E7E34BF984" box="[987,1059,1614,1640]" captionStart="FIGURE 7" captionStartId="6.[151,244,1774,1795]" captionTargetBox="[153,1429,525,1731]" captionTargetId="figure@6.[151,1436,503,1750]" captionTargetPageId="6" captionText="FIGURE 7. Distribution of Mesopodagrion." httpUri="https://zenodo.org/record/189652/files/figure.png" pageId="5" pageNumber="64">Fig. 7</figureCitation>
). This subspecies is believed by
<bibRefCitation id="D832DEC4FFA9D97DFF6BF9DDE63FF962" author="Kalkman" box="[151,343,1652,1678]" pageId="5" pageNumber="68" refString="Kalkman, V. J., (2008) Taxonomy, behaviour, and habitat of Mesopodagrion and Sinocnemis. Notes on old world Megapodagrionidae 3 (Odonata). International Journal of Odonatology 11 (2): 185 - 193." type="journal article" year="2008">Kalkman (2008)</bibRefCitation>
to occur in Kambaiti, NW
<collectingCountry id="C4B4E3A5FFA9D97DFD6BF9DDE583F962" box="[663,747,1652,1678]" name="Myanmar" pageId="5" pageNumber="64">Burma</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>