143 lines
14 KiB
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143 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.147.1898" ID-GBIF-Dataset="5207e87b-8044-498d-a80d-7275387282d2" ID-PMC="PMC3286249" ID-Pensoft-Pub="1313-2970-147-277" ID-PubMed="22371667" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-147-277" ModsDocOrigin="ZooKeys 147" ModsDocTitle="Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n. from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae)" checkinTime="1451249665881" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2011" docId="8E5316E54C210E90F38A52F7DFB2BB65" docLanguage="en" docName="ZooKeys 147: 277-335" docOrigin="ZooKeys 147" docSource="http://dx.doi.org/10.3897/zookeys.147.1898" docTitle="Meonochilus rectus Liebherr, 2011, sp. n." docType="treatment" docVersion="4" lastPageNumber="308" masterDocId="554A3015B71F903E5450EF01FFFDC71C" masterDocTitle="Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n. from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae)" masterLastPageNumber="335" masterPageNumber="277" pageNumber="307" updateTime="1668152665929" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n. from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liebherr, James K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>147</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>277</mods:start>
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<mods:end>335</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.147.1898</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.147.1898</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-147-277</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152032013" LSID="urn:lsid:zoobank.org:act:B3B1A69B-D519-4DC2-8055-44D04BB4ECCE" httpUri="http://treatment.plazi.org/id/8E5316E54C210E90F38A52F7DFB2BB65" lastPageId="31" lastPageNumber="308" pageId="30" pageNumber="307">
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<subSubSection pageId="30" pageNumber="307" type="nomenclature">
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<paragraph pageId="30" pageNumber="307">
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3.
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<taxonomicName LSID="urn:lsid:zoobank.org:act:B3B1A69B-D519-4DC2-8055-44D04BB4ECCE" class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus rectus" order="Coleoptera" pageId="30" pageNumber="307" phylum="Arthropoda" rank="species" species="rectus">Meonochilus rectus</taxonomicName>
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<taxonomicNameLabel pageId="30" pageNumber="307">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="30" pageNumber="307" type="diagnosis">
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<paragraph pageId="30" pageNumber="307">Diagnosis.</paragraph>
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<paragraph pageId="30" pageNumber="307">
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Individuals of this species look like a broad-bodied version of
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus spiculatus" order="Coleoptera" pageId="30" pageNumber="307" phylum="Arthropoda" rank="species" species="spiculatus">Meonochilus spiculatus</taxonomicName>
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, with the broadly based, subparallel elytra exhibiting deep, distinctly punctate striae and convex, glossy intervals (Fig. 9C). The antennae are not as elongate as those of
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus spiculatus" order="Coleoptera" pageId="30" pageNumber="307" phylum="Arthropoda" rank="species" species="spiculatus">Meonochilus spiculatus</taxonomicName>
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, with the ninth antennomere length 1.8
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<normalizedToken originalValue="–2.0×">-2.0x</normalizedToken>
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its maximum breadth. Like
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus spiculatus" order="Coleoptera" pageId="30" pageNumber="307" phylum="Arthropoda" rank="species" species="spiculatus">Meonochilus spiculatus</taxonomicName>
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, the internal sac of the male aedeagus bears a spikelike flagellum (Fig. 2F, 2G), however the flagellum of
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus rectus" order="Coleoptera" pageId="30" pageNumber="307" phylum="Arthropoda" rank="species" species="rectus">Meonochilus rectus</taxonomicName>
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males is stouter and more sinuously curved, the aedeagal median lobe is broader (Fig. 2G), and it terminates in a downwardly expanded apex. Standardized body length 5.4-5.7 mm.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="30" pageNumber="307" type="male genitalia">
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<paragraph pageId="30" pageNumber="307">Male Genitalia.</paragraph>
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<paragraph pageId="30" pageNumber="307">
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(n = 1). Aedeagal median lobe with basally deep sagittal crest, the lobe shaft broad and robust with ventral surface recurved before downturned and broadly rounded apex (Fig. 2G); internal sac elongate, with stout spikelike flagellum inserted on ventral surface about half the distance from ostium to apex of apical membranous lobe; flagellar sheath closely appressed to flagellum, length about half the length of flagellum; convex roll of sac membrane extended on right side of flagellar articulation toward sac apex, the roll covered with broad shingle-like sculpticells, the exposed shingle edges lining the distal edges of sculpticells (brush sclerite;
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<bibRefCitation author="Lindroth, CH" journalOrPublisher="New Zealand Journal of Zoology" pageId="41" pageNumber="318" pagination="161 - 198" title="Genus Bembidion Latreille (Coleoptera: Carabidae) in New Zealand: a revision." url="doi: 10.1080/03014223.1976.9517912" volume="3" year="1976">Lindroth 1976</bibRefCitation>
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); ventral right paramere parallel-sided and apically rounded, length equal to
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<normalizedToken originalValue="½">1/2</normalizedToken>
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distance from parameral articulation to external face of apex, 10 long setae lining the apical half of ventral edge, 12 long setae arrayed on convex parameral apex, and 6 long, thinner setae in apical half of dorsal edge; dorsal left paramere broad basally, subconchoid, apex narrowly rounded, 2 long setae on dorsal surface near tip, with 2 short setae immediately basad to those.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="31" lastPageNumber="308" pageId="30" pageNumber="307" type="female reproductive tract">
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<paragraph pageId="30" pageNumber="307">Female Reproductive Tract.</paragraph>
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<paragraph lastPageId="31" lastPageNumber="308" pageId="30" pageNumber="307">
