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<document id="469EBEF9B4DB85B67878BD5760F9D4CA" ID-DOI="10.11646/zootaxa.5032.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5495554" ID-ZooBank="65894101-2D60-426D-B149-4CF6E1C561BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631115156823" checkinUser="plazi" docAuthor="Enuschenko, Ilya V. &amp; Shavrin, Alexey V." docDate="2021" docId="0C1987ABFFE0297DFF7EA367B0A0F81B" docLanguage="en" docName="zootaxa.5032.3.2.pdf" docOrigin="Zootaxa 5032 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gyrophaena (Gyrophaena) immatura Kraatz 1859" docType="treatment" docVersion="3" lastPageNumber="337" masterDocId="F020FFD3FFE6297BFFE9A646B131FFA5" masterDocTitle="On some species of Gyrophaenina Kraatz 1856 of Sri Lanka and India (Coleoptera: Staphylinidae: Aleocharinae: Homalotini)" masterLastPageNumber="356" masterPageNumber="331" pageNumber="337" updateTime="1699405676542" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F621358E84D6ADAF798C8C5018641715">On some species of Gyrophaenina Kraatz 1856 of Sri Lanka and India (Coleoptera: Staphylinidae: Aleocharinae: Homalotini)</mods:title>
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<mods:namePart id="E78603E0EAB4D777430EA0C429C35016">Enuschenko, Ilya V.</mods:namePart>
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<mods:affiliation id="A4FB0D3DC974D6974DC146715103393B">Limnological Institute SB RAS, Ulan-Batorskaya str., 3, Irkutsk, 664033, Russia.</mods:affiliation>
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<mods:namePart id="0FE73C430CEC5CE826C8AFFB57F7189B">Shavrin, Alexey V.</mods:namePart>
<mods:nameIdentifier id="20FDD76955D58D2DB9874E0A62998840" type="ORCID">0000-0001-6487-4530</mods:nameIdentifier>
<mods:affiliation id="80EDEA58D2EB131E024B08402A9B1CEE">Institute of Life Sciences and Technologies, Daugavpils Universit, Vienibas 13, Daugavpils, LV- 5401, Latvia. ashavrin @ hotmail. com</mods:affiliation>
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<mods:title id="41E5DBF79E5F8273DE587CFC87A7EC95">Zootaxa</mods:title>
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<mods:date id="BD46714D86FBA57E75F1A062626BD2BF">2021</mods:date>
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<mods:number id="8CD193A7793660B8977581FBCB72089E">5032</mods:number>
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<treatment id="0C1987ABFFE0297DFF7EA367B0A0F81B" LSID="urn:lsid:plazi:treatment:0C1987ABFFE0297DFF7EA367B0A0F81B" httpUri="http://treatment.plazi.org/id/0C1987ABFFE0297DFF7EA367B0A0F81B" lastPageNumber="337" pageId="6" pageNumber="337">
<subSubSection id="CCAA6536FFE0297DFF7EA367B234FA99" box="[151,773,1313,1340]" pageId="6" pageNumber="337" type="nomenclature">
<paragraph id="840F36BDFFE0297DFF7EA367B234FA99" blockId="6.[151,773,1313,1377]" box="[151,773,1313,1340]" pageId="6" pageNumber="337">
<heading id="DF4781D1FFE0297DFF7EA367B234FA99" bold="true" box="[151,773,1313,1340]" fontSize="11" level="1" pageId="6" pageNumber="337" reason="1">
<taxonomicName id="43B04D3EFFE0297DFF7EA367B234FA99" ID-CoL="8Y465" authority="Kraatz, 1859" authorityName="Kraatz" authorityYear="1859" box="[151,773,1313,1340]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="immatura" subGenus="Gyrophaena">
<emphasis id="B6C4EAAFFFE0297DFF7EA367B234FA99" bold="true" box="[151,773,1313,1340]" pageId="6" pageNumber="337">
<emphasis id="B6C4EAAFFFE0297DFF7EA367B01CFA99" bold="true" box="[151,301,1313,1340]" italics="true" pageId="6" pageNumber="337">Gyrophaena</emphasis>
(
<emphasis id="B6C4EAAFFFE0297DFED7A367B0E5FA99" bold="true" box="[318,468,1313,1340]" italics="true" pageId="6" pageNumber="337">Gyrophaena</emphasis>
)
<emphasis id="B6C4EAAFFFE0297DFE0CA364B36DFA99" bold="true" box="[485,604,1314,1340]" italics="true" pageId="6" pageNumber="337">immatura</emphasis>
<bibRefCitation id="E0214B4CFFE0297DFD8DA364B234FA99" author="Kraatz, G." box="[612,773,1314,1340]" pageId="6" pageNumber="337" pagination="1 - 193" refId="ref14558" refString="Kraatz, G. (1859) Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. Archiv fur Naturgeschichte, 25 (1), 1 - 193, pls. 1 - 3. https: // doi. org / 10.5962 / bhl. title. 66002" type="journal article" year="1859">Kraatz, 1859</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCAA6536FFE0297DFF7EA300B016FAC4" box="[151,295,1350,1377]" pageId="6" pageNumber="337" type="description">
<paragraph id="840F36BDFFE0297DFF7EA300B016FAC4" blockId="6.[151,773,1313,1377]" box="[151,295,1350,1377]" pageId="6" pageNumber="337">
(
<figureCitation id="1C8B2A38FFE0297DFF76A300B02EFAC4" box="[159,287,1350,1377]" captionStart="FIGURES 22–26" captionStartId="7.[151,264,1076,1100]" captionTargetBox="[168,1418,188,1041]" captionTargetId="figure-19@7.[151,1435,181,1052]" captionTargetPageId="7" captionText="FIGURES 22–26. Gyrophaena immatura: 22–habitus, 23–aedeagus, lateral view, 24–male abdominal tergite VIII, dorsal view, 25–male abdominal sternite VIII, ventral view, 26–labels. Scale bars: 1.0 mm (Fig. 22), 0.10 mm (Figs 23–25)." figureDoi="http://doi.org/10.5281/zenodo.5495638" httpUri="https://zenodo.org/record/5495638/files/figure.png" pageId="6" pageNumber="337">Figs 22–26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CCAA6536FFE0297DFF7EA3CCB45DFA07" box="[151,1388,1418,1442]" pageId="6" pageNumber="337" type="reference_group">
<paragraph id="840F36BDFFE0297DFF7EA3CCB45DFA07" blockId="6.[151,1388,1418,1442]" box="[151,1388,1418,1442]" pageId="6" pageNumber="337">
<treatmentCitationGroup id="A4A01193FFE0297DFF7EA3CCB45DFA07" box="[151,1388,1418,1442]" pageId="6" pageNumber="337">
<treatmentCitation id="051110ACFFE0297DFF7EA3CCB301FA07" author="Kraatz, G." box="[151,560,1418,1442]" page="47" pageId="6" pageNumber="337" year="1859">
<taxonomicName id="43B04D3EFFE0297DFF7EA3CCB301FA07" ID-CoL="8Y465" authority="Kraatz, 1859: 47" authorityName="Kraatz" authorityPageNumber="47" authorityYear="1859" box="[151,560,1418,1442]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="immatura">
<emphasis id="B6C4EAAFFFE0297DFF7EA3CCB04FFA07" box="[151,382,1418,1442]" italics="true" pageId="6" pageNumber="337">Gyrophaena immatura</emphasis>
<bibRefCitation id="E0214B4CFFE0297DFE6DA3CDB301FA07" author="Kraatz, G." box="[388,560,1418,1442]" pageId="6" pageNumber="337" pagination="1 - 193" refId="ref14558" refString="Kraatz, G. (1859) Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. Archiv fur Naturgeschichte, 25 (1), 1 - 193, pls. 1 - 3. https: // doi. org / 10.5962 / bhl. title. 66002" type="journal article" year="1859">Kraatz, 1859: 47</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="051110ACFFE0297DFDD2A3CCB21DFA07" author="Motschulsky, V. I." box="[571,812,1418,1442]" page="150" pageId="6" pageNumber="337" year="1861">
<bibRefCitation id="E0214B4CFFE0297DFDD2A3CCB21DFA07" author="Motschulsky, V. I." box="[571,812,1418,1442]" pageId="6" pageNumber="337" pagination="95 - 155" refId="ref14675" refString="Motschulsky, V. I. (1861) Essai d'un catalogue des insects de l'ile Ceylan. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 34 (1), 95 - 155." type="journal article" year="1861">Motschulsky 1861: 150</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="051110ACFFE0297DFCDEA3CCB594FA07" author="Bernhauer, M. &amp; Scheerpeltz, O." box="[823,1189,1418,1442]" page="531" pageId="6" pageNumber="337" year="1926">
<bibRefCitation id="E0214B4CFFE0297DFCDEA3CCB594FA07" author="Bernhauer, M. &amp; Scheerpeltz, O." box="[823,1189,1418,1442]" pageId="6" pageNumber="337" pagination="499 - 988" refId="ref13956" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. (Pars 82). In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Volumen 5. Staphylinidae. W. Junk, Berlin, pp. 499 - 988." type="book chapter" year="1926">Bernhauer &amp; Scheerpeltz 1926: 531</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="051110ACFFE0297DFB59A3CCB45DFA07" author="Cameron, M." box="[1200,1388,1418,1442]" page="81" pageId="6" pageNumber="337" year="1939">
<bibRefCitation id="E0214B4CFFE0297DFB59A3CCB45DFA07" author="Cameron, M." box="[1200,1388,1418,1442]" pageId="6" pageNumber="337" pagination="1 - 3" refId="ref14209" refString="Cameron, M. (1939) Coleoptera, Staphylinidae. Vol. IV. Parts I &amp; II. In: Sewell, R. B. S. (Ed.), The Fauna of British India, including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xviii + 1 - 691, pls. 1 - 3." type="book chapter" year="1939">Cameron 1939: 81</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CCAA6536FFE0297DFF7EA396B008F9AA" pageId="6" pageNumber="337" type="materials_examined">
<paragraph id="840F36BDFFE0297DFF7EA396B008F9AA" blockId="6.[151,1437,1488,1982]" pageId="6" pageNumber="337">
<emphasis id="B6C4EAAFFFE0297DFF7EA396B04CFA4F" bold="true" box="[151,381,1488,1514]" pageId="6" pageNumber="337">Material examined</emphasis>
:
<materialsCitation id="34D83CE0FFE0297DFE66A397B004F9AA" collectionCode="ZIN" country="Sri Lanka" location="Kraatz'" pageId="6" pageNumber="337" specimenCount="5" specimenCount-male="4">
<specimenCount id="92B6FD34FFE0297DFE66A397B08AFA4E" box="[399,443,1489,1515]" pageId="6" pageNumber="337" type="male">4 ♂</specimenCount>
# [
<specimenCount id="92B6FD34FFE0297DFE32A397B3B6FA4F" box="[475,647,1488,1515]" pageId="6" pageNumber="337" type="generic">two specimens</specimenCount>
dissected; labels as in
<figureCitation id="1C8B2A38FFE0297DFC7FA396B2DEFA4E" box="[918,1007,1488,1515]" captionStart="FIGURES 22–26" captionStartId="7.[151,264,1076,1100]" captionTargetBox="[168,1418,188,1041]" captionTargetId="figure-19@7.[151,1435,181,1052]" captionTargetPageId="7" captionText="FIGURES 22–26. Gyrophaena immatura: 22–habitus, 23–aedeagus, lateral view, 24–male abdominal tergite VIII, dorsal view, 25–male abdominal sternite VIII, ventral view, 26–labels. Scale bars: 1.0 mm (Fig. 22), 0.10 mm (Figs 23–25)." figureDoi="http://doi.org/10.5281/zenodo.5495638" httpUri="https://zenodo.org/record/5495638/files/figure.png" pageId="6" pageNumber="337">Fig. 26</figureCitation>
]: ‘
<collectingCountry id="FCA7762DFFE0297DFBE0A396B56CFA4F" box="[1033,1117,1488,1514]" name="Sri Lanka" pageId="6" pageNumber="337">Ceylan</collectingCountry>
6515–1.’, ‘6515.
