280 lines
38 KiB
XML
280 lines
38 KiB
XML
<document id="E7D4A533EABBD28F15295B64ABD13BDC" ID-CLB-Dataset="37554" ID-DOI="10.5281/zenodo.275634" ID-GBIF-Dataset="60a5304a-95a2-4b2e-9258-17806aff8197" ID-ISSN="1175-5326" ID-Zenodo-Dep="275634" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481580325892" checkinUser="plazi" docAuthor="JIŘÍ KOLIBÁČ" docDate="2010" docId="425A8793076DFFE89DD5FC1FFB82FE8D" docLanguage="en" docName="zt02353p042.pdf.imf" docOrigin="Zootaxa 2353" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Trichocateres fasciculifer KOLIBÁČ, 2010, n. sp." docType="treatment" docVersion="7" lastPageNumber="42" masterDocId="BE63FFEB076FFFE09D42FFA5FFEEFFCA" masterDocTitle="Trichocateres fasciculifer, a new genus and species of Trogossitidae: Lophocaterini (Coleoptera)" masterLastPageNumber="42" masterPageNumber="34" pageNumber="36" updateTime="1698433707470" updateUser="ExternalLinkService">
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<mods:mods id="911061AB3DA2473B40C7B062AD14C989" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8CBA5C5BF96C8F93267550C5DDB1B762">
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<mods:title id="7365747AE50D8372DA8C22236D8BEE8F">Trichocateres fasciculifer, a new genus and species of Trogossitidae: Lophocaterini (Coleoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="5D792A3C9CAC2E172C6B0EB4EC8D25AC" type="personal">
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<mods:role id="1ED8A9156CE2A448FD26890A4B70FC49">
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<mods:roleTerm id="71E098E0C9AE1FF3E8FDB3C5AB982ED6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FEBF04E904713070CDA922EC536549AE">JIŘÍ KOLIBÁČ</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1837BF2C4A64D7B927E177D0A6A89F45">text</mods:typeOfResource>
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<mods:titleInfo id="DBD994371ED27FE5F2AD789D8AB134B6">
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<mods:title id="CAFBE5F65B66572B429F92C49128A82C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="93410DDD1B86125540B05107F9B8FCD2">
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<mods:date id="774DC4C227CD12BC2BF735049B8BC8B1">2010</mods:date>
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<mods:detail id="5EAC3F7F56378D219B9AFF956B17D265" type="volume">
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<mods:number id="DF8098B1AB6624840488E141F200355E">2353</mods:number>
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</mods:detail>
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<mods:extent id="C02FEDF6317F41BE71E0E187F934C61D" unit="page">
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<mods:start id="C094F0360E4B1B7E84B46B3B890A0987">34</mods:start>
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<mods:end id="365E4DC518C57B7D0FE6C7513D967FE4">42</mods:end>
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</mods:extent>
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</mods:relatedItem>
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<mods:classification id="0C96AC636C2D4F0DAD3A52C5CF8CD932">journal article</mods:classification>
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<mods:identifier id="026FE06E3D42FA5578EF67909C5AE5CC" type="CLB-Dataset">37554</mods:identifier>
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<mods:identifier id="99EAADC0845336D3B96A43CDEA52BBA8" type="DOI">10.5281/zenodo.275634</mods:identifier>
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<mods:identifier id="02E4A60EE52FFF0A98D0B575A012958E" type="GBIF-Dataset">60a5304a-95a2-4b2e-9258-17806aff8197</mods:identifier>
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<mods:identifier id="7397A7D79C37AA603E96F46628826722" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="CBB7181A1E13E5A28764DF99470AFD9C" type="Zenodo-Dep">275634</mods:identifier>
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</mods:mods>
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<treatment id="425A8793076DFFE89DD5FC1FFB82FE8D" ID-DOI="http://doi.org/10.5281/zenodo.6196977" ID-GBIF-Taxon="125107178" ID-Zenodo-Dep="6196977" LSID="urn:lsid:plazi:treatment:425A8793076DFFE89DD5FC1FFB82FE8D" httpUri="http://treatment.plazi.org/id/425A8793076DFFE89DD5FC1FFB82FE8D" lastPageId="8" lastPageNumber="42" pageId="2" pageNumber="36">
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<subSubSection id="82E9650E076DFFE29DD5FC1FFDFBFC1E" box="[151,533,954,980]" pageId="2" pageNumber="36" type="nomenclature">
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<paragraph id="CA4C3685076DFFE29DD5FC1FFDFBFC1E" blockId="2.[151,533,954,980]" box="[151,533,954,980]" pageId="2" pageNumber="36">
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<heading id="910481E9076DFFE29DD5FC1FFDFBFC1E" bold="true" box="[151,533,954,980]" fontSize="11" level="1" pageId="2" pageNumber="36" reason="1">
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<emphasis id="F887EA97076DFFE29DD5FC1FFDFBFC1E" bold="true" box="[151,533,954,980]" pageId="2" pageNumber="36">
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<taxonomicName id="0DF34D06076DFFE29DD5FC1FFE22FC1E" box="[151,460,954,980]" class="Insecta" family="Trogossitidae" genus="Trichocateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="36" phylum="Arthropoda" rank="species" species="fasciculifer" status="sp. nov.">
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<emphasis id="F887EA97076DFFE29DD5FC1FFE22FC1E" bold="true" box="[151,460,954,980]" italics="true" pageId="2" pageNumber="36">Trichocateres fasciculifer</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E3B457EC076DFFE29C91FC1FFDFBFC1E" box="[467,533,954,980]" pageId="2" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="82E9650E076DFFE29DD5FC5FFED1FB60" pageId="2" pageNumber="36" type="materials_examined">
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<paragraph id="CA4C3685076DFFE29DD5FC5FFED1FB60" blockId="2.[151,1437,1018,2015]" pageId="2" pageNumber="36">
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<emphasis id="F887EA97076DFFE29DD5FC5FFE81FBDE" bold="true" box="[151,367,1018,1044]" pageId="2" pageNumber="36">
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<typeStatus id="15488827076DFFE29DD5FC5FFF39FBDE" box="[151,215,1018,1044]" pageId="2" pageNumber="36">Type</typeStatus>
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specimens:
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</emphasis>
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<typeStatus id="15488827076DFFE29C3AFC5FFE05FBDE" box="[376,491,1018,1044]" pageId="2" pageNumber="36" type="holotype">Holotype</typeStatus>
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(not dissected): “NE
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<collectionCode id="ACE2AE40076DFFE29FBBFC5FFCBAFBDE" box="[761,852,1018,1044]" pageId="2" pageNumber="36">
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<collectingCountry id="B2E47615076DFFE29FBBFC5FFCBAFBDE" box="[761,852,1018,1044]" name="India" pageId="2" pageNumber="36">INDIA</collectingCountry>
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</collectionCode>
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;