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(n = 1) Bursa copulatrix broadest medially, extended to broadly rounded apex that constitutes spermatheca (based on insertion of spermathecal gland duct), overall bursal plus spermathecal length 1.1
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<normalizedToken originalValue="×">x</normalizedToken>
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maximal bursal breadth (Fig. 14B); common oviduct broadest distad juncture with ventral surface of bursa; spermathecal gland duct short, length approximately equal to basal stem of spermathecal gland, the narrow duct base joining bursal apex at short, stump-like protuberance; basal gonocoxite 1 with lateroapical series of 3-4 setae, a single additional seta basal to the series of 4 on one side of the lone dissected female specimen, 3-4 smaller setae along apical half of dorsomedial surface (Fig. 14C); apical gonocoxite 2 with 2 broad
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<pageBreakToken pageId="31" pageNumber="308" start="start">lateral</pageBreakToken>
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ensiform setae and 1 dorsal ensiform seta on dorsal surface; apical sensory furrow bearing 2 nematiform setae at 0.70
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<normalizedToken originalValue="×">x</normalizedToken>
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coxite length.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="308" type="holotype">
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<paragraph pageId="31" pageNumber="308">Holotype.</paragraph>
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<paragraph pageId="31" pageNumber="308">
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Male (NZAC): NEW ZEALAND CL / Great Barrier I / Mt. Hobson Tk. / 24 Mar- / 27 Mar 1978 // J.C. Watt + / J. Murcer / Pit trap 9 // N Z Arthropod Collection / [bar code] / NZAC04007299 // HOLOTYPE ♂ / Meonochilus / rectus / det. J.K.
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<bibRefCitation author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="41" pageNumber="318" pagination="15 - 76" title="The Mecyclothorax beetles (Coleoptera: Carabidae: Moriomorphini) of West Maui, Hawaii: taxonomy, biogeography, and conservation." url="doi: 10.1002/mmnd.201100005" volume="58" year="2011">Liebherr 2011</bibRefCitation>
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(black-bordered red label).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="308" type="paratypes">
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<paragraph pageId="31" pageNumber="308">Paratypes.</paragraph>
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<paragraph pageId="31" pageNumber="308">
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CL: Great Barrier Island, Mt. Hobson, leaf litter, RL805,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-36.18783">36°11.27'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="175.4115">175°24.69'E</geoCoordinate>
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, 500 m el., 17-xii-2003, Leschen (AMNZ, 1), pitfall trap, 9-xi-17-xii-2003, Early (AMNZ, 2), 17-xii-2003-07-iv-2004, Early (AMNZ, 1), Te Paparahi, pitfall trap, G1,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-35.4345">35°26.07'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="174.3905">174°23.43'E</geoCoordinate>
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, iii-2002, Warren (AMNZ, 1), Windy Hill, pitfall trap,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-36.302">36°18.12'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="175.53517">175°32.11'E</geoCoordinate>
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, 200 m el., 30-iv-11-vi-2003, Parsons (AMNZ, 1); Mt. Moehau, under log,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-36.53417">36°32.05'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="175.3995">175°23.97'E</geoCoordinate>
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, 765 m el., 19-iii-1980, Holloway (NZAC, 1); Tapu-Coroglen Rd., Tk. to Crosbies,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-36.990334">36°59.42'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="175.58766">175°35.26'E</geoCoordinate>
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, 13-xi-1996, Gleeson (NZAC, 1).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="308" type="etymology">
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<paragraph pageId="31" pageNumber="308">Etymology.</paragraph>
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<paragraph pageId="31" pageNumber="308">
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The species epithet
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<normalizedToken originalValue="rectus–Latin">rectus-Latin</normalizedToken>
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adjective with male
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<normalizedToken originalValue="ending–denotes">ending-denotes</normalizedToken>
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the short, straight and parallel to slightly converging basolateral margins of the pronotum present immediately anterad the pronotal hind angles.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="308" type="distribution and habitat">
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<paragraph pageId="31" pageNumber="308">Distribution and Habitat.</paragraph>
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<paragraph pageId="31" pageNumber="308">
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The distribution of
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus rectus" order="Coleoptera" pageId="31" pageNumber="308" phylum="Arthropoda" rank="species" species="rectus">Meonochilus rectus</taxonomicName>
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includes the Coromandel Peninsula and Great Barrier Island to the north (Fig. 13). This range lies disjunctly north of the Kaimai Range distribution of the sister species,
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus spiculatus" order="Coleoptera" pageId="31" pageNumber="308" phylum="Arthropoda" rank="species" species="spiculatus">Meonochilus spiculatus</taxonomicName>
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, suggesting that a vicariant barrier separating the mountains of Coromandel and Bay of Plenty facilitated allopatric speciation. Nonetheless, eustatic sea level changes that isolated Great Barrier Island from the mainland have not resulted in speciation of the island population. Most specimens of this species have been collected via pitfall trapping, with singletons found in leaf litter and under a log. Localities range in elevation from 200-765 m el.
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</paragraph>
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<caption pageId="31" pageNumber="308">
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<paragraph pageId="31" pageNumber="308">
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Figure 15. Distribution map for
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus" order="Coleoptera" pageId="31" pageNumber="308" phylum="Arthropoda" rank="genus">Meonochilus</taxonomicName>
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spp.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |