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<document ID-DOI="10.11646/zootaxa.5270.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7850611" ID-ZooBank="751612E7-95DB-428C-8B18-E5E3A6A0CFB2" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.tables_approvedBy="diego" checkinTime="1681978884392" checkinUser="plazi" docAuthor="Perkins, Michael A., Williams, Bronwyn W. &amp; Russ, William T." docDate="2023" docId="360887F0FF92A34FFF0F5735FDBFF98C" docLanguage="en" docName="zootaxa.5270.2.3.pdf" docOrigin="Zootaxa 5270 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5270.2.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cambarus lapidosus Perkins, Williams &amp; Russ 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="218" masterDocId="CA31FF88FF96A344FF985141FFE3FF85" masterDocTitle="Integrative taxonomy reveals two new narrowly-endemic crayfish species (Decapoda: Cambaridae) from the Yadkin River Basin in western North Carolina, USA" masterLastPageNumber="230" masterPageNumber="207" pageNumber="211" updateTime="1682449342342" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Integrative taxonomy reveals two new narrowly-endemic crayfish species (Decapoda: Cambaridae) from the Yadkin River Basin in western North Carolina, USA</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Perkins, Michael A.</mods:namePart>
<mods:affiliation>North Carolina Wildlife Resources Commission, 645 Fish Hatchery Road, Marion NC, 28752</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Williams, Bronwyn W.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-6403-112X</mods:nameIdentifier>
<mods:affiliation>North Carolina Museum of Natural Sciences, Research Laboratory, 1671 Gold Star Drive, Raleigh NC, 27669 bronwyn. williams @ naturalsciences. org; https: // orcid. org / 0000 - 0001 - 6403 - 112 X</mods:affiliation>
<mods:nameIdentifier type="email">bronwyn.williams@naturalsciences.org</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russ, William T.</mods:namePart>
<mods:nameIdentifier type="ORCID">0009-0000-5135-4309</mods:nameIdentifier>
<mods:affiliation>North Carolina Wildlife Resources Commission, 645 Fish Hatchery Road, Marion NC, 28752 &amp; William. russ @ ncwildlife. org; https: // orcid. org / 0009 - 0000 - 5135 - 4309</mods:affiliation>
<mods:nameIdentifier type="email">illiam.russ@ncwildlife.org</mods:nameIdentifier>
</mods:name>
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<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5270</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>207</mods:start>
<mods:end>230</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5270.2.3</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5270.2.3</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7850611</mods:identifier>
<mods:identifier type="ZooBank">751612E7-95DB-428C-8B18-E5E3A6A0CFB2</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7864644" ID-Zenodo-Dep="7864644" LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C" httpUri="http://treatment.plazi.org/id/360887F0FF92A34FFF0F5735FDBFF98C" lastPageId="11" lastPageNumber="218" pageId="4" pageNumber="211">
<subSubSection box="[151,835,1652,1679]" pageId="4" pageNumber="211" type="nomenclature">
<paragraph blockId="4.[151,835,1652,1716]" box="[151,835,1652,1679]" pageId="4" pageNumber="211">
<heading bold="true" box="[151,835,1652,1679]" fontSize="11" level="1" pageId="4" pageNumber="211" reason="1">
<emphasis bold="true" box="[151,835,1652,1679]" pageId="4" pageNumber="211">
<taxonomicName authority="Perkins, Williams &amp; Russ" authorityName="Perkins, Williams &amp; Russ" authorityYear="2023" box="[151,729,1652,1679]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="211" phylum="Arthropoda" rank="species" species="lapidosus" status="sp. nov.">
<emphasis bold="true" box="[151,398,1652,1679]" italics="true" pageId="4" pageNumber="211">Cambarus lapidosus</emphasis>
Perkins, Williams &amp; Russ
</taxonomicName>
,
<taxonomicNameLabel box="[742,835,1653,1679]" pageId="4" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,391,1689,1716]" pageId="4" pageNumber="211" type="description">
<paragraph blockId="4.[151,835,1652,1716]" box="[151,391,1689,1716]" pageId="4" pageNumber="211">
<figureCitation box="[151,291,1689,1716]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="6.[151,250,1945,1969]" captionStartId-1="8.[151,250,1800,1824]" captionStartId-2="9.[151,250,915,939]" captionTargetBox-0="[169,1419,669,1922]" captionTargetBox-1="[192,1396,309,1776]" captionTargetBox-2="[190,1398,181,892]" captionTargetId-0="figure-243@6.[169,1419,669,1922]" captionTargetId-1="figure-78@8.[192,1396,306,1776]" captionTargetId-2="figure-18@9.[190,1398,181,892]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 3. Cambarus lapidosus, new species. (A.) Lateral view of carapace; (B.) dorsal view of carapace; (C.) right antennal scale; (D.) dorsal view of carpus and chela; (E.) epistome; (F.) lateral and (G.) mesial view of form I gonopod; (H.) lateral and (I.) mesial view of form II gonopod; (J.) annulus ventralis.AE, FG from holotype; HI from morphotype; J from allotype." captionText-1="FIGURE 4. Cambarus lapidosus, holotype (NCSM 90222) in life (photo credit: Michael A. Perkins, NCWRC)." captionText-2="FIGURE 5. Cambarus lapidosus, (left) second form male morphotype (NCSM 90224) in life, Stony Fork, Wilkes County, NC; (right) female allotype (NCSM 90223) in life, Orchard Creek, Watauga County, NC (note: regenerated left chela with single row of mesial tubercles); (photo credit: Michael A. Perkins, NCWRC)." figureDoi-0="http://doi.org/10.5281/zenodo.7850617" figureDoi-1="http://doi.org/10.5281/zenodo.7850619" figureDoi-2="http://doi.org/10.5281/zenodo.7850621" httpUri-0="https://zenodo.org/record/7850617/files/figure.png" httpUri-1="https://zenodo.org/record/7850619/files/figure.png" httpUri-2="https://zenodo.org/record/7850621/files/figure.png" pageId="4" pageNumber="211">Figures 35</figureCitation>
,
<tableCitation box="[304,391,1689,1715]" captionStart="TABLE 3" captionStartId="7.[151,239,151,175]" captionTargetPageId="7" captionText="TABLE 3. Morphological measurements (mm) of the primary type specimens of Cambarus lapidosus (NCSM 90222, Holotype; NCSM 90223, Allotype; NCSM 90224, Morphotype)." httpUri="http://table.plazi.org/id/EADE666EFF91A343FF0F51D6FCA8FF56" pageId="4" pageNumber="211" tableUuid="EADE666EFF91A343FF0F51D6FCA8FF56">Table 3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="214" pageId="4" pageNumber="211" type="diagnosis">
<paragraph blockId="4.[151,1437,1761,2039]" pageId="4" pageNumber="211">
<emphasis bold="true" box="[151,274,1761,1787]" pageId="4" pageNumber="211">Diagnosis.</emphasis>
Body and eyes pigmented. Base of rostrum broad, mid and anterior portions subparallel, weakly excavated, ventrally deflected, margins thickened to acumen, without spines or tubercles; median carina not present. Acumen well-defined with prominent terminal, dorsally-deflected, spiniform tubercle. Rostrum 1.011.44 (x = 1.21, n=26) times long as wide.
</paragraph>
<paragraph blockId="4.[151,1437,1761,2039]" pageId="4" pageNumber="211">Areola 1.23.7 (x = 2.73, n=26) times long as wide, with 57 punctations through narrowest point. Carapace compressed dorsoventrally, subovate in cross section; lateral area of branchiostegal region heavily tuberculate; 14 well-defined cervical tubercles, rarely spiniform. Branchiostegal spine present but greatly reduced. Postorbital ridge well-developed, not terminating in tubercle or spine. Suborbital angle strongly acute, not terminating in tubercle or spine.</paragraph>
<paragraph blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="212">Mesial palm of chelae with two prominent rows of well-developed cristiform tubercles; both rows with 58 tubercles. The majority of specimens measured (84%, n=21) possess one subpalmar tubercle; 4% (n=1) possess 2 subpalmar tubercles, and 12% (n=3) had none. Opposable margin of propodus typically with 57 well-developed denticles, from base consisting of 34 tubercles, 23 reduced tubercles, terminating in sharply pointed fourth tubercle; denticles extending an average of 73.2% of distal length. Opposable margin of dactyl with 58 denticles, from base consisting of 45 tubercles, tapering to 23 reduced tubercles; denticles extending an average of 58.0% distal length. Dactyl with 59 tubercles along dorsal surface, tapering in count and prominence towards terminus. Palm length to dactyl length ratio averaging 0.69. Sharp corneous tips present on dactyl and fixed finger of propodus. Carpus with one large well-developed spine on mesial margin, with one additional spiniform tubercle towards posterior margin of carpus, occasionally expressed as well-developed spine in some individuals (31%, n=23).</paragraph>
<paragraph blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="212">First pleopod of form I male with long terminal elements; central projection not tapered, curved greater than 90° to shaft of gonopod, with conspicuous subapical notch, length extending greater than the marginal terminus of the mesial process; mesial process recurved&gt; 90° to main shaft, conically-shaped at base, tapering to a rounded terminus projecting beyond margin of main shaft, directed caudolaterally. Form II male central projection rounded, curved 90° to mesial process, approximately equal in length; mesial process conical, tapering to a rounded terminus, directed caudolaterally, terminal length greater than margin of shaft. Hooks on ischium of third pereiopods in form I and II males. Female annulus ventralis asymmetrical and appearing sclerotized caudally; distinct “S”-shaped bend, terminating in caudal portion; cephalic portion consists of median trough leading to central fossa, not sclerotized.</paragraph>
<paragraph blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="212">
<emphasis bold="true" box="[199,715,798,825]" pageId="5" pageNumber="212">
Holotypic Male, Form I (
<figureCitation box="[501,597,798,824]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="6.[151,250,1945,1969]" captionStartId-1="8.[151,250,1800,1824]" captionTargetBox-0="[169,1419,669,1922]" captionTargetBox-1="[192,1396,309,1776]" captionTargetId-0="figure-243@6.[169,1419,669,1922]" captionTargetId-1="figure-78@8.[192,1396,306,1776]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="FIGURE 3. Cambarus lapidosus, new species. (A.) Lateral view of carapace; (B.) dorsal view of carapace; (C.) right antennal scale; (D.) dorsal view of carpus and chela; (E.) epistome; (F.) lateral and (G.) mesial view of form I gonopod; (H.) lateral and (I.) mesial view of form II gonopod; (J.) annulus ventralis.AE, FG from holotype; HI from morphotype; J from allotype." captionText-1="FIGURE 4. Cambarus lapidosus, holotype (NCSM 90222) in life (photo credit: Michael A. Perkins, NCWRC)." figureDoi-0="http://doi.org/10.5281/zenodo.7850617" figureDoi-1="http://doi.org/10.5281/zenodo.7850619" httpUri-0="https://zenodo.org/record/7850617/files/figure.png" httpUri-1="https://zenodo.org/record/7850619/files/figure.png" pageId="5" pageNumber="212">Fig. 3-4</figureCitation>
;
<tableCitation box="[611,701,798,825]" captionStart="TABLE 3" captionStartId="7.[151,239,151,175]" captionTargetPageId="7" captionText="TABLE 3. Morphological measurements (mm) of the primary type specimens of Cambarus lapidosus (NCSM 90222, Holotype; NCSM 90223, Allotype; NCSM 90224, Morphotype)." httpUri="http://table.plazi.org/id/EADE666EFF91A343FF0F51D6FCA8FF56" pageId="5" pageNumber="212" tableUuid="EADE666EFF91A343FF0F51D6FCA8FF56">Table 3</tableCitation>
).
</emphasis>
Body somewhat compressed dorsoventrally; carapace posterior to cervical groove wider than abdomen. Carapace width greater than carapace depth at caudodorsal margin of cervical groove. Total carapace length
<quantity box="[577,678,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="4.97" pageId="5" pageNumber="212" unit="mm" value="49.7">49.7 mm</quantity>
, PCL
<quantity box="[744,846,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="4.21" pageId="5" pageNumber="212" unit="mm" value="42.1">42.1 mm</quantity>
. Areola 2.6 times longer than wide, moderately broad with 6 punctations across the narrowest section. Rostrum excavated; margins moderately thickened, subparallel and tapering at acumen. Rostrum 1.3 times longer than width at base, moderately truncated, acumen well-defined and terminating in deflected corneous tip; floor of rostrum with numerous punctations. Postorbital ridge well-developed but lacking well-defined terminal tubercle. Suborbital angle acute, lacking terminal tubercle. Cluster of 4 tubercles present at lateral margin of cervical groove. Orbital, branchiostegal, and mandibular regions of carapace with well-developed tubercles; greatest tubercle density in hepatic region. Branchiostegal spine present but reduced. Abdomen subequal in length to carapace, 1.7 times longer than wide, pleura rounded cephaloventrally and angled caudoventrally. Lateral margin of terga weakly angulate, lateral margin of second pleuron with deep furrow. Cephalic section of telson with 2 conspicuous spines in each caudolateral corner. Proximal podomere of uropod with distal spine on mesial lobe. Mesial ramus of uropod with median ridge tapering distally, terminating in median spine not extending beyond margin of ramus; laterodistal spine absent. Distal margin of proximal segment of lateral ramus of right uropod with 1 large fixed lateral spine, 1 reduced movable spine, and numerous small immovable spines. Cephalomedian lobe of epistome subtriangular, thin, lateral margins weakly inflated, tapering medially to conspicuous anterior terminus; zygoma moderately arched; cephalolateral margins thickened, rounded at junction with endostyle; body of epistome possessing prominent median fovea.Antennal scale broadest proximally, setiferous along mesial margin, lateral margin thickened and terminating in large spine. Right antennal scale 2.1 times longer than wide. Antennae extending to anterior margin of rami when adpressed. All following descriptions from right chela: mesial surface with two rows of 7 well-defined tubercles; palm length 69.7% palm width; dorsal longitudinal ridge of dactyl with 12 scattered well-defined tubercles, dactyl terminating in corneous spine; moderately pronounced dorsomedian ridge on fixed finger of propodus, scattered and numerous punctations throughout, absent of elongated setae, terminating in corneous tip; opposable margin of propodus from base consisting of 5 well-defined tubercles, one smaller tubercle, ending in sharply pointed tubercle; opposable margin of dactyl from base consisting of 4 well-defined tubercles and 4 smaller tubercles tapering in size to terminus; small subpalmar tubercle absent on right chelae but present on left chelae; carpus with prominent dorsal furrow, surface with scattered punctuations, mesial margin with large procurved spine and reduced anterior proximal spine; distodorsal surface of merus with 14 spines and spiniform tubercles, ventromesial ridge with 2 well-developed spiniform tubercles, ventrolateral ridge with 4 small spines. Well-developed hooks on ischium of third pereiopods. Form I gonopod as described in Diagnosis.
</paragraph>
<paragraph blockId="5.[151,1437,150,2013]" lastBlockId="6.[151,1437,150,645]" lastPageId="6" lastPageNumber="213" pageId="5" pageNumber="212">
<emphasis bold="true" box="[199,601,1878,1905]" pageId="5" pageNumber="212">
Allotypic Female (
<figureCitation box="[417,486,1878,1904]" captionStart="FIGURE 3" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[169,1419,669,1922]" captionTargetId="figure-243@6.[169,1419,669,1922]" captionTargetPageId="6" captionText="FIGURE 3. Cambarus lapidosus, new species. (A.) Lateral view of carapace; (B.) dorsal view of carapace; (C.) right antennal scale; (D.) dorsal view of carpus and chela; (E.) epistome; (F.) lateral and (G.) mesial view of form I gonopod; (H.) lateral and (I.) mesial view of form II gonopod; (J.) annulus ventralis.AE, FG from holotype; HI from morphotype; J from allotype." figureDoi="http://doi.org/10.5281/zenodo.7850617" httpUri="https://zenodo.org/record/7850617/files/figure.png" pageId="5" pageNumber="212">Fig. 3</figureCitation>
,
<tableCitation box="[498,587,1878,1905]" captionStart="TABLE 3" captionStartId="7.[151,239,151,175]" captionTargetPageId="7" captionText="TABLE 3. Morphological measurements (mm) of the primary type specimens of Cambarus lapidosus (NCSM 90222, Holotype; NCSM 90223, Allotype; NCSM 90224, Morphotype)." httpUri="http://table.plazi.org/id/EADE666EFF91A343FF0F51D6FCA8FF56" pageId="5" pageNumber="212" tableUuid="EADE666EFF91A343FF0F51D6FCA8FF56">Table 3</tableCitation>
).
</emphasis>
Differing from
<typeStatus box="[779,877,1879,1905]" pageId="5" pageNumber="212">holotype</typeStatus>
in the following respects: carapace damaged with dorsolongitudinal crack; TCL
<quantity box="[488,589,1915,1941]" metricMagnitude="-2" metricUnit="m" metricValue="3.27" pageId="5" pageNumber="212" unit="mm" value="32.7">32.7 mm</quantity>
, PCL
<quantity box="[657,760,1915,1941]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="5" pageNumber="212" unit="mm" value="26.6">26.6 mm</quantity>
; areola 2.4 times longer than wide; rostrum length and width approximately equal at base; abdomen 1.5 times longer than wide. All following descriptions from right chela; mesial margin with one row of 7 well-developed tubercles and a second row of 6 well-developed tubercles; palm length 75% palm width; single small subpalmar tubercle present on both chelae. Antennae extending to fourth terga when adpressed. Antennal scale 1.7 times longer than wide. Specimen lacks all male secondary sexual traits. Annulus ventralis as described in Diagnosis.
</paragraph>
<paragraph blockId="6.[151,1437,150,645]" pageId="6" pageNumber="213">
<emphasis bold="true" box="[199,749,258,285]" pageId="6" pageNumber="213">
Morphotypic Male, Form II (
<figureCitation box="[557,629,258,284]" captionStart="FIGURE 3" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[169,1419,669,1922]" captionTargetId="figure-243@6.[169,1419,669,1922]" captionTargetPageId="6" captionText="FIGURE 3. Cambarus lapidosus, new species. (A.) Lateral view of carapace; (B.) dorsal view of carapace; (C.) right antennal scale; (D.) dorsal view of carpus and chela; (E.) epistome; (F.) lateral and (G.) mesial view of form I gonopod; (H.) lateral and (I.) mesial view of form II gonopod; (J.) annulus ventralis.AE, FG from holotype; HI from morphotype; J from allotype." figureDoi="http://doi.org/10.5281/zenodo.7850617" httpUri="https://zenodo.org/record/7850617/files/figure.png" pageId="6" pageNumber="213">Fig. 3</figureCitation>
,
<tableCitation box="[643,735,258,285]" captionStart="TABLE 3" captionStartId="7.[151,239,151,175]" captionTargetPageId="7" captionText="TABLE 3. Morphological measurements (mm) of the primary type specimens of Cambarus lapidosus (NCSM 90222, Holotype; NCSM 90223, Allotype; NCSM 90224, Morphotype)." httpUri="http://table.plazi.org/id/EADE666EFF91A343FF0F51D6FCA8FF56" pageId="6" pageNumber="213" tableUuid="EADE666EFF91A343FF0F51D6FCA8FF56">Table 3</tableCitation>
).
</emphasis>
Differing from the
<typeStatus box="[978,1076,259,285]" pageId="6" pageNumber="213">holotype</typeStatus>
in the following respects: TCL
<quantity box="[151,254,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="3.27" pageId="6" pageNumber="213" unit="mm" value="32.7">32.7 mm</quantity>
, PCL
<quantity box="[325,429,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="6" pageNumber="213" unit="mm" value="26.6">26.6 mm</quantity>
; areola 2.7 times longer than wide, rostrum 1.1 times longer than wide at base; abdomen 1.7 times longer than wide. All following descriptions from right chela: mesial margin with one row of 7 well-developed tubercles and a second row of 8 well-developed tubercles; palm length 68% palm width; single reduced subpalmar tubercle present. Antennae extending to posterior margin of second terga when adpressed. Antennal scale 1.9 times longer than wide. Hooks on ischium of third pereiopods weekly developed. Form II gonopod as described in Diagnosis.
</paragraph>
<paragraph blockId="6.[151,1437,150,645]" pageId="6" pageNumber="213">
<emphasis bold="true" box="[199,254,510,536]" pageId="6" pageNumber="213">Size.</emphasis>
Form I male (n = 9) TCL ranging in size from
<quantity box="[776,875,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="3.79" pageId="6" pageNumber="213" unit="mm" value="37.9">37.9 mm</quantity>
to
<quantity box="[910,1009,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="4.97" pageId="6" pageNumber="213" unit="mm" value="49.7">49.7 mm</quantity>
(x= 40.3, SD = 3.76). Form II male (n = 7) TCL ranging in size from
<quantity box="[498,598,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="3.1399999999999997" pageId="6" pageNumber="213" unit="mm" value="31.4">31.4 mm</quantity>
to
<quantity box="[634,734,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="4.42" pageId="6" pageNumber="213" unit="mm" value="44.2">44.2 mm</quantity>
(x = 36.2, SD = 4.81). Female (n = 10) TCL ranging
<quantity box="[1336,1437,546,573]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="6" pageNumber="213" unit="mm" value="25.7">25.7 mm</quantity>
to
<quantity box="[181,283,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="4.32" pageId="6" pageNumber="213" unit="mm" value="43.2">43.2 mm</quantity>
(x = 35.3, SD = 6.22). The largest specimen observed was the holotypic form I male measuring
<quantity metricMagnitude="-2" metricUnit="m" metricValue="4.97" pageId="6" pageNumber="213" unit="mm" value="49.7">49.7 mm</quantity>
TCL.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7850617" ID-Zenodo-Dep="7850617" httpUri="https://zenodo.org/record/7850617/files/figure.png" pageId="6" pageNumber="213" startId="6.[151,250,1945,1969]" targetBox="[169,1419,669,1922]" targetPageId="6" targetType="figure">
<paragraph blockId="6.[151,1437,1945,2042]" pageId="6" pageNumber="213">
<emphasis bold="true" box="[151,636,1945,1970]" pageId="6" pageNumber="213">
FIGURE 3.
<taxonomicName authority="Perkins &amp; Williams &amp; Russ, 2023" authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[281,494,1945,1969]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="213" phylum="Arthropoda" rank="species" species="lapidosus" status="new species">
<emphasis bold="true" box="[281,494,1945,1969]" italics="true" pageId="6" pageNumber="213">Cambarus lapidosus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[506,632,1945,1970]" pageId="6" pageNumber="213" rank="species">new species</taxonomicNameLabel>
.
</emphasis>
(A.) Lateral view of carapace; (B.) dorsal view of carapace; (C.) right antennal scale; (D.) dorsal view of carpus and chela; (E.) epistome; (F.) lateral and (G.) mesial view of form I gonopod; (H.) lateral and (I.) mesial view of form II gonopod; (J.) annulus ventralis. AE, FG from holotype; HI from morphotype; J from allotype.
</paragraph>
</caption>
<caption ID-Table-UUID="EADE666EFF91A343FF0F51D6FCA8FF56" httpUri="http://table.plazi.org/id/EADE666EFF91A343FF0F51D6FCA8FF56" pageId="7" pageNumber="214" startId="7.[151,239,151,175]" targetBox="[166,1319,227,1378]" targetIsTable="true" targetPageId="7" targetType="table">
<paragraph blockId="7.[151,1436,151,211]" pageId="7" pageNumber="214">
<emphasis bold="true" box="[151,268,151,175]" pageId="7" pageNumber="214">TABLE 3.</emphasis>
Morphological measurements (mm) of the primary type specimens of
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[1040,1261,151,175]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="214" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[1040,1261,151,175]" italics="true" pageId="7" pageNumber="214">Cambarus lapidosus</emphasis>
</taxonomicName>
(NCSM 90222, Holotype; NCSM 90223, Allotype; NCSM 90224, Morphotype).
</paragraph>
</caption>
<paragraph pageId="7" pageNumber="214">
<table box="[166,1319,227,1378]" gridcols="4" gridrows="29" pageId="7" pageNumber="214">
<tr box="[166,1319,227,251]" gridrow="0" pageId="7" pageNumber="214">
<th box="[166,426,227,251]" gridcol="0" gridrow="0" pageId="7" pageNumber="214">
<emphasis bold="true" box="[166,277,227,251]" pageId="7" pageNumber="214">Character</emphasis>
</th>
<th box="[613,768,227,251]" gridcol="1" gridrow="0" pageId="7" pageNumber="214">
<emphasis bold="true" box="[613,768,227,251]" pageId="7" pageNumber="214">Holotype (MI)</emphasis>
</th>
<th box="[888,1019,227,251]" gridcol="2" gridrow="0" pageId="7" pageNumber="214">
<emphasis bold="true" box="[888,1019,227,251]" pageId="7" pageNumber="214">Allotype (F)</emphasis>
</th>
<th box="[1117,1319,227,251]" gridcol="3" gridrow="0" pageId="7" pageNumber="214">
<emphasis bold="true" box="[1117,1319,227,251]" pageId="7" pageNumber="214">Morphotype (MII)</emphasis>
</th>
</tr>
<tr box="[166,1319,266,290]" gridrow="1" pageId="7" pageNumber="214">
<th box="[166,1319,266,290]" colspan="4" colspanRight="3" gridcol="0" gridrow="1" pageId="7" pageNumber="214">
<emphasis bold="true" box="[166,270,266,290]" pageId="7" pageNumber="214">Carapace</emphasis>
</th>
</tr>
<tr box="[166,1319,307,332]" gridrow="2" pageId="7" pageNumber="214">
<th box="[166,426,307,332]" gridcol="0" gridrow="2" pageId="7" pageNumber="214">Height</th>
<td box="[613,768,307,332]" gridcol="1" gridrow="2" pageId="7" pageNumber="214">21.0</td>
<td box="[888,1019,307,332]" gridcol="2" gridrow="2" pageId="7" pageNumber="214">14.0</td>
<td box="[1117,1319,307,332]" gridcol="3" gridrow="2" pageId="7" pageNumber="214">13.7</td>
</tr>
<tr box="[166,1319,348,372]" gridrow="3" pageId="7" pageNumber="214">
<th box="[166,426,348,372]" gridcol="0" gridrow="3" pageId="7" pageNumber="214">Width</th>
<td box="[613,768,348,372]" gridcol="1" gridrow="3" pageId="7" pageNumber="214">26.2</td>
<td box="[888,1019,348,372]" gridcol="2" gridrow="3" pageId="7" pageNumber="214">19.0</td>
<td box="[1117,1319,348,372]" gridcol="3" gridrow="3" pageId="7" pageNumber="214">18.1</td>
</tr>
<tr box="[166,1319,388,412]" gridrow="4" pageId="7" pageNumber="214">
<th box="[166,426,388,412]" gridcol="0" gridrow="4" pageId="7" pageNumber="214">Length (TCL)</th>
<td box="[613,768,388,412]" gridcol="1" gridrow="4" pageId="7" pageNumber="214">49.7</td>
<td box="[888,1019,388,412]" gridcol="2" gridrow="4" pageId="7" pageNumber="214">36.7</td>
<td box="[1117,1319,388,412]" gridcol="3" gridrow="4" pageId="7" pageNumber="214">32.7</td>
</tr>
<tr box="[166,1319,427,451]" gridrow="5" pageId="7" pageNumber="214">
<th box="[166,1319,427,451]" colspan="4" colspanRight="3" gridcol="0" gridrow="5" pageId="7" pageNumber="214">
<emphasis bold="true" box="[166,239,427,451]" pageId="7" pageNumber="214">Areola</emphasis>
</th>
</tr>
<tr box="[166,1319,468,492]" gridrow="6" pageId="7" pageNumber="214">
<th box="[166,426,468,492]" gridcol="0" gridrow="6" pageId="7" pageNumber="214">Length</th>
<td box="[613,768,468,492]" gridcol="1" gridrow="6" pageId="7" pageNumber="214">17.6</td>
<td box="[888,1019,468,492]" gridcol="2" gridrow="6" pageId="7" pageNumber="214">12.0</td>
<td box="[1117,1319,468,492]" gridcol="3" gridrow="6" pageId="7" pageNumber="214">10.8</td>
</tr>
<tr box="[166,1319,509,533]" gridrow="7" pageId="7" pageNumber="214">
<th box="[166,426,509,533]" gridcol="0" gridrow="7" pageId="7" pageNumber="214">Width</th>
<td box="[613,768,509,533]" gridcol="1" gridrow="7" pageId="7" pageNumber="214">6.9</td>
<td box="[888,1019,509,533]" gridcol="2" gridrow="7" pageId="7" pageNumber="214">4.9</td>
<td box="[1117,1319,509,533]" gridcol="3" gridrow="7" pageId="7" pageNumber="214">4.0</td>
</tr>
<tr box="[166,1319,549,573]" gridrow="8" pageId="7" pageNumber="214">
<th box="[166,1319,549,573]" colspan="4" colspanRight="3" gridcol="0" gridrow="8" pageId="7" pageNumber="214">
<emphasis bold="true" box="[166,262,549,573]" pageId="7" pageNumber="214">Rostrum</emphasis>
</th>
</tr>
<tr box="[166,1319,589,613]" gridrow="9" pageId="7" pageNumber="214">
<th box="[166,426,589,613]" gridcol="0" gridrow="9" pageId="7" pageNumber="214">Width at base</th>
<td box="[613,768,589,613]" gridcol="1" gridrow="9" pageId="7" pageNumber="214">6.0</td>
<td box="[888,1019,589,613]" gridcol="2" gridrow="9" pageId="7" pageNumber="214">6.2</td>
<td box="[1117,1319,589,613]" gridcol="3" gridrow="9" pageId="7" pageNumber="214">5.8</td>
</tr>
<tr box="[166,1319,629,654]" gridrow="10" pageId="7" pageNumber="214">
<th box="[166,426,629,654]" gridcol="0" gridrow="10" pageId="7" pageNumber="214">Width at eyes</th>
<td box="[613,768,629,654]" gridcol="1" gridrow="10" pageId="7" pageNumber="214">5.5</td>
<td box="[888,1019,629,654]" gridcol="2" gridrow="10" pageId="7" pageNumber="214">4.3</td>
<td box="[1117,1319,629,654]" gridcol="3" gridrow="10" pageId="7" pageNumber="214">4.1</td>
</tr>
<tr box="[166,1319,669,694]" gridrow="11" pageId="7" pageNumber="214">
<th box="[166,426,669,694]" gridcol="0" gridrow="11" pageId="7" pageNumber="214">Length</th>
<td box="[613,768,669,694]" gridcol="1" gridrow="11" pageId="7" pageNumber="214">7.6</td>
<td box="[888,1019,669,694]" gridcol="2" gridrow="11" pageId="7" pageNumber="214">6.3</td>
<td box="[1117,1319,669,694]" gridcol="3" gridrow="11" pageId="7" pageNumber="214">6.1</td>
</tr>
<tr box="[166,1319,710,734]" gridrow="12" pageId="7" pageNumber="214">
<th box="[166,1319,710,734]" colspan="4" colspanRight="3" gridcol="0" gridrow="12" pageId="7" pageNumber="214">
<emphasis bold="true" box="[166,207,710,734]" pageId="7" pageNumber="214">Eye</emphasis>
</th>
</tr>
<tr box="[166,1319,750,774]" gridrow="13" pageId="7" pageNumber="214">
<th box="[166,426,750,774]" gridcol="0" gridrow="13" pageId="7" pageNumber="214">Diameter</th>
<td box="[613,768,750,774]" gridcol="1" gridrow="13" pageId="7" pageNumber="214">1.7</td>
<td box="[888,1019,750,774]" gridcol="2" gridrow="13" pageId="7" pageNumber="214">2.1</td>
<td box="[1117,1319,750,774]" gridcol="3" gridrow="13" pageId="7" pageNumber="214">1.6</td>
</tr>
<tr box="[166,1319,790,814]" gridrow="14" pageId="7" pageNumber="214">
<th box="[166,1319,790,814]" colspan="4" colspanRight="3" gridcol="0" gridrow="14" pageId="7" pageNumber="214">
<emphasis bold="true" box="[166,306,790,814]" pageId="7" pageNumber="214">Chela (right)</emphasis>
</th>
</tr>
<tr box="[166,1319,830,855]" gridrow="15" pageId="7" pageNumber="214">
<th box="[166,426,830,855]" gridcol="0" gridrow="15" pageId="7" pageNumber="214">Length lateral margin</th>
<td box="[613,768,830,855]" gridcol="1" gridrow="15" pageId="7" pageNumber="214">48.0</td>
<td box="[888,1019,830,855]" gridcol="2" gridrow="15" pageId="7" pageNumber="214">30.0</td>
<td box="[1117,1319,830,855]" gridcol="3" gridrow="15" pageId="7" pageNumber="214">25.3</td>
</tr>
<tr box="[166,1319,871,895]" gridrow="16" pageId="7" pageNumber="214">
<th box="[166,426,871,895]" gridcol="0" gridrow="16" pageId="7" pageNumber="214">Length mesial palm</th>
<td box="[613,768,871,895]" gridcol="1" gridrow="16" pageId="7" pageNumber="214">13.6</td>
<td box="[888,1019,871,895]" gridcol="2" gridrow="16" pageId="7" pageNumber="214">9.0</td>
<td box="[1117,1319,871,895]" gridcol="3" gridrow="16" pageId="7" pageNumber="214">7.2</td>
</tr>
<tr box="[166,1319,911,935]" gridrow="17" pageId="7" pageNumber="214">
<th box="[166,426,911,935]" gridcol="0" gridrow="17" pageId="7" pageNumber="214">Palm width</th>
<td box="[613,768,911,935]" gridcol="1" gridrow="17" pageId="7" pageNumber="214">19.5</td>
<td box="[888,1019,911,935]" gridcol="2" gridrow="17" pageId="7" pageNumber="214">12.0</td>
<td box="[1117,1319,911,935]" gridcol="3" gridrow="17" pageId="7" pageNumber="214">10.6</td>
</tr>
<tr box="[166,1319,951,975]" gridrow="18" pageId="7" pageNumber="214">
<th box="[166,426,951,975]" gridcol="0" gridrow="18" pageId="7" pageNumber="214">Palm depth</th>
<td box="[613,768,951,975]" gridcol="1" gridrow="18" pageId="7" pageNumber="214">11.7</td>
<td box="[888,1019,951,975]" gridcol="2" gridrow="18" pageId="7" pageNumber="214">7.8</td>
<td box="[1117,1319,951,975]" gridcol="3" gridrow="18" pageId="7" pageNumber="214">7.7</td>
</tr>
<tr box="[166,1319,991,1016]" gridrow="19" pageId="7" pageNumber="214">
<th box="[166,426,991,1016]" gridcol="0" gridrow="19" pageId="7" pageNumber="214">Dactyl length</th>
<td box="[613,768,991,1016]" gridcol="1" gridrow="19" pageId="7" pageNumber="214">32.2</td>
<td box="[888,1019,991,1016]" gridcol="2" gridrow="19" pageId="7" pageNumber="214">18.2</td>
<td box="[1117,1319,991,1016]" gridcol="3" gridrow="19" pageId="7" pageNumber="214">16</td>
</tr>
<tr box="[166,1319,1032,1056]" gridrow="20" pageId="7" pageNumber="214">
<th box="[166,426,1032,1056]" gridcol="0" gridrow="20" pageId="7" pageNumber="214">Subpalmar tubercle count</th>
<td box="[613,768,1032,1056]" gridcol="1" gridrow="20" pageId="7" pageNumber="214">1 (reduced)</td>
<td box="[888,1019,1032,1056]" gridcol="2" gridrow="20" pageId="7" pageNumber="214">1</td>
<td box="[1117,1319,1032,1056]" gridcol="3" gridrow="20" pageId="7" pageNumber="214">1</td>
</tr>
<tr box="[166,1319,1071,1095]" gridrow="21" pageId="7" pageNumber="214">
<th box="[166,1319,1071,1095]" colspan="4" colspanRight="3" gridcol="0" gridrow="21" pageId="7" pageNumber="214">
<emphasis bold="true" box="[166,272,1071,1095]" pageId="7" pageNumber="214">Abdomen</emphasis>
</th>
</tr>
<tr box="[166,1319,1112,1136]" gridrow="22" pageId="7" pageNumber="214">
<th box="[166,426,1112,1136]" gridcol="0" gridrow="22" pageId="7" pageNumber="214">Length</th>
<td box="[613,768,1112,1136]" gridcol="1" gridrow="22" pageId="7" pageNumber="214">35.1</td>
<td box="[888,1019,1112,1136]" gridcol="2" gridrow="22" pageId="7" pageNumber="214">27.6</td>
<td box="[1117,1319,1112,1136]" gridcol="3" gridrow="22" pageId="7" pageNumber="214">26.8</td>
</tr>
<tr box="[166,1319,1152,1177]" gridrow="23" pageId="7" pageNumber="214">
<th box="[166,426,1152,1177]" gridcol="0" gridrow="23" pageId="7" pageNumber="214">Width</th>
<td box="[613,768,1152,1177]" gridcol="1" gridrow="23" pageId="7" pageNumber="214">21.2</td>
<td box="[888,1019,1152,1177]" gridcol="2" gridrow="23" pageId="7" pageNumber="214">18.7</td>
<td box="[1117,1319,1152,1177]" gridcol="3" gridrow="23" pageId="7" pageNumber="214">15.6</td>
</tr>
<tr box="[166,1319,1192,1216]" gridrow="24" pageId="7" pageNumber="214">
<th box="[166,1319,1192,1216]" colspan="4" colspanRight="3" gridcol="0" gridrow="24" pageId="7" pageNumber="214">
<emphasis bold="true" box="[166,266,1192,1216]" pageId="7" pageNumber="214">Gonopod</emphasis>
</th>
</tr>
<tr box="[166,1319,1233,1257]" gridrow="25" pageId="7" pageNumber="214">
<th box="[166,426,1233,1257]" gridcol="0" gridrow="25" pageId="7" pageNumber="214">Length</th>
<td box="[613,768,1233,1257]" gridcol="1" gridrow="25" pageId="7" pageNumber="214">11.8</td>
<td box="[888,1019,1233,1257]" gridcol="2" gridrow="25" pageId="7" pageNumber="214">n/a</td>
<td box="[1117,1319,1233,1257]" gridcol="3" gridrow="25" pageId="7" pageNumber="214">7.6</td>
</tr>
<tr box="[166,1319,1272,1297]" gridrow="26" pageId="7" pageNumber="214">
<th box="[166,1319,1272,1297]" colspan="4" colspanRight="3" gridcol="0" gridrow="26" pageId="7" pageNumber="214">
<emphasis bold="true" box="[166,412,1272,1297]" pageId="7" pageNumber="214">Antennal Scale (Right)</emphasis>
</th>
</tr>
<tr box="[166,1319,1313,1338]" gridrow="27" pageId="7" pageNumber="214">
<th box="[166,426,1313,1338]" gridcol="0" gridrow="27" pageId="7" pageNumber="214">Length</th>
<td box="[613,768,1313,1338]" gridcol="1" gridrow="27" pageId="7" pageNumber="214">8.4</td>
<td box="[888,1019,1313,1338]" gridcol="2" gridrow="27" pageId="7" pageNumber="214">6.3</td>
<td box="[1117,1319,1313,1338]" gridcol="3" gridrow="27" pageId="7" pageNumber="214">6.5</td>
</tr>
<tr box="[166,1319,1354,1378]" gridrow="28" pageId="7" pageNumber="214">
<th box="[166,426,1354,1378]" gridcol="0" gridrow="28" pageId="7" pageNumber="214">Width</th>
<td box="[613,768,1354,1378]" gridcol="1" gridrow="28" pageId="7" pageNumber="214">4.0</td>
<td box="[888,1019,1354,1378]" gridcol="2" gridrow="28" pageId="7" pageNumber="214">3.8</td>
<td box="[1117,1319,1354,1378]" gridcol="3" gridrow="28" pageId="7" pageNumber="214">3.4</td>
</tr>
</table>
</paragraph>
<paragraph blockId="7.[151,1437,1425,2028]" pageId="7" pageNumber="214">
<emphasis bold="true" box="[199,350,1425,1451]" pageId="7" pageNumber="214">Color Notes.</emphasis>
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[359,592,1425,1451]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="214" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[359,592,1425,1451]" italics="true" pageId="7" pageNumber="214">Cambarus lapidosus</emphasis>
</taxonomicName>
(
<figureCitation box="[610,708,1425,1452]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="8.[151,250,1800,1824]" captionStartId-1="9.[151,250,915,939]" captionTargetBox-0="[192,1396,309,1776]" captionTargetBox-1="[190,1398,181,892]" captionTargetId-0="figure-78@8.[192,1396,306,1776]" captionTargetId-1="figure-18@9.[190,1398,181,892]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 4. Cambarus lapidosus, holotype (NCSM 90222) in life (photo credit: Michael A. Perkins, NCWRC)." captionText-1="FIGURE 5. Cambarus lapidosus, (left) second form male morphotype (NCSM 90224) in life, Stony Fork, Wilkes County, NC; (right) female allotype (NCSM 90223) in life, Orchard Creek, Watauga County, NC (note: regenerated left chela with single row of mesial tubercles); (photo credit: Michael A. Perkins, NCWRC)." figureDoi-0="http://doi.org/10.5281/zenodo.7850619" figureDoi-1="http://doi.org/10.5281/zenodo.7850621" httpUri-0="https://zenodo.org/record/7850619/files/figure.png" httpUri-1="https://zenodo.org/record/7850621/files/figure.png" pageId="7" pageNumber="214">Fig. 45</figureCitation>
) carapace ground color mottled, olivaceous to brown. Cephalic portion of carapace immediately anterior to cervical groove dark, forming narrow saddle; mandibular adductor scars mottled, light brown to olivaceous. Posterior portion of carapace, immediately anterior to abdomen, with rust-colored dorsal patch; sides of carapace darkened with heavy mottling. Hepatic region olive, punctuated with cream-colored tubercles. Rostrum ground color olive, margins of acumen ochre to tan. Lateral margin of antennal scale olive to brown, body light brown to cream. Antennae and antennules olive to brown. Dorsal surface of chelae mottled, olive to brown, tubercles on mesial margin and surface of dactyl cream-colored; ventral surface of chelae cream. Denticles on opposable surfaces of chelae cream-colored. Dorsal surface of carpus olivaceous, carpus spine and tubercles cream-colored. Ventral surface of carpus tan to cream. Dorsal surface of merus olive to brown, ventral surface tan to cream, tubercles cream-colored. Dorsal surface of abdomen ground color mottled, deeper olive to brown, posterior margins of terga light brown to tan, pleura olivaceous. Ventral surface of abdomen olive to light brown, swimmerets brown to tan. Dorsal ridge of form I gonopod amber, body of gonopod, mesial process, shaft cream-colored to tan. Form II process and shaft cream-colored or tan. Annulus ventralis cream-colored to brown. Juvenile coloration is similar to that observed in adults.
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="217" pageId="7" pageNumber="214" type="materials_examined">
<paragraph blockId="7.[151,1437,1425,2028]" lastBlockId="8.[151,1436,150,285]" lastPageId="8" lastPageNumber="215" pageId="7" pageNumber="214">
<emphasis bold="true" box="[199,370,1929,1955]" pageId="7" pageNumber="214">Type Locality.</emphasis>
<materialsCitation county="Watauga County" elevation="645" lastPageId="8" lastPageNumber="215" latitude="36.1949" location="Mt. Zion Road" longLatPrecision="6" longitude="-81.466" municipality="Stony Fork" pageId="7" pageNumber="214" specimenCount="1" stateProvince="North Carolina" typeStatus="holotype">
<collectingMunicipality box="[378,508,1929,1955]" pageId="7" pageNumber="214">Stony Fork</collectingMunicipality>
, in the vicinity of the SR 1167 (Mt. Zion Road) bridge crossing, approximately
<locationDeviation location="Laxon" pageId="7" pageNumber="214">
<quantity metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="214" unit="km" value="8.0">8 km</quantity>
SW of Laxon
</locationDeviation>
and 1 km upstream of the Flowers Branch confluence (
<geoCoordinate box="[965,1088,1965,1991]" degrees="36.1949" direction="north" orientation="latitude" pageId="7" pageNumber="214" precision="5" value="36.1949">N 36.1949</geoCoordinate>
,
<geoCoordinate box="[1098,1226,1965,1992]" degrees="81.4660" direction="west" orientation="longitude" pageId="7" pageNumber="214" precision="5" value="-81.466">W 81.4660</geoCoordinate>
),
<collectingCounty box="[1244,1433,1965,1992]" pageId="7" pageNumber="214">Watauga County</collectingCounty>
,
<collectingRegion box="[151,320,2001,2027]" country="United States of America" name="North Carolina" pageId="7" pageNumber="214">North Carolina</collectingRegion>
(
<figureCitation box="[336,405,2001,2027]" captionStart="FIGURE 6" captionStartId="9.[151,250,1969,1993]" captionTargetBox="[188,1400,1036,1945]" captionTargetId="figure-68@9.[188,1400,1036,1945]" captionTargetPageId="9" captionText="FIGURE 6. Stony Fork, facing south near the type locality of Cambarus lapidosus, in Watauga County, NC—25 October 2019 (photo credit: Michael A. Perkins, NCWRC)." figureDoi="http://doi.org/10.5281/zenodo.7850623" httpUri="https://zenodo.org/record/7850623/files/figure.png" pageId="7" pageNumber="214">Fig. 6</figureCitation>
). This section of Stony Fork is approximately 645 m elevation above sea level, buffered by a secondary forest of intact hardwoods and high canopy cover, approximately 5 m width, 0.25 m in depth, with a clean heterogenous stream bottom composed primarily of gravel, cobble, and boulder; secondarily composed of bedrock, sand, and woody debris/organic material. The
<typeStatus box="[782,880,223,249]" pageId="8" pageNumber="215">holotype</typeStatus>
was collected 29 October 2019
</materialsCitation>
, and the
<typeStatus box="[1346,1436,223,249]" pageId="8" pageNumber="215">allotype</typeStatus>
and morphotype were collected
<date box="[510,717,259,285]" pageId="8" pageNumber="215" value="2017-11-06">6 November 2017</date>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7850619" ID-Zenodo-Dep="7850619" box="[151,1289,1800,1824]" httpUri="https://zenodo.org/record/7850619/files/figure.png" pageId="8" pageNumber="215" startId="8.[151,250,1800,1824]" targetBox="[192,1396,309,1776]" targetPageId="8" targetType="figure">
<paragraph blockId="8.[151,1289,1800,1824]" box="[151,1289,1800,1824]" pageId="8" pageNumber="215">
<emphasis bold="true" box="[151,275,1800,1824]" pageId="8" pageNumber="215">FIGURE 4.</emphasis>
<emphasis box="[281,497,1800,1824]" italics="true" pageId="8" pageNumber="215">
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[281,493,1800,1824]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="215" phylum="Arthropoda" rank="species" species="lapidosus">Cambarus lapidosus</taxonomicName>
,
</emphasis>
holotype (NCSM 90222) in life (photo credit: Michael A. Perkins, NCWRC).
</paragraph>
</caption>
<paragraph blockId="8.[151,1436,1870,2040]" pageId="8" pageNumber="215">
<emphasis bold="true" box="[199,449,1870,1896]" pageId="8" pageNumber="215">Disposition of types.</emphasis>
The
<typeStatus box="[516,617,1870,1896]" pageId="8" pageNumber="215">holotype</typeStatus>
,
<typeStatus box="[633,726,1870,1896]" pageId="8" pageNumber="215">allotype</typeStatus>
, morphotype, and
<specimenCount box="[948,1114,1870,1897]" pageId="8" pageNumber="215" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
are deposited in the
<collectingRegion country="United States of America" name="North Carolina" pageId="8" pageNumber="215">North Carolina</collectingRegion>
<collectingCounty box="[258,1059,1906,1933]" pageId="8" pageNumber="215">Museum of Natural Sciences Non-molluscan Invertebrates Collection</collectingCounty>
, catalog numbers
<specimenCode box="[1270,1432,1906,1932]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="8" pageNumber="215" type="Museum">NCSM 90222</specimenCode>
,
<specimenCode box="[151,224,1942,1968]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="8" pageNumber="215" type="Museum">90223</specimenCode>
,
<specimenCode box="[236,309,1942,1968]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="8" pageNumber="215" type="Museum">90224</specimenCode>
, and 90241, respectively.
<collectingMunicipality box="[603,652,1943,1969]" pageId="8" pageNumber="215">Two</collectingMunicipality>
additional
<typeStatus box="[781,891,1943,1969]" pageId="8" pageNumber="215">paratypes</typeStatus>
, 1 MI and 1 F, are deposited in the
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9EA34CFA8856D7FC2FF851" county="Museum of Natural Sciences Non-molluscan Invertebrates Collection" municipality="Two" name="Smithsonian National Museum of Natural History Invertebrate Zoology Collection" pageId="8" pageNumber="215" stateProvince="North Carolina">Smithsonian National Museum of Natural History Invertebrate Zoology Collection</location>
,
<collectionCode box="[988,1069,1978,2004]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="215" type="Museum">USNM</collectionCode>
<collectionCode box="[1081,1223,1979,2005]" pageId="8" pageNumber="215">XXXXXXX</collectionCode>
and
<collectionCode box="[1288,1432,1979,2005]" pageId="8" pageNumber="215">XXXXXXX</collectionCode>
, respectively.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7850621" ID-Zenodo-Dep="7850621" httpUri="https://zenodo.org/record/7850621/files/figure.png" pageId="9" pageNumber="216" startId="9.[151,250,915,939]" targetBox="[190,1398,181,892]" targetPageId="9" targetType="figure">
<paragraph blockId="9.[151,1436,915,1012]" pageId="9" pageNumber="216">
<emphasis bold="true" box="[151,274,915,940]" pageId="9" pageNumber="216">FIGURE 5.</emphasis>
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[280,489,916,940]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="216" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[280,489,916,940]" italics="true" pageId="9" pageNumber="216">Cambarus lapidosus</emphasis>
</taxonomicName>
, (left) second form male morphotype (NCSM 90224) in life, Stony Fork, Wilkes County, NC; (right) female allotype (NCSM 90223) in life, Orchard Creek, Watauga County, NC (note: regenerated left chela with single row of mesial tubercles); (photo credit: Michael A. Perkins, NCWRC).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7850623" ID-Zenodo-Dep="7850623" httpUri="https://zenodo.org/record/7850623/files/figure.png" pageId="9" pageNumber="216" startId="9.[151,250,1969,1993]" targetBox="[188,1400,1036,1945]" targetPageId="9" targetType="figure">
<paragraph blockId="9.[151,1437,1969,2029]" pageId="9" pageNumber="216">
<emphasis bold="true" box="[151,274,1969,1993]" pageId="9" pageNumber="216">FIGURE 6.</emphasis>
Stony Fork, facing south near the type locality of
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[781,989,1969,1993]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="216" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[781,989,1969,1993]" italics="true" pageId="9" pageNumber="216">Cambarus lapidosus</emphasis>
</taxonomicName>
, in Watauga County, NC—25 October 2019 (photo credit: Michael A. Perkins, NCWRC).
</paragraph>
</caption>
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="217">
<emphasis bold="true" box="[199,596,150,176]" pageId="10" pageNumber="217">Range and specimens examined.</emphasis>
<materialsCitation box="[606,1097,150,177]" county="Watauga County" location="Watauga County" pageId="10" pageNumber="217" specimenCount="1" stateProvince="North Carolina">
<collectingRegion box="[606,859,150,176]" country="United States of America" name="North Carolina" pageId="10" pageNumber="217">NORTH CAROLINA</collectingRegion>
,
<emphasis box="[873,1072,151,177]" italics="true" pageId="10" pageNumber="217">
<collectingCounty box="[873,1066,151,177]" pageId="10" pageNumber="217">Watauga County</collectingCounty>
:
</emphasis>
<emphasis bold="true" box="[1083,1097,150,176]" pageId="10" pageNumber="217">1</emphasis>
</materialsCitation>
<emphasis bold="true" box="[1097,1107,150,176]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectingDate="2019-10-29" collectionCode="NCSM" collectorName="M. A. Perkins" latitude="36.1949" location="Mt. Zion Rd." longLatPrecision="6" longitude="-81.466" municipality="Stony Fork" pageId="10" pageNumber="217" specimenCode="NCSM 90236" specimenCount="1" stateProvince="NORTH CAROLINA">
<emphasis bold="true" box="[1117,1280,150,176]" pageId="10" pageNumber="217">
<specimenCode box="[1117,1280,150,176]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90236</specimenCode>
</emphasis>
<collectingMunicipality box="[1308,1436,150,176]" pageId="10" pageNumber="217">Stony Fork</collectingMunicipality>
at SR 1500 bridge,
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFEEE51FAFDEDFF51" box="[374,526,186,213]" latitude="36.1949" longLatPrecision="6" longitude="-81.466" municipality="Stony Fork" name="Mt. Zion Rd." pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Mt. Zion Rd.</location>
,
<geoCoordinate box="[539,659,186,213]" degrees="36.1949" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.1949">N 36.1949</geoCoordinate>
<geoCoordinate box="[667,796,186,213]" degrees="81.4660" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.466">W 81.4660</geoCoordinate>
, 1 F,
<date box="[860,1054,186,212]" pageId="10" pageNumber="217" value="2019-10-29">
<collectingDate box="[860,1054,186,212]" pageId="10" pageNumber="217" value="2019-10-29">29 October 2019</collectingDate>
</date>
,
<collectorName box="[1066,1221,186,212]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
, W
</materialsCitation>
.
<materialsCitation collectionCode="T" collectorName="W. J. Russ" location="Russ" pageId="10" pageNumber="217" specimenCount="1" stateProvince="NORTH CAROLINA">
<collectionCode box="[1265,1283,187,213]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="217">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFA9751FAFAA6FF50" box="[1295,1349,187,213]" name="Russ" pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Russ</location>
&amp;
<collectorName pageId="10" pageNumber="217">W.J. Russ</collectorName>
;
<emphasis bold="true" box="[221,236,222,248]" pageId="10" pageNumber="217">2</emphasis>
</materialsCitation>
<emphasis bold="true" box="[236,244,222,248]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectingDate="2019-10-29" collectionCode="NCSM" collectorName="M. A. Perkins" latitude="36.1949" location="Mt. Zion Rd." longLatPrecision="6" longitude="-81.466" municipality="Stony Fork" pageId="10" pageNumber="217" specimenCode="NCSM 90222" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="holotype">
<emphasis bold="true" box="[252,529,222,248]" pageId="10" pageNumber="217">
<specimenCode box="[252,412,222,248]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90222</specimenCode>
<typeStatus box="[420,529,222,248]" pageId="10" pageNumber="217">Holotype</typeStatus>
</emphasis>
<collectingMunicipality box="[558,683,222,248]" pageId="10" pageNumber="217">Stony Fork</collectingMunicipality>
at SR 1500 bridge,
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFC16519EFBC7FF7D" box="[910,1060,222,249]" latitude="36.1949" longLatPrecision="6" longitude="-81.466" municipality="Stony Fork" name="Mt. Zion Rd." pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Mt. Zion Rd.</location>
,
<geoCoordinate box="[1072,1191,222,249]" degrees="36.1949" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.1949">N 36.1949</geoCoordinate>
<geoCoordinate box="[1198,1326,222,249]" degrees="81.4660" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.466">W 81.4660</geoCoordinate>
, 1 MI,
<date pageId="10" pageNumber="217" value="2019-10-29">
<collectingDate pageId="10" pageNumber="217" value="2019-10-29">29 October 2019</collectingDate>
</date>
,
<collectorName box="[323,479,258,284]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
, W
</materialsCitation>
.
<materialsCitation box="[523,793,258,285]" collectionCode="T" collectorName="W. J. Russ" location="Russ" pageId="10" pageNumber="217" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="holotype">
<collectionCode box="[523,541,259,285]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="217">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFDB25042FD83FE98" box="[554,608,259,285]" name="Russ" pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Russ</location>
&amp;
<collectorName box="[648,764,259,285]" pageId="10" pageNumber="217">W.J. Russ</collectorName>
;
<emphasis bold="true" box="[778,793,258,284]" pageId="10" pageNumber="217">3</emphasis>
</materialsCitation>
<emphasis bold="true" box="[793,802,258,284]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectingDate="2020-11-19" collectionCode="NCSM" collectorName="M. A. Perkins" county="Orchard Creek" latitude="36.2157" location="Orchard Creek Rd." longLatPrecision="6" longitude="-81.4735" municipality="Bridge" pageId="10" pageNumber="217" specimenCode="NCSM 90241" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="paratype">
<emphasis bold="true" box="[811,1102,258,284]" pageId="10" pageNumber="217">
<specimenCode box="[811,973,258,284]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90241</specimenCode>
<typeStatus box="[982,1102,258,284]" pageId="10" pageNumber="217">Paratypes</typeStatus>
</emphasis>
<collectingCounty box="[1130,1298,258,284]" pageId="10" pageNumber="217">Orchard Creek</collectingCounty>
at SR 1362
<collectingMunicipality box="[151,230,294,320]" pageId="10" pageNumber="217">Bridge</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFF695067FE2AFEC5" box="[241,457,294,320]" county="Orchard Creek" latitude="36.2157" longLatPrecision="6" longitude="-81.4735" municipality="Bridge" name="Orchard Creek Rd." pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Orchard Creek Rd.</location>
,
<geoCoordinate box="[468,586,294,321]" degrees="36.2157" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.2157">N 36.2157</geoCoordinate>
<geoCoordinate box="[593,720,294,321]" degrees="81.4735" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.4735">W 81.4735</geoCoordinate>
, 1 MI, 1 MII, 2 F,
<date box="[928,1147,294,320]" pageId="10" pageNumber="217" value="2020-11-19">
<collectingDate box="[928,1147,294,320]" pageId="10" pageNumber="217" value="2020-11-19">19 November 2020</collectingDate>
</date>
,
<collectorName box="[1158,1311,294,320]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
, W
</materialsCitation>
.
<materialsCitation collectionCode="T" collectorName="W. J. Russ" location="Russ" pageId="10" pageNumber="217" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="paratype">
<collectionCode box="[1354,1372,295,321]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="217">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFAFE5066FA7FFEC4" box="[1382,1436,295,321]" name="Russ" pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Russ</location>
&amp;
<collectorName box="[180,295,331,357]" pageId="10" pageNumber="217">W.J. Russ</collectorName>
;
<emphasis bold="true" box="[307,322,330,356]" pageId="10" pageNumber="217">4</emphasis>
</materialsCitation>
<emphasis bold="true" box="[322,330,330,356]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectingDate="2020-11-19" collectionCode="NCSM" collectorName="M. A. Perkins" county="Orchard Creek" latitude="36.2157" location="Orchard Creek Rd." longLatPrecision="6" longitude="-81.4735" municipality="Bridge" pageId="10" pageNumber="217" specimenCode="NCSM 90242" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="paratype">
<emphasis bold="true" box="[339,499,330,356]" pageId="10" pageNumber="217">
<specimenCode box="[339,499,330,356]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90242</specimenCode>
</emphasis>
<collectingCounty box="[527,694,330,356]" pageId="10" pageNumber="217">Orchard Creek</collectingCounty>
at SR 1362
<collectingMunicipality box="[836,915,330,356]" pageId="10" pageNumber="217">Bridge</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFC07500BFB94FEE1" box="[927,1143,330,356]" county="Orchard Creek" latitude="36.2157" longLatPrecision="6" longitude="-81.4735" municipality="Bridge" name="Orchard Creek Rd." pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Orchard Creek Rd.</location>
,
<geoCoordinate box="[1156,1275,330,357]" degrees="36.2157" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.2157">N 36.2157</geoCoordinate>
<geoCoordinate box="[1282,1410,330,357]" degrees="81.4735" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.4735">W 81.4735</geoCoordinate>
, 1 M juv;
<date box="[233,452,366,392]" pageId="10" pageNumber="217" value="2020-11-19">
<collectingDate box="[233,452,366,392]" pageId="10" pageNumber="217" value="2020-11-19">19 November 2020</collectingDate>
</date>
,
<collectorName box="[463,615,366,392]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
, W
</materialsCitation>
.
<materialsCitation box="[658,914,366,393]" collectionCode="T" collectorName="W. J. Russ" location="Russ" pageId="10" pageNumber="217" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="paratype">
<collectionCode box="[658,676,367,393]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="217">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFD35502EFD00FE0C" box="[685,739,367,393]" name="Russ" pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Russ</location>
&amp;
<collectorName box="[775,888,367,393]" pageId="10" pageNumber="217">W.J. Russ</collectorName>
;
<emphasis bold="true" box="[899,914,366,392]" pageId="10" pageNumber="217">5</emphasis>
</materialsCitation>
<emphasis bold="true" box="[914,923,366,392]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectingDate="2020-11-19" collectionCode="NCSM" collectorName="M. A. Perkins" latitude="36.2109" location="Stony Fork Rd." longLatPrecision="6" longitude="-81.4819" municipality="Stony Fork" pageId="10" pageNumber="217" specimenCode="NCSM 90237" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="paratype">
<emphasis bold="true" box="[929,1088,366,392]" pageId="10" pageNumber="217">
<specimenCode box="[929,1088,366,392]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90237</specimenCode>
</emphasis>
<collectingMunicipality box="[1095,1220,366,392]" pageId="10" pageNumber="217">Stony Fork</collectingMunicipality>
at SR 1505 bridge,
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFF0F50D3FEA4FE29" box="[151,327,402,428]" latitude="36.2109" longLatPrecision="6" longitude="-81.4819" municipality="Stony Fork" name="Stony Fork Rd." pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Stony Fork Rd.</location>
,
<geoCoordinate box="[338,457,402,429]" degrees="36.2109" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.2109">N 36.2109</geoCoordinate>
<geoCoordinate box="[464,591,402,429]" degrees="81.4819" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.4819">W 81.4819</geoCoordinate>
, 2 MI, 1 MII, 1 F, 1 F juv;
<date box="[896,1116,402,428]" pageId="10" pageNumber="217" value="2020-11-19">
<collectingDate box="[896,1116,402,428]" pageId="10" pageNumber="217" value="2020-11-19">19 November 2020</collectingDate>
</date>
,
<collectorName box="[1128,1281,402,428]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
, W
</materialsCitation>
.
<materialsCitation collectionCode="T" collectorName="W. J. Russ" location="Russ" pageId="10" pageNumber="217" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="paratype">
<collectionCode box="[1324,1342,403,429]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="217">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFAD050D2FA9DFE28" box="[1352,1406,403,429]" name="Russ" pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Russ</location>
&amp;
<collectorName box="[151,266,439,465]" pageId="10" pageNumber="217">W.J. Russ</collectorName>
;
<emphasis bold="true" box="[279,294,438,464]" pageId="10" pageNumber="217">6</emphasis>
</materialsCitation>
<emphasis bold="true" box="[294,303,438,464]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectingDate="2020-11-19" collectionCode="NCSM" collectorName="M. A. Perkins" latitude="36.1939" location="Flowers Branch Rd." longLatPrecision="6" longitude="-81.4754" municipality="Flowers Branch" pageId="10" pageNumber="217" specimenCode="NCSM 90238" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="paratype">
<emphasis bold="true" box="[311,499,438,464]" pageId="10" pageNumber="217">
<specimenCode box="[311,472,438,464]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90238</specimenCode>
</emphasis>
<collectingMunicipality box="[500,679,438,464]" pageId="10" pageNumber="217">Flowers Branch</collectingMunicipality>
at SR 1501 bridge,
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFC1750F7FB96FE55" box="[911,1141,438,464]" latitude="36.1939" longLatPrecision="6" longitude="-81.4754" municipality="Flowers Branch" name="Flowers Branch Rd." pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Flowers Branch Rd.</location>
,
<geoCoordinate box="[1154,1273,438,465]" degrees="36.1939" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.1939">N 36.1939</geoCoordinate>
<geoCoordinate box="[1281,1410,438,465]" degrees="81.4754" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.4754">W 81.4754</geoCoordinate>
, 1 MI, 1 MII, 1 F,
<date box="[324,542,474,500]" pageId="10" pageNumber="217" value="2020-11-19">
<collectingDate box="[324,542,474,500]" pageId="10" pageNumber="217" value="2020-11-19">19 November 2020</collectingDate>
</date>
,
<collectorName box="[552,705,474,500]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
, W
</materialsCitation>
.
<materialsCitation box="[747,1002,474,501]" collectionCode="T" collectorName="W. J. Russ" location="Russ" pageId="10" pageNumber="217" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="paratype">
<collectionCode box="[747,765,475,501]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="217">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFC9E509AFCDFFE70" box="[774,828,475,501]" name="Russ" pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Russ</location>
&amp;
<collectorName box="[863,976,475,501]" pageId="10" pageNumber="217">W.J. Russ</collectorName>
;
<emphasis bold="true" box="[987,1002,474,500]" pageId="10" pageNumber="217">7</emphasis>
</materialsCitation>
<emphasis bold="true" box="[1002,1010,474,500]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectingDate="2020-11-19" collectionCode="USNM, XXXXXXX" collectorName="M. A. Perkins" latitude="36.1992" location="Wildcat Rd." longLatPrecision="6" longitude="-81.5013" municipality="Wildcat Creek" pageId="10" pageNumber="217" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="paratype">
<emphasis bold="true" box="[1017,1248,474,500]" pageId="10" pageNumber="217">
<collectionCode box="[1017,1099,474,500]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="217" type="Museum">USNM</collectionCode>
<collectionCode box="[1106,1248,474,500]" pageId="10" pageNumber="217">XXXXXXX</collectionCode>
</emphasis>
<collectingMunicipality box="[1275,1436,474,500]" pageId="10" pageNumber="217">Wildcat Creek</collectingMunicipality>
at SR 1505 bridge,
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFEEE50BFFDE3FD9D" box="[374,512,510,536]" latitude="36.1992" longLatPrecision="6" longitude="-81.5013" municipality="Wildcat Creek" name="Wildcat Rd." pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Wildcat Rd.</location>
,
<geoCoordinate box="[525,645,510,537]" degrees="36.1992" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.1992">N 36.1992</geoCoordinate>
<geoCoordinate box="[653,782,510,537]" degrees="81.5013" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.5013">W 81.5013</geoCoordinate>
, 1 MI, 1 F,
<date box="[918,1141,510,536]" pageId="10" pageNumber="217" value="2020-11-19">
<collectingDate box="[918,1141,510,536]" pageId="10" pageNumber="217" value="2020-11-19">19 November 2020</collectingDate>
</date>
,
<collectorName box="[1153,1308,510,536]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
, W
</materialsCitation>
.
<materialsCitation collectionCode="T" collectorName="W. J. Russ" location="Russ" pageId="10" pageNumber="217" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="paratype">
<collectionCode box="[1352,1370,511,537]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="217">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFAFE50BEFA7FFD9C" box="[1382,1436,511,537]" name="Russ" pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Russ</location>
&amp;
<collectorName box="[180,295,547,573]" pageId="10" pageNumber="217">W.J. Russ</collectorName>
;
<emphasis bold="true" box="[307,322,546,572]" pageId="10" pageNumber="217">8</emphasis>
</materialsCitation>
<emphasis bold="true" box="[322,331,546,572]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectionCode="NCSM" collectorName="F, B. W. Williams, M. A &amp; Perkins, S. G &amp; Anderson" county="Stony Fork" elevation="50" latitude="36.1895" location="Mt. Zion Church Rd." longLatPrecision="6" longitude="-81.4661" municipality="Bridge" pageId="10" pageNumber="217" specimenCode="NCSM 90223" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="allotype">
<emphasis bold="true" box="[338,633,546,572]" pageId="10" pageNumber="217">
<specimenCode box="[338,498,546,572]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90223</specimenCode>
<typeStatus box="[505,606,546,572]" pageId="10" pageNumber="217">Allotype</typeStatus>
</emphasis>
<collectingCounty box="[634,760,546,572]" pageId="10" pageNumber="217">Stony Fork</collectingCounty>
at SR 1500
<collectingMunicipality box="[902,981,546,572]" pageId="10" pageNumber="217">Bridge</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFC795362FB2CFDB9" box="[993,1231,546,573]" county="Stony Fork" latitude="36.1895" longLatPrecision="6" longitude="-81.4661" municipality="Bridge" name="Mt. Zion Church Rd." pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Mt. Zion Church Rd.</location>
approximately
<quantity metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="217" unit="m" value="50.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="217" unit="m" value="50.0">50 m</elevation>
</quantity>
upstream of the
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFEED5307FDC9FDE5" box="[373,554,582,608]" county="Stony Fork" latitude="36.1895" longLatPrecision="6" longitude="-81.4661" municipality="Bridge" name="Flowers Branch" pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Flowers Branch</location>
confluence,
<geoCoordinate box="[703,825,582,609]" degrees="36.1895" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.1895">N 36.1895</geoCoordinate>
<geoCoordinate box="[834,964,582,609]" degrees="81.4661" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.4661">W 81.4661</geoCoordinate>
, 1
<collectorName box="[1002,1272,582,609]" pageId="10" pageNumber="217">F, B.W. Williams, M.A</collectorName>
.
<collectorName box="[1285,1432,582,608]" pageId="10" pageNumber="217">Perkins, S.G</collectorName>
.
<collectorName box="[151,264,618,644]" pageId="10" pageNumber="217">Anderson</collectorName>
;
<emphasis bold="true" box="[274,289,618,644]" pageId="10" pageNumber="217">9</emphasis>
</materialsCitation>
<emphasis bold="true" box="[289,298,618,644]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectionCode="NCSM" collectorName="B. W. Williams &amp; M. A. Perkins &amp; S. G. Anderson" county="Morphotype - Stony Fork" elevation="50" latitude="36.1895" location="Mt. Zion Church Rd." longLatPrecision="6" longitude="-81.4661" municipality="Bridge" pageId="10" pageNumber="217" specimenCode="NCSM 90224" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="allotype">
<emphasis bold="true" box="[304,647,618,644]" pageId="10" pageNumber="217">
<specimenCode box="[304,463,618,644]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90224</specimenCode>
<collectingCounty box="[470,647,618,644]" pageId="10" pageNumber="217">Morphotype—</collectingCounty>
</emphasis>
Stony Fork at SR 1500
<collectingMunicipality box="[909,988,618,644]" pageId="10" pageNumber="217">Bridge</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFC7F532AFB31FD01" box="[999,1234,618,645]" county="Morphotype - Stony Fork" latitude="36.1895" longLatPrecision="6" longitude="-81.4661" municipality="Bridge" name="Mt. Zion Church Rd." pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Mt. Zion Church Rd.</location>
approximately
<quantity metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="217" unit="m" value="50.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="217" unit="m" value="50.0">50 m</elevation>
</quantity>
upstream of the
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFEF653CFFDC2FD2D" box="[366,545,654,680]" county="Morphotype - Stony Fork" latitude="36.1895" longLatPrecision="6" longitude="-81.4661" municipality="Bridge" name="Flowers Branch" pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Flowers Branch</location>
confluence,
<geoCoordinate box="[691,810,654,681]" degrees="36.1895" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.1895">N 36.1895</geoCoordinate>
<geoCoordinate box="[818,946,654,681]" degrees="81.4661" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.4661">W 81.4661</geoCoordinate>
, 1 MII,
<collectorName box="[1039,1207,654,681]" pageId="10" pageNumber="217">B.W. Williams</collectorName>
,
<collectorName box="[1220,1374,654,680]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
,
<collectorName pageId="10" pageNumber="217">S.G. Anderson</collectorName>
;
<emphasis bold="true" box="[275,304,690,716]" pageId="10" pageNumber="217">10</emphasis>
</materialsCitation>
<emphasis bold="true" box="[304,312,690,716]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectionCode="NCSM" collectorName="B. W. Williams &amp; M. A. Perkins &amp; S. G. Anderson" county="Stony Fork" elevation="50" latitude="36.1895" location="Mt. Zion Church Rd." longLatPrecision="6" longitude="-81.4661" municipality="Bridge" pageId="10" pageNumber="217" specimenCode="NCSM 90244" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="allotype">
<emphasis bold="true" box="[319,478,690,716]" pageId="10" pageNumber="217">
<specimenCode box="[319,478,690,716]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90244</specimenCode>
</emphasis>
<collectingCounty box="[507,631,690,716]" pageId="10" pageNumber="217">Stony Fork</collectingCounty>
at SR 1500
<collectingMunicipality box="[769,848,690,716]" pageId="10" pageNumber="217">Bridge</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFCC353F2FBA4FD49" box="[859,1095,690,717]" county="Stony Fork" latitude="36.1895" longLatPrecision="6" longitude="-81.4661" municipality="Bridge" name="Mt. Zion Church Rd." pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Mt. Zion Church Rd.</location>
approximately
<quantity box="[1270,1327,690,717]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="217" unit="m" value="50.0">
<elevation box="[1270,1327,690,717]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="217" unit="m" value="50.0">50 m</elevation>
</quantity>
upstream of the
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFF475397FE72FD75" box="[223,401,726,752]" county="Stony Fork" latitude="36.1895" longLatPrecision="6" longitude="-81.4661" municipality="Bridge" name="Flowers Branch" pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Flowers Branch</location>
confluence,
<geoCoordinate box="[545,664,726,753]" degrees="36.1895" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.1895">N 36.1895</geoCoordinate>
<geoCoordinate box="[670,797,726,753]" degrees="81.4661" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.4661">W 81.4661</geoCoordinate>
, 1 MI,
<collectorName box="[878,1045,726,753]" pageId="10" pageNumber="217">B.W. Williams</collectorName>
,
<collectorName box="[1057,1210,726,752]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
,
<collectorName box="[1221,1390,726,752]" pageId="10" pageNumber="217">S.G. Anderson</collectorName>
;
<emphasis bold="true" box="[1400,1428,726,752]" pageId="10" pageNumber="217">11</emphasis>
</materialsCitation>
<emphasis bold="true" box="[1428,1436,726,752]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectingDate="1996-07-23" collectionCode="NCSM" collectorName="L. Eaton &amp; N. Medlin" county="Stony Fork" location="Hayes Welborn Rd" municipality="Mt. Zion Rd" pageId="10" pageNumber="217" specimenCode="NCSM 3314" specimenCount="1" stateProvince="NORTH CAROLINA" typeStatus="allotype">
<emphasis bold="true" box="[151,298,762,788]" pageId="10" pageNumber="217">
<specimenCode box="[151,298,762,788]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 3314</specimenCode>
</emphasis>
<collectingCounty box="[327,452,762,788]" pageId="10" pageNumber="217">Stony Fork</collectingCounty>
at intersection of
<collectingMunicipality box="[659,801,762,789]" pageId="10" pageNumber="217">Mt. Zion Rd</collectingMunicipality>
(SR 1500) and
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFC4F53BAFB4DFC91" box="[983,1198,762,789]" county="Stony Fork" municipality="Mt. Zion Rd" name="Hayes Welborn Rd" pageId="10" pageNumber="217" stateProvince="NORTH CAROLINA">Hayes Welborn Rd</location>
(SR 1505), 2 F, 2 M juv,
<date box="[199,347,798,824]" pageId="10" pageNumber="217" value="1996-07-23">
<collectingDate box="[199,347,798,824]" pageId="10" pageNumber="217" value="1996-07-23">23 July 1996</collectingDate>
</date>
,
<collectorName box="[358,454,799,825]" pageId="10" pageNumber="217">L. Eaton</collectorName>
&amp;
<collectorName box="[490,609,798,825]" pageId="10" pageNumber="217">N. Medlin</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="217">
<emphasis box="[199,361,834,861]" italics="true" pageId="10" pageNumber="217">Wilkes County</emphasis>
:
<emphasis bold="true" box="[377,414,834,860]" pageId="10" pageNumber="217">12)</emphasis>
<materialsCitation collectingDate="2020-11-19" collectionCode="NCSM" collectorName="M. A. Perkins" latitude="36.1773" location="Stony Fork" longLatPrecision="6" longitude="-81.4654" municipality="Stony Fork Rd." pageId="10" pageNumber="217" specimenCode="NCSM 90232" specimenCount="1">
<emphasis bold="true" box="[422,582,834,860]" pageId="10" pageNumber="217">
<specimenCode box="[422,582,834,860]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90232</specimenCode>
</emphasis>
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFDFB5203FD02FCD9" box="[611,737,834,860]" latitude="36.1773" longLatPrecision="6" longitude="-81.4654" municipality="Stony Fork Rd." name="Stony Fork" pageId="10" pageNumber="217">Stony Fork</location>
at CR 1167 bridge,
<collectingMunicipality box="[967,1144,834,860]" pageId="10" pageNumber="217">Stony Fork Rd.</collectingMunicipality>
,
<geoCoordinate box="[1156,1275,834,861]" degrees="36.1773" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.1773">N 36.1773</geoCoordinate>
<geoCoordinate box="[1282,1410,834,861]" degrees="81.4654" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.4654">W 81.4654</geoCoordinate>
, 1 MI, 1 F, 1 M juv,
<date box="[347,565,870,896]" pageId="10" pageNumber="217" value="2020-11-19">
<collectingDate box="[347,565,870,896]" pageId="10" pageNumber="217" value="2020-11-19">19 November 2020</collectingDate>
</date>
,
<collectorName box="[576,729,870,896]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
, W
</materialsCitation>
.
<materialsCitation box="[771,1041,870,897]" collectionCode="T" collectorName="W. J. Russ" location="Russ" pageId="10" pageNumber="217" specimenCount="1">
<collectionCode box="[771,789,871,897]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="217">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFC875226FCB6FC04" box="[799,853,871,897]" name="Russ" pageId="10" pageNumber="217">Russ</location>
&amp;
<collectorName box="[888,1001,871,897]" pageId="10" pageNumber="217">W.J. Russ</collectorName>
;
<emphasis bold="true" box="[1012,1041,870,896]" pageId="10" pageNumber="217">13</emphasis>
</materialsCitation>
<emphasis bold="true" box="[1041,1049,870,896]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectingDate="2020-11-19" collectionCode="NCSM" collectorName="M. A. Perkins" county="Stony Fork Falls" elevation="50" latitude="36.1695" location="Stony Fork" longLatPrecision="6" longitude="-81.4659" municipality="Stony Fork Rd." pageId="10" pageNumber="217" specimenCode="NCSM 90233" specimenCount="1">
<emphasis bold="true" box="[1056,1215,870,896]" pageId="10" pageNumber="217">
<specimenCode box="[1056,1215,870,896]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90233</specimenCode>
</emphasis>
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFB445227FABBFC05" box="[1244,1368,870,896]" county="Stony Fork Falls" latitude="36.1695" longLatPrecision="6" longitude="-81.4659" municipality="Stony Fork Rd." name="Stony Fork" pageId="10" pageNumber="217">Stony Fork</location>
at SR 1167 bridge,
<collectingMunicipality box="[299,473,906,932]" pageId="10" pageNumber="217">Stony Fork Rd.</collectingMunicipality>
, approximately
<quantity box="[653,710,906,933]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="217" unit="m" value="50.0">
<elevation box="[653,710,906,933]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="217" unit="m" value="50.0">50 m</elevation>
</quantity>
downstream of
<collectingCounty box="[890,1080,906,932]" pageId="10" pageNumber="217">Stony Fork Falls</collectingCounty>
,
<geoCoordinate box="[1091,1209,906,933]" degrees="36.1695" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.1695">N 36.1695</geoCoordinate>
<geoCoordinate box="[1215,1342,906,933]" degrees="81.4659" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.4659">W 81.4659</geoCoordinate>
, 1 MI, 2 F, 2 M juv,
<date box="[280,500,942,968]" pageId="10" pageNumber="217" value="2020-11-19">
<collectingDate box="[280,500,942,968]" pageId="10" pageNumber="217" value="2020-11-19">19 November 2020</collectingDate>
</date>
,
<collectorName box="[511,664,942,968]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
, W
</materialsCitation>
.
<materialsCitation box="[707,980,942,969]" collectionCode="T" collectorName="W. J. Russ" location="Russ" pageId="10" pageNumber="217" specimenCount="1">
<collectionCode box="[707,725,943,969]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="217">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFD4752EEFCF6FC4C" box="[735,789,943,969]" name="Russ" pageId="10" pageNumber="217">Russ</location>
&amp;
<collectorName box="[825,939,943,969]" pageId="10" pageNumber="217">W.J. Russ</collectorName>
;
<emphasis bold="true" box="[951,980,942,968]" pageId="10" pageNumber="217">14</emphasis>
</materialsCitation>
<emphasis bold="true" box="[980,988,942,968]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectingDate="2020-11-19" collectionCode="NCSM" collectorName="M. A. Perkins" latitude="36.1504" location="Left Prong Stony Fork" longLatPrecision="6" longitude="-81.4652" municipality="Lee Mountain Rd." pageId="10" pageNumber="217" specimenCode="NCSM 90234" specimenCount="1">
<emphasis bold="true" box="[995,1155,942,968]" pageId="10" pageNumber="217">
<specimenCode box="[995,1155,942,968]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90234</specimenCode>
</emphasis>
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFB0752EFFA7FFC4D" box="[1183,1436,942,969]" latitude="36.1504" longLatPrecision="6" longitude="-81.4652" municipality="Lee Mountain Rd." name="Left Prong Stony Fork" pageId="10" pageNumber="217">Left Prong Stony Fork</location>
at SR 1155,
<collectingMunicipality box="[289,497,978,1005]" pageId="10" pageNumber="217">Lee Mountain Rd.</collectingMunicipality>
,
<geoCoordinate box="[509,627,978,1005]" degrees="36.1504" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.1504">N 36.1504</geoCoordinate>
<geoCoordinate box="[634,761,978,1005]" degrees="81.4652" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.4652">W 81.4652</geoCoordinate>
, 2 MI,
<date box="[842,1062,978,1004]" pageId="10" pageNumber="217" value="2020-11-19">
<collectingDate box="[842,1062,978,1004]" pageId="10" pageNumber="217" value="2020-11-19">19 November 2020</collectingDate>
</date>
,
<collectorName box="[1073,1226,978,1004]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
, W
</materialsCitation>
.
<materialsCitation collectionCode="T" collectorName="W. J. Russ" location="Russ" pageId="10" pageNumber="217" specimenCount="1">
<collectionCode box="[1269,1287,979,1005]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="217">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFA895292FAA4FC68" box="[1297,1351,979,1005]" name="Russ" pageId="10" pageNumber="217">Russ</location>
&amp;
<collectorName pageId="10" pageNumber="217">W.J. Russ</collectorName>
;
<emphasis bold="true" box="[221,250,1014,1040]" pageId="10" pageNumber="217">15</emphasis>
</materialsCitation>
<emphasis bold="true" box="[250,258,1014,1040]" pageId="10" pageNumber="217">)</emphasis>
<materialsCitation collectingDate="2020-11-20" collectionCode="NCSM" collectorName="M. A. Perkins &amp; B. W. Williams" latitude="36.1561" location="Stony Fork" longLatPrecision="6" longitude="-81.4563" municipality="Stony Fork Rd." pageId="10" pageNumber="217" specimenCode="NCSM 90231" specimenCount="1">
<emphasis bold="true" box="[266,426,1014,1040]" pageId="10" pageNumber="217">
<specimenCode box="[266,426,1014,1040]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="10" pageNumber="217" type="Museum">NCSM 90231</specimenCode>
</emphasis>
<location LSID="urn:lsid:plazi:treatment:360887F0FF92A34FFF0F5735FDBFF98C:BB7E603DFF9CA34EFE5F52B7FDA7FB95" box="[455,580,1014,1040]" latitude="36.1561" longLatPrecision="6" longitude="-81.4563" municipality="Stony Fork Rd." name="Stony Fork" pageId="10" pageNumber="217">Stony Fork</location>
at SR 1167,
<collectingMunicipality box="[727,903,1014,1040]" pageId="10" pageNumber="217">Stony Fork Rd.</collectingMunicipality>
,
<geoCoordinate box="[915,1033,1014,1041]" degrees="36.1561" direction="north" orientation="latitude" pageId="10" pageNumber="217" precision="5" value="36.1561">N 36.1561</geoCoordinate>
<geoCoordinate box="[1041,1168,1014,1041]" degrees="81.4563" direction="west" orientation="longitude" pageId="10" pageNumber="217" precision="5" value="-81.4563">W 81.4563</geoCoordinate>
, 1 M juv,
<date pageId="10" pageNumber="217" value="2020-11-20">
<collectingDate pageId="10" pageNumber="217" value="2020-11-20">20 November 2020</collectingDate>
</date>
,
<collectorName box="[221,371,1050,1076]" pageId="10" pageNumber="217">M.A. Perkins</collectorName>
&amp;
<collectorName box="[407,575,1050,1077]" pageId="10" pageNumber="217">B.W. Williams</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="217">
<emphasis bold="true" box="[199,444,1086,1112]" pageId="10" pageNumber="217">Conservation status.</emphasis>
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[452,683,1086,1112]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="217" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[452,683,1086,1112]" italics="true" pageId="10" pageNumber="217">Cambarus lapidosus</emphasis>
</taxonomicName>
is locally common but narrowly distributed in the headwaters of a single small watershed in western
<collectingRegion box="[541,713,1122,1148]" country="United States of America" name="North Carolina" pageId="10" pageNumber="217">North Carolina</collectingRegion>
, with an estimated current extent of approximately
<quantity box="[1298,1370,1122,1149]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="10" pageNumber="217" unit="km" value="85.0">85 km</quantity>
<superScript attach="left" box="[1370,1378,1122,1136]" fontSize="6" pageId="10" pageNumber="217">2</superScript>
. The population size for
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[368,507,1158,1185]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="217" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[368,507,1158,1185]" italics="true" pageId="10" pageNumber="217">C. lapidosus</emphasis>
</taxonomicName>
is currently unknown. While an assessment of threats to the stability of the species is incomplete at this time, habitat degradation and diminished water quality from agriculture and silviculture impact some areas of the watershed, and necessitate future monitoring. Because of these threats, coupled with the species narrow distribution, we suggest that the species be assigned the status of G1/S1 (Endangered) based on NatureServe criteria (NatureServe 2020;) and Endangered (
<collectionCode box="[682,722,1302,1328]" pageId="10" pageNumber="217">EN</collectionCode>
) based on International Union for the Conservation of Nature criteria (
<bibRefCitation author="IUCN" box="[245,381,1338,1364]" pageId="10" pageNumber="217" refId="ref13170" refString="IUCN (2001) Categories and Criteria. Version 3.1. Available from: http: // www. iucnredlist. org / static / categories _ criteria _ 3 _ 1 (accessed 9 November 2022)" type="url" year="2001">IUCN 2001</bibRefCitation>
; B2 a,b
<emphasis box="[473,510,1339,1364]" italics="true" pageId="10" pageNumber="217">i-iv</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="218" pageId="10" pageNumber="217" type="description">
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="217">
<emphasis bold="true" box="[199,319,1374,1400]" pageId="10" pageNumber="217">Variation.</emphasis>
Approximately 88% of the 26 individuals examined possess at least one subpalmar tubercle on the ventral palm of each chelae; 4% of individuals examined possess two subpalmar tubercles. Juveniles and sub-adults lack well-developed chelae making detection of subpalmar tubercles—a key character in identification of this species—difficult. As with other species in the genus
<taxonomicName authorityName="Erichson" authorityYear="1846" box="[812,929,1482,1508]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="217" phylum="Arthropoda" rank="genus">
<emphasis box="[812,929,1482,1508]" italics="true" pageId="10" pageNumber="217">Cambarus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[944,1085,1482,1509]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="217" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[944,1085,1482,1509]" italics="true" pageId="10" pageNumber="217">C. lapidosus</emphasis>
</taxonomicName>
exhibits sexual dimorphism in reproductive structures. We have not observed any obvious spatial variation in this species.
</paragraph>
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="217">
<emphasis bold="true" box="[199,347,1554,1580]" pageId="10" pageNumber="217">Life history.</emphasis>
Collections of
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[527,669,1554,1581]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="217" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[527,669,1554,1581]" italics="true" pageId="10" pageNumber="217">C. lapidosus</emphasis>
</taxonomicName>
have been limited to the months of July, October, and November. First and second form males have been observed in October and November. Females in glare have been observed in November and December. Young-of-year have not been observed. The species appears to prefer cool, clean 2
<superScript attach="left" box="[1392,1408,1626,1640]" fontSize="6" pageId="10" pageNumber="217">nd</superScript>
to 3
<superScript attach="left" box="[165,179,1662,1676]" fontSize="6" pageId="10" pageNumber="217">rd</superScript>
order streams and has been collected almost exclusively from under large cobbles (approx.
<quantity box="[1220,1290,1662,1689]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="217" unit="cm" value="25.0">25 cm</quantity>
in length) to small boulders (approx.
<quantity box="[422,505,1698,1725]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="217" unit="cm" value="100.0">100 cm</quantity>
in length) in moderately deep riffles or runs (
<quantity box="[1012,1077,1698,1725]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="217" unit="m" value="0.1">0.1 m</quantity>
to
<quantity box="[1112,1176,1698,1725]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="217" unit="m" value="0.5">0.5 m</quantity>
) with moderate to high flow and high canopy cover.
</paragraph>
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="217">
<emphasis bold="true" box="[199,437,1770,1796]" pageId="10" pageNumber="217">Crayfish associates.</emphasis>
Several native stream-dwelling associate crayfish species are thought to occur with
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="217" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis italics="true" pageId="10" pageNumber="217">C. lapidosus</emphasis>
</taxonomicName>
including
<emphasis box="[383,409,1807,1833]" italics="true" pageId="10" pageNumber="217">C.</emphasis>
species C, and members of the
<taxonomicName authority="Girard, 1852" authorityName="Girard" authorityYear="1852" box="[783,1160,1806,1833]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="217" phylum="Arthropoda" rank="species" species="longulus">
<emphasis box="[783,1005,1806,1832]" italics="true" pageId="10" pageNumber="217">Cambarus longulus</emphasis>
Girard, 1852
</taxonomicName>
and
<taxonomicName authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="217" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis box="[1220,1436,1806,1832]" italics="true" pageId="10" pageNumber="217">Cambarus bartonii</emphasis>
(Fabricius, 1798)
</taxonomicName>
species complexes.
<taxonomicName authority="Faxon, 1884" authorityName="Faxon" authorityYear="1884" box="[581,1001,1842,1869]" class="Malacostraca" family="Cambaridae" genus="Cambarus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="217" phylum="Arthropoda" rank="species" species="dubius">
<emphasis box="[581,698,1842,1868]" italics="true" pageId="10" pageNumber="217">Cambarus</emphasis>
[aff.]
<emphasis box="[772,847,1842,1868]" italics="true" pageId="10" pageNumber="217">dubius</emphasis>
Faxon, 1884
</taxonomicName>
, a primary burrower, was observed in burrows along stream banks.
</paragraph>
<paragraph blockId="10.[151,1437,150,2013]" lastBlockId="11.[151,1437,151,1545]" lastPageId="11" lastPageNumber="218" pageId="10" pageNumber="217">
<emphasis bold="true" box="[199,578,1914,1940]" pageId="10" pageNumber="217">Relationships and comparisons.</emphasis>
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[585,815,1914,1940]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="217" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[585,815,1914,1940]" italics="true" pageId="10" pageNumber="217">Cambarus lapidosus</emphasis>
</taxonomicName>
is superficially similar to several described species and yet-diagnosed members of the genus
<taxonomicName authorityName="Erichson" authorityYear="1846" box="[574,691,1950,1976]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="217" phylum="Arthropoda" rank="genus">
<emphasis box="[574,691,1950,1976]" italics="true" pageId="10" pageNumber="217">Cambarus</emphasis>
</taxonomicName>
in Atlantic Slope and interior basin streams.
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[1204,1436,1950,1976]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="217" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[1204,1436,1950,1976]" italics="true" pageId="10" pageNumber="217">Cambarus lapidosus</emphasis>
</taxonomicName>
was previously subsumed within
<emphasis box="[526,552,1987,2013]" italics="true" pageId="10" pageNumber="217">C.</emphasis>
species C, the dominant species complex occuring in Atlantic Slope drainages in the Mountain and Piedmont ecoregions of NC.
<taxonomicName box="[703,931,151,177]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="undefined-C">
<emphasis box="[703,820,151,177]" italics="true" pageId="11" pageNumber="218">Cambarus</emphasis>
species C
</taxonomicName>
from the upper Yadkin and adjacent Catawba basins in NC represents at least one morphologically distinct species and can be differentiated from
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[1262,1401,187,213]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[1262,1401,187,213]" italics="true" pageId="11" pageNumber="218">C. lapidosus</emphasis>
</taxonomicName>
by the absence of subpalmar tubercles on the palm of the chelae and the presence of conspicuous cervical spines. The lower reaches and small tributaries of the Stony Fork watershed are dominated by
<emphasis box="[1063,1089,259,285]" italics="true" pageId="11" pageNumber="218">C.</emphasis>
species C, from approximately downstream of Stony Fork Falls to near the confluence with the mainstem Yadkin River; however, the two taxa do co-occur in some reaches downstream, and in the proximity, of Stony Fork Falls, and in the lower Left Prong Stony Fork. Stony Fork Falls appears to be a barrier to upstream migration of
<emphasis box="[1030,1055,367,393]" italics="true" pageId="11" pageNumber="218">C.</emphasis>
species C, but not to downstream movement of
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[307,446,403,429]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[307,446,403,429]" italics="true" pageId="11" pageNumber="218">C. lapidosus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="11.[151,1437,151,1545]" pageId="11" pageNumber="218">
Despite its historic affiliation with
<emphasis box="[584,609,439,465]" italics="true" pageId="11" pageNumber="218">C.</emphasis>
species C,
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[733,872,439,465]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[733,872,439,465]" italics="true" pageId="11" pageNumber="218">C. lapidosus</emphasis>
</taxonomicName>
is more morphologically and genetically similar to undiagnosed forms of
<taxonomicName box="[399,568,475,501]" class="Malacostraca" family="Cambaridae" genus="Cambarus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="robustus">
<emphasis box="[399,424,475,501]" italics="true" pageId="11" pageNumber="218">C.</emphasis>
aff.
<emphasis box="[473,568,475,501]" italics="true" pageId="11" pageNumber="218">robustus</emphasis>
</taxonomicName>
occurring in interior basins of the NC mountain region. These yet undescribed forms—from the New and Watauga River basins—both possess a more widely-punctate areola (810 punctations) and sometimes lack the subpalmar tubercles on the palm of the chelae more often-exhibited by
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[1285,1429,547,573]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[1285,1429,547,573]" italics="true" pageId="11" pageNumber="218">C. lapidosus</emphasis>
</taxonomicName>
. Futhermore,
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[300,444,583,609]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[300,444,583,609]" italics="true" pageId="11" pageNumber="218">C. lapidosus</emphasis>
</taxonomicName>
is not known to occur in any interior basin streams, and is therefore geographically isolated from other taxa currently subsumed within the
<taxonomicName box="[772,899,619,645]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="robustus">
<emphasis box="[772,899,619,645]" italics="true" pageId="11" pageNumber="218">C. robustus</emphasis>
</taxonomicName>
complex.
</paragraph>
<paragraph blockId="11.[151,1437,151,1545]" pageId="11" pageNumber="218">
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[199,436,655,681]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[199,436,655,681]" italics="true" pageId="11" pageNumber="218">Cambarus lapidosus</emphasis>
</taxonomicName>
is genetically most similar to
<emphasis box="[809,981,655,681]" italics="true" pageId="11" pageNumber="218">
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[809,976,655,681]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="burchfielae">C. burchfielae</taxonomicName>
,
</emphasis>
described below, which occurs in the neighboring Lewis Fork drainage. These species are easily morphologically distinguished from each other despite their genetic and geographic affinities.
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[614,870,727,753]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="burchfielae">
<emphasis box="[614,870,727,753]" italics="true" pageId="11" pageNumber="218">Cambarus burchfielae</emphasis>
</taxonomicName>
typically lacks a double row of well-developed cristiform tubercles on the mesial margin of the chelae and tubercles on the dorsal surface of the dactyl are absent or greatly reduced, possesses a narrower and more-sparsely punctate areola (45 punctations), and typically lacks the rusty pigment patch on the dorsal surface of the carapace immediately anterior of the abdomen common to
<emphasis box="[151,334,871,897]" italics="true" pageId="11" pageNumber="218">
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[151,295,871,897]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="lapidosus">C. lapidosus</taxonomicName>
, C.
</emphasis>
species C, and
<taxonomicName authority="Cooper, 2006" authorityName="Cooper" authorityYear="2006" box="[515,859,871,897]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="johni">
<emphasis box="[515,698,871,897]" italics="true" pageId="11" pageNumber="218">Cambarus johni</emphasis>
Cooper, 2006
</taxonomicName>
. These species can also be differentiated by subtle differences in gonopod morphology (see
<figureCitation box="[613,690,907,933]" captionStart="FIGURE 3" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[169,1419,669,1922]" captionTargetId="figure-243@6.[169,1419,669,1922]" captionTargetPageId="6" captionText="FIGURE 3. Cambarus lapidosus, new species. (A.) Lateral view of carapace; (B.) dorsal view of carapace; (C.) right antennal scale; (D.) dorsal view of carpus and chela; (E.) epistome; (F.) lateral and (G.) mesial view of form I gonopod; (H.) lateral and (I.) mesial view of form II gonopod; (J.) annulus ventralis.AE, FG from holotype; HI from morphotype; J from allotype." figureDoi="http://doi.org/10.5281/zenodo.7850617" httpUri="https://zenodo.org/record/7850617/files/figure.png" pageId="11" pageNumber="218">Figs. 3</figureCitation>
&amp;
<figureCitation box="[726,742,907,933]" captionStart="FIGURE 7" captionStartId="13.[151,250,1888,1912]" captionTargetBox="[151,1436,525,1865]" captionTargetId="figure-176@13.[151,1436,525,1865]" captionTargetPageId="13" captionText="FIGURE 7. Cambarus burchfielae, new species. (A.) Lateral view of carapace; (B.) dorsal view of carapace; (C.) right antennal scale; (D.) dorsal view of carpus and chela; (E.) epistome; (F.) lateral and (G.) mesial view of form I gonopod; (H.) lateral and (I.) mesial view of form II gonopod; (J.) annulus ventralis.AE, FG from holotype; HI from morphotype; J from allotype." figureDoi="http://doi.org/10.5281/zenodo.7850625" httpUri="https://zenodo.org/record/7850625/files/figure.png" pageId="11" pageNumber="218">7</figureCitation>
); the central projection of the form I gonopod of
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[1296,1436,907,933]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[1296,1436,907,933]" italics="true" pageId="11" pageNumber="218">C. lapidosus</emphasis>
</taxonomicName>
is longer and more strongly-curved than
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[612,772,943,969]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="burchfielae">
<emphasis box="[612,772,943,969]" italics="true" pageId="11" pageNumber="218">C. burchfielae</emphasis>
</taxonomicName>
. Conversely, the central projection of the form II gonopod of
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[182,343,979,1005]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="burchfielae">
<emphasis box="[182,343,979,1005]" italics="true" pageId="11" pageNumber="218">C. burchfielae</emphasis>
</taxonomicName>
extends beyond the margin of the mesial process whereas the processes are approximately equal in length in
<emphasis box="[286,433,1015,1041]" italics="true" pageId="11" pageNumber="218">
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[286,429,1015,1041]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="lapidosus">C. lapidosus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph blockId="11.[151,1437,151,1545]" pageId="11" pageNumber="218">
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[199,434,1051,1077]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[199,434,1051,1077]" italics="true" pageId="11" pageNumber="218">Cambarus lapidosus</emphasis>
</taxonomicName>
is easily distinguished from other congeners in the upper Yadkin River basin.
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1798" class="Malacostraca" family="Cambaridae" genus="Cambarus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="bartonii">
<emphasis box="[1362,1388,1051,1077]" italics="true" pageId="11" pageNumber="218">C.</emphasis>
aff.
<emphasis box="[151,241,1087,1113]" italics="true" pageId="11" pageNumber="218">bartonii</emphasis>
</taxonomicName>
is a small species characterized by a wide and truncate rostrum, single row of tubercles on the mesial margin of the chelae, and narrow sparsely punctate areola (34 punctations).
<taxonomicName authorityName="Girard" authorityYear="1852" box="[1045,1315,1123,1149]" class="Malacostraca" family="Cambaridae" genus="Cambarus" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="longulus">
<emphasis box="[1045,1162,1123,1149]" italics="true" pageId="11" pageNumber="218">Cambarus</emphasis>
aff.
<emphasis box="[1219,1315,1123,1149]" italics="true" pageId="11" pageNumber="218">longulus</emphasis>
</taxonomicName>
is a small species easily characterized by conspicuous tufts of setae at the base of the opposable margin of the propodus and a narrow rostrum with an elongated acumen; the species is not known to occur upstream of Stony Fork Falls.
<taxonomicName authorityName="Cooper" authorityYear="2006" box="[151,334,1231,1257]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="johni">
<emphasis box="[151,334,1231,1257]" italics="true" pageId="11" pageNumber="218">Cambarus johni</emphasis>
</taxonomicName>
lacks subpalmar tubercles on the chelae and possesses a wider and more-heavily punctate areola (&gt;10 punctations), conspicuous abdominal stripes, a much wider zygoma, and smaller chelae with an enlarged first denticle on the opposable margin of the dactyl; the species has not been found with
<taxonomicName authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" box="[1092,1232,1303,1329]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[1092,1232,1303,1329]" italics="true" pageId="11" pageNumber="218">C. lapidosus</emphasis>
</taxonomicName>
but may co-occur downstream of Stony Fork Falls.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="218" type="etymology">
<paragraph blockId="11.[151,1437,151,1545]" pageId="11" pageNumber="218">
<emphasis bold="true" box="[199,334,1375,1401]" pageId="11" pageNumber="218">Etymology.</emphasis>
The species epithet “
<taxonomicName authority="(Lat.)" authorityName="Perkins &amp; Williams &amp; Russ" authorityYear="2023" baseAuthorityName="Lat." box="[572,679,1375,1401]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="218" phylum="Arthropoda" rank="species" species="lapidosus">
<emphasis box="[572,679,1375,1401]" italics="true" pageId="11" pageNumber="218">lapidosus</emphasis>
</taxonomicName>
” (Lat.) as an adjective in the nominative case means stony or rocky, which we use in reference to the Stony Fork, the small tributary of the Yadkin River in western
<collectingRegion box="[1265,1436,1411,1437]" country="United States of America" name="North Carolina" pageId="11" pageNumber="218">North Carolina</collectingRegion>
which harbors the entire known range of this species.
</paragraph>
<paragraph blockId="11.[151,1437,151,1545]" pageId="11" pageNumber="218">
<emphasis bold="true" box="[199,392,1482,1509]" pageId="11" pageNumber="218">Common name.</emphasis>
We propose Stony Fork Crayfish as the common name for this species in reference to its occurrence in the Stony Fork watershed.
</paragraph>
</subSubSection>
</treatment>
</document>