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<document id="37965D7478149FD1FFFC1F42B76BEE0E" ID-CLB-Dataset="9554" ID-DOI="10.5281/zenodo.4272467" ID-GBIF-Dataset="59fc10fe-e8c3-4537-abb9-049fbda4c7ed" ID-ISSN="0374-1036" ID-Zenodo-Dep="4272467" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1605298451113" checkinUser="felipe" docAuthor="Hoshina, Hideto" docDate="2012" docId="03E982243225074C76954108FBA5A25F" docLanguage="en" docName="ActaEntMusNatPra.52.suppl.1-168.pdf.imf" docOrigin="Acta Entomologica Musei Nationalis Pragae (suppl. 1) 52" docStyle="DocumentStyle:CE2601992B79F4B08764287CD82450D1.3:ActaEntMusNatPra.2010-2016.monograph" docStyleId="CE2601992B79F4B08764287CD82450D1" docStyleName="ActaEntMusNatPra.2010-2016.monograph" docStyleVersion="3" docTitle="Leiodes circinipes" docType="treatment" docVersion="11" lastPageNumber="30" masterDocId="FFD0FA5C3200076977334233FFA5A45C" masterDocTitle="Review of the tribes Sogdini and Leiodini from Japan and North Chishima Islands. Part II. Genera Hydnobius and Leiodes (Coleoptera: Leiodidae)" masterLastPageNumber="168" masterPageNumber="1" pageNumber="30" updateTime="1698846259177" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="1E5CED3A38868385A1491FA6635A9B65" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="08BCF8783F019968EDA0368D23DF4A0D">Review of the tribes Sogdini and Leiodini from Japan and North Chishima Islands. Part II. Genera Hydnobius and Leiodes (Coleoptera: Leiodidae)</mods:title>
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<mods:name id="FA88B53E616022C63A9CD8736833C6CE" type="personal">
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<mods:namePart id="5E1D4A9CED50FA6FB0F0B006A11D3593">Hoshina, Hideto</mods:namePart>
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<mods:title id="9DE9424A3419099A295065C57F25D1CE">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="BB36731B1B368DDDE8E1DC6AC954622F">2012</mods:date>
<mods:detail id="128C92748FED0B0D77273A52FECBCF8E" type="series">
<mods:title id="93653935130E2D7DC28F8701120E57C5">suppl. 1</mods:title>
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<mods:number id="46353D0E0089B29C98825ABBF5C0EE66">2012-09-15</mods:number>
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<mods:detail id="BCFF56CA7C617CE96A61FD67EA4C26A2" type="volume">
<mods:number id="7628EC77CA42F78E90AF1E2C0BBBDA60">52</mods:number>
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<mods:classification id="9BAA104D0FC052B6500CDEDA31178C20">journal article</mods:classification>
<mods:identifier id="DF2EAD77A372ECF2226A62CB8EDFA478" type="CLB-Dataset">9554</mods:identifier>
<mods:identifier id="61C2DA6B56EF4BE3EC0CDF3FBE46693A" type="DOI">10.5281/zenodo.4272467</mods:identifier>
<mods:identifier id="D46938E84FD3C6275BF2BEB45E04A875" type="GBIF-Dataset">59fc10fe-e8c3-4537-abb9-049fbda4c7ed</mods:identifier>
<mods:identifier id="B5CBECDC1A2962DFC2868CA3344FC5CE" type="ISSN">0374-1036</mods:identifier>
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<treatment id="03E982243225074C76954108FBA5A25F" ID-DOI="http://doi.org/10.5281/zenodo.4339467" ID-GBIF-Taxon="170447931" ID-Zenodo-Dep="4339467" LSID="urn:lsid:plazi:treatment:03E982243225074C76954108FBA5A25F" httpUri="http://treatment.plazi.org/id/03E982243225074C76954108FBA5A25F" lastPageNumber="30" pageId="37" pageNumber="30">
<subSubSection id="C35A60B93225074C76954108FCA9A708" box="[422,780,827,852]" pageId="37" pageNumber="30" type="nomenclature">
<paragraph id="8BFF33323225074C76954108FCA9A708" blockId="37.[422,780,827,852]" box="[422,780,827,852]" pageId="37" pageNumber="30">
<heading id="D0B7845E3225074C76954108FCA9A708" bold="true" box="[422,780,827,852]" centered="true" fontSize="10" level="2" pageId="37" pageNumber="30" reason="2">
<emphasis id="B934EF203225074C76954108FCA9A708" bold="true" box="[422,780,827,852]" pageId="37" pageNumber="30">
<taxonomicName id="4C4048B13225074C76954108FDC9A708" ID-CoL="3SYC7" baseAuthorityName="Rye" baseAuthorityYear="1873" box="[422,620,827,852]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="30" phylum="Arthropoda" rank="species" species="circinipes">
<emphasis id="B934EF203225074C76954108FDC9A708" bold="true" box="[422,620,827,852]" italics="true" pageId="37" pageNumber="30">Leiodes circinipes</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35A60B93225074C77424158FE92A7FF" pageId="37" pageNumber="30" type="discussion">
<paragraph id="8BFF33323225074C77424158FE92A7FF" blockId="37.[113,1089,875,1539]" pageId="37" pageNumber="30">
<emphasis id="B934EF203225074C77424158FE90A7DF" bold="true" box="[113,309,875,899]" pageId="37" pageNumber="30">Species included.</emphasis>
<taxonomicName id="4C4048B13225074C760E415FFD2FA7DF" authority="(Rye, 1873)" baseAuthorityName="Rye" baseAuthorityYear="1873" box="[317,650,875,899]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="30" phylum="Arthropoda" rank="species" species="circinipes">
<emphasis id="B934EF203225074C760E415FFDA5A7DF" box="[317,512,876,899]" italics="true" pageId="37" pageNumber="30">Leiodes circinipes</emphasis>
(
<bibRefCitation id="EFD14EC33225074C753C4158FD27A7DF" author="RYE E. C." box="[527,642,875,899]" pageId="37" pageNumber="30" pagination="131 - 135" refId="ref74051" refString="RYE E. C. 1873: Notes on Anisotomidae, with description of three new species (two from Japan, and one from Gt. Britain [sic!]). No. 1. Entomologist's Monthly Magazine 10: 131 - 135." type="journal article" year="1873">Rye, 1873</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C4048B13225074C75A5415FFD42A7DF" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[662,743,876,899]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="30" phylum="Arthropoda" rank="species" species="juzoi" status="sp. nov.">
<emphasis id="B934EF203225074C75A5415FFD42A7DF" box="[662,743,876,899]" italics="true" pageId="37" pageNumber="30">L. juzoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207525B3225074C75DC415FFCE0A7DF" box="[751,837,876,899]" pageId="37" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C4048B13225074C7462415FFC64A7DF" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[849,961,876,899]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="30" phylum="Arthropoda" rank="species" species="yasudai" status="sp. nov.">
<emphasis id="B934EF203225074C7462415FFC64A7DF" box="[849,961,876,899]" italics="true" pageId="37" pageNumber="30">L. yasudai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207525B3225074C74FA415FFBBAA7DF" box="[969,1055,876,899]" pageId="37" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C4048B13225074C7318415FFF78A7FF" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="30" phylum="Arthropoda" rank="species" species="yoshitakei" status="sp. nov.">
<emphasis id="B934EF203225074C7318415FFF78A7FF" italics="true" pageId="37" pageNumber="30">L. yoshitakei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207525B3225074C77D741BFFE92A7FF" box="[228,311,908,931]" pageId="37" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C35A60B93225074C77424198FBA5A25F" pageId="37" pageNumber="30" type="diagnosis">
<paragraph id="8BFF33323225074C77424198FE19A17F" blockId="37.[113,1089,875,1539]" pageId="37" pageNumber="30">
<emphasis id="B934EF203225074C77424198FF43A79F" bold="true" box="[113,230,939,963]" pageId="37" pageNumber="30">Diagnosis.</emphasis>
Elytra unicolor or bicolored, with densely arranged large punctures between rows of punctures, and therefore superficially appearing not to bear rows of punctures (
<figureCitation id="137B2FB73225074C74E041F8FB99A7BF" box="[979,1084,971,995]" captionStart="Fig" captionStartId="38.[151,182,1544,1563]" captionTargetBox="[251,1026,769,1525]" captionTargetId="figure-252@38.[244,1031,760,1529]" captionTargetPageId="38" captionText="Fig. 16. Leiodes circinipes (Rye, 1873).A – body, dorsal view; B – ditto, lateral view; C – dorsal color; D – antenna; E – elytral punctures; F – mesoventrite, lateral view. Scale I: 1 mm for A and B; II: 0.5 mm for D; III: 0.5 mm for F." figureDoi="http://doi.org/10.5281/zenodo.4272501" httpUri="https://zenodo.org/record/4272501/files/figure.png" pageId="37" pageNumber="30">Figs. 16E</figureCitation>
,
<figureCitation id="137B2FB73225074C774241D8FF3BA05F" box="[113,158,1003,1027]" captionStart="Fig" captionStartId="42.[151,182,1518,1537]" captionTargetBox="[209,1078,504,1492]" captionTargetId="figure-181@42.[199,1079,490,1493]" captionTargetPageId="42" captionText="Fig. 19. Leiodes juzoi sp. nov.A – body, dorsal view; B – ditto, lateral view; C – dorsal color; D – antenna; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for D; III: 0.5 mm for F; IV: 0.5 mm for G." figureDoi="http://doi.org/10.5281/zenodo.4272507" httpUri="https://zenodo.org/record/4272507/files/figure.png" pageId="37" pageNumber="30">19E</figureCitation>
,
<figureCitation id="137B2FB73225074C779A41D8FF7CA05F" box="[169,217,1003,1027]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="37" pageNumber="30">21D</figureCitation>
,
<figureCitation id="137B2FB73225074C77D741D8FEB6A05F" box="[228,275,1003,1027]" captionStart="Fig" captionStartId="49.[113,144,1516,1535]" captionTargetBox="[222,978,550,1490]" captionTargetId="figure-135@49.[218,983,542,1502]" captionTargetPageId="49" captionText="Fig. 24. Leiodes yoshitakei sp. nov. A – body, dorsal view; B – ditto, lateral view; C – dorsal color; D – antenna; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for D; III: 0.5 mm for F; IV: 1 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272517" httpUri="https://zenodo.org/record/4272517/files/figure.png" pageId="37" pageNumber="30">24E</figureCitation>
); mesoventrite without distinct excavation between median carina and transverse carina (
<figureCitation id="137B2FB73225074C77CE4638FEC7A07F" box="[253,354,1035,1059]" captionStart="Fig" captionStartId="38.[151,182,1544,1563]" captionTargetBox="[251,1026,769,1525]" captionTargetId="figure-252@38.[244,1031,760,1529]" captionTargetPageId="38" captionText="Fig. 16. Leiodes circinipes (Rye, 1873).A – body, dorsal view; B – ditto, lateral view; C – dorsal color; D – antenna; E – elytral punctures; F – mesoventrite, lateral view. Scale I: 1 mm for A and B; II: 0.5 mm for D; III: 0.5 mm for F." figureDoi="http://doi.org/10.5281/zenodo.4272501" httpUri="https://zenodo.org/record/4272501/files/figure.png" pageId="37" pageNumber="30">Figs. 16F</figureCitation>
,
<figureCitation id="137B2FB73225074C76584638FE30A07F" box="[363,405,1035,1059]" captionStart="Fig" captionStartId="42.[151,182,1518,1537]" captionTargetBox="[209,1078,504,1492]" captionTargetId="figure-181@42.[199,1079,490,1493]" captionTargetPageId="42" captionText="Fig. 19. Leiodes juzoi sp. nov.A – body, dorsal view; B – ditto, lateral view; C – dorsal color; D – antenna; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for D; III: 0.5 mm for F; IV: 0.5 mm for G." figureDoi="http://doi.org/10.5281/zenodo.4272507" httpUri="https://zenodo.org/record/4272507/files/figure.png" pageId="37" pageNumber="30">19F</figureCitation>
,
<figureCitation id="137B2FB73225074C76AC4638FE69A07F" box="[415,460,1035,1059]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="37" pageNumber="30">21E</figureCitation>
,
<figureCitation id="137B2FB73225074C76E54638FDA6A07F" box="[470,515,1035,1059]" captionStart="Fig" captionStartId="49.[113,144,1516,1535]" captionTargetBox="[222,978,550,1490]" captionTargetId="figure-135@49.[218,983,542,1502]" captionTargetPageId="49" captionText="Fig. 24. Leiodes yoshitakei sp. nov. A – body, dorsal view; B – ditto, lateral view; C – dorsal color; D – antenna; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for D; III: 0.5 mm for F; IV: 1 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272517" httpUri="https://zenodo.org/record/4272517/files/figure.png" pageId="37" pageNumber="30">24F</figureCitation>
); protarsi sexually dimorphic, male protarsomeres 2–4 expanded (
<figureCitation id="137B2FB73225074C77D04618FEE9A01F" box="[227,332,1067,1091]" captionStart="Fig" captionStartId="39.[113,144,1452,1471]" captionTargetBox="[119,1082,224,1375]" captionTargetId="figure-15@39.[113,1088,219,1384]" captionTargetPageId="39" captionText="Fig. 17. Leiodes circinipes (Rye, 1873).A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – ditto; G – female hind leg, ventral view; H – male metafemur, dorsal view; I – female metafemur, dorsal view; J – male protibiae, dorsal view; K – female protibiae, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E; III: 0.5 mm for F and G; IV: 0.2 mm for H and I; V: 0.2 mm for J and K." figureDoi="http://doi.org/10.5281/zenodo.4272503" httpUri="https://zenodo.org/record/4272503/files/figure.png" pageId="37" pageNumber="30">Figs. 17A</figureCitation>
,
<figureCitation id="137B2FB73225074C76664618FE20A01F" box="[341,389,1067,1091]" captionStart="Fig" captionStartId="43.[113,144,1486,1505]" captionTargetBox="[159,1040,441,1464]" captionTargetId="figure-15@43.[154,1048,418,1482]" captionTargetPageId="43" captionText="Fig. 20. Leiodes juzoi sp. nov. A – male fore leg, ventral view; B – male middle leg, ventral view; C – male hind leg, ventral view; D – male protibia, dorsal view; E – male metafemur, dorsal view; F – aedeagus, dorsal view; G – ditto, lateral view; H – male abdominal sternite 8. Scale I: 0.5 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for D; IV: 0.2 mm for E; V: 0.5 mm for F and G; VI: 0.2 mm for H." figureDoi="http://doi.org/10.5281/zenodo.4272509" httpUri="https://zenodo.org/record/4272509/files/figure.png" pageId="37" pageNumber="30">20A</figureCitation>
,
<figureCitation id="137B2FB73225074C76BE4618FE18A01F" box="[397,445,1067,1091]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="37" pageNumber="30">22A</figureCitation>
,
<figureCitation id="137B2FB73225074C76F54618FE52A01F" box="[454,503,1067,1091]" captionStart="Fig" captionStartId="50.[151,182,1463,1482]" captionTargetBox="[217,1059,677,1455]" captionTargetId="figure-167@50.[213,1064,665,1455]" captionTargetPageId="50" captionText="Fig. 25. Leiodes yoshitakei sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272519" httpUri="https://zenodo.org/record/4272519/files/figure.png" pageId="37" pageNumber="30">25A</figureCitation>
); male mesotibiae strongly protuberant at an interoapical corner, female mesotibiae simply quadrate interoapically (
<figureCitation id="137B2FB73225074C75D04678FC2EA03F" box="[739,907,1099,1123]" captionStart="Fig" captionStartId="39.[113,144,1452,1471]" captionTargetBox="[119,1082,224,1375]" captionTargetId="figure-15@39.[113,1088,219,1384]" captionTargetPageId="39" captionText="Fig. 17. Leiodes circinipes (Rye, 1873).A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – ditto; G – female hind leg, ventral view; H – male metafemur, dorsal view; I – female metafemur, dorsal view; J – male protibiae, dorsal view; K – female protibiae, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E; III: 0.5 mm for F and G; IV: 0.2 mm for H and I; V: 0.2 mm for J and K." figureDoi="http://doi.org/10.5281/zenodo.4272503" httpUri="https://zenodo.org/record/4272503/files/figure.png" pageId="37" pageNumber="30">Figs. 17C, 17D</figureCitation>
,
<figureCitation id="137B2FB73225074C74A44678FBA7A03F" box="[919,1026,1099,1123]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="37" pageNumber="30">22C, 22D</figureCitation>
,
<figureCitation id="137B2FB73225074C733D4678FF06A0DF" captionStart="Fig" captionStartId="50.[151,182,1463,1482]" captionTargetBox="[217,1059,677,1455]" captionTargetId="figure-167@50.[213,1064,665,1455]" captionTargetPageId="50" captionText="Fig. 25. Leiodes yoshitakei sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272519" httpUri="https://zenodo.org/record/4272519/files/figure.png" pageId="37" pageNumber="30">25C, 25D</figureCitation>
); dorsal posteroapical projection of metafemur sexually dimorphic (
<figureCitation id="137B2FB73225074C745B4658FC5DA0DF" box="[872,1016,1131,1155]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="37" pageNumber="30">Figs. 22I, 22J</figureCitation>
) or not (
<figureCitation id="137B2FB73225074C774A46B8FEBDA0FF" box="[121,280,1163,1187]" captionStart="Fig" captionStartId="39.[113,144,1452,1471]" captionTargetBox="[119,1082,224,1375]" captionTargetId="figure-15@39.[113,1088,219,1384]" captionTargetPageId="39" captionText="Fig. 17. Leiodes circinipes (Rye, 1873).A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – ditto; G – female hind leg, ventral view; H – male metafemur, dorsal view; I – female metafemur, dorsal view; J – male protibiae, dorsal view; K – female protibiae, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E; III: 0.5 mm for F and G; IV: 0.2 mm for H and I; V: 0.2 mm for J and K." figureDoi="http://doi.org/10.5281/zenodo.4272503" httpUri="https://zenodo.org/record/4272503/files/figure.png" pageId="37" pageNumber="30">Figs. 17H, 17I</figureCitation>
,
<figureCitation id="137B2FB73225074C761046B8FEDBA0FF" box="[291,382,1163,1187]" captionStart="Fig" captionStartId="50.[151,182,1463,1482]" captionTargetBox="[217,1059,677,1455]" captionTargetId="figure-167@50.[213,1064,665,1455]" captionTargetPageId="50" captionText="Fig. 25. Leiodes yoshitakei sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272519" httpUri="https://zenodo.org/record/4272519/files/figure.png" pageId="37" pageNumber="30">25I, 25J</figureCitation>
); pubescence of metaventrite showing distinct sexual dimorphism (
<figureCitation id="137B2FB73225074C774A4698FEBCA09F" box="[121,281,1195,1219]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="37" pageNumber="30">Figs. 21F, 21G</figureCitation>
,
<figureCitation id="137B2FB73225074C76174698FE2AA09F" box="[292,399,1195,1219]" captionStart="Fig" captionStartId="49.[113,144,1516,1535]" captionTargetBox="[222,978,550,1490]" captionTargetId="figure-135@49.[218,983,542,1502]" captionTargetPageId="49" captionText="Fig. 24. Leiodes yoshitakei sp. nov. A – body, dorsal view; B – ditto, lateral view; C – dorsal color; D – antenna; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for D; III: 0.5 mm for F; IV: 1 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272517" httpUri="https://zenodo.org/record/4272517/files/figure.png" pageId="37" pageNumber="30">24G, 24H</figureCitation>
) or not; aedeagus slender, without distinct large sclerites in inner sac (
<figureCitation id="137B2FB73225074C779246F8FEA8A0BF" box="[161,269,1227,1251]" captionStart="Fig" captionStartId="40.[151,182,1507,1526]" captionTargetBox="[152,1117,792,1470]" captionTargetId="figure-232@40.[151,1126,778,1480]" captionTargetPageId="40" captionText="Fig. 18. Leiodes circinipes (Rye, 1873). A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272505" httpUri="https://zenodo.org/record/4272505/files/figure.png" pageId="37" pageNumber="30">Figs. 18A</figureCitation>
,
<figureCitation id="137B2FB73225074C762B46F8FEE4A0BF" box="[280,321,1227,1251]" captionStart="Fig" captionStartId="42.[151,182,1518,1537]" captionTargetBox="[209,1078,504,1492]" captionTargetId="figure-181@42.[199,1079,490,1493]" captionTargetPageId="42" captionText="Fig. 19. Leiodes juzoi sp. nov.A – body, dorsal view; B – ditto, lateral view; C – dorsal color; D – antenna; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for D; III: 0.5 mm for F; IV: 0.5 mm for G." figureDoi="http://doi.org/10.5281/zenodo.4272507" httpUri="https://zenodo.org/record/4272507/files/figure.png" pageId="37" pageNumber="30">19F</figureCitation>
,
<figureCitation id="137B2FB73225074C767F46F8FED9A0BF" box="[332,380,1227,1251]" captionStart="Fig" captionStartId="47.[113,144,1191,1210]" captionTargetBox="[136,1062,228,1156]" captionTargetId="figure-106@47.[121,1080,219,1171]" captionTargetPageId="47" captionText="Fig. 23. Leiodes yasudai sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272515" httpUri="https://zenodo.org/record/4272515/files/figure.png" pageId="37" pageNumber="30">23A</figureCitation>
,
<figureCitation id="137B2FB73225074C76B546F8FE1DA0BF" box="[390,440,1227,1251]" captionStart="Fig" captionStartId="51.[113,144,1537,1556]" captionTargetBox="[213,968,890,1530]" captionTargetId="figure-271@51.[209,994,881,1539]" captionTargetPageId="51" captionText="Fig. 26. Leiodes yoshitakei sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272521" httpUri="https://zenodo.org/record/4272521/files/figure.png" pageId="37" pageNumber="30">26A</figureCitation>
); male abdominal sternite 8 strongly curved (
<figureCitation id="137B2FB73225074C74A446F8FBA7A0BF" box="[919,1026,1227,1251]" captionStart="Fig" captionStartId="40.[151,182,1507,1526]" captionTargetBox="[152,1117,792,1470]" captionTargetId="figure-232@40.[151,1126,778,1480]" captionTargetPageId="40" captionText="Fig. 18. Leiodes circinipes (Rye, 1873). A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272505" httpUri="https://zenodo.org/record/4272505/files/figure.png" pageId="37" pageNumber="30">Figs. 18C</figureCitation>
,
<figureCitation id="137B2FB73225074C733F46F8FB99A0BF" box="[1036,1084,1227,1251]" captionStart="Fig" captionStartId="43.[113,144,1486,1505]" captionTargetBox="[159,1040,441,1464]" captionTargetId="figure-15@43.[154,1048,418,1482]" captionTargetPageId="43" captionText="Fig. 20. Leiodes juzoi sp. nov. A – male fore leg, ventral view; B – male middle leg, ventral view; C – male hind leg, ventral view; D – male protibia, dorsal view; E – male metafemur, dorsal view; F – aedeagus, dorsal view; G – ditto, lateral view; H – male abdominal sternite 8. Scale I: 0.5 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for D; IV: 0.2 mm for E; V: 0.5 mm for F and G; VI: 0.2 mm for H." figureDoi="http://doi.org/10.5281/zenodo.4272509" httpUri="https://zenodo.org/record/4272509/files/figure.png" pageId="37" pageNumber="30">20H</figureCitation>
,
<figureCitation id="137B2FB73225074C774246D8FF05A15F" box="[113,160,1259,1283]" captionStart="Fig" captionStartId="47.[113,144,1191,1210]" captionTargetBox="[136,1062,228,1156]" captionTargetId="figure-106@47.[121,1080,219,1171]" captionTargetPageId="47" captionText="Fig. 23. Leiodes yasudai sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272515" httpUri="https://zenodo.org/record/4272515/files/figure.png" pageId="37" pageNumber="30">23C</figureCitation>
,
<figureCitation id="137B2FB73225074C779946D8FF7FA15F" box="[170,218,1259,1283]" captionStart="Fig" captionStartId="51.[113,144,1537,1556]" captionTargetBox="[213,968,890,1530]" captionTargetId="figure-271@51.[209,994,881,1539]" captionTargetPageId="51" captionText="Fig. 26. Leiodes yoshitakei sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272521" httpUri="https://zenodo.org/record/4272521/files/figure.png" pageId="37" pageNumber="30">26C</figureCitation>
); female abdominal sternite 8 with a spiculum ventrale at a central point of anterior margin (
<figureCitation id="137B2FB73225074C77F84738FE92A17F" box="[203,311,1291,1315]" captionStart="Fig" captionStartId="40.[151,182,1507,1526]" captionTargetBox="[152,1117,792,1470]" captionTargetId="figure-232@40.[151,1126,778,1480]" captionTargetPageId="40" captionText="Fig. 18. Leiodes circinipes (Rye, 1873). A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272505" httpUri="https://zenodo.org/record/4272505/files/figure.png" pageId="37" pageNumber="30">Figs. 18D</figureCitation>
,
<figureCitation id="137B2FB73225074C76714738FED6A17F" box="[322,371,1291,1315]" captionStart="Fig" captionStartId="47.[113,144,1191,1210]" captionTargetBox="[136,1062,228,1156]" captionTargetId="figure-106@47.[121,1080,219,1171]" captionTargetPageId="47" captionText="Fig. 23. Leiodes yasudai sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272515" httpUri="https://zenodo.org/record/4272515/files/figure.png" pageId="37" pageNumber="30">23D</figureCitation>
,
<figureCitation id="137B2FB73225074C764D4738FE15A17F" box="[382,432,1291,1315]" captionStart="Fig" captionStartId="51.[113,144,1537,1556]" captionTargetBox="[213,968,890,1530]" captionTargetId="figure-271@51.[209,994,881,1539]" captionTargetPageId="51" captionText="Fig. 26. Leiodes yoshitakei sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272521" httpUri="https://zenodo.org/record/4272521/files/figure.png" pageId="37" pageNumber="30">26D</figureCitation>
).
</paragraph>
<paragraph id="8BFF33323225074C77A34718FEE4A1FF" blockId="37.[113,1089,875,1539]" pageId="37" pageNumber="30">
Representatives of the genus
<taxonomicName id="4C4048B13225074C76FA471FFDB9A11F" authorityName=": HATCH" authorityYear="1929" box="[457,540,1324,1347]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B934EF203225074C76FA471FFDB9A11F" box="[457,540,1324,1347]" italics="true" pageId="37" pageNumber="30">Leiodes</emphasis>
</taxonomicName>
occasionally show sexual dimorphism in the shape of the spines at the interoapical corner of the mesotibiae (
<bibRefCitation id="EFD14EC33225074C75F54778FC2CA13F" author="BARANOWSKI R." box="[710,905,1355,1379]" pageId="37" pageNumber="30" pagination="1 - 149" refId="ref71522" refString="BARANOWSKI R. 1993: Revision of the genus Leiodes Latreille of North and Central America (Coleoptera: Leiodidae). Entomologica Scandinavica 42 (supplement): 1 - 149." type="journal article" year="1993">BARANOWSKI 1993</bibRefCitation>
). Among
<taxonomicName id="4C4048B13225074C74DD477FFBE4A13F" authorityName=": HATCH" authorityYear="1929" box="[1006,1089,1356,1379]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B934EF203225074C74DD477FFBE4A13F" box="[1006,1089,1356,1379]" italics="true" pageId="37" pageNumber="30">Leiodes</emphasis>
</taxonomicName>
from
<collectingCountry id="F35773A23225074C7798475FFF4DA1DF" box="[171,232,1388,1411]" name="Japan" pageId="37" pageNumber="30">Japan</collectingCountry>
and the North Chishima Islands,
<taxonomicName id="4C4048B13225074C757B475FFD6EA1DF" baseAuthorityName="Rye" baseAuthorityYear="1873" box="[584,715,1388,1411]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="30" phylum="Arthropoda" rank="species" species="circinipes">
<emphasis id="B934EF203225074C757B475FFD6EA1DF" box="[584,715,1388,1411]" italics="true" pageId="37" pageNumber="30">L. circinipes</emphasis>
</taxonomicName>
group is the only group having this character (
<tableCitation id="C6C206893225074C77D247B8FE94A1FF" box="[225,305,1419,1443]" captionStart="Table 1" captionStartId="19.[55,102,450,470]" captionTargetId="graphics-27@19.[54,1469,483,1415]" captionTargetPageId="19" captionText="Table 1. List of important morphological features for identification of species" httpUri="http://table.plazi.org/id/DF3F63BA3213077A770443F1FD6EA58A" pageId="37" pageNumber="30" tableUuid="DF3F63BA3213077A770443F1FD6EA58A">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BFF33323225074C77424798FBA5A25F" blockId="37.[113,1089,875,1539]" pageId="37" pageNumber="30">
<emphasis id="B934EF203225074C77424798FECBA19F" bold="true" box="[113,366,1451,1475]" pageId="37" pageNumber="30">Differential diagnosis.</emphasis>
The
<taxonomicName id="4C4048B13225074C7694479FFDCEA19F" baseAuthorityName="Rye" baseAuthorityYear="1873" box="[423,619,1452,1475]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="30" phylum="Arthropoda" rank="species" species="circinipes">
<emphasis id="B934EF203225074C7694479FFDCEA19F" box="[423,619,1452,1475]" italics="true" pageId="37" pageNumber="30">Leiodes circinipes</emphasis>
</taxonomicName>
species group can be separated from other species groups by having elytra with densely arranged large punctures between rows of punctures and having the male mesotibiae strongly protuberant at the interoapical corner.
</paragraph>
</subSubSection>
</treatment>
</document>
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