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<mods:title id="903F7BD98DFD7DB4B02CCCF9BAD505F9">A new phylogenetic classification for the gymnophthalmid genera Cercosaura, Pantodactylus and Prionodactylus (Reptilia: Squamata)</mods:title>
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<heading id="392481E3FF84FFB1FC97FCA4463AFC64" box="[824,1404,891,915]" centered="true" fontSize="9" level="2" pageId="6" pageNumber="107" reason="2">
<taxonomicName id="A5D34D0CFF84FFB1FC97FCA4463AFC64" authority="(O'SHAUGHNESSY)" baseAuthorityName="O'SHAUGHNESSY" box="[824,1404,891,915]" class="Reptilia" family="Gymnophthalmidae" genus="Prionodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="50A7EA9DFF84FFB1FC97FCA447E2FC66" box="[824,1188,891,914]" italics="true" pageId="6" pageNumber="107">PRIONODACTYLUS VERTEBRALIS</emphasis>
(OSHAUGHNESSY)
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<subSubSection id="2AC96504FF84FFB1FC89FC7B4670F94F" pageId="6" pageNumber="107" type="reference_group">
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<treatmentCitation id="E372109EFF84FFB1FC89FC7B40F7FC20" author="O'Shaughnessy AWE" page="298" pageId="6" pageNumber="107" year="1879">
<taxonomicName id="A5D34D0CFF84FFB1FC89FC7B40F7FC20" authority="O'Shaughnessy, 1879: 298" authorityName="O'Shaughnessy" authorityPageNumber="298" authorityYear="1879" class="Reptilia" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="vertebralis" subGenus="Pantodactylus">
<emphasis id="50A7EA9DFF84FFB1FC89FC7B40E0FC4E" box="[806,934,932,953]" italics="true" pageId="6" pageNumber="107">Cercosaura</emphasis>
(
<emphasis id="50A7EA9DFF84FFB1FC1DFC7C4715FC4F" box="[946,1107,931,952]" italics="true" pageId="6" pageNumber="107">Pantodactylus</emphasis>
)
<emphasis id="50A7EA9DFF84FFB1FBF0FC7C479EFC4F" box="[1119,1240,931,952]" italics="true" pageId="6" pageNumber="107">vertebralis</emphasis>
<bibRefCitation id="06424B7EFF84FFB1FB74FC7C40F7FC20" author="O'Shaughnessy AWE" pageId="6" pageNumber="107" pagination="295 - 303" refId="ref9083" refString="O'Shaughnessy AWE. 1879. Descriptions of new species of lizards in the collection of the British Museum. Annals and Magazine of Natural History. Series 5 4: 295 - 303." type="journal article" year="1879">OShaughnessy, 1879: 298</bibRefCitation>
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.
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</paragraph>
<footnote id="01C82A81FF84FFB1FD9BFC624197FC3D" box="[564,721,957,970]" pageId="6" pageNumber="107">
<paragraph id="626C368FFF84FFB1FD9BFC624197FC3D" blockId="6.[564,754,893,1611]" box="[564,721,957,970]" pageId="6" pageNumber="107">(O) Madre de Dios, Peru</paragraph>
</footnote>
<footnote id="01C82A81FF84FFB1FD9BFC0241EBFC1D" box="[564,685,989,1002]" pageId="6" pageNumber="107">
<paragraph id="626C368FFF84FFB1FD9BFC0241EBFC1D" blockId="6.[564,754,893,1611]" box="[564,685,989,1002]" pageId="6" pageNumber="107">(O) Napo, Ecuador</paragraph>
</footnote>
<paragraph id="626C368FFF84FFB1FC89FC3E461EFC01" blockId="6.[806,1422,931,1720]" box="[806,1368,993,1015]" pageId="6" pageNumber="107">
<treatmentCitationGroup id="42C311A1FF84FFB1FC89FC3E461EFC01" box="[806,1368,993,1015]" pageId="6" pageNumber="107">
<treatmentCitation id="E372109EFF84FFB1FC89FC3E4612FC01" author="Boulenger GA" box="[806,1364,993,1015]" page="394" pageId="6" pageNumber="107" year="1885">
<taxonomicName id="A5D34D0CFF84FFB1FC89FC3E4612FC01" authority=": Boulenger, 1885: 394" authorityName=": Boulenger" authorityPageNumber="394" authorityYear="1885" box="[806,1364,993,1015]" class="Reptilia" family="Gymnophthalmidae" genus="Prionodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="50A7EA9DFF84FFB1FC89FC3E4714FC01" box="[806,1106,993,1014]" italics="true" pageId="6" pageNumber="107">Prionodactylus vertebralis</emphasis>
:
<bibRefCitation id="06424B7EFF84FFB1FBCEFC3E4612FC01" author="Boulenger GA" box="[1121,1364,993,1015]" pageId="6" pageNumber="107" refId="ref8308" refString="Boulenger GA. 1885. Catalogue of the Lizards in the British Museum (Natural History) I - III. London: British Natural History Museum." type="book" year="1885">Boulenger, 1885: 394</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<footnote id="01C82A81FF84FFB1FD9BFC2141F4FBFC" box="[564,690,1022,1035]" pageId="6" pageNumber="107">
<paragraph id="626C368FFF84FFB1FD9BFC2141F4FBFC" blockId="6.[564,754,893,1611]" box="[564,690,1022,1035]" pageId="6" pageNumber="107">(O) Meta, Colombia</paragraph>
</footnote>
<paragraph id="626C368FFF84FFB1FC89FC204674FBE2" blockId="6.[806,1422,931,1720]" box="[806,1330,1023,1045]" pageId="6" pageNumber="107">
<treatmentCitationGroup id="42C311A1FF84FFB1FC89FC204674FBE2" box="[806,1330,1023,1045]" pageId="6" pageNumber="107">
<treatmentCitation id="E372109EFF84FFB1FC89FC204668FBE2" author="Boulenger GA" box="[806,1326,1023,1045]" page="518" pageId="6" pageNumber="107" year="1908">
<taxonomicName id="A5D34D0CFF84FFB1FC89FC204668FBE2" authority="Boulenger, 1908: 518" authorityName="Boulenger" authorityPageNumber="518" authorityYear="1908" box="[806,1326,1023,1045]" class="Reptilia" family="Gymnophthalmidae" genus="Prionodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="palmeri">
<emphasis id="50A7EA9DFF84FFB1FC89FC204774FBE3" box="[806,1074,1023,1044]" italics="true" pageId="6" pageNumber="107">Prionodactylus palmeri</emphasis>
<bibRefCitation id="06424B7EFF84FFB1FB94FC204668FBE2" author="Boulenger GA" box="[1083,1326,1023,1045]" pageId="6" pageNumber="107" pagination="515 - 522" refId="ref8336" refString="Boulenger GA. 1908. Descriptions of new batrachians and reptiles discovered by Mr M. G. Palmer in south-western Colombia. Annals and Magazine of Natural History, Series 8 2: 515 - 522." type="journal article" year="1908">Boulenger, 1908: 518</bibRefCitation>
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.
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</paragraph>
<footnote id="01C82A81FF84FFB1FD9BFBC141F4FBDC" box="[564,690,1054,1067]" pageId="6" pageNumber="107">
<paragraph id="626C368FFF84FFB1FD9BFBC141F4FBDC" blockId="6.[564,754,893,1611]" box="[564,690,1054,1067]" pageId="6" pageNumber="107">(O) Meta, Colombia</paragraph>
</footnote>
<paragraph id="626C368FFF84FFB1FC89FBC1465AFBC4" blockId="6.[806,1422,931,1720]" box="[806,1308,1054,1076]" pageId="6" pageNumber="107">
<treatmentCitationGroup id="42C311A1FF84FFB1FC89FBC1465AFBC4" box="[806,1308,1054,1076]" pageId="6" pageNumber="107">
<treatmentCitation id="E372109EFF84FFB1FC89FBC14650FBC4" author="Ruthven AG" box="[806,1302,1054,1076]" page="1" pageId="6" pageNumber="107" year="1921">
<taxonomicName id="A5D34D0CFF84FFB1FC89FBC14650FBC4" authority="Ruthven, 1921: 1" authorityName="Ruthven" authorityPageNumber="1" authorityYear="1921" box="[806,1302,1054,1076]" class="Reptilia" family="Gymnophthalmidae" genus="Prionodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="marianus">
<emphasis id="50A7EA9DFF84FFB1FC89FBC1470EFBC3" box="[806,1096,1054,1076]" italics="true" pageId="6" pageNumber="107">Prionodactylus marianus</emphasis>
<bibRefCitation id="06424B7EFF84FFB1FBFEFBC14650FBC4" author="Ruthven AG" box="[1105,1302,1054,1076]" pageId="6" pageNumber="107" pagination="1 - 4" refId="ref9409" refString="Ruthven AG. 1921. Description of an apparently new lizard from Colombia. Occasional Papers of the Museum of Zoology University of Michigan 103: 1 - 4." type="journal article" year="1921">Ruthven, 1921: 1</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<footnote id="01C82A81FF84FFB1FD9BFBE241F4FBBD" box="[564,690,1085,1098]" pageId="6" pageNumber="107">
<paragraph id="626C368FFF84FFB1FD9BFBE241F4FBBD" blockId="6.[564,754,893,1611]" box="[564,690,1085,1098]" pageId="6" pageNumber="107">(A) Meta, Colombia</paragraph>
</footnote>
<paragraph id="626C368FFF84FFB1FC89FBE247B8FBA5" blockId="6.[806,1422,931,1720]" box="[806,1278,1085,1107]" pageId="6" pageNumber="107">
<treatmentCitationGroup id="42C311A1FF84FFB1FC89FBE247B8FBA5" box="[806,1278,1085,1107]" pageId="6" pageNumber="107">
<treatmentCitation id="E372109EFF84FFB1FC89FBE247BDFBA5" author="Lancini AR" box="[806,1275,1085,1107]" page="4" pageId="6" pageNumber="107" year="1968">
<taxonomicName id="A5D34D0CFF84FFB1FC89FBE247BDFBA5" authority="Lancini, 1968: 4" authorityName="Lancini" authorityPageNumber="4" authorityYear="1968" box="[806,1275,1085,1107]" class="Reptilia" family="Gymnophthalmidae" genus="Euspondylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="ampuedae">
<emphasis id="50A7EA9DFF84FFB1FC89FBE24770FBA5" box="[806,1078,1085,1106]" italics="true" pageId="6" pageNumber="107">Euspondylus ampuedae</emphasis>
<bibRefCitation id="06424B7EFF84FFB1FB90FBE247BDFBA5" author="Lancini AR" box="[1087,1275,1085,1107]" pageId="6" pageNumber="107" pagination="1 - 8" refId="ref8883" refString="Lancini AR. 1968. El genero Euspondylus (Sauria: Teiidae) en Venezuela. Publicaciones Ocasionales del Museo de Ciencias Naturales, Zoologia 12: 1 - 8." type="journal article" year="1968">Lancini, 1968: 4</bibRefCitation>
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.
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</paragraph>
<footnote id="01C82A81FF84FFB1FD9BFB824197FB9D" box="[564,721,1117,1130]" pageId="6" pageNumber="107">
<paragraph id="626C368FFF84FFB1FD9BFB824197FB9D" blockId="6.[564,754,893,1611]" box="[564,721,1117,1130]" pageId="6" pageNumber="107">(A) Madre de Dios, Peru</paragraph>
</footnote>
<paragraph id="626C368FFF84FFB1FC89FB8440F2FB78" blockId="6.[806,1422,931,1720]" pageId="6" pageNumber="107">
<treatmentCitationGroup id="42C311A1FF84FFB1FC89FB8440F2FB78" pageId="6" pageNumber="107">
<treatmentCitation id="E372109EFF84FFB1FC89FB8440F6FB78" author="La Marca E &amp; Garcia-Perez JE" page="111" pageId="6" pageNumber="107" year="1990">
<taxonomicName id="A5D34D0CFF84FFB1FC89FB8440F6FB78" authority=": La Marca &amp; Garcia-Perez, 1990: 111" authorityName=": La Marca &amp; Garcia-Perez" authorityPageNumber="111" authorityYear="1990" class="Reptilia" family="Gymnophthalmidae" genus="Prionodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="ampuedai">
<emphasis id="50A7EA9DFF84FFB1FC89FB844714FB87" box="[806,1106,1115,1136]" italics="true" pageId="6" pageNumber="107">Prionodactylus ampuedai:</emphasis>
<bibRefCitation id="06424B7EFF84FFB1FBF3FB8340F6FB78" author="La Marca E &amp; Garcia-Perez JE" pageId="6" pageNumber="107" pagination="111 - 115" refId="ref8840" refString="La Marca E, Garcia-Perez JE. 1990. Prionodactylus ampuedai (Sauria: Teiidae): a new lizard record for Colombia, with taxonomic comments on the species. Bulletin of the Maryland Herpetological Society 26: 111 - 115." type="journal article" year="1990">La Marca &amp; García-Pérez, 1990: 111</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<footnote id="01C82A81FF84FFB1FD9BFBA24191FB7D" box="[564,727,1149,1162]" pageId="6" pageNumber="107">
<paragraph id="626C368FFF84FFB1FD9BFBA24191FB7D" blockId="6.[564,754,893,1611]" box="[564,727,1149,1162]" pageId="6" pageNumber="107">(A) Cochabamba, Bolivia</paragraph>
</footnote>
<footnote id="01C82A81FF84FFB1FD9BFB424197FB5D" box="[564,721,1181,1194]" pageId="6" pageNumber="107">
<paragraph id="626C368FFF84FFB1FD9BFB424197FB5D" blockId="6.[564,754,893,1611]" box="[564,721,1181,1194]" pageId="6" pageNumber="107">(A) Cochabamba,Bolivia</paragraph>
</footnote>
<paragraph id="626C368FFF84FFB1FC90FB464670F94F" blockId="6.[806,1422,931,1720]" pageId="6" pageNumber="107">
When
<bibRefCitation id="06424B7EFF84FFB1FC38FB46477AFB58" author="Uzzell TM" box="[919,1084,1177,1199]" pageId="6" pageNumber="107" pagination="1 - 67" refId="ref9618" refString="Uzzell TM. 1973. A revision of lizards of the genus Prionodactylus, with a new genus for P. leucostictus and notes on the genus Euspondylus (Sauria, Teiidae). Postilla 159: 1 - 67." type="journal article" year="1973">Uzzell (1973)</bibRefCitation>
synonymized
<taxonomicName id="A5D34D0CFF84FFB1FB54FB4640DCFB3B" authorityName="Lancini" authorityYear="1968" class="Reptilia" family="Gymnophthalmidae" genus="Euspondylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="ampuedae">
<emphasis id="50A7EA9DFF84FFB1FB54FB4640DCFB3B" italics="true" pageId="6" pageNumber="107">Euspondylus ampuedae</emphasis>
</taxonomicName>
with
<taxonomicName id="A5D34D0CFF84FFB1FC4CFB684654FB3B" authorityName=": Boulenger" authorityYear="1885" box="[995,1298,1207,1228]" class="Reptilia" family="Gymnophthalmidae" genus="Prionodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="50A7EA9DFF84FFB1FC4CFB684654FB3B" box="[995,1298,1207,1228]" italics="true" pageId="6" pageNumber="107">Prionodactylus vertebralis</emphasis>
</taxonomicName>
, he did so without examining any specimens of
<taxonomicName id="A5D34D0CFF84FFB1FB41FB0846C1FB1C" authorityName="Lancini" authorityYear="1968" box="[1262,1415,1238,1260]" class="Reptilia" family="Gymnophthalmidae" genus="Euspondylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="ampuedae">
<emphasis id="50A7EA9DFF84FFB1FB41FB0846C1FB1C" box="[1262,1415,1238,1260]" italics="true" pageId="6" pageNumber="107">E. ampuedae</emphasis>
</taxonomicName>
. There were few specimens known at that time.
<bibRefCitation id="06424B7EFF84FFB1FADFFB2A47D3FADD" author="La Marca E &amp; Garcia-Perez JE" pageId="6" pageNumber="107" pagination="111 - 115" refId="ref8840" refString="La Marca E, Garcia-Perez JE. 1990. Prionodactylus ampuedai (Sauria: Teiidae): a new lizard record for Colombia, with taxonomic comments on the species. Bulletin of the Maryland Herpetological Society 26: 111 - 115." type="journal article" year="1990">La Marca &amp; García-Pérez (1990)</bibRefCitation>
reported additional specimens and resurrected the species as
<taxonomicName id="A5D34D0CFF84FFB1FABDFAED409AFA91" authorityName=": La Marca &amp; Garcia-Perez" authorityYear="1990" class="Reptilia" family="Gymnophthalmidae" genus="Prionodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="ampuedai">
<emphasis id="50A7EA9DFF84FFB1FABDFAED409AFA91" italics="true" pageId="6" pageNumber="107">Prionodactylus ampuedai</emphasis>
</taxonomicName>
(correcting the latinized name ending). They based their conclusion on five differences, three of which are based on colour pattern. Similar to
<bibRefCitation id="06424B7EFF84FFB1FC89FA724086FA34" author="Uzzell TM" box="[806,960,1453,1475]" pageId="6" pageNumber="107" pagination="1 - 67" refId="ref9618" refString="Uzzell TM. 1973. A revision of lizards of the genus Prionodactylus, with a new genus for P. leucostictus and notes on the genus Euspondylus (Sauria, Teiidae). Postilla 159: 1 - 67." type="journal article" year="1973">Uzzell (1973)</bibRefCitation>
, I have not been able to examine any of the specimens designated as
<taxonomicName id="A5D34D0CFF84FFB1FB23FA134659FA17" authorityName=": La Marca &amp; Garcia-Perez" authorityYear="1990" box="[1164,1311,1483,1505]" class="Reptilia" family="Gymnophthalmidae" genus="Prionodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="ampuedai">
<emphasis id="50A7EA9DFF84FFB1FB23FA134659FA17" box="[1164,1311,1483,1505]" italics="true" pageId="6" pageNumber="107">P. ampuedai</emphasis>
</taxonomicName>
(most of which are held in private collections), but I tend to follow
<bibRefCitation id="06424B7EFF84FFB1FCFBF9D640ADF9E8" author="Uzzell TM" box="[852,1003,1545,1567]" pageId="6" pageNumber="107" pagination="1 - 67" refId="ref9618" refString="Uzzell TM. 1973. A revision of lizards of the genus Prionodactylus, with a new genus for P. leucostictus and notes on the genus Euspondylus (Sauria, Teiidae). Postilla 159: 1 - 67." type="journal article" year="1973">Uzzell (1973)</bibRefCitation>
because the differences presented in the
<bibRefCitation id="06424B7EFF84FFB1FCFBF9F74789F9C9" author="La Marca E &amp; Garcia-Perez JE" box="[852,1231,1575,1598]" pageId="6" pageNumber="107" pagination="111 - 115" refId="ref8840" refString="La Marca E, Garcia-Perez JE. 1990. Prionodactylus ampuedai (Sauria: Teiidae): a new lizard record for Colombia, with taxonomic comments on the species. Bulletin of the Maryland Herpetological Society 26: 111 - 115." type="journal article" year="1990">La Marca &amp; García-Pérez (1990)</bibRefCitation>
table are poorly delineated and not convincing (i.e. shape of pale lip line and overlapping femoral pore counts). Until more definitive evidence is presented, the status and relationships of
<taxonomicName id="A5D34D0CFF84FFB1FC1BF97C4704F940" authorityName=": La Marca &amp; Garcia-Perez" authorityYear="1990" box="[948,1090,1698,1720]" class="Reptilia" family="Gymnophthalmidae" genus="Prionodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="ampuedai">
<emphasis id="50A7EA9DFF84FFB1FC1BF97C4704F940" box="[948,1090,1698,1720]" italics="true" pageId="6" pageNumber="107">P. ampuedai</emphasis>
</taxonomicName>
will not be assessed.
</paragraph>
</subSubSection>
</treatment>
</document>