<taxonomicName id="43B04D3EFFE0297DFAC6A397B4ADFA4F" authorityName="Kraatz" authorityYear="1859" box="[1327,1436,1489,1514]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="immatura">
<emphasis id="B6C4EAAFFFE0297DFAC6A397B4ADFA4F" box="[1327,1436,1489,1514]" italics="true" pageId="6" pageNumber="337">immatura</emphasis>
</taxonomicName>
<location id="816F6066FFE0297DFF7EA3B3B1D8F9AA" LSID="urn:lsid:plazi:treatment:0C1987ABFFE0297DFF7EA367B0A0F81B:816F6066FFE0297DFF7EA3B3B1D8F9AA" box="[151,233,1525,1551]" country="Sri Lanka" name="Kraatz'" pageId="6" pageNumber="337">Kraatz’</location>
(
<collectionCode id="E2A1AE78FFE0297DFF11A3B3B01DF9AA" box="[248,300,1525,1551]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="337" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCAA6536FFE0297DFF2EA05EB0ECF997" box="[199,477,1560,1586]" pageId="6" pageNumber="337" type="distribution">
<paragraph id="840F36BDFFE0297DFF2EA05EB0ECF997" blockId="6.[151,1437,1488,1982]" box="[199,477,1560,1586]" pageId="6" pageNumber="337">
<emphasis id="B6C4EAAFFFE0297DFF2EA05EB050F997" bold="true" box="[199,353,1560,1586]" pageId="6" pageNumber="337">Distribution.</emphasis>
<collectingCountry id="FCA7762DFFE0297DFE8EA05EB0E8F997" box="[359,473,1560,1586]" name="Sri Lanka" pageId="6" pageNumber="337">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CCAA6536FFE0297DFF2EA07BB0A0F81B" pageId="6" pageNumber="337" type="discussion">
<paragraph id="840F36BDFFE0297DFF2EA07BB34FF93B" blockId="6.[151,1437,1488,1982]" pageId="6" pageNumber="337">
<emphasis id="B6C4EAAFFFE0297DFF2EA07BB005F9F2" bold="true" box="[199,308,1597,1623]" pageId="6" pageNumber="337">Remarks</emphasis>
.
<taxonomicName id="43B04D3EFFE0297DFEA8A07AB371F9F3" authorityName="Kraatz" authorityYear="1859" box="[321,576,1596,1622]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="immatura">
<emphasis id="B6C4EAAFFFE0297DFEA8A07AB371F9F3" box="[321,576,1596,1622]" italics="true" pageId="6" pageNumber="337">Gyrophaena immatura</emphasis>
</taxonomicName>
was originally described from “insula
<collectingCountry id="FCA7762DFFE0297DFC06A07AB570F9F3" box="[1007,1089,1596,1622]" name="Sri Lanka" pageId="6" pageNumber="337">Ceylan</collectingCountry>
” based on an unspecified number of
<typeStatus id="5B0B881FFFE0297DFF0BA027B079F9DE" box="[226,328,1633,1659]" pageId="6" pageNumber="337" type="syntype">syntypes</typeStatus>
.
<bibRefCitation id="E0214B4CFFE0297DFEBAA027B33DF9DE" author="Cameron, M." box="[339,524,1633,1659]" pageId="6" pageNumber="337" pagination="1 - 3" refId="ref14209" refString="Cameron, M. (1939) Coleoptera, Staphylinidae. Vol. IV. Parts I &amp; II. In: Sewell, R. B. S. (Ed.), The Fauna of British India, including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xviii + 1 - 691, pls. 1 - 3." type="book chapter" year="1939">Cameron (1939)</bibRefCitation>
redescribed it. During the study of collection of ZIN, the first author found
<specimenCount id="92B6FD34FFE0297DFA84A026B1E9F93B" pageId="6" pageNumber="337" type="male">four males</specimenCount>
possible belonging to the type series.
</paragraph>
<paragraph id="840F36BDFFE0297DFF2EA0EEB086F966" blockId="6.[151,1437,1488,1982]" box="[199,439,1704,1731]" pageId="6" pageNumber="337">
Habitus as in
<figureCitation id="1C8B2A38FFE0297DFE89A0EEB082F966" box="[352,435,1704,1731]" captionStart="FIGURES 22–26" captionStartId="7.[151,264,1076,1100]" captionTargetBox="[168,1418,188,1041]" captionTargetId="figure-19@7.[151,1435,181,1052]" captionTargetPageId="7" captionText="FIGURES 22–26. Gyrophaena immatura: 22–habitus, 23–aedeagus, lateral view, 24–male abdominal tergite VIII, dorsal view, 25–male abdominal sternite VIII, ventral view, 26–labels. Scale bars: 1.0 mm (Fig. 22), 0.10 mm (Figs 23–25)." figureDoi="http://doi.org/10.5281/zenodo.5495638" httpUri="https://zenodo.org/record/5495638/files/figure.png" pageId="6" pageNumber="337">Fig. 22</figureCitation>
.
</paragraph>
<paragraph id="840F36BDFFE0297DFF2EA08AB1F5F83E" blockId="6.[151,1437,1488,1982]" pageId="6" pageNumber="337">
Male. Dorsal plate of aedeagus wide and moderately short, with very long spirally folded flagellum; ventral projection narrower than dorsal plate, bifurcate in apical portion into ventral elongate projection and dorsal projection, curved in apical part (
<figureCitation id="1C8B2A38FFE0297DFE2AA152B326F88A" box="[451,535,1812,1839]" captionStart="FIGURES 22–26" captionStartId="7.[151,264,1076,1100]" captionTargetBox="[168,1418,188,1041]" captionTargetId="figure-19@7.[151,1435,181,1052]" captionTargetPageId="7" captionText="FIGURES 22–26. Gyrophaena immatura: 22–habitus, 23–aedeagus, lateral view, 24–male abdominal tergite VIII, dorsal view, 25–male abdominal sternite VIII, ventral view, 26–labels. Scale bars: 1.0 mm (Fig. 22), 0.10 mm (Figs 23–25)." figureDoi="http://doi.org/10.5281/zenodo.5495638" httpUri="https://zenodo.org/record/5495638/files/figure.png" pageId="6" pageNumber="337">Fig. 23</figureCitation>
). Apical margin of abdominal tergite VIII with three projections: two moderately long projections, curved apically and very short median projection, with moderately wide and superficial concavities between it and each lateral projection (
<figureCitation id="1C8B2A38FFE0297DFD9EA11AB3FDF8D2" box="[631,716,1884,1911]" captionStart="FIGURES 22–26" captionStartId="7.[151,264,1076,1100]" captionTargetBox="[168,1418,188,1041]" captionTargetId="figure-19@7.[151,1435,181,1052]" captionTargetPageId="7" captionText="FIGURES 22–26. Gyrophaena immatura: 22–habitus, 23–aedeagus, lateral view, 24–male abdominal tergite VIII, dorsal view, 25–male abdominal sternite VIII, ventral view, 26–labels. Scale bars: 1.0 mm (Fig. 22), 0.10 mm (Figs 23–25)." figureDoi="http://doi.org/10.5281/zenodo.5495638" httpUri="https://zenodo.org/record/5495638/files/figure.png" pageId="6" pageNumber="337">Fig. 24</figureCitation>
). Apical margin of abdominal sternite VIII widely rounded (
<figureCitation id="1C8B2A38FFE0297DFA86A11AB186F83E" captionStart="FIGURES 22–26" captionStartId="7.[151,264,1076,1100]" captionTargetBox="[168,1418,188,1041]" captionTargetId="figure-19@7.[151,1435,181,1052]" captionTargetPageId="7" captionText="FIGURES 22–26. Gyrophaena immatura: 22–habitus, 23–aedeagus, lateral view, 24–male abdominal tergite VIII, dorsal view, 25–male abdominal sternite VIII, ventral view, 26–labels. Scale bars: 1.0 mm (Fig. 22), 0.10 mm (Figs 23–25)." figureDoi="http://doi.org/10.5281/zenodo.5495638" httpUri="https://zenodo.org/record/5495638/files/figure.png" pageId="6" pageNumber="337">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="840F36BDFFE0297DFF2EA1E2B0A0F81B" blockId="6.[151,1437,1488,1982]" box="[199,401,1956,1982]" pageId="6" pageNumber="337">Female unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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