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<collectionCode id="ACE2AE40076DFFE29E23FC5FFC20FBDE" box="[865,974,1018,1044]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12694" name="Botanical Survey of India, Eastern Regional Centre" pageId="2" pageNumber="36">
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<collectingRegion id="0837F867076DFFE29E23FC5FFC20FBDE" box="[865,974,1018,1044]" country="India" name="Assam" pageId="2" pageNumber="36">ASSAM</collectingRegion>
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</collectionCode>
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; 1999 /
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<quantity id="0D0B9B60076DFFE29971FC5FFB87FBDE" box="[1075,1129,1018,1044]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="36" unit="km" value="5.0">5km</quantity>
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N of Umrongso;
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<quantity id="0D0B9B60076DFFE29807FC5FFA65FBDE" box="[1349,1419,1018,1044]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="36" unit="m" value="700.0">700m</quantity>
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/
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<geoCoordinate id="AFC75042076DFFE29DD5FB85FF13FBF0" box="[151,253,1056,1082]" direction="north" orientation="latitude" pageId="2" pageNumber="36" precision="925" value="25.45">25°27’N</geoCoordinate>
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<geoCoordinate id="AFC75042076DFFE29C47FB85FE85FBF0" box="[261,363,1056,1082]" direction="east" orientation="longitude" pageId="2" pageNumber="36" precision="925" value="92.71667">92°43’E</geoCoordinate>
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;
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<date id="BE4D1045076DFFE29C3AFB85FE0EFBF0" box="[376,480,1056,1082]" pageId="2" pageNumber="36">17.–25.v</date>
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/ Dembický & Pacholátko leg.”. Two
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<typeStatus id="15488827076DFFE29EE9FB85FBF2FBF0" box="[939,1052,1056,1082]" pageId="2" pageNumber="36" type="paratype">paratypes</typeStatus>
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(males): same data as
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<typeStatus id="15488827076DFFE2986EFB85FA79FBF0" box="[1324,1431,1056,1082]" pageId="2" pageNumber="36" type="holotype">holotype</typeStatus>
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; “
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<collectingCountry id="B2E47615076DFFE29DE3FBE0FF1DFB95" box="[161,243,1093,1119]" name="Laos" pageId="2" pageNumber="36">LAOS</collectingCountry>
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north,
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<date id="BE4D1045076DFFE29C0EFBE0FE10FB95" box="[332,510,1093,1119]" pageId="2" pageNumber="36" value="1997-05-13" valueMax="1997-05-24" valueMin="1997-05-13">13.–24.V.1997</date>
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/
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<quantity id="0D0B9B60076DFFE29F55FBE0FD8AFB95" box="[535,612,1093,1119]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="36" unit="km" value="15.0">15 km</quantity>
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NW
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<collectingRegion id="0837F867076DFFE29FEAFBE0FC83FB95" box="[680,877,1093,1119]" country="Laos" name="Louangnamtha" pageId="2" pageNumber="36">Louang Namtha</collectingRegion>
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/ N 21°07.5, S 101°21.0 / at. 750±
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<quantity id="0D0B9B60076DFFE2985EFBE0FA86FB95" box="[1308,1384,1093,1119]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="36" unit="m" value="100.0">100 m</quantity>
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/ E. Jendek & O. Šauša leg.” (all three
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<typeStatus id="15488827076DFFE29F01FBCFFD99FB4E" box="[579,631,1130,1156]" pageId="2" pageNumber="36">type</typeStatus>
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specimens deposited in the Moravian Museum, Brno,
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<collectingCountry id="B2E47615076DFFE29854FBCFFA92FB4E" box="[1302,1404,1130,1156]" name="Czech Republic" pageId="2" pageNumber="36">Czechia</collectingCountry>
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; J. Kolibáč coll.).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="82E9650E076DFFE89D86FB10FCB6FEE8" lastPageId="8" lastPageNumber="42" pageId="2" pageNumber="36" type="description">
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||
<paragraph id="CA4C3685076DFFE29D86FB10FBE9FAD0" blockId="2.[151,1437,1018,2015]" pageId="2" pageNumber="36">
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||
<emphasis id="F887EA97076DFFE29D86FB10FE8FFB05" bold="true" box="[196,353,1205,1231]" pageId="2" pageNumber="36">Description:</emphasis>
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Measurements:
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||
<typeStatus id="15488827076DFFE29F6AFB10FD79FB05" box="[552,663,1205,1231]" pageId="2" pageNumber="36" type="holotype">Holotype</typeStatus>
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||
– body length (from elytral apex to clypeus)
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<quantity id="0D0B9B60076DFFE299FEFB10FAF5FB05" box="[1212,1307,1205,1231]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="36" unit="mm" value="5.2">5.2 mm</quantity>
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, pronotum length
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<quantity id="0D0B9B60076DFFE29DA5FB7FFEAAFB3E" box="[231,324,1242,1268]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="36" unit="mm" value="1.3">1.3 mm</quantity>
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||
, elytra length
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||
<quantity id="0D0B9B60076DFFE29CABFB7FFDABFB3E" box="[489,581,1242,1268]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="36" unit="mm" value="3.5">3.5 mm</quantity>
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||
, pronotum width
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||
<quantity id="0D0B9B60076DFFE29E51FB7FFC9EFB3E" box="[787,880,1242,1268]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="36" unit="mm" value="3.1">3.1 mm</quantity>
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||
, elytra width (together)
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<quantity id="0D0B9B60076DFFE299CBFB7FFB0BFB3E" box="[1161,1253,1242,1268]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="36" unit="mm" value="3.3">3.3 mm</quantity>
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||
, antenna length
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<emphasis id="F887EA97076DFFE29DD5FAA5FF42FAD3" box="[151,172,1280,1305]" italics="true" pageId="2" pageNumber="36">c.</emphasis>
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||
<quantity id="0D0B9B60076DFFE29DF1FAA5FEE1FAD0" box="[179,271,1280,1306]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="36" unit="mm" value="0.8">0.8 mm</quantity>
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||
.
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<typeStatus id="15488827076DFFE29C59FAA5FE6CFAD0" box="[283,386,1280,1306]" pageId="2" pageNumber="36" type="paratype">Paratype</typeStatus>
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from
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<collectingCountry id="B2E47615076DFFE29C8BFAA5FDECFAD0" box="[457,514,1280,1306]" name="Laos" pageId="2" pageNumber="36">Laos</collectingCountry>
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– body length
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<quantity id="0D0B9B60076DFFE29FF3FAA5FCE0FAD0" box="[689,782,1280,1306]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="36" unit="mm" value="5.5">5.5 mm</quantity>
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, body width
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<quantity id="0D0B9B60076DFFE29EE4FAA5FBECFAD0" box="[934,1026,1280,1306]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="36" unit="mm" value="3.5">3.5 mm</quantity>
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||
.
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</paragraph>
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||
<paragraph id="CA4C3685076DFFE29D86FA80FC00F910" blockId="2.[151,1437,1018,2015]" pageId="2" pageNumber="36">
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Coloration and sculpture (
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<figureCitation id="52C82A00076DFFE29F41FA80FD5FFAF5" box="[515,689,1317,1343]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,272,1014,1038]" captionTargetBox="[171,1404,301,988]" captionTargetId="figure@3.[151,1436,291,993]" captionTargetPageId="3" captionText="FIGURES 1 – 5. (1) Adult habitus dorsally (holotype); (2) adult habitus laterally (holotype); (3) head and pronotum dorsally (paratype from Laos); (4 ab) head and prothorax ventrally, right leg removed (paratype from Assam); (5 ab) meso-, metathorax and abdomen ventrally, right legs removed (paratype from Assam)." httpUri="https://zenodo.org/record/275635/files/figure.png" pageId="2" pageNumber="36">Figs 1, 2, 3, 4</figureCitation>
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a, 5a): Body oval, conspicuously convex; dorsal surface dark brown to black in
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<collectingRegion id="0837F867076DFFE29C36FAEFFE2BFAAE" box="[372,453,1354,1380]" country="India" name="Assam" pageId="2" pageNumber="36">Assam</collectingRegion>
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specimens, brown in
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<collectingCountry id="B2E47615076DFFE29F8CFAEFFCE6FAAE" box="[718,776,1354,1380]" name="Laos" pageId="2" pageNumber="36">Laos</collectingCountry>
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specimen; margins of pronotum and elytra red-brown, colour lighter in
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<collectingCountry id="B2E47615076DFFE29C25FAD5FE4CFA40" box="[359,418,1392,1418]" name="Laos" pageId="2" pageNumber="36">Laos</collectingCountry>
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specimen. Head finely, densely granulate, with short, decumbent, yellowish setae. Pronotum granulate at centre, sparsely punctate along explanate lateral margins, with semi-erect yellowish pubescence; longitudinal tuft of long, erect, yellow-orange hairs situated at centre, small patches of decumbent hairs occurring along it. Each elytron with eight tufts of yellow-orange hairs (same pattern of tufts in all three specimens), a number of other long, erect hairs situated along mesonotum and elytral suture; explanate lateral part of elytra with semi-erect yellowish pubescence, finely and sparsely punctate; five elytral carinae present; one regular row of deep punctures running along each side of carina, the last two rows of punctures (in lateral direction) not accompanied by carina. Ventral surface of body dark brown to black; sparsely punctate, punctures minute; pubescence formed by short, sparse, decumbent pale hairs. Legs without sculpture, sparsely pubescent in the same way as ventral surface of body.
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||
</paragraph>
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<paragraph id="CA4C3685076DFFE29D86F940FCA6F85E" blockId="2.[151,1437,1018,2015]" pageId="2" pageNumber="36">
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||
Head (
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<figureCitation id="52C82A00076DFFE29C55F940FE65F935" box="[279,395,1765,1791]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,272,1014,1038]" captionTargetBox="[171,1404,301,988]" captionTargetId="figure@3.[151,1436,291,993]" captionTargetPageId="3" captionText="FIGURES 1 – 5. (1) Adult habitus dorsally (holotype); (2) adult habitus laterally (holotype); (3) head and pronotum dorsally (paratype from Laos); (4 ab) head and prothorax ventrally, right leg removed (paratype from Assam); (5 ab) meso-, metathorax and abdomen ventrally, right legs removed (paratype from Assam)." httpUri="https://zenodo.org/record/275635/files/figure.png" pageId="2" pageNumber="36">Figs 3–4</figureCitation>
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): gular sutures widely separated, extending to approximately midpoint of cranium, strongly convergent; frontoclypeal suture well-developed, deep, arcuate; antennal grooves in ventral side of cranium short, shallow; antennal sockets partly visible from above; two eyes present, relatively large (space between them twice as wide as eye diameter), distinctly elevated, not emarginate, elliptic, greater part situated dorsally; epicranial acumination conspicuous but not deep.
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</paragraph>
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<paragraph id="CA4C3685076DFFE39D86F805FBFCFF36" blockId="2.[151,1437,1018,2015]" lastBlockId="3.[151,1437,152,252]" lastPageId="3" lastPageNumber="37" pageId="2" pageNumber="36">
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Maxilla (
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<figureCitation id="52C82A00076DFFE29C6DF805FE99F870" box="[303,375,1952,1978]" captionStart="FIGURES 6 – 8" captionStartId="4.[151,269,1959,1983]" captionTargetBox="[212,1379,189,1931]" captionTargetId="figure@4.[191,1396,165,1937]" captionTargetPageId="4" captionText="FIGURES 6 – 8. (6) labrum (left side with dorsal, outer setae; right side inner, epipharyngeal setae); (7) labium (left site outer setae of prementum, right side inner setae of ligula and hypopharynx), right palpus removed; (8) maxilla." httpUri="https://zenodo.org/record/275636/files/figure.png" pageId="2" pageNumber="36">Fig. 8</figureCitation>
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): lacinia relatively robust, with three dark spines at apex in pattern 1+2 and
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<emphasis id="F887EA97076DFFE299B5F805FAE2F873" box="[1271,1292,1952,1977]" italics="true" pageId="2" pageNumber="36">c.</emphasis>
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3 pale setae between them, apical spine hook-like and two spines beneath it shorter; basigalea well-developed, pigmented; galea very weakly clavate, with numerous elongate setae, without spines and ciliate setae; mediostipes perfectly free, not fused with lacinia; basistipes elongate, rectangular; palpifer large, not denticulate along outer margin; maxillary palps 4-segmented, terminal joint conical, elongate.
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</paragraph>
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||
<caption id="9E8C660D076CFFE39DD5FC53FBDFFB84" httpUri="https://zenodo.org/record/275635/files/figure.png" pageId="3" pageNumber="37" targetBox="[171,1404,301,988]" targetPageId="3">
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||
<paragraph id="CA4C3685076CFFE39DD5FC53FBDFFB84" blockId="3.[151,1436,1014,1102]" pageId="3" pageNumber="37">
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<emphasis id="F887EA97076CFFE39DD5FC53FEA6FBC4" bold="true" box="[151,328,1014,1038]" pageId="3" pageNumber="37">FIGURES 1–5.</emphasis>
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(1) Adult habitus dorsally (holotype); (2) adult habitus laterally (holotype); (3) head and pronotum dorsally (paratype from Laos); (4ab) head and prothorax ventrally, right leg removed (paratype from Assam); (5ab) meso-, metathorax and abdomen ventrally, right legs removed (paratype from Assam).
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</paragraph>
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</caption>
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<paragraph id="CA4C3685076CFFE39D86FBD9FD55FACC" blockId="3.[151,1437,1148,2033]" pageId="3" pageNumber="37">
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Mandible (
|
||
<figureCitation id="52C82A00076CFFE39C08FBD9FE48FB5C" box="[330,422,1148,1174]" captionStart="FIGURES 9 – 13" captionStartId="5.[151,271,1964,1988]" captionTargetBox="[214,1375,219,1923]" captionTargetId="figure@5.[202,1389,194,1939]" captionTargetPageId="5" captionText="FIGURES 9 – 13. (9) tergite VIII (pygidium); (10) antenna; (11) middle tibia and tarsus; (12) mandible ventrally; (13) sternite (ventrite) VIII." httpUri="https://zenodo.org/record/275637/files/figure.png" pageId="3" pageNumber="37">Fig. 12</figureCitation>
|
||
): with two apical teeth situated side by side (in horizontal axis); medial tooth very small, double; mola present, denticulate along margin; membranous, finely but densely pubescent appendage situated above mola (penicillus?); prostheca situated below medial tooth, formed by tuft of long, robust setae; ventral ciliate furrow absent; basal notch deep.
|
||
</paragraph>
|
||
<paragraph id="CA4C3685076CFFE39D86FAB7FDC0FABC" blockId="3.[151,1437,1148,2033]" pageId="3" pageNumber="37">
|
||
Labrum (
|
||
<figureCitation id="52C82A00076CFFE39C71FAB7FE90FAE6" box="[307,382,1298,1324]" captionStart="FIGURES 6 – 8" captionStartId="4.[151,269,1959,1983]" captionTargetBox="[212,1379,189,1931]" captionTargetId="figure@4.[191,1396,165,1937]" captionTargetPageId="4" captionText="FIGURES 6 – 8. (6) labrum (left side with dorsal, outer setae; right side inner, epipharyngeal setae); (7) labium (left site outer setae of prementum, right side inner setae of ligula and hypopharynx), right palpus removed; (8) maxilla." httpUri="https://zenodo.org/record/275636/files/figure.png" pageId="3" pageNumber="37">Fig. 6</figureCitation>
|
||
): free (not fused with cranium); oval, not emarginate in apical part; epipharynx without distinct sclerite; lateral tormal processes not connected in middle; V-shaped sclerite with dense pubescence situated between tormal processes.
|
||
</paragraph>
|
||
<paragraph id="CA4C3685076CFFE39D86FA27FB43F9FB" blockId="3.[151,1437,1148,2033]" pageId="3" pageNumber="37">
|
||
Labium (
|
||
<figureCitation id="52C82A00076CFFE39C74FA27FE6BFA56" box="[310,389,1410,1436]" captionStart="FIGURES 6 – 8" captionStartId="4.[151,269,1959,1983]" captionTargetBox="[212,1379,189,1931]" captionTargetId="figure@4.[191,1396,165,1937]" captionTargetPageId="4" captionText="FIGURES 6 – 8. (6) labrum (left side with dorsal, outer setae; right side inner, epipharyngeal setae); (7) labium (left site outer setae of prementum, right side inner setae of ligula and hypopharynx), right palpus removed; (8) maxilla." httpUri="https://zenodo.org/record/275636/files/figure.png" pageId="3" pageNumber="37">Fig. 7</figureCitation>
|
||
): submentum without tuft of setae; mentum trapezoidal, anterior corners extended; prementum without notch and not divided into two parts; ligula membranous, deeply emarginate, without ciliate setae, with long pale setae; mental apodeme composed of two elongate sclerites; hypopharyngeal bar absent; hypopharyngeal sclerite absent; hypopharynx with two strips of short setae along ligula, strips fused into one in mental area; terminal joint of labial palps shorter than the labial ones, conical.
|
||
</paragraph>
|
||
<paragraph id="CA4C3685076CFFE39D86F999FD8CF90C" blockId="3.[151,1437,1148,2033]" pageId="3" pageNumber="37">
|
||
Antennae (
|
||
<figureCitation id="52C82A00076CFFE39C0CF999FE43F99C" box="[334,429,1596,1622]" captionStart="FIGURES 9 – 13" captionStartId="5.[151,271,1964,1988]" captionTargetBox="[214,1375,219,1923]" captionTargetId="figure@5.[202,1389,194,1939]" captionTargetPageId="5" captionText="FIGURES 9 – 13. (9) tergite VIII (pygidium); (10) antenna; (11) middle tibia and tarsus; (12) mandible ventrally; (13) sternite (ventrite) VIII." httpUri="https://zenodo.org/record/275637/files/figure.png" pageId="3" pageNumber="37">Fig. 10</figureCitation>
|
||
): 11-segmented; antennal club 3-segmented, relatively compact; antennal joints symmetrical or very weakly asymmetrical (joints 9–10), without conspicuous sensorial fields, with short pubescence; scapus robust, pedicellus smaller, but larger than joint 3; antennae relatively short, extending backwards to midway along pronotum.
|
||
</paragraph>
|
||
<paragraph id="CA4C3685076CFFE39D86F977FB4AF84B" blockId="3.[151,1437,1148,2033]" pageId="3" pageNumber="37">
|
||
Prothorax (
|
||
<figureCitation id="52C82A00076CFFE39C0AF977FE5AF926" box="[328,436,1746,1772]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,272,1014,1038]" captionTargetBox="[171,1404,301,988]" captionTargetId="figure@3.[151,1436,291,993]" captionTargetPageId="3" captionText="FIGURES 1 – 5. (1) Adult habitus dorsally (holotype); (2) adult habitus laterally (holotype); (3) head and pronotum dorsally (paratype from Laos); (4 ab) head and prothorax ventrally, right leg removed (paratype from Assam); (5 ab) meso-, metathorax and abdomen ventrally, right legs removed (paratype from Assam)." httpUri="https://zenodo.org/record/275635/files/figure.png" pageId="3" pageNumber="37">Figs 3, 4</figureCitation>
|
||
): pronotum transverse; anterior margin deeply emarginate (arcuate), anterior corners projecting; lateral margins excavate; lateral edge present, finely undulating but not denticulate; prosternal process dilated at apex, with two sharp grooves running along margins of process and procoxal cavities; procoxal cavities externally open to halfway, internally closed; postcoxal projections pubescent along margins; trochantin well-developed; inner part of hypomeron with fine longitudinal line.
|
||
</paragraph>
|
||
<paragraph id="CA4C3685076CFFE39D86F829FCACF83B" blockId="3.[151,1437,1148,2033]" pageId="3" pageNumber="37">
|
||
Mesothorax (
|
||
<figureCitation id="52C82A00076CFFE39C2DF829FE09F86C" box="[367,487,1932,1958]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,272,1014,1038]" captionTargetBox="[171,1404,301,988]" captionTargetId="figure@3.[151,1436,291,993]" captionTargetPageId="3" captionText="FIGURES 1 – 5. (1) Adult habitus dorsally (holotype); (2) adult habitus laterally (holotype); (3) head and pronotum dorsally (paratype from Laos); (4 ab) head and prothorax ventrally, right leg removed (paratype from Assam); (5 ab) meso-, metathorax and abdomen ventrally, right legs removed (paratype from Assam)." httpUri="https://zenodo.org/record/275635/files/figure.png" pageId="3" pageNumber="37">Figs 3, 5</figureCitation>
|
||
): prepectus present in anterior part of mesepisternum only, absent above mesosternum; mesocoxal cavities externally widely open, internally closed; trochantin minute; mesonotum wide, scutum with extended basal corners, scutellum oval.
