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<document id="8A6948924E7F7B7606B286AFB193DC37" ID-CLB-Dataset="21808" ID-DOI="10.5852/ejt.2014.100" ID-GBIF-Dataset="5329bfc4-61b2-48f5-a96a-caf4ba31039b" ID-ISSN="2118-9773" ID-Zenodo-Dep="3860450" ID-ZooBank="B3E6C489-6D96-4AF5-A33D-EE8329A9321B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1590595554426" checkinUser="carolina" docAuthor="Enghoff, Henrik" docDate="2014" docId="334F2769C170FF971C6AF339FBCCFAE6" docLanguage="en" docName="ejt-2014-100.pdf.imf" docOrigin="European Journal of Taxonomy 100" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chaleponcus nikolajscharffi Enghoff 2014, sp. nov." docType="treatment" docUuid="FA4BA44E-0547-4E9C-A4A4-781E2298A650" docUuidSource="ZooBank" docVersion="7" lastPageNumber="32" masterDocId="CF765F11C16DFF881E57F137FFA0FFA3" masterDocTitle="A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="30" updateTime="1698841637397" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FC75824F2ADE13AE76D0ADE710C51D18">A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:namePart id="3B01912B7C652516B9C81567993C012F">Enghoff, Henrik</mods:namePart>
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<mods:title id="9F65E4A573870FBFB93E73405C1C8BAE">European Journal of Taxonomy</mods:title>
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<mods:date id="3746F3C3D7277973557EC80B2B4E10D3">2014</mods:date>
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<mods:number id="9A6108A64AEA5D4F0E768A4664F42E8B">2014-10-24</mods:number>
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<treatment id="334F2769C170FF971C6AF339FBCCFAE6" ID-DOI="http://doi.org/10.5281/zenodo.3861171" ID-GBIF-Taxon="164362771" ID-Zenodo-Dep="3861171" LSID="urn:lsid:zoobank.org:act:FA4BA44E-0547-4E9C-A4A4-781E2298A650" httpUri="http://treatment.plazi.org/id/334F2769C170FF971C6AF339FBCCFAE6" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
<subSubSection id="F3FCC5F4C170FF951C6AF339FC56FD8A" box="[573,1014,526,553]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="BB59967FC170FF951C6AF339FC56FD8A" blockId="29.[367,1221,526,623]" box="[573,1014,526,553]" pageId="29" pageNumber="30">
<heading id="E0112113C170FF951C6AF339FC56FD8A" box="[573,1014,526,553]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="30" reason="2">
<taxonomicName id="7CE6EDFCC170FF951C6AF339FC35FD8A" ID-CoL="TH76" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[573,917,526,553]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="nikolajscharffi" status="sp. nov.">
<emphasis id="89924A6DC170FF951C6AF339FC35FD8A" bold="true" box="[573,917,526,553]" italics="true" pageId="29" pageNumber="30">Chaleponcus nikolajscharffi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C170FF951DCBF338FC56FD8A" box="[924,1014,527,553]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C170FF951F38F306FCCAFDCC" pageId="29" pageNumber="30" type="description">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="BB59967FC170FF951C9EF363FCCAFDCC" blockId="29.[367,1221,526,623]" box="[713,874,596,623]" pageId="29" pageNumber="30">
<figureCitation id="23DD8AFAC170FF951C9EF363FCB4FDCC" box="[713,788,596,623]" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The transverse plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="29" pageNumber="30">Figs 9</figureCitation>
,
<figureCitation id="23DD8AFAC170FF951D76F363FCE1FDCD" box="[801,833,596,622]" captionStart="Fig" captionStartId="30.[189,232,1896,1922]" captionTargetBox="[189,1399,435,1852]" captionTargetId="figure@30.[189,1399,435,1852]" captionTargetPageId="30" captionText="Fig. 16. Chaleponcus nikolajscharffi sp. nov., right gonopod. A. Mesal view. B. Anterior view. C. Ventral view. D. Posterior view. al = anterior lamella of telomere, cu = cucullus, lp = lateral coxal process, mf = metaplical flange, mp = mesal metaplical process, msl = metaplical shelf-forming lobe, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860486" httpUri="https://zenodo.org/record/3860486/files/figure.png" pageId="29" pageNumber="30">16</figureCitation>
,
<figureCitation id="23DD8AFAC170FF951D1AF362FCCAFDCC" box="[845,874,597,623]" captionStart="Fig" captionStartId="71.[189,232,1966,1992]" captionTargetBox="[189,1398,556,1921]" captionTargetId="figure@71.[189,1398,556,1921]" captionTargetPageId="71" captionText="Fig. 34. Intercalary cuticular microscutes in Chaleponcus spp. A. C. netus sp. nov. B. C. tintin sp. nov. CD. C. nikolajscharffi sp. nov. Fig. D shows fungus-like outgrowths from scutes. Scales 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3860522" httpUri="https://zenodo.org/record/3860522/files/figure.png" pageId="29" pageNumber="30">34</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C170FF951EEAF3A1FDADFC2C" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="BB59967FC170FF951EEAF3A1FE96FD13" blockId="29.[189,310,662,688]" box="[189,310,662,688]" pageId="29" pageNumber="30">
<heading id="E0112113C170FF951EEAF3A1FE96FD13" bold="true" box="[189,310,662,688]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="89924A6DC170FF951EEAF3A1FE96FD13" bold="true" box="[189,310,662,688]" pageId="29" pageNumber="30">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC170FF951EEAF3F3FDADFC2C" blockId="29.[189,1399,708,911]" pageId="29" pageNumber="30">
Medium-sized. Differs from all other group members by the profile of the gonopod coxal tip (
<figureCitation id="23DD8AFAC170FF951B10F3F3FF6EFCA1" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The transverse plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="29" pageNumber="30">Fig. 9</figureCitation>
). Very similar to
<taxonomicName id="7CE6EDFCC170FF951FC8F3DFFDB2FCA2" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[415,530,744,770]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="scopus" status="sp. nov.">
<emphasis id="89924A6DC170FF951FC8F3DFFDB2FCA2" box="[415,530,744,770]" italics="true" pageId="29" pageNumber="30">C. scopus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C170FF951C4CF3DFFDD7FCA1" box="[539,631,744,770]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
, sharing, i.a., a non-transverse metaplical shelf and a groove for accommodating the solenomere formed by two telomere lobes; differing from
<taxonomicName id="7CE6EDFCC170FF951A24F23CFB48FC86" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1139,1256,779,805]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="scopus" status="sp. nov.">
<emphasis id="89924A6DC170FF951A24F23CFB48FC86" box="[1139,1256,779,805]" italics="true" pageId="29" pageNumber="30">C. scopus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C170FF951AA5F23CFAEFFC86" box="[1266,1359,779,805]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
by apparently having the solenomere-conducting groove formed by the anterior and the posterior telomeral lamellae, instead of by the anterior lamella and a process from the latter, as well as by having a differently shaped lateral coxal process.
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C170FF951EEAF281FDA2FB81" pageId="29" pageNumber="30" type="etymology">
<paragraph id="BB59967FC170FF951EEAF281FEE4FC73" blockId="29.[189,324,950,976]" box="[189,324,950,976]" pageId="29" pageNumber="30">
<heading id="E0112113C170FF951EEAF281FEE4FC73" bold="true" box="[189,324,950,976]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="89924A6DC170FF951EEAF281FEE4FC73" bold="true" box="[189,324,950,976]" pageId="29" pageNumber="30">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC170FF951EEAF2D3FDA2FB81" blockId="29.[189,1399,996,1058]" pageId="29" pageNumber="30">The name honours my colleague, arachnologist Nikolaj Scharff (ZMUC), collector of this and numerous other Tanzanian millipedes.</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C170FF951EEAF57EFE09F965" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="BB59967FC170FF951EEAF57EFD9BFBC7" blockId="29.[189,571,1097,1124]" box="[189,571,1097,1124]" pageId="29" pageNumber="30">
<emphasis id="89924A6DC170FF951EEAF57EFE2CFBC0" bold="true" box="[189,396,1097,1123]" pageId="29" pageNumber="30">Material studied</emphasis>
(total:
<specimenCount id="ADE05DF6C170FF951FB6F57EFD92FBC7" box="[481,562,1097,1124]" pageId="29" pageNumber="30" type="male">11 ♂♂</specimenCount>
)
</paragraph>
<materialsCitation id="0B8E9C22C170FF951EBDF54FFB8CFB7B" ID-GBIF-Occurrence="2626747304" collectingDate="2 - 13 Nov." collectingMethod="pitfall traps" collectionCode="ZMUC" collectorName="N. Scharff" country="Tanzania" county="Udzungwa Mts" elevation="1500" location="above Chita village" municipality="Udzungwa Scarp" pageId="29" pageNumber="30" specimenCount="1" specimenCount-male="1" stateProvince="Iringa Region" typeStatus="holotype">
<paragraph id="BB59967FC170FF951EBDF54FFEFCFB31" blockId="29.[189,1399,1144,1241]" box="[234,348,1144,1170]" pageId="29" pageNumber="30">
<emphasis id="89924A6DC170FF951EBDF54FFEFCFB31" bold="true" box="[234,348,1144,1170]" pageId="29" pageNumber="30">
<typeStatus id="645D28DDC170FF951EBDF54FFEFCFB31" box="[234,348,1144,1170]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="BB59967FC170FF951EEAF5ACFB8CFB7B" blockId="29.[189,1399,1144,1241]" pageId="29" pageNumber="30">
<collectingCountry id="C3F1D6EFC170FF951EEAF5ACFEF7FB16" box="[189,343,1179,1205]" name="Tanzania" pageId="29" pageNumber="30">TANZANIA</collectingCountry>
:
<specimenCount id="ADE05DF6C170FF951F32F5ACFEDDFB16" box="[357,381,1179,1205]" pageId="29" pageNumber="30" type="male"></specimenCount>
,
<collectingRegion id="7922589DC170FF951FDBF5ACFD95FB16" box="[396,565,1179,1205]" country="Tanzania" name="Iringa" pageId="29" pageNumber="30">Iringa Region</collectingRegion>
,
<collectingCounty id="5238EEF3C170FF951C14F5ACFD5CFB16" box="[579,764,1179,1205]" pageId="29" pageNumber="30">Udzungwa Mts</collectingCounty>
,
<collectingMunicipality id="5B3D0C05C170FF951D5BF5ACFC78FB16" box="[780,984,1179,1205]" pageId="29" pageNumber="30">Udzungwa Scarp</collectingMunicipality>
FR,
<location id="BE39C0A4C170FF951A4FF5ACFAA5FB16" LSID="urn:lsid:plazi:treatment:334F2769C170FF971C6AF339FBCCFAE6:BE39C0A4C170FF951A4FF5ACFAA5FB16" box="[1048,1285,1179,1205]" country="Tanzania" county="Udzungwa Mts" municipality="Udzungwa Scarp" name="above Chita village" pageId="29" pageNumber="30" stateProvince="Iringa Region">above Chita village</location>
,
<quantity id="7C1E3B9AC170FF951B44F5ACFAD1FB16" box="[1299,1393,1179,1206]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">
<elevation id="30CB714CC170FF951B44F5ACFAD1FB16" box="[1299,1393,1179,1206]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="CF58B0BFC170FF951EEAF589FE97FB7A" box="[189,311,1214,1241]" pageId="29" pageNumber="30">
<collectingDate id="DF1C4957C170FF951EEAF589FE97FB7A" box="[189,311,1214,1241]" pageId="29" pageNumber="30">213 Nov.</collectingDate>
</date>
l984,
<collectingMethod id="62A7EE68C170FF951FD6F589FDAAFB7A" box="[385,522,1214,1241]" pageId="29" pageNumber="30">pitfall traps</collectingMethod>
, montane rain forest.
<collectorName id="1613F3A9C170FF951D5EF588FC23FB7B" box="[777,899,1214,1241]" pageId="29" pageNumber="30">N. Scharff</collectorName>
leg. (
<collectionCode id="DDF70EBAC170FF951D92F589FB81FB7B" box="[965,1057,1214,1240]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="29" pageNumber="30">ZMUC</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="BB59967FC170FF951EBDF437FEC7FAB9" blockId="29.[189,1399,1280,1588]" box="[234,359,1280,1306]" pageId="29" pageNumber="30">
<emphasis id="89924A6DC170FF951EBDF437FEC7FAB9" bold="true" box="[234,359,1280,1306]" pageId="29" pageNumber="30">
<typeStatus id="645D28DDC170FF951EBDF437FEC7FAB9" box="[234,359,1280,1306]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="BB59967FC170FF951EEAF414FB30F997" blockId="29.[189,1399,1280,1588]" pageId="29" pageNumber="30">
All from
<collectingCountry id="C3F1D6EFC170FF951F7CF414FE63FA9E" box="[299,451,1315,1341]" name="Tanzania" pageId="29" pageNumber="30">TANZANIA</collectingCountry>
,
<collectingRegion id="7922589DC170FF951F87F414FDD6FA9E" box="[464,630,1315,1341]" country="Tanzania" name="Iringa" pageId="29" pageNumber="30">Iringa Region</collectingRegion>
, Udzungwa Mts, Udzungwa Scarp FR:
<specimenCount id="ADE05DF6C170FF951A1AF414FBDCFA9E" box="[1101,1148,1315,1341]" pageId="29" pageNumber="30" type="male">1 ♂</specimenCount>
, above Chita village,
<quantity id="7C1E3B9AC170FF951EEAF471FEC2FAC3" box="[189,354,1350,1377]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.65" metricValueMin="1.45" pageId="29" pageNumber="30" unit="m" value="1550.0" valueMax="1650.0" valueMin="1450.0">14501650 m</quantity>
, 49 Nov. l984, pitfall traps, montane rain forest, N. Scharff leg. (ZMUC);
<specimenCount id="ADE05DF6C170FF951A89F471FA83FAC3" box="[1246,1315,1350,1376]" pageId="29" pageNumber="30" type="male">2 ♂♂</specimenCount>
, above Chita village,
<quantity id="7C1E3B9AC170FF951F36F45EFDA4FA27" box="[353,516,1385,1412]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.65" metricValueMin="1.6" pageId="29" pageNumber="30" unit="m" value="1625.0" valueMax="1650.0" valueMin="1600.0">16001650 m</quantity>
, 813 Nov. l984, pitfall traps, montane rain forest, N. Scharff leg. (ZMUC);
<specimenCount id="ADE05DF6C170FF951EEAF4BBFEA3FA04" box="[189,259,1420,1447]" pageId="29" pageNumber="30" type="male">2 ♂♂</specimenCount>
, above Chita village,
<quantity id="7C1E3B9AC170FF951C5CF4BBFDC8FA04" box="[523,616,1420,1447]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="30" unit="m" value="1400.0">1400 m</quantity>
, 45 Nov. l984, pitfall traps, montane rain forest, N. Scharff leg. (ZMUC);
<specimenCount id="ADE05DF6C170FF951F6EF487FECBFA69" box="[313,363,1456,1482]" pageId="29" pageNumber="30" type="male">1 ♂</specimenCount>
, above Chita village,
<quantity id="7C1E3B9AC170FF951C2BF498FD7BFA69" box="[636,731,1455,1482]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">1500 m</quantity>
, 213 Nov. l984, pitfall traps, montane rain forest, N. Scharff leg. (ZMUC);
<specimenCount id="ADE05DF6C170FF951F91F4E4FDAAFA4E" box="[454,522,1491,1517]" pageId="29" pageNumber="30" type="male">2 ♂♂</specimenCount>
,
<quantity id="7C1E3B9AC170FF951C4FF4E4FDC1FA4E" box="[536,609,1491,1517]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="30" unit="km" value="11.0">11 km</quantity>
SE of Masisiwe village, Kihanga Stream,
<quantity id="7C1E3B9AC170FF951A0FF4E4FB14FA4E" box="[1112,1204,1491,1517]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="30" unit="m" value="1800.0">1800 m</quantity>
,
<geoCoordinate id="DED2F0B8C170FF951A96F4E4FAD1FA4E" box="[1217,1393,1490,1517]" degrees="08" direction="south" minutes="22" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="05.7" value="-8.36825">08°22 05.7” S</geoCoordinate>
,
<geoCoordinate id="DED2F0B8C170FF951EEAF4C1FECFF9B3" box="[189,367,1526,1553]" degrees="35" direction="east" minutes="58" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="41.6" value="35.97822">35°58 41.6” E</geoCoordinate>
,
<date id="CF58B0BFC170FF951F2AF4C1FDE9F9B3" box="[381,585,1526,1552]" pageId="29" pageNumber="30" value="1997-05-17" valueMax="1997-05-27" valueMin="1997-05-17">1727 May 1997</date>
, ZMUC &amp; SI Exp. leg. (ZMUC);
<specimenCount id="ADE05DF6C170FF951D89F4C1FB83F9B3" box="[990,1059,1526,1552]" pageId="29" pageNumber="30" type="male">2 ♂♂</specimenCount>
, nr. Masisiwe,
<geoCoordinate id="DED2F0B8C170FF951A81F4C1FAD1F9B3" box="[1238,1393,1526,1552]" degrees="08" direction="south" minutes="20" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="32" value="-8.342222">08°20 32” S</geoCoordinate>
,
<geoCoordinate id="DED2F0B8C170FF951EEAF72EFEF3F997" box="[189,339,1561,1588]" degrees="35" direction="east" minutes="58" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="03" value="35.9675">35°5803” E</geoCoordinate>
,
<date id="CF58B0BFC170FF951F08F72EFDBFF990" box="[351,543,1561,1588]" pageId="29" pageNumber="30" value="2004-07-12" valueMax="2004-07-15" valueMin="2004-07-12">1215 Jul. 2004</date>
, A. Sforzi &amp; L. Bartolozzi leg. (MZF,
<superScript id="4C933B37C170FF951DB6F72DFC58F985" attach="left" box="[993,1016,1560,1588]" fontSize="6" pageId="29" pageNumber="30">no</superScript>
<date id="CF58B0BFC170FF951DA8F72EFB24F997" box="[1023,1156,1561,1588]" pageId="29" pageNumber="30" value="2695-05">Mag. 2695</date>
).
</paragraph>
<paragraph id="BB59967FC170FF951EEAF76CFEFDF9D7" blockId="29.[189,349,1626,1653]" box="[189,349,1626,1653]" pageId="29" pageNumber="30">
<heading id="E0112113C170FF951EEAF76CFEFDF9D7" bold="true" box="[189,349,1626,1653]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="89924A6DC170FF951EEAF76CFEFDF9D7" bold="true" box="[189,349,1626,1653]" pageId="29" pageNumber="30">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC170FF951EEAF7BEFE09F965" blockId="29.[189,1399,1673,1735]" pageId="29" pageNumber="30">
<collectingCountry id="C3F1D6EFC170FF951EEAF7BEFEF7F900" box="[189,343,1673,1699]" name="Tanzania" pageId="29" pageNumber="30">TANZANIA</collectingCountry>
:
<collectingRegion id="7922589DC170FF951F3FF7BEFDB4F900" box="[360,532,1673,1699]" country="Tanzania" name="Iringa" pageId="29" pageNumber="30">Iringa Region</collectingRegion>
, Udzungwa Mts, Udzungwa Scarp
<collectionCode id="DDF70EBAC170FF951D85F7BEFC59F900" box="[978,1017,1673,1699]" pageId="29" pageNumber="30">FR</collectionCode>
, above Chita village,
<quantity id="7C1E3B9AC170FF951B47F7BEFAD1F900" box="[1296,1393,1673,1700]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">1500 m</quantity>
, montane rain forest.
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C170FF971EEAF7DAFB5EFCCE" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30" type="description">
<paragraph id="BB59967FC170FF951EEAF7DAFE02F8AA" blockId="29.[189,418,1773,1801]" box="[189,418,1773,1801]" pageId="29" pageNumber="30">
<emphasis id="89924A6DC170FF951EEAF7DAFEEEF8A4" bold="true" box="[189,334,1773,1799]" pageId="29" pageNumber="30">Description</emphasis>
(male)
</paragraph>
<paragraph id="BB59967FC170FF951EEAF62AFD67F895" blockId="29.[189,711,1820,1847]" box="[189,711,1820,1847]" pageId="29" pageNumber="30">
DIAMETER.
<quantity id="7C1E3B9AC170FF951F6BF62BFE6CF894" box="[316,460,1820,1847]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="29" pageNumber="30" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.22.4 mm</quantity>
, 4849 podous rings.
</paragraph>
<paragraph id="BB59967FC170FF951EEAF669FE0CF81D" blockId="29.[189,1398,1885,1982]" pageId="29" pageNumber="30">COLOUR. After 10 years in alcohol with a broad light middorsal stripe formed by an hourglass-shaped spot on each body ring; stripe flanked by brownish paramedian bands; older specimens unicolourous light brown/pinkish.</paragraph>
<paragraph id="BB59967FC173FF961EEAF03BFC59FEEB" blockId="30.[189,1399,267,329]" pageId="30" pageNumber="31">ANAL VALVES. Each with a long, slightly curving spine and a well-developed, triangular ventral one; marginal rim raised, setiferous tubercles well developed, on ravelins.</paragraph>
<caption id="EF99C6F7C173FF961EEAF65FFA9EF84F" ID-DOI="http://doi.org/10.5281/zenodo.3860486" ID-Zenodo-Dep="3860486" httpUri="https://zenodo.org/record/3860486/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1896,1922]" targetBox="[189,1399,435,1852]" targetPageId="30">
<paragraph id="BB59967FC173FF961EEAF65FFA9EF84F" blockId="30.[189,1399,1896,2028]" pageId="30" pageNumber="31">
<emphasis id="89924A6DC173FF961EEAF65FFEB8F821" bold="true" box="[189,280,1896,1922]" pageId="30" pageNumber="31">Fig. 16.</emphasis>
<taxonomicName id="7CE6EDFCC173FF961F4AF65FFDC5F821" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[285,613,1896,1922]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nikolajscharffi" status="sp. nov.">
<emphasis id="89924A6DC173FF961F4AF65FFDC5F821" box="[285,613,1896,1922]" italics="true" pageId="30" pageNumber="31">Chaleponcus nikolajscharffi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C173FF961C3CF65EFD64F820" box="[619,708,1897,1923]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
, right gonopod.
<emphasis id="89924A6DC173FF961DD6F65FFC36F821" bold="true" box="[897,918,1896,1922]" pageId="30" pageNumber="31">A</emphasis>
. Mesal view.
<emphasis id="89924A6DC173FF961A63F65EFBE8F820" bold="true" box="[1076,1096,1897,1923]" pageId="30" pageNumber="31">B</emphasis>
. Anterior view.
<emphasis id="89924A6DC173FF961B57F65FFAB5F821" bold="true" box="[1280,1301,1896,1922]" pageId="30" pageNumber="31">C</emphasis>
. Ventral view.
<emphasis id="89924A6DC173FF961F55F6BBFEB7F805" bold="true" box="[258,279,1932,1958]" pageId="30" pageNumber="31">D</emphasis>
. Posterior view.
<emphasis id="89924A6DC173FF961F8BF6BBFE53F805" box="[476,499,1932,1958]" italics="true" pageId="30" pageNumber="31">al</emphasis>
= anterior lamella of telomere,
<emphasis id="89924A6DC173FF961D32F6BBFC21F806" box="[869,897,1932,1957]" italics="true" pageId="30" pageNumber="31">cu</emphasis>
= cucullus,
<emphasis id="89924A6DC173FF961A58F6BBFB86F805" box="[1039,1062,1932,1958]" italics="true" pageId="30" pageNumber="31">lp</emphasis>
= lateral coxal process,
<emphasis id="89924A6DC173FF961B15F6BBFAFFF805" box="[1346,1375,1932,1958]" italics="true" pageId="30" pageNumber="31">mf</emphasis>
= metaplical flange,
<emphasis id="89924A6DC173FF961FC4F687FE17F86A" box="[403,439,1968,1993]" italics="true" pageId="30" pageNumber="31">mp</emphasis>
= mesal metaplical process,
<emphasis id="89924A6DC173FF961D52F698FC8EF86A" box="[773,814,1967,1993]" italics="true" pageId="30" pageNumber="31">msl</emphasis>
= metaplical shelf-forming lobe,
<emphasis id="89924A6DC173FF961AE4F687FB7FF86A" box="[1203,1247,1968,1993]" italics="true" pageId="30" pageNumber="31">mss</emphasis>
= metaplical shelf-spine,
<emphasis id="89924A6DC173FF961F1BF6E5FEC3F84F" box="[332,355,2002,2028]" italics="true" pageId="30" pageNumber="31">pl</emphasis>
= posterior lamella of telomere,
<emphasis id="89924A6DC173FF961CB2F6E4FD5FF84F" box="[741,767,2003,2028]" italics="true" pageId="30" pageNumber="31">ps</emphasis>
= proximal spine of solenomere. Scales 0.1 mm.
</paragraph>
</caption>
<paragraph id="BB59967FC172FF971EEAF03BFE53FEEB" blockId="31.[189,1398,266,329]" pageId="31" pageNumber="32">
LIMBUS. With long, slender triangular lobes; lobes
<emphasis id="89924A6DC172FF971D40F03BFC93FE86" box="[791,819,268,293]" italics="true" pageId="31" pageNumber="32">ca</emphasis>
. 2½ × as long as broad, almost meeting at base, striate on external surface.
</paragraph>
<paragraph id="BB59967FC172FF971EEAF046FE4FFE29" blockId="31.[189,495,368,395]" box="[189,495,368,395]" pageId="31" pageNumber="32">TARSAL SETATION. Normal.</paragraph>
<paragraph id="BB59967FC172FF971EEAF085FB19FD45" blockId="31.[189,1399,433,742]" pageId="31" pageNumber="32">
GONOPOD COXA (
<figureCitation id="23DD8AFAC172FF971F2CF086FE66FE6F" box="[379,454,433,460]" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The transverse plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="31" pageNumber="32">Figs 9</figureCitation>
,
<figureCitation id="23DD8AFAC172FF971F84F086FE54FE6F" box="[467,500,433,460]" captionStart="Fig" captionStartId="30.[189,232,1896,1922]" captionTargetBox="[189,1399,435,1852]" captionTargetId="figure@30.[189,1399,435,1852]" captionTargetPageId="30" captionText="Fig. 16. Chaleponcus nikolajscharffi sp. nov., right gonopod. A. Mesal view. B. Anterior view. C. Ventral view. D. Posterior view. al = anterior lamella of telomere, cu = cucullus, lp = lateral coxal process, mf = metaplical flange, mp = mesal metaplical process, msl = metaplical shelf-forming lobe, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860486" httpUri="https://zenodo.org/record/3860486/files/figure.png" pageId="31" pageNumber="32">16</figureCitation>
). About 4 × as long as wide. Lateral margin basally slightly convex, curving strongly laterad
<emphasis id="89924A6DC172FF971FDDF0E2FE0DFE4D" box="[394,429,469,494]" italics="true" pageId="31" pageNumber="32">ca.</emphasis>
at level of
<emphasis id="89924A6DC172FF971C1AF0E2FDCFFE4C" box="[589,623,469,495]" italics="true" pageId="31" pageNumber="32">prl</emphasis>
, forming semicircular incision, and meeting apical margin in irregularly knobbed laterad process (
<emphasis id="89924A6DC172FF971C22F0CFFD2CFDB1" box="[629,652,504,530]" italics="true" pageId="31" pageNumber="32">lp</emphasis>
); apical margin sloping apico-mesad from process, irregularly undulate; cucullus (
<emphasis id="89924A6DC172FF971FF4F32BFE1FFD96" box="[419,447,540,565]" italics="true" pageId="31" pageNumber="32">cu</emphasis>
) transverse, mesally projecting as rounded lobe with irregularly undulate margin. Metaplical flange (
<emphasis id="89924A6DC172FF971FCEF309FE16FDFB" box="[409,438,574,600]" italics="true" pageId="31" pageNumber="32">mf</emphasis>
) ending in low, rounded process; metaplical mesal margin basally straight, further distally overlaid by lamelloid process (
<emphasis id="89924A6DC172FF971CD0F355FD0BFDD8" box="[647,683,610,635]" italics="true" pageId="31" pageNumber="32">mp</emphasis>
) covering arculus. Metaplical shelf not transverse as in most other species: posterior surface of metaplica with large lobe (
<emphasis id="89924A6DC172FF971DFCF3B2FC74FD3C" box="[939,980,645,671]" italics="true" pageId="31" pageNumber="32">msl</emphasis>
) roofing arculus; lobe concave on posterior face, folded anteriad and on anterior face giving rise to metaplical shelf-spine (
<emphasis id="89924A6DC172FF971A8DF39EFAA6FD61" box="[1242,1286,681,706]" italics="true" pageId="31" pageNumber="32">mss</emphasis>
).
<emphasis id="89924A6DC172FF971B77F39EFAF6FD61" box="[1312,1366,681,706]" italics="true" pageId="31" pageNumber="32">mms</emphasis>
of medium length, curving first anteriad, then mesad, then basad, reaching level of torsus.
</paragraph>
<paragraph id="BB59967FC172FF971EEAF23AFB5EFCCE" blockId="31.[189,1399,780,877]" pageId="31" pageNumber="32">
GONOPOD TELOPODITE (
<figureCitation id="23DD8AFAC172FF971FE9F23BFDB5FC84" box="[446,533,780,807]" captionStart="Fig" captionStartId="30.[189,232,1896,1922]" captionTargetBox="[189,1399,435,1852]" captionTargetId="figure@30.[189,1399,435,1852]" captionTargetPageId="30" captionText="Fig. 16. Chaleponcus nikolajscharffi sp. nov., right gonopod. A. Mesal view. B. Anterior view. C. Ventral view. D. Posterior view. al = anterior lamella of telomere, cu = cucullus, lp = lateral coxal process, mf = metaplical flange, mp = mesal metaplical process, msl = metaplical shelf-forming lobe, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860486" httpUri="https://zenodo.org/record/3860486/files/figure.png" pageId="31" pageNumber="32">Fig. 16</figureCitation>
). Solenomere with a long, straight, mesad proximal spine (
<emphasis id="89924A6DC172FF971A9DF23AFB44FC85" box="[1226,1252,781,806]" italics="true" pageId="31" pageNumber="32">ps</emphasis>
) originating from a wrinkled area. Telomere distally with two branches,
<emphasis id="89924A6DC172FF971D22F207FC2CFCE9" box="[885,908,816,842]" italics="true" pageId="31" pageNumber="32">al</emphasis>
and
<emphasis id="89924A6DC172FF971D93F207FC7BFCE9" box="[964,987,816,842]" italics="true" pageId="31" pageNumber="32">pl</emphasis>
, both curving anteriad, both broad, in part with irregularly serrate edges, lying closely together and accommodating solenomere.
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C172FF971EEAF2A3FAD7FC7E" pageId="31" pageNumber="32" type="multiple">
<paragraph id="BB59967FC172FF971EEAF2A3FE4EFC0D" blockId="31.[189,494,916,943]" box="[189,494,916,943]" pageId="31" pageNumber="32">
<heading id="E0112113C172FF971EEAF2A3FE4EFC0D" bold="true" box="[189,494,916,943]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="89924A6DC172FF971EEAF2A3FE4EFC0D" bold="true" box="[189,494,916,943]" pageId="31" pageNumber="32">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC172FF971EEAF2F4FAD7FC7E" blockId="31.[189,1399,963,989]" box="[189,1399,963,989]" pageId="31" pageNumber="32">
Known only from Udzungwa Scarp FR. Altitudinal rang:
<quantity id="7C1E3B9AC172FF971D01F2F4FC57FC7E" box="[854,1015,963,989]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="31" pageNumber="32" unit="m" value="1600.0" valueMax="1800.0" valueMin="1400.0">14001800 m</quantity>
asl. Habitat: montane rain forest.
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C172FF971EEAF533FBCCFAE6" pageId="31" pageNumber="32" type="discussion">
<paragraph id="BB59967FC172FF971EEAF533FE3EFBBD" blockId="31.[189,414,1028,1054]" box="[189,414,1028,1054]" pageId="31" pageNumber="32">
<heading id="E0112113C172FF971EEAF533FE3EFBBD" bold="true" box="[189,414,1028,1054]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="89924A6DC172FF971EEAF533FE3EFBBD" bold="true" box="[189,414,1028,1054]" pageId="31" pageNumber="32">Coexisting species</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC172FF971EEAF504FCCEFBD2" blockId="31.[189,1398,1075,1137]" pageId="31" pageNumber="32">
<taxonomicName id="7CE6EDFCC172FF971EEAF504FEEBFBEE" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[189,331,1075,1101]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="hamerae" status="sp. nov.">
<emphasis id="89924A6DC172FF971EEAF504FEEBFBEE" box="[189,331,1075,1101]" italics="true" pageId="31" pageNumber="32">C. hamerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C172FF971F00F504FE17FBEE" box="[343,439,1075,1101]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
was found in the same samples as
<taxonomicName id="7CE6EDFCC172FF971D2EF504FBE9FBEE" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[889,1097,1075,1101]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nikolajscharffi" status="sp. nov.">
<emphasis id="89924A6DC172FF971D2EF504FBE9FBEE" box="[889,1097,1075,1101]" italics="true" pageId="31" pageNumber="32">C. nikolajscharffi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C172FF971A01F504FB16FBEE" box="[1110,1206,1075,1101]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
In addition,
<taxonomicName id="7CE6EDFCC172FF971B0BF504FECAFBD3" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="circumvallatus" status="sp. nov.">
<emphasis id="89924A6DC172FF971B0BF504FECAFBD3" italics="true" pageId="31" pageNumber="32">C. circumvallatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C172FF971F25F560FE6CFBD2" box="[370,460,1111,1137]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
was found in Udzungwa Scarp FR.
</paragraph>
<paragraph id="BB59967FC172FF971EEAF5AFFEA3FB11" blockId="31.[189,259,1176,1202]" box="[189,259,1176,1202]" pageId="31" pageNumber="32">
<heading id="E0112113C172FF971EEAF5AFFEA3FB11" bold="true" box="[189,259,1176,1202]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="89924A6DC172FF971EEAF5AFFEA3FB11" bold="true" box="[189,259,1176,1202]" pageId="31" pageNumber="32">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC172FF971EEAF5F0FBCCFB42" blockId="31.[189,1132,1222,1249]" box="[189,1132,1222,1249]" pageId="31" pageNumber="32">
Very similar and probably closely related to
<taxonomicName id="7CE6EDFCC172FF971C92F5F0FC98FB43" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[709,824,1223,1249]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="scopus" status="sp. nov.">
<emphasis id="89924A6DC172FF971C92F5F0FC98FB43" box="[709,824,1223,1249]" italics="true" pageId="31" pageNumber="32">C. scopus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C172FF971D68F5F0FC3AFB42" box="[831,922,1223,1249]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
from Kigogo FR.
</paragraph>
<paragraph id="BB59967FC172FF971EEAF43FFBCCFAE6" blockId="31.[189,1398,1287,1349]" pageId="31" pageNumber="32">
Very peculiar, mushroom-like inter-cytoscutal structures were observed with the scanning electron microscope in this species; see “Two notes on general morphology” and
<figureCitation id="23DD8AFAC172FF971A46F41CFBC8FAE6" box="[1041,1128,1323,1349]" captionStart="Fig" captionStartId="71.[189,232,1966,1992]" captionTargetBox="[189,1398,556,1921]" captionTargetId="figure@71.[189,1398,556,1921]" captionTargetPageId="71" captionText="Fig. 34. Intercalary cuticular microscutes in Chaleponcus spp. A. C. netus sp. nov. B. C. tintin sp. nov. CD. C. nikolajscharffi sp. nov. Fig. D shows fungus-like outgrowths from scutes. Scales 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3860522" httpUri="https://zenodo.org/record/3860522/files/figure.png" pageId="31" pageNumber="32">Fig. 34</figureCitation>
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