|
||
</paragraph>
|
||
<caption id="9E8C660D076BFFE49DD5F802FAD8F814" httpUri="https://zenodo.org/record/275636/files/figure.png" pageId="4" pageNumber="38" targetBox="[212,1379,189,1931]" targetPageId="4">
|
||
<paragraph id="CA4C3685076BFFE49DD5F802FAD8F814" blockId="4.[151,1436,1959,2014]" pageId="4" pageNumber="38">
|
||
<emphasis id="F887EA97076BFFE49DD5F802FEADF875" bold="true" box="[151,323,1959,1983]" pageId="4" pageNumber="38">FIGURES 6–8.</emphasis>
|
||
(6) labrum (left side with dorsal, outer setae; right side inner, epipharyngeal setae); (7) labium (left site outer setae of prementum, right side inner setae of ligula and hypopharynx), right palpus removed; (8) maxilla.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="9E8C660D076AFFE59DD5F809FE65F829" httpUri="https://zenodo.org/record/275637/files/figure.png" pageId="5" pageNumber="39" targetBox="[214,1375,219,1923]" targetPageId="5">
|
||
<paragraph id="CA4C3685076AFFE59DD5F809FE65F829" blockId="5.[151,1436,1964,2019]" pageId="5" pageNumber="39">
|
||
<emphasis id="F887EA97076AFFE59DD5F809FEBDF80E" bold="true" box="[151,339,1964,1988]" pageId="5" pageNumber="39">FIGURES 9–13.</emphasis>
|
||
(9) tergite VIII (pygidium); (10) antenna; (11) middle tibia and tarsus; (12) mandible ventrally; (13) sternite (ventrite) VIII.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="9E8C660D0769FFE69DD5F80AFAB6F80C" box="[151,1368,1967,1991]" httpUri="https://zenodo.org/record/275638/files/figure.png" pageId="6" pageNumber="40" targetBox="[226,1367,218,1939]" targetPageId="6">
|
||
<paragraph id="CA4C36850769FFE69DD5F80AFAB6F80C" blockId="6.[151,1368,1967,1991]" box="[151,1368,1967,1991]" pageId="6" pageNumber="40">
|
||
<emphasis id="F887EA970769FFE69DD5F80AFEB3F80D" bold="true" box="[151,349,1967,1991]" pageId="6" pageNumber="40">FIGURES 14–17.</emphasis>
|
||
(14) wing; (15) aedeagus ventrally; (16) abdominal segment IX dorsally; (17) metendosternite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CA4C36850768FFE79D84FF3DFD00FF3D" blockId="7.[151,1437,152,916]" pageId="7" pageNumber="41">
|
||
Wing (
|
||
<figureCitation id="52C82A000768FFE79C5AFF3DFE9CFF78" box="[280,370,152,178]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,269,1967,1991]" captionTargetBox="[226,1367,218,1939]" captionTargetId="figure@6.[198,1389,194,1944]" captionTargetPageId="6" captionText="FIGURES 14 – 17. (14) wing; (15) aedeagus ventrally; (16) abdominal segment IX dorsally; (17) metendosternite." httpUri="https://zenodo.org/record/275638/files/figure.png" pageId="7" pageNumber="41">Fig. 14</figureCitation>
|
||
): radial cell displaced downwards; r4 relatively short; pigmented fleck (below Rc) small, pale, not complete; medial field with all four veins, cross-veins absent (wedge cell absent); MP4 shortly bifurcate at apex; wing pigmented above vein Rp2.
|
||
</paragraph>
|
||
<paragraph id="CA4C36850768FFE79D86FEA5FB70FE40" blockId="7.[151,1437,152,916]" pageId="7" pageNumber="41">
|
||
Metathorax (
|
||
<figureCitation id="52C82A000768FFE79C19FEA5FE49FED0" box="[347,423,256,282]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,272,1014,1038]" captionTargetBox="[171,1404,301,988]" captionTargetId="figure@3.[151,1436,291,993]" captionTargetPageId="3" captionText="FIGURES 1 – 5. (1) Adult habitus dorsally (holotype); (2) adult habitus laterally (holotype); (3) head and pronotum dorsally (paratype from Laos); (4 ab) head and prothorax ventrally, right leg removed (paratype from Assam); (5 ab) meso-, metathorax and abdomen ventrally, right legs removed (paratype from Assam)." httpUri="https://zenodo.org/record/275635/files/figure.png" pageId="7" pageNumber="41">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="52C82A000768FFE79CF7FEA5FE38FED0" box="[437,470,256,282]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,269,1967,1991]" captionTargetBox="[226,1367,218,1939]" captionTargetId="figure@6.[198,1389,194,1944]" captionTargetPageId="6" captionText="FIGURES 14 – 17. (14) wing; (15) aedeagus ventrally; (16) abdominal segment IX dorsally; (17) metendosternite." httpUri="https://zenodo.org/record/275638/files/figure.png" pageId="7" pageNumber="41">17</figureCitation>
|
||
): metasternum flat and wide, distinctly narrowed towards anterior portion, widely bordered along lateral sides; discriminal line (discrimen) short; paracoxal sutures well-demarcated, parallel with coxae; metepisternum distinctly triangular; metanotum rectangular, extremely wide; metendosternite with broad stalk, laminae narrowed at apex, anterior tendons long and widely separated.
|
||
</paragraph>
|
||
<paragraph id="CA4C36850768FFE79D86FE30FCBCFE1E" blockId="7.[151,1437,152,916]" pageId="7" pageNumber="41">
|
||
Elytra (
|
||
<figureCitation id="52C82A000768FFE79C58FE30FE4FFE65" box="[282,417,405,431]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,272,1014,1038]" captionTargetBox="[171,1404,301,988]" captionTargetId="figure@3.[151,1436,291,993]" captionTargetPageId="3" captionText="FIGURES 1 – 5. (1) Adult habitus dorsally (holotype); (2) adult habitus laterally (holotype); (3) head and pronotum dorsally (paratype from Laos); (4 ab) head and prothorax ventrally, right leg removed (paratype from Assam); (5 ab) meso-, metathorax and abdomen ventrally, right legs removed (paratype from Assam)." httpUri="https://zenodo.org/record/275635/files/figure.png" pageId="7" pageNumber="41">Figs 1, 2, 5</figureCitation>
|
||
): regularly punctate, with extremely wide epipleura; elytral locking mechanism absent; five elytral carinae present; lateral sides of elytra explanate.
|
||
</paragraph>
|
||
<paragraph id="CA4C36850768FFE79D86FE45FCBDFD7E" blockId="7.[151,1437,152,916]" pageId="7" pageNumber="41">
|
||
Legs (
|
||
<figureCitation id="52C82A000768FFE79C4FFE45FE97FE30" box="[269,377,480,506]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,272,1014,1038]" captionTargetBox="[171,1404,301,988]" captionTargetId="figure@3.[151,1436,291,993]" captionTargetPageId="3" captionText="FIGURES 1 – 5. (1) Adult habitus dorsally (holotype); (2) adult habitus laterally (holotype); (3) head and pronotum dorsally (paratype from Laos); (4 ab) head and prothorax ventrally, right leg removed (paratype from Assam); (5 ab) meso-, metathorax and abdomen ventrally, right legs removed (paratype from Assam)." httpUri="https://zenodo.org/record/275635/files/figure.png" pageId="7" pageNumber="41">Figs 4, 5</figureCitation>
|
||
,
|
||
<figureCitation id="52C82A000768FFE79CC5FE45FE49FE30" box="[391,423,480,506]" captionStart="FIGURES 9 – 13" captionStartId="5.[151,271,1964,1988]" captionTargetBox="[214,1375,219,1923]" captionTargetId="figure@5.[202,1389,194,1939]" captionTargetPageId="5" captionText="FIGURES 9 – 13. (9) tergite VIII (pygidium); (10) antenna; (11) middle tibia and tarsus; (12) mandible ventrally; (13) sternite (ventrite) VIII." httpUri="https://zenodo.org/record/275637/files/figure.png" pageId="7" pageNumber="41">11</figureCitation>
|
||
): procoxae not projecting, transverse; mesocoxae oval, metacoxae extended to lateral surface of metathorax; trochanters relatively small, triangular; femora very weakly clavate; tibiae with row of small spines along outer margin; tibial apical spur pattern 1–1–1; protibial spur hooked; apices of all tibiae with row of
|
||
<emphasis id="F887EA970768FFE79C6EFDF5FEAFFDA3" box="[300,321,592,617]" italics="true" pageId="7" pageNumber="41">c.</emphasis>
|
||
5 spines; tarsomere
|
||
<quantity id="0D0B9B600768FFE79F7AFDF5FD88FDA0" box="[568,614,592,618]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="41" unit="in" value="1.0">1 in</quantity>
|
||
all pairs of legs conspicuous, approximately as long as tarsomere 2; tarsomere 5 as long as, or slightly longer than, 1–4 combined; tarsal lobes absent; claws without denticles; empodium large, projecting, bisetose; tarsal formula 5–5–5.
|
||
</paragraph>
|
||
<paragraph id="CA4C36850768FFE79D86FD65FBBCFC5E" blockId="7.[151,1437,152,916]" pageId="7" pageNumber="41">
|
||
Abdomen (
|
||
<figureCitation id="52C82A000768FFE79C0AFD65FE78FD10" box="[328,406,704,730]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,272,1014,1038]" captionTargetBox="[171,1404,301,988]" captionTargetId="figure@3.[151,1436,291,993]" captionTargetPageId="3" captionText="FIGURES 1 – 5. (1) Adult habitus dorsally (holotype); (2) adult habitus laterally (holotype); (3) head and pronotum dorsally (paratype from Laos); (4 ab) head and prothorax ventrally, right leg removed (paratype from Assam); (5 ab) meso-, metathorax and abdomen ventrally, right legs removed (paratype from Assam)." httpUri="https://zenodo.org/record/275635/files/figure.png" pageId="7" pageNumber="41">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="52C82A000768FFE79CE1FD65FE0FFD10" box="[419,481,704,730]" captionStart="FIGURES 9 – 13" captionStartId="5.[151,271,1964,1988]" captionTargetBox="[214,1375,219,1923]" captionTargetId="figure@5.[202,1389,194,1939]" captionTargetPageId="5" captionText="FIGURES 9 – 13. (9) tergite VIII (pygidium); (10) antenna; (11) middle tibia and tarsus; (12) mandible ventrally; (13) sternite (ventrite) VIII." httpUri="https://zenodo.org/record/275637/files/figure.png" pageId="7" pageNumber="41">9, 13</figureCitation>
|
||
,
|
||
<figureCitation id="52C82A000768FFE79CACFD65FDD2FD10" box="[494,572,704,730]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,269,1967,1991]" captionTargetBox="[226,1367,218,1939]" captionTargetId="figure@6.[198,1389,194,1944]" captionTargetPageId="6" captionText="FIGURES 14 – 17. (14) wing; (15) aedeagus ventrally; (16) abdominal segment IX dorsally; (17) metendosternite." httpUri="https://zenodo.org/record/275638/files/figure.png" pageId="7" pageNumber="41">15, 16</figureCitation>
|
||
): five sternites visible, the last one (sternite VII) with row of short hairs; male sternite VIII without spiculum gastrale, with longitudinal unpigmented area in centre (sternite VIII in the second
|
||
<typeStatus id="154888270768FFE79DB0FCAFFEB7FCEE" box="[242,345,778,804]" pageId="7" pageNumber="41" type="paratype">paratype</typeStatus>
|
||
shorter); male segment IX complete; tegmen composed of two parts (parameres separated from phallobase, dorsal and ventral pieces fused), inverted (phallobase ventrally open); phallobasic apodeme well-developed; tegminal struts coalescent with base of phallobasic apodeme; parameres densely pubescent; phallus stout, slightly shorter than phallobase, with several minute spines at apex.
|
||
</paragraph>
|
||
<caption id="9E8C660D0768FFE79DD5F880FBE7F8F6" box="[151,1033,1829,1853]" httpUri="https://zenodo.org/record/275639/files/figure.png" pageId="7" pageNumber="41" targetBox="[151,1436,968,1808]" targetPageId="7">
|
||
<paragraph id="CA4C36850768FFE79DD5F880FBE7F8F6" blockId="7.[151,1033,1829,1853]" box="[151,1033,1829,1853]" pageId="7" pageNumber="41">
|
||
<emphasis id="F887EA970768FFE79DD5F880FEC8F8F7" bold="true" box="[151,294,1829,1853]" pageId="7" pageNumber="41">FIGURE 18.</emphasis>
|
||
The type locality: the forest clearing near Umrongso (India: Assam).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CA4C36850768FFE89D86F820FCB6FEE8" blockId="7.[151,1437,1925,2026]" lastBlockId="8.[151,1437,152,327]" lastPageId="8" lastPageNumber="42" pageId="7" pageNumber="41">
|
||
<emphasis id="F887EA970768FFE79D86F820FD93F855" bold="true" box="[196,637,1925,1951]" pageId="7" pageNumber="41">Note on distribution and biology:</emphasis>
|
||
Two specimens were knocked down from fallen logs in a forest clearing (
|
||
<collectingCountry id="B2E476150768FFE79C46F80EFEA8F80F" box="[260,326,1963,1989]" name="India" pageId="7" pageNumber="41">India</collectingCountry>
|
||
:
|
||
<collectingRegion id="0837F8670768FFE79C13F80EFE4CF80F" box="[337,418,1963,1989]" country="India" name="Assam" pageId="7" pageNumber="41">Assam</collectingRegion>
|
||
) (
|
||
<figureCitation id="52C82A000768FFE79CF9F80EFDFFF80F" box="[443,529,1963,1989]" captionStart="FIGURE 18" captionStartId="7.[151,255,1829,1853]" captionTargetBox="[151,1436,968,1808]" captionTargetId="figure@7.[151,1436,968,1808]" captionTargetPageId="7" captionText="FIGURE 18. The type locality: the forest clearing near Umrongso (India: Assam)." httpUri="https://zenodo.org/record/275639/files/figure.png" pageId="7" pageNumber="41">Fig. 18</figureCitation>
|
||
); one specimen from northern
|
||
<collectingCountry id="B2E476150768FFE79E3BF80EFC5CF80F" box="[889,946,1963,1989]" name="Laos" pageId="7" pageNumber="41">Laos</collectingCountry>
|
||
was probably caught in the same manner. The species may also be present in
|
||
<collectingCountry id="B2E476150768FFE79F11F875FD24F820" box="[595,714,2000,2026]" name="Myanmar" pageId="7" pageNumber="41">Myanmar</collectingCountry>
|
||
(
|
||
<collectingCountry id="B2E476150768FFE79F99F875FCDCF820" box="[731,818,2000,2026]" name="Myanmar" pageId="7" pageNumber="41">Burma</collectingCountry>
|
||
) between the two known localities but is perhaps relatively rare (numerous coleopterists, including myself, have collected beetles in the surroundings of
|
||
<collectingRegion id="0837F8670767FFE89DD5FF18FEBBFF1D" box="[151,341,189,215]" country="Laos" name="Louangnamtha" pageId="8" pageNumber="42">Louang Namtha</collectingRegion>
|
||
in recent years but only the single specimen is known from
|
||
<collectingCountry id="B2E476150767FFE89954FF18FBA1FF1D" box="[1046,1103,189,215]" name="Laos" pageId="8" pageNumber="42">Laos</collectingCountry>
|
||
to date). Gut content of the both dissected
|
||
<typeStatus id="154888270767FFE89F09FF47FD55FF36" box="[587,699,226,252]" pageId="8" pageNumber="42" type="paratype">paratypes</typeStatus>
|
||
with small remnants of insect cuticle,
|
||
<typeStatus id="154888270767FFE8993FFF47FB0CFF36" box="[1149,1250,226,252]" pageId="8" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
from
|
||
<collectingCountry id="B2E476150767FFE8986AFF47FA8FFF36" box="[1320,1377,226,252]" name="Laos" pageId="8" pageNumber="42">Laos</collectingCountry>
|
||
with the remains of a small insect larva. The species is predatory.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="82E9650E0767FFE89D84FE88FB82FE8D" box="[198,1132,301,327]" pageId="8" pageNumber="42" type="etymology">
|
||
<paragraph id="CA4C36850767FFE89D84FE88FB82FE8D" blockId="8.[151,1437,152,327]" box="[198,1132,301,327]" pageId="8" pageNumber="42">
|
||
<emphasis id="F887EA970767FFE89D84FE88FEB9FE8D" bold="true" box="[198,343,301,327]" pageId="8" pageNumber="42">Etymology:</emphasis>
|
||
<emphasis id="F887EA970767FFE89C1CFE88FE3CFE8C" box="[350,466,301,326]" italics="true" pageId="8" pageNumber="42">Fascicula</emphasis>
|
||
= small beam, bundle, sheaf;
|
||
<emphasis id="F887EA970767FFE89E70FE88FC84FE8C" box="[818,874,301,326]" italics="true" pageId="8" pageNumber="42">ferre</emphasis>
|
||
= to carry, bear, hold.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |