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<mods:title id="DADAA626986A6485B719C5462EDABD92">Identities of some names in Gomphrena (Amaranthaceae)</mods:title>
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<mods:namePart id="5D03B82F9325A00B1710134763947C00">Sandoval-Ortega, Manuel Higinio</mods:namePart>
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<mods:namePart id="591A91C1E3D7E3B5D54BAF1690341B81">Zumaya-Mendoza, Silvia</mods:namePart>
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<mods:affiliation id="696FC014F71FB3A371461309C84BE8A0">Universidad Nacional Autónoma de México, Facultad de Ciencias, Ciudad Universitaria, Coyoacán, Ciudad de México, MÉXICO, C. P. 04510. &amp; szumaya @ ciencias. unam. mx; https: // orcid. org / 0000 - 0003 - 0337 - 4924</mods:affiliation>
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<treatment id="33056301B73BFFDDFF75DEEBD1DDFE70" LSID="urn:lsid:plazi:treatment:33056301B73BFFDDFF75DEEBD1DDFE70" httpUri="http://treatment.plazi.org/id/33056301B73BFFDDFF75DEEBD1DDFE70" lastPageId="11" lastPageNumber="89" pageId="5" pageNumber="83">
<subSubSection id="F3B6819CB73BFFD3FF75DEEBD26EFF18" pageId="5" pageNumber="83" type="nomenclature">
<paragraph id="BB13D217B73BFFD3FF75DEEBD26EFF18" blockId="5.[136,1452,157,327]" pageId="5" pageNumber="83">
<emphasis id="89D80E05B73BFFD3FF75DEEBD3B8FF7C" bold="true" box="[136,157,158,184]" pageId="5" pageNumber="83">3.</emphasis>
<taxonomicName id="7CACA994B73BFFD3FF58DEE8D19BFF7D" ID-CoL="3GTL7" authority="Standley (1917: 149 - 150)" authorityName="Standley" authorityPageNumber="149 - 150" authorityYear="1917" box="[165,702,157,185]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FF58DEE8D2B5FF7C" bold="true" box="[165,400,157,184]" italics="true" pageId="5" pageNumber="83">Gomphrena palmeri</emphasis>
<bibRefCitation id="DF3DAFE6B73BFFD3FE65DEEBD19BFF7D" author="Standley, P. C." box="[408,702,158,185]" pageId="5" pageNumber="83" pagination="95 - 169" refId="ref6564" refString="Standley, P. C. (1917) Amaranthaceae. In: Britton, N. L., Murrill, W. A. &amp; Barnhart (eds.) North American Flora, vol. 21 (2). The New York Botanical Garden, New York, pp. 95 - 169." type="book chapter" year="1917">Standley (1917: 149150)</bibRefCitation>
</taxonomicName>
<taxonomicName id="7CACA994B73BFFD3FD20DEEBD262FF18" authority="Moquin-Tandon (1849: 409 - 410)" authorityName="Moquin-Tandon" authorityPageNumber="409 - 410" authorityYear="1849" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B73BFFD3FD20DEEBD07CFF7C" box="[733,857,158,184]" italics="true" pageId="5" pageNumber="83">G. globosa</emphasis>
<bibRefCitation id="DF3DAFE6B73BFFD3FC9CDEEBD776FF7C" author="Linnaeus, C." box="[865,1107,158,185]" pageId="5" pageNumber="83" refId="ref5668" refString="Linnaeus, C. (1753) Species plantarum, vol. 1. Salvius, Stockholm, 560 pp." type="book" year="1753">Linnaeus (1753: 224)</bibRefCitation>
var.
<emphasis id="89D80E05B73BFFD3FB71DEEBD7CBFF7C" box="[1164,1262,158,184]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
<bibRefCitation id="DF3DAFE6B73BFFD3FB0BDEEBD262FF18" author="Moquin-Tandon, C. H." pageId="5" pageNumber="83" pagination="231 - 424" refId="ref5819" refString="Moquin-Tandon, C. H. (1849) Amaranthaeae Juss. In: Candolle, A. P. de (ed.) Prodromus systematis naturalis regni vegetabilis, Vol. 13, part. 2. Victor Masson, Paris, pp. 231 - 424." type="book chapter" year="1849">Moquin-Tandon (1849: 409410)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F3B6819CB73BFFD3FF75DE9CD03FF900" pageId="5" pageNumber="83" type="description">
<paragraph id="BB13D217B73BFFD3FF75DE9CD10BFE82" blockId="5.[136,1452,157,327]" pageId="5" pageNumber="83">
<emphasis id="89D80E05B73BFFD3FF75DE9CD3C8FF3A" bold="true" box="[136,237,233,254]" pageId="5" pageNumber="83">
<typeStatus id="64176CB5B73BFFD3FF75DE9CD3C8FF3A" box="[136,237,233,254]" pageId="5" pageNumber="83" type="lectotype">Lectotype</typeStatus>
</emphasis>
(designated by
<bibRefCitation id="DF3DAFE6B73BFFD3FE70DE9CD15BFF3A" author="Holzhammer, E." box="[397,638,233,255]" pageId="5" pageNumber="83" pagination="178 - 249" refId="ref5184" refString="Holzhammer, E. (1956) Die amerikanischen Arten der Gattung Gomphrena L. 2 Teil. Mitteilungen der Botanischen Staatssammlung Munchen 14 - 15: 178 - 249." type="journal article" year="1956">Holzhammer (1956: 193</bibRefCitation>
[first-step typification]; second-step typification by
<bibRefCitation id="DF3DAFE6B73BFFD3FB7EDE9FD63AFF3B" author="Bena, M. J." box="[1155,1311,233,255]" pageId="5" pageNumber="83" pagination="131 - 146" refId="ref4410" refString="Bena, M. J. (2017) Typification and nomenclatural notes on twenty-seven names of Gomphrena L. (Amaranthaceae) linked to the South American flora. Phytotaxa 296: 131 - 146. https: // doi. org / 10.11646 / phytotaxa. 296.2.3" type="journal article" year="2017">Bena 2017: 135</bibRefCitation>
):—
<collectingCountry id="C3BB9287B73BFFD3FABEDE9CD68CFF3B" box="[1347,1449,233,255]" name="Mexico" pageId="5" pageNumber="83">MEXICO</collectingCountry>
.
<collectingRegion id="79681CF5B73BFFD3FF40DF78D22DFEE6" box="[189,264,269,290]" country="Mexico" name="Oaxaca" pageId="5" pageNumber="83">Oaxaca</collectingRegion>
. Plages de locean pacifique,
<emphasis id="89D80E05B73BFFD3FDB7DF78D149FEE6" box="[586,620,269,290]" italics="true" pageId="5" pageNumber="83">s.d.</emphasis>
,
<emphasis id="89D80E05B73BFFD3FD82DF78D1DAFEE6" box="[639,767,269,290]" italics="true" pageId="5" pageNumber="83">Galeotti 429</emphasis>
(P00622632, image available at http://mediaphoto.mnhn.fr/media/ 1442830899193xGp2dsSd4co2oyCn).
</paragraph>
<paragraph id="BB13D217B73BFFD3FF75DF03D2BEFC84" blockId="5.[136,1452,373,1985]" pageId="5" pageNumber="83">
<emphasis id="89D80E05B73BFFD3FF75DF03D054FE54" bold="true" box="[136,881,373,400]" pageId="5" pageNumber="83">
Notes on
<taxonomicName id="7CACA994B73BFFD3FF01DF00D1ABFE54" authorityName="Moquin-Tandon" authorityYear="1849" box="[252,654,373,400]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B73BFFD3FF01DF00D2CCFE54" bold="true" box="[252,489,373,400]" italics="true" pageId="5" pageNumber="83">Gomphrena globosa</emphasis>
var.
<emphasis id="89D80E05B73BFFD3FDD7DF00D1ABFE54" bold="true" box="[554,654,373,400]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
</taxonomicName>
and
<taxonomicName id="7CACA994B73BFFD3FD33DF03D069FE54" authorityName="Standley" authorityYear="1917" box="[718,844,373,400]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FD33DF03D069FE54" bold="true" box="[718,844,373,400]" italics="true" pageId="5" pageNumber="83">G. palmeri</emphasis>
</taxonomicName>
:—
</emphasis>
<taxonomicName id="7CACA994B73BFFD3FC8CDF03D7ABFE54" authorityName="Moquin-Tandon" authorityYear="1849" box="[881,1166,374,400]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B73BFFD3FC8CDF03D0CAFE54" box="[881,1007,374,400]" italics="true" pageId="5" pageNumber="83">G. globosa</emphasis>
var.
<emphasis id="89D80E05B73BFFD3FBD1DF03D7ABFE54" box="[1068,1166,374,400]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
</taxonomicName>
was cited as a synonym for various other names in
<taxonomicName id="7CACA994B73BFFD3FE36DFEFD175FE70" authorityName="Linnaeus" authorityYear="1753" box="[459,592,410,436]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="genus">
<emphasis id="89D80E05B73BFFD3FE36DFEFD175FE70" box="[459,592,410,436]" italics="true" pageId="5" pageNumber="83">Gomphrena</emphasis>
</taxonomicName>
that correspond to different species. For example, when
<bibRefCitation id="DF3DAFE6B73BFFD3FB08DFEFD39EFE1C" author="Standley, P. C." pageId="5" pageNumber="83" pagination="95 - 169" refId="ref6564" refString="Standley, P. C. (1917) Amaranthaceae. In: Britton, N. L., Murrill, W. A. &amp; Barnhart (eds.) North American Flora, vol. 21 (2). The New York Botanical Garden, New York, pp. 95 - 169." type="book chapter" year="1917">Standley (1917: 149)</bibRefCitation>
published
<taxonomicName id="7CACA994B73BFFD3FEBCDFCBD13BFE1C" authority="Standl." authorityName="Standley" authorityYear="1917" box="[321,542,446,472]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FEBCDFCBD2E5FE1C" box="[321,448,446,472]" italics="true" pageId="5" pageNumber="83">G. palmeri</emphasis>
Standl.
</taxonomicName>
, he cited as synonym
<taxonomicName id="7CACA994B73BFFD3FCCDDFCBD770FE1C" authorityName="Moquin-Tandon" authorityYear="1849" box="[816,1109,446,472]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B73BFFD3FCCDDFCBD094FE1C" box="[816,945,446,472]" italics="true" pageId="5" pageNumber="83">G. globosa</emphasis>
var.
<emphasis id="89D80E05B73BFFD3FC0EDFCBD770FE1C" box="[1011,1109,446,472]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
</taxonomicName>
and included the legend “
<emphasis id="89D80E05B73BFFD3FA6BDFCAD392FE38" italics="true" pageId="5" pageNumber="83">in part</emphasis>
”.
<bibRefCitation id="DF3DAFE6B73BFFD3FF2EDF97D109FE39" author="Holzhammer, E." box="[211,556,482,509]" pageId="5" pageNumber="83" pagination="178 - 249" refId="ref5184" refString="Holzhammer, E. (1956) Die amerikanischen Arten der Gattung Gomphrena L. 2 Teil. Mitteilungen der Botanischen Staatssammlung Munchen 14 - 15: 178 - 249." type="journal article" year="1956">Holzhammer (1956: 195196)</bibRefCitation>
considered
<taxonomicName id="7CACA994B73BFFD3FD4ADF97D0F4FE38" authorityName="Moquin-Tandon" authorityYear="1849" box="[695,977,482,508]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B73BFFD3FD4ADF97D011FE38" box="[695,820,482,508]" italics="true" pageId="5" pageNumber="83">G. globosa</emphasis>
var.
<emphasis id="89D80E05B73BFFD3FC92DF97D0F4FE38" box="[879,977,482,508]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
</taxonomicName>
as a synonym of both
<taxonomicName id="7CACA994B73BFFD3FB27DF97D673FE38" authorityName="Standley" authorityYear="1917" box="[1242,1366,482,508]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FB27DF97D673FE38" box="[1242,1366,482,508]" italics="true" pageId="5" pageNumber="83">G. palmeri</emphasis>
</taxonomicName>
and
<taxonomicName id="7CACA994B73BFFD3FA6DDF97D229FDE4" authorityName="Martens &amp; Galeotti" authorityYear="1843" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="filaginoides">
<emphasis id="89D80E05B73BFFD3FA6DDF97D229FDE4" italics="true" pageId="5" pageNumber="83">G. filaginoides</emphasis>
</taxonomicName>
.
<bibRefCitation id="DF3DAFE6B73BFFD3FEE1DC73D127FDE4" author="Pedersen, T. M." box="[284,514,518,544]" pageId="5" pageNumber="83" pagination="69 - 97" refId="ref5994" refString="Pedersen, T. M. (1990) Studies in South American Amaranthaceae III (including one amphi-Atlantic species). Bulletin du Museum National d'Histoire Naturelle Section B 12: 69 - 97. https: // doi. org / 10.5962 / p. 326734" type="journal article" year="1990">Pedersen (1990: 74)</bibRefCitation>
regarded
<taxonomicName id="7CACA994B73BFFD3FD8BDC73D0B5FDE4" authorityName="Moquin-Tandon" authorityYear="1849" box="[630,912,518,544]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B73BFFD3FD8BDC73D1D6FDE4" box="[630,755,518,544]" italics="true" pageId="5" pageNumber="83">G. globosa</emphasis>
var.
<emphasis id="89D80E05B73BFFD3FCD3DC73D0B5FDE4" box="[814,912,518,544]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="7CACA994B73BFFD3FBA2DC73D22AFD80" authority="Moquin-Tandon (1849: 401)" authorityName="Moquin-Tandon" authorityPageNumber="401" authorityYear="1849" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="boliviana">
<emphasis id="89D80E05B73BFFD3FBA2DC73D7C8FDE4" box="[1119,1261,518,544]" italics="true" pageId="5" pageNumber="83">G. boliviana</emphasis>
<bibRefCitation id="DF3DAFE6B73BFFD3FB0BDC73D22AFD80" author="Moquin-Tandon, C. H." pageId="5" pageNumber="83" pagination="231 - 424" refId="ref5819" refString="Moquin-Tandon, C. H. (1849) Amaranthaeae Juss. In: Candolle, A. P. de (ed.) Prodromus systematis naturalis regni vegetabilis, Vol. 13, part. 2. Victor Masson, Paris, pp. 231 - 424." type="book chapter" year="1849">Moquin-Tandon (1849: 401)</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="DF3DAFE6B73BFFD3FEE4DC5FD128FD81" author="Clemants, S. E." box="[281,525,554,581]" pageId="5" pageNumber="83" pagination="451 - 454" refId="ref4825" refString="Clemants, S. E. (2003) Gomphrena. In: Flora of North America Editorial Committee (eds.) Flora of North America North of Mexico (Magnoliophyta: Caryophyllidae, part 1), vol. 4. Oxford University Press, Oxford, pp. 451 - 454." type="book chapter" year="2003">Clemants (2003: 453)</bibRefCitation>
accepted
<taxonomicName id="7CACA994B73BFFD3FD80DC5FD0B4FD80" authorityName="Moquin-Tandon" authorityYear="1849" box="[637,913,554,580]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B73BFFD3FD80DC5FD1DDFD80" box="[637,760,554,580]" italics="true" pageId="5" pageNumber="83">G. globosa</emphasis>
var.
<emphasis id="89D80E05B73BFFD3FCD2DC5FD0B4FD80" box="[815,913,554,580]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="7CACA994B73BFFD3FBA8DC5FD79DFD80" authorityName="Rothrock" authorityYear="1878" box="[1109,1208,554,580]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="nitida">
<emphasis id="89D80E05B73BFFD3FBA8DC5FD79DFD80" box="[1109,1208,554,580]" italics="true" pageId="5" pageNumber="83">G. nitida</emphasis>
</taxonomicName>
. Finally,
<bibRefCitation id="DF3DAFE6B73BFFD3FADFDC5FD39EFDAC" author="Bena, M. J." pageId="5" pageNumber="83" pagination="131 - 146" refId="ref4410" refString="Bena, M. J. (2017) Typification and nomenclatural notes on twenty-seven names of Gomphrena L. (Amaranthaceae) linked to the South American flora. Phytotaxa 296: 131 - 146. https: // doi. org / 10.11646 / phytotaxa. 296.2.3" type="journal article" year="2017">Bena (2017: 136)</bibRefCitation>
synonymized Moquin-Tandons variety as synonym of
<taxonomicName id="7CACA994B73BFFD3FCDADC3BD096FDAC" authorityName="Moquin-Tandon" authorityYear="1849" box="[807,947,590,616]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="boliviana">
<emphasis id="89D80E05B73BFFD3FCDADC3BD096FDAC" box="[807,947,590,616]" italics="true" pageId="5" pageNumber="83">G. boliviana</emphasis>
</taxonomicName>
. These different views of variety by
<bibRefCitation id="DF3DAFE6B73BFFD3FAB4DC3BD2BEFD48" author="Moquin-Tandon, C. H." pageId="5" pageNumber="83" pagination="231 - 424" refId="ref5819" refString="Moquin-Tandon, C. H. (1849) Amaranthaeae Juss. In: Candolle, A. P. de (ed.) Prodromus systematis naturalis regni vegetabilis, Vol. 13, part. 2. Victor Masson, Paris, pp. 231 - 424." type="book chapter" year="1849">Moquin-Tandon (1849: 409410)</bibRefCitation>
are probably related to the protologue of
<taxonomicName id="7CACA994B73BFFD3FC98DC07D750FD48" authorityName="Moquin-Tandon" authorityYear="1849" box="[869,1141,626,652]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B73BFFD3FC98DC07D0FAFD48" box="[869,991,626,652]" italics="true" pageId="5" pageNumber="83">G. globosa</emphasis>
var.
<emphasis id="89D80E05B73BFFD3FBEEDC07D750FD48" box="[1043,1141,626,652]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
</taxonomicName>
, where the listed specimens (
<typeStatus id="64176CB5B73BFFD3FF6DDCE3D3D1FD74" box="[144,244,662,688]" pageId="5" pageNumber="83" type="syntype">syntypes</typeStatus>
according to the Art. 9.6 of ICN) correspond to more than one currently accepted species.
<bibRefCitation id="DF3DAFE6B73BFFD3FB0BDCE3D266FD10" author="Moquin-Tandon, C. H." pageId="5" pageNumber="83" pagination="231 - 424" refId="ref5819" refString="Moquin-Tandon, C. H. (1849) Amaranthaeae Juss. In: Candolle, A. P. de (ed.) Prodromus systematis naturalis regni vegetabilis, Vol. 13, part. 2. Victor Masson, Paris, pp. 231 - 424." type="book chapter" year="1849">Moquin-Tandon (1849: 409410)</bibRefCitation>
mentioned some specimens from
<collectingCountry id="C3BB9287B73BFFD3FD3CDCCFD03CFD10" box="[705,793,698,724]" name="Mexico" pageId="5" pageNumber="83">Mexico</collectingCountry>
, i.e.: “in Regni Mexicani province
<collectingRegion id="79681CF5B73BFFD3FB5CDCCFD7D0FD10" box="[1185,1269,698,724]" country="Mexico" name="Oaxaca" pageId="5" pageNumber="83">Oaxaca</collectingRegion>
(Galeotti n. 429 et 443)”, “inter Victoria et Tula (Berlandier n. 2228)”, “prope
<collectingRegion id="79681CF5B73BFFD3FCB1DCABD0E8FD3C" box="[844,973,734,760]" country="Mexico" name="Tamaulipas" pageId="5" pageNumber="83">Tamaulipas</collectingRegion>
(Id.! [= ibidem, so refers to Berlandier as collector] n. 2130), as well as a collection from South America “prope Mendozam (Gillies)”, and even one from Java (“Zoll. 2944 an culta?”).
</paragraph>
<paragraph id="BB13D217B73BFFD3FF40DD3FD70FFAFC" blockId="5.[136,1452,373,1985]" pageId="5" pageNumber="83">
Among the specimens collected in
<collectingRegion id="79681CF5B73BFFD3FDB0DD3FD1E2FCA0" box="[589,711,842,868]" country="Mexico" name="Distrito Federal" pageId="5" pageNumber="83">Mexico by</collectingRegion>
Galeotti and cited by
<bibRefCitation id="DF3DAFE6B73BFFD3FC38DD3FD666FCA0" author="Moquin-Tandon, C. H." box="[965,1347,842,868]" pageId="5" pageNumber="83" pagination="231 - 424" refId="ref5819" refString="Moquin-Tandon, C. H. (1849) Amaranthaeae Juss. In: Candolle, A. P. de (ed.) Prodromus systematis naturalis regni vegetabilis, Vol. 13, part. 2. Victor Masson, Paris, pp. 231 - 424." type="book chapter" year="1849">Moquin-Tandon (1849: 409410)</bibRefCitation>
, one was designated by
<bibRefCitation id="DF3DAFE6B73BFFD3FECFDD1BD175FC4C" author="Holzhammer, E." box="[306,592,878,905]" pageId="5" pageNumber="83" pagination="178 - 249" refId="ref5184" refString="Holzhammer, E. (1956) Die amerikanischen Arten der Gattung Gomphrena L. 2 Teil. Mitteilungen der Botanischen Staatssammlung Munchen 14 - 15: 178 - 249." type="journal article" year="1956">Holzhammer (1956: 193</bibRefCitation>
, first-step typification) and
<bibRefCitation id="DF3DAFE6B73BFFD3FC64DD1BD77BFC4D" author="Bena, M. J." box="[921,1118,878,905]" pageId="5" pageNumber="83" pagination="131 - 146" refId="ref4410" refString="Bena, M. J. (2017) Typification and nomenclatural notes on twenty-seven names of Gomphrena L. (Amaranthaceae) linked to the South American flora. Phytotaxa 296: 131 - 146. https: // doi. org / 10.11646 / phytotaxa. 296.2.3" type="journal article" year="2017">Bena (2017: 135</bibRefCitation>
, second-step typificaiton) as a
<typeStatus id="64176CB5B73BFFD3FF62DDE7D223FC68" box="[159,262,914,940]" pageId="5" pageNumber="83" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="7CACA994B73BFFD3FECFDDE7D19EFC68" authorityName="Moquin-Tandon" authorityYear="1849" box="[306,699,914,940]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B73BFFD3FECFDDE7D13EFC68" box="[306,539,914,940]" italics="true" pageId="5" pageNumber="83">Gomphrena globosa</emphasis>
var.
<emphasis id="89D80E05B73BFFD3FDA4DDE7D19EFC68" box="[601,699,914,940]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
</taxonomicName>
(
<emphasis id="89D80E05B73BFFD3FD32DDE7D07AFC68" box="[719,863,914,940]" italics="true" pageId="5" pageNumber="83">Galeotti 429</emphasis>
; P00622632,
<figureCitation id="2397CE92B73BFFD3FBFEDDE7D76EFC68" box="[1027,1099,914,940]" captionStart="FIGURE 4" captionStartId="6.[136,229,2028,2050]" captionTargetBox="[210,1377,190,2004]" captionTargetId="figure-17@6.[210,1377,190,2004]" captionTargetPageId="6" captionText="FIGURE 4. Lectotype of Gomphrena globosa var. albiflora (P00622632)." figureDoi="http://doi.org/10.5281/zenodo.10184563" httpUri="https://zenodo.org/record/10184563/files/figure.png" pageId="5" pageNumber="83">Fig. 4</figureCitation>
); four duplicates were located (BR0000021450990, BR0000021451003, P00622633, and P00622631); Concerning the collection
<emphasis id="89D80E05B73BFFD3FB10DDC3D65EFC14" box="[1261,1403,950,976]" italics="true" pageId="5" pageNumber="83">Galeotti 443</emphasis>
, we found one duplicate deposited in BR (BR0000027814574). All this material (BR0000021450990, BR0000021451003, BR0000027814574, P00622633, and P00622631) corresponds to the description provided by
<bibRefCitation id="DF3DAFE6B73BFFD3FB47DD8BD68EFBDC" author="Standley, P. C." box="[1210,1451,1022,1048]" pageId="5" pageNumber="83" pagination="95 - 169" refId="ref6564" refString="Standley, P. C. (1917) Amaranthaceae. In: Britton, N. L., Murrill, W. A. &amp; Barnhart (eds.) North American Flora, vol. 21 (2). The New York Botanical Garden, New York, pp. 95 - 169." type="book chapter" year="1917">Standley (1917: 149)</bibRefCitation>
for
<taxonomicName id="7CACA994B73BFFD3FF4CDA57D209FBF8" authorityName="Standley" authorityYear="1917" box="[177,300,1058,1084]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FF4CDA57D209FBF8" box="[177,300,1058,1084]" italics="true" pageId="5" pageNumber="83">G. palmeri</emphasis>
</taxonomicName>
. Actually
<bibRefCitation id="DF3DAFE6B73BFFD3FE5EDA57D1B7FBF8" author="Standley, P. C." box="[419,658,1058,1084]" pageId="5" pageNumber="83" pagination="95 - 169" refId="ref6564" refString="Standley, P. C. (1917) Amaranthaceae. In: Britton, N. L., Murrill, W. A. &amp; Barnhart (eds.) North American Flora, vol. 21 (2). The New York Botanical Garden, New York, pp. 95 - 169." type="book chapter" year="1917">Standley (1917: 149)</bibRefCitation>
listed
<taxonomicName id="7CACA994B73BFFD3FD1DDA57D051FBF8" authorityName="Moquin-Tandon" authorityYear="1849" box="[736,884,1058,1084]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
var.
<emphasis id="89D80E05B73BFFD3FCEFDA57D051FBF8" box="[786,884,1058,1084]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
</taxonomicName>
by
<bibRefCitation id="DF3DAFE6B73BFFD3FC5CDA57D639FBF8" author="Moquin-Tandon, C. H." box="[929,1308,1058,1084]" pageId="5" pageNumber="83" pagination="231 - 424" refId="ref5819" refString="Moquin-Tandon, C. H. (1849) Amaranthaeae Juss. In: Candolle, A. P. de (ed.) Prodromus systematis naturalis regni vegetabilis, Vol. 13, part. 2. Victor Masson, Paris, pp. 231 - 424." type="book chapter" year="1849">Moquin-Tandon (1849: 409410)</bibRefCitation>
as synonym of his
<taxonomicName id="7CACA994B73BFFD3FF2FDA33D268FBA4" authorityName="Standley" authorityYear="1917" box="[210,333,1094,1120]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FF2FDA33D268FBA4" box="[210,333,1094,1120]" italics="true" pageId="5" pageNumber="83">G. palmeri</emphasis>
</taxonomicName>
and cited a specimen collected by Palmer in Acapulco,
<collectingCountry id="C3BB9287B73BFFD3FC29DA33D70FFBA4" box="[980,1066,1094,1120]" name="Mexico" pageId="5" pageNumber="83">Mexico</collectingCountry>
and deposited in the
<collectingCountry id="C3BB9287B73BFFD3FADEDA33D662FBA4" box="[1315,1351,1094,1120]" name="United States of America" pageId="5" pageNumber="83">US</collectingCountry>
(
<emphasis id="89D80E05B73BFFD3FAA4DA33D397FB40" italics="true" pageId="5" pageNumber="83">Palmer 269</emphasis>
;
<collectingCountry id="C3BB9287B73BFFD3FF3FDA1FD3C0FB40" box="[194,229,1130,1156]" name="United States of America" pageId="5" pageNumber="83">US</collectingCountry>
00102797,
<figureCitation id="2397CE92B73BFFD3FE98DA1FD28CFB41" box="[357,425,1130,1157]" captionStart="FIGURE 5" captionStartId="7.[136,229,1933,1955]" captionTargetBox="[151,1436,190,1909]" captionTargetId="figure-24@7.[151,1436,190,1909]" captionTargetPageId="7" captionText="FIGURE 5. Specimen of Gomphrena palmeri (US00102797)." figureDoi="http://doi.org/10.5281/zenodo.10184565" httpUri="https://zenodo.org/record/10184565/files/figure.png" pageId="5" pageNumber="83">Fig. 5</figureCitation>
). The name
<taxonomicName id="7CACA994B73BFFD3FDCADA1FD06BFB40" authorityName="Moquin-Tandon" authorityYear="1849" box="[567,846,1130,1156]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B73BFFD3FDCADA1FD196FB40" box="[567,691,1130,1156]" italics="true" pageId="5" pageNumber="83">G. globosa</emphasis>
var.
<emphasis id="89D80E05B73BFFD3FD11DA1FD06BFB40" box="[748,846,1130,1156]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
</taxonomicName>
is, therefore, the basionym of
<taxonomicName id="7CACA994B73BFFD3FB54DA1FD601FB40" authorityName="Standley" authorityYear="1917" box="[1193,1316,1130,1156]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FB54DA1FD601FB40" box="[1193,1316,1130,1156]" italics="true" pageId="5" pageNumber="83">G. palmeri</emphasis>
</taxonomicName>
. So, Benas synonymization (with
<taxonomicName id="7CACA994B73BFFD3FE7DDAFBD12FFB6C" authorityName="Moquin-Tandon" authorityYear="1849" box="[384,522,1166,1192]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="boliviana">
<emphasis id="89D80E05B73BFFD3FE7DDAFBD12FFB6C" box="[384,522,1166,1192]" italics="true" pageId="5" pageNumber="83">G. boliviana</emphasis>
</taxonomicName>
) cannot be retained, unless
<taxonomicName id="7CACA994B73BFFD3FCCADAFBD0E4FB6C" authorityName="Moquin-Tandon" authorityYear="1849" box="[823,961,1166,1192]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="boliviana">
<emphasis id="89D80E05B73BFFD3FCCADAFBD0E4FB6C" box="[823,961,1166,1192]" italics="true" pageId="5" pageNumber="83">G. boliviana</emphasis>
</taxonomicName>
is treated as heterotypic synonym. However,
<taxonomicName id="7CACA994B73BFFD3FF75DAC7D221FB08" authorityName="Standley" authorityYear="1917" box="[136,260,1202,1228]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FF75DAC7D221FB08" box="[136,260,1202,1228]" italics="true" pageId="5" pageNumber="83">G. palmeri</emphasis>
</taxonomicName>
and
<taxonomicName id="7CACA994B73BFFD3FEBDDAC7D2EAFB08" authorityName="Moquin-Tandon" authorityYear="1849" box="[320,463,1202,1228]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="boliviana">
<emphasis id="89D80E05B73BFFD3FEBDDAC7D2EAFB08" box="[320,463,1202,1228]" italics="true" pageId="5" pageNumber="83">G. boliviana</emphasis>
</taxonomicName>
are two different species easily distinguished each other by the bracteoles, which are broadly cristate above in
<taxonomicName id="7CACA994B73BFFD3FE5CDAA3D13FFB34" authorityName="Standley" authorityYear="1917" box="[417,538,1238,1264]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FE5CDAA3D13FFB34" box="[417,538,1238,1264]" italics="true" pageId="5" pageNumber="83">G. palmeri</emphasis>
</taxonomicName>
(see e.g.,
<bibRefCitation id="DF3DAFE6B73BFFD3FD76DAA3D009FB34" author="Standley, P. C." box="[651,812,1238,1264]" pageId="5" pageNumber="83" pagination="95 - 169" refId="ref6564" refString="Standley, P. C. (1917) Amaranthaceae. In: Britton, N. L., Murrill, W. A. &amp; Barnhart (eds.) North American Flora, vol. 21 (2). The New York Botanical Garden, New York, pp. 95 - 169." type="book chapter" year="1917">Standley 1917</bibRefCitation>
) and not crested in
<taxonomicName id="7CACA994B73BFFD3FBFADAA3D7B6FB34" authorityName="Moquin-Tandon" authorityYear="1849" box="[1031,1171,1238,1264]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="boliviana">
<emphasis id="89D80E05B73BFFD3FBFADAA3D7B6FB34" box="[1031,1171,1238,1264]" italics="true" pageId="5" pageNumber="83">G. boliviana</emphasis>
</taxonomicName>
(see e.g.,
<bibRefCitation id="DF3DAFE6B73BFFD3FAFEDAA3D3E0FAD0" author="Bena, M. J. &amp; Acosta, J. M." pageId="5" pageNumber="83" pagination="72 - 100" refId="ref4459" refString="Bena, M. J. &amp; Acosta, J. M. (2020) Gomphrena. In: Anton, A. M., Zuloaga, F. O., Zuloaga, F. O. &amp; Belgrano, M. J. (eds.) Flora vascular de la Republica Argentina, vol. 19 (1). Instituto de Botanica Darwinion, Buenos Aires, pp. 72 - 100. https: // doi. org / 10.2307 / j. ctv 1 gwqr 79" type="book chapter" year="2020">Bena &amp; Acosta 2020</bibRefCitation>
). Furthermore,
<taxonomicName id="7CACA994B73BFFD3FE89DA8FD124FAD0" authorityName="Moquin-Tandon" authorityYear="1849" box="[372,513,1274,1300]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="boliviana">
<emphasis id="89D80E05B73BFFD3FE89DA8FD124FAD0" box="[372,513,1274,1300]" italics="true" pageId="5" pageNumber="83">G. boliviana</emphasis>
</taxonomicName>
is only distributed in South America, in
<collectingCountry id="C3BB9287B73BFFD3FC32DA8FD764FAD0" box="[975,1089,1274,1300]" name="Argentina" pageId="5" pageNumber="83">Argentina</collectingCountry>
,
<collectingCountry id="C3BB9287B73BFFD3FBB0DA8FD787FAD0" box="[1101,1186,1274,1300]" name="Bolivia" pageId="5" pageNumber="83">Bolivia</collectingCountry>
, and
<collectingCountry id="C3BB9287B73BFFD3FB22DA8FD662FAD0" box="[1247,1351,1274,1300]" name="Paraguay" pageId="5" pageNumber="83">Paraguay</collectingCountry>
(
<bibRefCitation id="DF3DAFE6B73BFFD3FAAADA8FD3E1FAFC" author="POWO" pageId="5" pageNumber="83" refId="ref6071" refString="POWO (2023) Plants of theWorld Online. Facilitated by the Royal Botanic Gardens, Kew. Available from: http: // www. plantsoftheworldonline. org / (accessed 30 October 2023)." type="url" year="2023">POWO 2023</bibRefCitation>
), whereas
<taxonomicName id="7CACA994B73BFFD3FEC7DB6BD296FAFC" authorityName="Standley" authorityYear="1917" box="[314,435,1310,1336]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FEC7DB6BD296FAFC" box="[314,435,1310,1336]" italics="true" pageId="5" pageNumber="83">G. palmeri</emphasis>
</taxonomicName>
is reported from
<collectingCountry id="C3BB9287B73BFFD3FD8BDB6BD1E9FAFC" box="[630,716,1310,1336]" name="Mexico" pageId="5" pageNumber="83">Mexico</collectingCountry>
to
<collectingCountry id="C3BB9287B73BFFD3FD12DB6BD04DFAFC" box="[751,872,1310,1336]" name="Guatemala" pageId="5" pageNumber="83">Guatemala</collectingCountry>
(
<bibRefCitation id="DF3DAFE6B73BFFD3FC8ADB6BD73BFAFC" author="Standley, P. C." box="[887,1054,1310,1336]" pageId="5" pageNumber="83" pagination="95 - 169" refId="ref6564" refString="Standley, P. C. (1917) Amaranthaceae. In: Britton, N. L., Murrill, W. A. &amp; Barnhart (eds.) North American Flora, vol. 21 (2). The New York Botanical Garden, New York, pp. 95 - 169." type="book chapter" year="1917">Standley 1917</bibRefCitation>
).
</paragraph>
<paragraph id="BB13D217B73BFFD3FF40DB37D7F8F9F0" blockId="5.[136,1452,373,1985]" pageId="5" pageNumber="83">
Concerning the material collected by J. Gillies, who explored South America, especially
<collectingCountry id="C3BB9287B73BFFD3FB44DB37D60DFA98" box="[1209,1320,1346,1372]" name="Argentina" pageId="5" pageNumber="83">Argentina</collectingCountry>
(see
<bibRefCitation id="DF3DAFE6B73BFFD3FA95DB37D3E7FA45" author="Gibbs, F. W." pageId="5" pageNumber="83" pagination="115 - 136" refId="ref5132" refString="Gibbs, F. W. (1951) John Gillies, M. D., traveller and botanist, 1792 - 1834. Notes and Records of the Royal Society of London 9: 115 - 136. https: // doi. org / 10.1098 / rsnr. 1951.0008" type="journal article" year="1951">Gibbs 1951</bibRefCitation>
, HUH-Index of Botanists 2023), a specimen from
<collectingRegion id="79681CF5B73BFFD3FD1ADB13D06AFA44" box="[743,847,1382,1408]" country="Argentina" name="Mendoza" pageId="5" pageNumber="83">Mendoza</collectingRegion>
is deposited at E (barcode E00334749, image available at http://data.rbge.org.uk/herb/E00334749). This specimen was originally identified as
<taxonomicName id="7CACA994B73BFFD3FB9FDBFFD65FFA60" authorityName="Moquin-Tandon" authorityYear="1849" box="[1122,1402,1418,1444]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B73BFFD3FB9FDBFFD7FAFA60" box="[1122,1247,1418,1444]" italics="true" pageId="5" pageNumber="83">G. globosa</emphasis>
var.
<emphasis id="89D80E05B73BFFD3FAE5DBFFD65FFA60" box="[1304,1402,1418,1444]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
</taxonomicName>
and later revised by Pedersen as
<taxonomicName id="7CACA994B73BFFD3FE3DDBDBD190FA0C" authorityName="Moquin-Tandon" authorityYear="1849" box="[448,693,1454,1480]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="boliviana">
<emphasis id="89D80E05B73BFFD3FE3DDBDBD190FA0C" box="[448,693,1454,1480]" italics="true" pageId="5" pageNumber="83">Gomphrena boliviana</emphasis>
</taxonomicName>
in 1973. Probably, this specimen (
<figureCitation id="2397CE92B73BFFD3FBCCDBDBD756FA0C" box="[1073,1139,1454,1480]" captionStart="FIGURE 6" captionStartId="8.[136,229,1985,2007]" captionTargetBox="[151,1436,190,1961]" captionTargetId="figure-17@8.[151,1436,190,1961]" captionTargetPageId="8" captionText="FIGURE 6. Specimen collected by Gillies in Mendoza (E00334749)." figureDoi="http://doi.org/10.5281/zenodo.10184567" httpUri="https://zenodo.org/record/10184567/files/figure.png" pageId="5" pageNumber="83">Fig. 6</figureCitation>
) is the reason why
<bibRefCitation id="DF3DAFE6B73BFFD3FABBDBDBD3F6FA28" author="Pedersen, T. M." pageId="5" pageNumber="83" pagination="69 - 97" refId="ref5994" refString="Pedersen, T. M. (1990) Studies in South American Amaranthaceae III (including one amphi-Atlantic species). Bulletin du Museum National d'Histoire Naturelle Section B 12: 69 - 97. https: // doi. org / 10.5962 / p. 326734" type="journal article" year="1990">Pedersen (1990)</bibRefCitation>
includes
<taxonomicName id="7CACA994B73BFFD3FEC2DBA7D173FA28" authorityName="Moquin-Tandon" authorityYear="1849" box="[319,598,1490,1516]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B73BFFD3FEC2DBA7D29EFA28" box="[319,443,1490,1516]" italics="true" pageId="5" pageNumber="83">G. globosa</emphasis>
var.
<emphasis id="89D80E05B73BFFD3FE09DBA7D173FA28" box="[500,598,1490,1516]" italics="true" pageId="5" pageNumber="83">albiflora</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="7CACA994B73BFFD3FCE3DBA7D08EFA28" authorityName="Moquin-Tandon" authorityYear="1849" box="[798,939,1490,1516]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="boliviana">
<emphasis id="89D80E05B73BFFD3FCE3DBA7D08EFA28" box="[798,939,1490,1516]" italics="true" pageId="5" pageNumber="83">G. boliviana</emphasis>
</taxonomicName>
and included the legend “
<emphasis id="89D80E05B73BFFD3FB29DBA6D6BDFA28" box="[1236,1432,1491,1516]" italics="true" pageId="5" pageNumber="83">pro minima parte</emphasis>
”. E00334749 has bracteoles without crest and an involucre of, at least, five reduced leaves subtending each inflorescence; these features correspond to
<taxonomicName id="7CACA994B73BFFD3FE35D86FD170F9F0" authorityName="Moquin-Tandon" authorityYear="1849" box="[456,597,1562,1588]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="boliviana">
<emphasis id="89D80E05B73BFFD3FE35D86FD170F9F0" box="[456,597,1562,1588]" italics="true" pageId="5" pageNumber="83">G. boliviana</emphasis>
</taxonomicName>
according to
<bibRefCitation id="DF3DAFE6B73BFFD3FD13D86FD0CCF9F0" author="Bena, M. J. &amp; Acosta, J. M." box="[750,1001,1562,1588]" pageId="5" pageNumber="83" pagination="72 - 100" refId="ref4459" refString="Bena, M. J. &amp; Acosta, J. M. (2020) Gomphrena. In: Anton, A. M., Zuloaga, F. O., Zuloaga, F. O. &amp; Belgrano, M. J. (eds.) Flora vascular de la Republica Argentina, vol. 19 (1). Instituto de Botanica Darwinion, Buenos Aires, pp. 72 - 100. https: // doi. org / 10.2307 / j. ctv 1 gwqr 79" type="book chapter" year="2020">Bena &amp; Acosta (2020)</bibRefCitation>
and
<bibRefCitation id="DF3DAFE6B73BFFD3FBE2D86FD7FCF9F0" author="Pedersen, T. M." box="[1055,1241,1562,1588]" pageId="5" pageNumber="83" pagination="217 - 251" refId="ref6042" refString="Pedersen, T. M. (1997) Studies in South American Amaranthaceae IV. Adansonia ser. 3 19 (2): 217 - 251." type="journal article" year="1997">Pedersen (1997)</bibRefCitation>
.
</paragraph>
<paragraph id="BB13D217B73BFFD3FF40D84BD7E2F964" blockId="5.[136,1452,373,1985]" pageId="5" pageNumber="83">
Finally, regarding Berlandier specimens, collection no. 2228 is deposited at
<collectingCountry id="C3BB9287B73BFFD3FBDED84BD762F99C" box="[1059,1095,1598,1624]" name="Philippines" pageId="5" pageNumber="83">PH</collectingCountry>
(barcode
<collectingCountry id="C3BB9287B73BFFD3FB47D84BD7FAF99C" box="[1210,1247,1598,1624]" name="Philippines" pageId="5" pageNumber="83">PH</collectingCountry>
00012667;
<figureCitation id="2397CE92B73BFFD3FAA2D84BD686F99C" box="[1375,1443,1598,1624]" captionStart="FIGURE 7" captionStartId="9.[136,229,2028,2050]" captionTargetBox="[157,1430,190,2004]" captionTargetId="figure-24@9.[157,1430,190,2004]" captionTargetPageId="9" captionText="FIGURE 7. Specimen collected by Berlandier (no. 2228) in Mexico (PH00012667)." figureDoi="http://doi.org/10.5281/zenodo.10184571" httpUri="https://zenodo.org/record/10184571/files/figure.png" pageId="5" pageNumber="83">Fig. 7</figureCitation>
) and is morphologically different from those collected by Galeotti in Oaxaca (nos. 429 and 443), matching instead
<taxonomicName id="7CACA994B73BFFD3FA6DD817D2DEF964" authority="Rothrock (1878: 233 - 234)" authorityName="Rothrock" authorityPageNumber="233 - 234" authorityYear="1878" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="nitida">
<emphasis id="89D80E05B73BFFD3FA6DD817D3ECF964" italics="true" pageId="5" pageNumber="83">G. nitida</emphasis>
<bibRefCitation id="DF3DAFE6B73BFFD3FF2DD8F3D2DEF964" author="Rothrock, J. T." box="[208,507,1670,1696]" pageId="5" pageNumber="83" pagination="53 - 351" refId="ref6196" refString="Rothrock, J. T. (1878) Plants collected in Nevada, Utah, Colorado, New Mexico and Arizona, description of those not contained in Gray's Manual of the Northern U. S., and Vol. V, Geological Exploration of the Fortieth Parallel. In: Wheeler, G. M. (Ed.) Report upon United States Geographical surveys west of the one hundredth meridian, vol. VI. Government printing office, Washington, pp. 53 - 351. https: // doi. org / 10.5962 / bhl. title. 44557" type="book chapter" year="1878">Rothrock (1878: 233234)</bibRefCitation>
</taxonomicName>
(see e.g.
<bibRefCitation id="DF3DAFE6B73BFFD3FD9BD8F3D034F964" author="Clemants, S. E." box="[614,785,1670,1696]" pageId="5" pageNumber="83" pagination="451 - 454" refId="ref4825" refString="Clemants, S. E. (2003) Gomphrena. In: Flora of North America Editorial Committee (eds.) Flora of North America North of Mexico (Magnoliophyta: Caryophyllidae, part 1), vol. 4. Oxford University Press, Oxford, pp. 451 - 454." type="book chapter" year="2003">Clemants 2003</bibRefCitation>
,
<bibRefCitation id="DF3DAFE6B73BFFD3FCE1D8F3D74FF965" author="Calderon de Rzedowski, G." box="[796,1130,1670,1697]" pageId="5" pageNumber="83" pagination="124 - 132" refId="ref4686" refString="Calderon de Rzedowski, G. (2005) Amaranthaceae. In: Calderon de Rzedowski, G. &amp; Rzedowski, J. (eds.) Flora fanerogamica del Valle de Mexico. Comision Nacional para el Conocimiento y Uso de la Biodiversidad and Instituto de Ecologia A. C., Mexico D. F., pp. 124 - 132." type="book chapter" year="2005">Calderón de Rzedowski 2005</bibRefCitation>
;
<figureCitation id="2397CE92B73BFFD3FB8BD8F3D79CF964" box="[1142,1209,1670,1696]" captionStart="FIGURE 3" captionStartId="4.[136,229,1925,1947]" captionTargetBox="[151,1436,190,1901]" captionTargetId="figure-17@4.[151,1436,190,1901]" captionTargetPageId="4" captionText="FIGURE 3. Lectotype of Gomphrena nitida (US00102795)." figureDoi="http://doi.org/10.5281/zenodo.10184561" httpUri="https://zenodo.org/record/10184561/files/figure.png" pageId="5" pageNumber="83">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="BB13D217B73BFFD3FF40D8DFD03FF900" blockId="5.[136,1452,373,1985]" box="[189,794,1706,1732]" pageId="5" pageNumber="83">
The specimen from Java (
<emphasis id="89D80E05B73BFFD3FE1DD8DFD176F900" box="[480,595,1706,1732]" italics="true" pageId="5" pageNumber="83">Zoll. 2944</emphasis>
) was not located.
</paragraph>
</subSubSection>
<subSubSection id="F3B6819CB73BFFDDFF40D8BBD2BEFEA9" lastPageId="11" lastPageNumber="89" pageId="5" pageNumber="83" type="reference_group">
<paragraph id="BB13D217B73BFFD3FF40D8BBD004F804" blockId="5.[136,1452,373,1985]" pageId="5" pageNumber="83">
<taxonomicName id="7CACA994B73BFFD3FF40D8BBD281F92C" ID-CoL="3GTL7" authorityName="Standley" authorityYear="1917" box="[189,420,1742,1768]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FF40D8BBD281F92C" box="[189,420,1742,1768]" italics="true" pageId="5" pageNumber="83">Gomphrena palmeri</emphasis>
</taxonomicName>
has been also considered a synonym of
<taxonomicName id="7CACA994B73BFFD3FC7ED8BBD708F92C" ID-CoL="3GTH5" authorityName="Martens &amp; Galeotti" authorityYear="1843" box="[899,1069,1742,1768]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="filaginoides">
<emphasis id="89D80E05B73BFFD3FC7ED8BBD708F92C" box="[899,1069,1742,1768]" italics="true" pageId="5" pageNumber="83">G. filaginoides</emphasis>
</taxonomicName>
(
<bibRefCitation id="DF3DAFE6B73BFFD3FBC2D8BBD7F3F92C" author="Borsch, T." box="[1087,1238,1742,1768]" pageId="5" pageNumber="83" pagination="56 - 83" refId="ref4553" refString="Borsch, T. (2001) Amaranthaceae. In: Stevens, W. D., Ulloa-Ulloa, C. U., Pool, A. &amp; Montiel, O. M. (eds.) Flora de Nicaragua, Tomo 1. Missouri Botanical Garden Press, St. Louis, pp. 56 - 83." type="book chapter" year="2001">Borsch 2001</bibRefCitation>
, Sánchez-del
<bibRefCitation id="DF3DAFE6B73BFFD3FA85D8BBD22CF8C8" author="Pino, I. &amp; Espadas, C. &amp; Pool, R." pageId="5" pageNumber="83" pagination="1 - 74" refId="ref6371" refString="Sanchez del Pino, I., Espadas, C. &amp; Pool, R. (2013) Taxonomy and richness of nine genera of Amaranthaceae s. s. (Caryophyllales) in the Yucatan Peninsula Biotic Province. Phytotaxa 107 (1): 1 - 74. https: // doi. org / 10.11646 / phytotaxa. 107.1.1" type="journal article" year="2013">
Pino
<emphasis id="89D80E05B73BFFD3FF75D886D399F8C8" box="[136,188,1778,1804]" italics="true" pageId="5" pageNumber="83">et al</emphasis>
. 2013
</bibRefCitation>
). This could be because Standley (1946: 165) demoted
<taxonomicName id="7CACA994B73BFFD3FC71D887D72DF8C8" ID-CoL="3GTL7" authorityName="Standley" authorityYear="1917" box="[908,1032,1778,1804]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FC71D887D72DF8C8" box="[908,1032,1778,1804]" italics="true" pageId="5" pageNumber="83">G. palmeri</emphasis>
</taxonomicName>
to a synonym of
<taxonomicName id="7CACA994B73BFFD3FB24D887D610F8C8" ID-CoL="C9HT" authorityName="Standley" authorityYear="1917" baseAuthorityName="Stuchlik" box="[1241,1333,1778,1804]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="nana">
<emphasis id="89D80E05B73BFFD3FB24D887D610F8C8" box="[1241,1333,1778,1804]" italics="true" pageId="5" pageNumber="83">G. nana</emphasis>
</taxonomicName>
, the latter in turn considered a synonym of
<taxonomicName id="7CACA994B73BFFD3FDF5D963D195F8F4" ID-CoL="3GTH5" authorityName="Martens &amp; Galeotti" authorityYear="1843" box="[520,688,1814,1840]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="filaginoides">
<emphasis id="89D80E05B73BFFD3FDF5D963D195F8F4" box="[520,688,1814,1840]" italics="true" pageId="5" pageNumber="83">G. filaginoides</emphasis>
</taxonomicName>
by
<bibRefCitation id="DF3DAFE6B73BFFD3FD1DD963D720F8F4" author="Holzhammer, E." box="[736,1029,1814,1841]" pageId="5" pageNumber="83" pagination="178 - 249" refId="ref5184" refString="Holzhammer, E. (1956) Die amerikanischen Arten der Gattung Gomphrena L. 2 Teil. Mitteilungen der Botanischen Staatssammlung Munchen 14 - 15: 178 - 249." type="journal article" year="1956">Holzhammer (1956: 196)</bibRefCitation>
. However, if the original material of both
<taxonomicName id="7CACA994B73BFFD3FF3FD94FD283F890" ID-CoL="3GTL7" authorityName="Standley" authorityYear="1917" box="[194,422,1850,1876]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FF3FD94FD283F890" box="[194,422,1850,1876]" italics="true" pageId="5" pageNumber="83">Gomphrena palmeri</emphasis>
</taxonomicName>
and
<taxonomicName id="7CACA994B73BFFD3FE23D94FD1A3F890" ID-CoL="3GTH5" authorityName="Martens &amp; Galeotti" authorityYear="1843" box="[478,646,1850,1876]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="filaginoides">
<emphasis id="89D80E05B73BFFD3FE23D94FD1A3F890" box="[478,646,1850,1876]" italics="true" pageId="5" pageNumber="83">G. filaginoides</emphasis>
</taxonomicName>
is observed (
<figureCitation id="2397CE92B73BFFD3FCE3D94FD047F890" box="[798,866,1850,1876]" captionStart="FIGURE 1" captionStartId="2.[136,229,2004,2026]" captionTargetBox="[151,1436,458,1977]" captionTargetId="figure-151@2.[151,1436,458,1977]" captionTargetPageId="2" captionText="FIGURE 1. Lectotype of Gomphrena filaginoides (BR0000006950835)." figureDoi="http://doi.org/10.5281/zenodo.10184553" httpUri="https://zenodo.org/record/10184553/files/figure.png" pageId="5" pageNumber="83">Fig. 1</figureCitation>
and
<figureCitation id="2397CE92B73BFFD3FC67D94FD08FF890" box="[922,938,1850,1876]" captionStart="FIGURE 4" captionStartId="6.[136,229,2028,2050]" captionTargetBox="[210,1377,190,2004]" captionTargetId="figure-17@6.[210,1377,190,2004]" captionTargetPageId="6" captionText="FIGURE 4. Lectotype of Gomphrena globosa var. albiflora (P00622632)." figureDoi="http://doi.org/10.5281/zenodo.10184563" httpUri="https://zenodo.org/record/10184563/files/figure.png" pageId="5" pageNumber="83">4</figureCitation>
), as well as the material that
<bibRefCitation id="DF3DAFE6B73BFFD3FB05D94FD3D6F8BD" author="Standley, P. C." pageId="5" pageNumber="83" pagination="95 - 169" refId="ref6564" refString="Standley, P. C. (1917) Amaranthaceae. In: Britton, N. L., Murrill, W. A. &amp; Barnhart (eds.) North American Flora, vol. 21 (2). The New York Botanical Garden, New York, pp. 95 - 169." type="book chapter" year="1917">Standley (1917: 149150)</bibRefCitation>
cited in the protologue of
<taxonomicName id="7CACA994B73BFFD3FDE9D92BD1A9F8BC" ID-CoL="3GTL7" authorityName="Standley" authorityYear="1917" box="[532,652,1886,1912]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FDE9D92BD1A9F8BC" box="[532,652,1886,1912]" italics="true" pageId="5" pageNumber="83">G. palmeri</emphasis>
</taxonomicName>
(
<figureCitation id="2397CE92B73BFFD3FD67D92BD1F9F8BD" box="[666,732,1886,1913]" captionStart="FIGURE 5" captionStartId="7.[136,229,1933,1955]" captionTargetBox="[151,1436,190,1909]" captionTargetId="figure-24@7.[151,1436,190,1909]" captionTargetPageId="7" captionText="FIGURE 5. Specimen of Gomphrena palmeri (US00102797)." figureDoi="http://doi.org/10.5281/zenodo.10184565" httpUri="https://zenodo.org/record/10184565/files/figure.png" pageId="5" pageNumber="83">Fig. 5</figureCitation>
), they refer to two different species.
<taxonomicName id="7CACA994B73BFFD3FB8DD92BD631F8BC" ID-CoL="3GTH5" authorityName="Martens &amp; Galeotti" authorityYear="1843" box="[1136,1300,1886,1912]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="filaginoides">
<emphasis id="89D80E05B73BFFD3FB8DD92BD631F8BC" box="[1136,1300,1886,1912]" italics="true" pageId="5" pageNumber="83">G. filaginoides</emphasis>
</taxonomicName>
is a cespitose plant with short internodes and short crested bracteoles, while
<taxonomicName id="7CACA994B73BFFD3FCA9D9F7D0EAF858" ID-CoL="3GTL7" authorityName="Standley" authorityYear="1917" box="[852,975,1922,1948]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="83" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B73BFFD3FCA9D9F7D0EAF858" box="[852,975,1922,1948]" italics="true" pageId="5" pageNumber="83">G. palmeri</emphasis>
</taxonomicName>
has procumbent to ascendant stems, with internodes longer than
<quantity id="7C547FF2B73BFFD3FE74D9D3D2E6F804" box="[393,451,1958,1985]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="83" unit="cm" value="5.0">5 cm</quantity>
, and widely crested bracteoles.
</paragraph>
<caption id="EFD3829FB738FFD0FF75D999D070F7C6" ID-DOI="http://doi.org/10.5281/zenodo.10184563" ID-Zenodo-Dep="10184563" box="[136,853,2028,2051]" httpUri="https://zenodo.org/record/10184563/files/figure.png" pageId="6" pageNumber="84" startId="6.[136,229,2028,2050]" targetBox="[210,1377,190,2004]" targetPageId="6" targetType="figure">
<paragraph id="BB13D217B738FFD0FF75D999D070F7C6" blockId="6.[136,853,2028,2051]" box="[136,853,2028,2051]" pageId="6" pageNumber="84">
<emphasis id="89D80E05B738FFD0FF75D999D3DBF7C5" bold="true" box="[136,254,2028,2050]" pageId="6" pageNumber="84">FIGURE 4.</emphasis>
Lectotype of
<taxonomicName id="7CACA994B738FFD0FE7ED998D1EFF7C6" authorityName="Moquin-Tandon" authorityYear="1849" box="[387,714,2029,2050]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="84" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B738FFD0FE7ED998D162F7C6" box="[387,583,2029,2050]" italics="true" pageId="6" pageNumber="84">Gomphrena globosa</emphasis>
var.
<emphasis id="89D80E05B738FFD0FD8BD998D1EFF7C6" box="[630,714,2029,2050]" italics="true" pageId="6" pageNumber="84">albiflora</emphasis>
</taxonomicName>
(P00622632).
</paragraph>
</caption>
<caption id="EFD3829FB739FFD1FF75D9F8D1C5F867" ID-DOI="http://doi.org/10.5281/zenodo.10184565" ID-Zenodo-Dep="10184565" box="[136,736,1933,1956]" httpUri="https://zenodo.org/record/10184565/files/figure.png" pageId="7" pageNumber="85" startId="7.[136,229,1933,1955]" targetBox="[151,1436,190,1909]" targetPageId="7" targetType="figure">
<paragraph id="BB13D217B739FFD1FF75D9F8D1C5F867" blockId="7.[136,736,1933,1956]" box="[136,736,1933,1956]" pageId="7" pageNumber="85">
<emphasis id="89D80E05B739FFD1FF75D9F8D3D8F867" bold="true" box="[136,253,1933,1955]" pageId="7" pageNumber="85">FIGURE 5.</emphasis>
Specimen of
<taxonomicName id="7CACA994B739FFD1FE7FD9FBD160F867" authorityName="Standley" authorityYear="1917" box="[386,581,1934,1955]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="7" pageNumber="85" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B739FFD1FE7FD9FBD160F867" box="[386,581,1934,1955]" italics="true" pageId="7" pageNumber="85">Gomphrena palmeri</emphasis>
</taxonomicName>
(US00102797).
</paragraph>
</caption>
<caption id="EFD3829FB736FFDEFF75D9B4D003F813" ID-DOI="http://doi.org/10.5281/zenodo.10184567" ID-Zenodo-Dep="10184567" box="[136,806,1985,2008]" httpUri="https://zenodo.org/record/10184567/files/figure.png" pageId="8" pageNumber="86" startId="8.[136,229,1985,2007]" targetBox="[151,1436,190,1961]" targetPageId="8" targetType="figure">
<paragraph id="BB13D217B736FFDEFF75D9B4D003F813" blockId="8.[136,806,1985,2008]" box="[136,806,1985,2008]" pageId="8" pageNumber="86">
<emphasis id="89D80E05B736FFDEFF75D9B4D3D8F812" bold="true" box="[136,253,1985,2007]" pageId="8" pageNumber="86">FIGURE 6.</emphasis>
Specimen collected by Gillies in Mendoza (E00334749).
</paragraph>
</caption>
<caption id="EFD3829FB737FFDFFF75D999D093F7C6" ID-DOI="http://doi.org/10.5281/zenodo.10184571" ID-Zenodo-Dep="10184571" box="[136,950,2028,2050]" httpUri="https://zenodo.org/record/10184571/files/figure.png" pageId="9" pageNumber="87" startId="9.[136,229,2028,2050]" targetBox="[157,1430,190,2004]" targetPageId="9" targetType="figure">
<paragraph id="BB13D217B737FFDFFF75D999D093F7C6" blockId="9.[136,950,2028,2050]" box="[136,950,2028,2050]" pageId="9" pageNumber="87">
<emphasis id="89D80E05B737FFDFFF75D999D3DBF7C6" bold="true" box="[136,254,2028,2050]" pageId="9" pageNumber="87">FIGURE 7.</emphasis>
Specimen collected by Berlandier (no. 2228) in Mexico (PH00012667).
</paragraph>
</caption>
<caption id="EFD3829FB734FFDCFF75D99FD1C6F83B" ID-DOI="http://doi.org/10.5281/zenodo.10184573" ID-Zenodo-Dep="10184573" box="[136,739,2026,2048]" httpUri="https://zenodo.org/record/10184573/files/figure.png" pageId="10" pageNumber="88" startId="10.[136,229,2026,2048]" targetBox="[151,1436,190,2002]" targetPageId="10" targetType="figure">
<paragraph id="BB13D217B734FFDCFF75D99FD1C6F83B" blockId="10.[136,739,2026,2048]" box="[136,739,2026,2048]" pageId="10" pageNumber="88">
<emphasis id="89D80E05B734FFDCFF75D99FD3D8F83B" bold="true" box="[136,253,2026,2048]" pageId="10" pageNumber="88">FIGURE 8.</emphasis>
Lectotype of
<taxonomicName id="7CACA994B734FFDCFE7ED99FD162F83B" authorityName="Coulter &amp; Fisher" authorityYear="1892" box="[387,583,2026,2047]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="88" phylum="Tracheophyta" rank="species" species="pringlei">
<emphasis id="89D80E05B734FFDCFE7ED99FD162F83B" box="[387,583,2026,2047]" italics="true" pageId="10" pageNumber="88">Gomphrena pringlei</emphasis>
</taxonomicName>
(US00102801).
</paragraph>
</caption>
<paragraph id="BB13D217B735FFDDFF40DEEBD2BEFEA9" blockId="11.[136,1452,158,437]" pageId="11" pageNumber="89">
Likewise, when
<bibRefCitation id="DF3DAFE6B735FFDDFE8ADEEBD17AFF7D" author="Stuchlik, V. J." box="[375,607,158,185]" pageId="11" pageNumber="89" pagination="151 - 162" refId="ref6665" refString="Stuchlik, V. J. (1912) Zur synonymik der Gattung Gomphrena. Repertorium Specierum Novarum Regni Vegetabilis 11: 151 - 162. https: // doi. org / 10.1002 / fedr. 19120110906" type="journal article" year="1912">Stuchlík (1912: 158)</bibRefCitation>
first described
<taxonomicName id="7CACA994B735FFDDFCF3DEEBD7A2FF7C" ID-CoL="6L267" baseAuthorityName="Stuchlik" baseAuthorityYear="1912" box="[782,1159,158,185]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="89" phylum="Tracheophyta" rank="variety" species="decumbens" variety="nana">
<emphasis id="89D80E05B735FFDDFCF3DEEBD733FF7C" box="[782,1046,158,184]" italics="true" pageId="11" pageNumber="89">Gomphrena decumbens</emphasis>
var.
<emphasis id="89D80E05B735FFDDFBB2DEEAD7A2FF7C" box="[1103,1159,159,184]" italics="true" pageId="11" pageNumber="89">nana</emphasis>
</taxonomicName>
, he refers to a small plant (“planta parva”), and, as it can be observed in the material cited in the protologue [Z000000292 (
<figureCitation id="2397CE92B735FFDDFB44DEB7D7D9FF18" box="[1209,1276,194,220]" captionStart="FIGURE 2" captionStartId="3.[136,229,1767,1789]" captionTargetBox="[282,1306,190,1743]" captionTargetId="figure-24@3.[282,1306,190,1743]" captionTargetPageId="3" captionText="FIGURE 2. Lectotype of Gomphrena decumbens var. nana (Z000000292 by United Herbaria Z+ZT / CC BY 4.0)." figureDoi="http://doi.org/10.5281/zenodo.10184559" httpUri="https://zenodo.org/record/10184559/files/figure.png" pageId="11" pageNumber="89">Fig. 2</figureCitation>
),
<collectingCountry id="C3BB9287B735FFDDFAECDEB7D611FF18" box="[1297,1332,194,220]" name="United States of America" pageId="11" pageNumber="89">US</collectingCountry>
00102786,
<collectingCountry id="C3BB9287B735FFDDFF75DE93D389FEC4" box="[136,172,230,256]" name="United States of America" pageId="11" pageNumber="89">US</collectingCountry>
00931477],
<taxonomicName id="7CACA994B735FFDDFEC8DE92D16FFEC4" ID-CoL="6L267" baseAuthorityName="Stuchlik" baseAuthorityYear="1912" box="[309,586,230,257]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="89" phylum="Tracheophyta" rank="variety" species="decumbens" variety="nana">
<emphasis id="89D80E05B735FFDDFEC8DE92D2F3FEC4" box="[309,470,230,257]" italics="true" pageId="11" pageNumber="89">G. decumbens</emphasis>
var.
<emphasis id="89D80E05B735FFDDFDEFDE92D16FFEC4" box="[530,586,231,256]" italics="true" pageId="11" pageNumber="89">nana</emphasis>
</taxonomicName>
corresponds to a small plant, more similar in habit and inflorescence to
<taxonomicName id="7CACA994B735FFDDFA6DDE92D229FEE0" ID-CoL="3GTH5" authorityName="Martens &amp; Galeotti" authorityYear="1843" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="89" phylum="Tracheophyta" rank="species" species="filaginoides">
<emphasis id="89D80E05B735FFDDFA6DDE92D229FEE0" italics="true" pageId="11" pageNumber="89">G. filaginoides</emphasis>
</taxonomicName>
(
<figureCitation id="2397CE92B735FFDDFEE3DF7FD247FEE0" box="[286,354,266,292]" captionStart="FIGURE 1" captionStartId="2.[136,229,2004,2026]" captionTargetBox="[151,1436,458,1977]" captionTargetId="figure-151@2.[151,1436,458,1977]" captionTargetPageId="2" captionText="FIGURE 1. Lectotype of Gomphrena filaginoides (BR0000006950835)." figureDoi="http://doi.org/10.5281/zenodo.10184553" httpUri="https://zenodo.org/record/10184553/files/figure.png" pageId="11" pageNumber="89">Fig. 1</figureCitation>
) (see also discussion in
<bibRefCitation id="DF3DAFE6B735FFDDFD7EDF7FD0B6FEE0" author="Holzhammer, E." box="[643,915,266,293]" pageId="11" pageNumber="89" pagination="178 - 249" refId="ref5184" refString="Holzhammer, E. (1956) Die amerikanischen Arten der Gattung Gomphrena L. 2 Teil. Mitteilungen der Botanischen Staatssammlung Munchen 14 - 15: 178 - 249." type="journal article" year="1956">Holzhammer 1956: 196</bibRefCitation>
) than to the material cited by
<bibRefCitation id="DF3DAFE6B735FFDDFB0BDF7FD3D6FE8D" author="Standley, P. C." pageId="11" pageNumber="89" pagination="95 - 169" refId="ref6564" refString="Standley, P. C. (1917) Amaranthaceae. In: Britton, N. L., Murrill, W. A. &amp; Barnhart (eds.) North American Flora, vol. 21 (2). The New York Botanical Garden, New York, pp. 95 - 169." type="book chapter" year="1917">Standley (1917: 149150)</bibRefCitation>
(
<figureCitation id="2397CE92B735FFDDFEF8DF5BD26EFE8D" box="[261,331,302,329]" captionStart="FIGURE 5" captionStartId="7.[136,229,1933,1955]" captionTargetBox="[151,1436,190,1909]" captionTargetId="figure-24@7.[151,1436,190,1909]" captionTargetPageId="7" captionText="FIGURE 5. Specimen of Gomphrena palmeri (US00102797)." figureDoi="http://doi.org/10.5281/zenodo.10184565" httpUri="https://zenodo.org/record/10184565/files/figure.png" pageId="11" pageNumber="89">Fig. 5</figureCitation>
), and the material designated by
<bibRefCitation id="DF3DAFE6B735FFDDFD31DF5AD07FFE8C" author="Bena, M. J." box="[716,858,302,329]" pageId="11" pageNumber="89" pagination="131 - 146" refId="ref4410" refString="Bena, M. J. (2017) Typification and nomenclatural notes on twenty-seven names of Gomphrena L. (Amaranthaceae) linked to the South American flora. Phytotaxa 296: 131 - 146. https: // doi. org / 10.11646 / phytotaxa. 296.2.3" type="journal article" year="2017">Bena (2017)</bibRefCitation>
as a
<typeStatus id="64176CB5B735FFDDFC64DF5BD725FE8C" box="[921,1024,302,328]" pageId="11" pageNumber="89" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="7CACA994B735FFDDFBD7DF5AD660FE8C" ID-CoL="6KPRW" authorityName="Moquin-Tandon" authorityYear="1849" box="[1066,1349,302,329]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="89" phylum="Tracheophyta" rank="variety" species="globosa" variety="albiflora">
<emphasis id="89D80E05B735FFDDFBD7DF5AD782FE8C" box="[1066,1191,302,329]" italics="true" pageId="11" pageNumber="89">G. globosa</emphasis>
var.
<emphasis id="89D80E05B735FFDDFB1EDF5BD660FE8C" box="[1251,1349,302,328]" italics="true" pageId="11" pageNumber="89">albiflora</emphasis>
</taxonomicName>
(
<figureCitation id="2397CE92B735FFDDFAABDF5BD6B8FE8C" box="[1366,1437,302,328]" captionStart="FIGURE 4" captionStartId="6.[136,229,2028,2050]" captionTargetBox="[210,1377,190,2004]" captionTargetId="figure-17@6.[210,1377,190,2004]" captionTargetPageId="6" captionText="FIGURE 4. Lectotype of Gomphrena globosa var. albiflora (P00622632)." figureDoi="http://doi.org/10.5281/zenodo.10184563" httpUri="https://zenodo.org/record/10184563/files/figure.png" pageId="11" pageNumber="89">Fig. 4</figureCitation>
), basionym of
<taxonomicName id="7CACA994B735FFDDFEE7DF26D2B6FEA8" ID-CoL="3GTL7" authorityName="Standley" authorityYear="1917" box="[282,403,338,365]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="89" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B735FFDDFEE7DF26D2B6FEA8" box="[282,403,338,365]" italics="true" pageId="11" pageNumber="89">G. palmeri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F3B6819CB735FFDDFF40DF02D1DDFE70" pageId="11" pageNumber="89" type="description">
<paragraph id="BB13D217B735FFDDFF40DF02D1DDFE70" blockId="11.[136,1452,158,437]" pageId="11" pageNumber="89">
Here, we agree with
<bibRefCitation id="DF3DAFE6B735FFDDFE5BDF03D1A9FE55" author="Holzhammer, E." box="[422,652,374,401]" pageId="11" pageNumber="89" pagination="178 - 249" refId="ref5184" refString="Holzhammer, E. (1956) Die amerikanischen Arten der Gattung Gomphrena L. 2 Teil. Mitteilungen der Botanischen Staatssammlung Munchen 14 - 15: 178 - 249." type="journal article" year="1956">Holzhammer (1956)</bibRefCitation>
, who includes
<taxonomicName id="7CACA994B735FFDDFCC9DF02D767FE54" baseAuthorityName="Stuchlik" baseAuthorityYear="1912" box="[820,1090,374,401]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="89" phylum="Tracheophyta" rank="variety" species="decumbens" variety="nana">
<emphasis id="89D80E05B735FFDDFCC9DF02D0F6FE54" box="[820,979,374,401]" italics="true" pageId="11" pageNumber="89">G. decumbens</emphasis>
var.
<emphasis id="89D80E05B735FFDDFBF7DF02D767FE54" box="[1034,1090,375,400]" italics="true" pageId="11" pageNumber="89">nana</emphasis>
</taxonomicName>
and
<taxonomicName id="7CACA994B735FFDDFB84DF02D7F6FE54" authorityName="Standley" authorityYear="1917" baseAuthorityName="Stuchlik" box="[1145,1235,375,401]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="89" phylum="Tracheophyta" rank="species" species="nana">
<emphasis id="89D80E05B735FFDDFB84DF02D7F6FE54" box="[1145,1235,375,401]" italics="true" pageId="11" pageNumber="89">G. nana</emphasis>
</taxonomicName>
as synonyms of
<taxonomicName id="7CACA994B735FFDDFA6DDF02D229FE70" authorityName="Martens &amp; Galeotti" authorityYear="1843" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="89" phylum="Tracheophyta" rank="species" species="filaginoides">
<emphasis id="89D80E05B735FFDDFA6DDF02D229FE70" italics="true" pageId="11" pageNumber="89">G. filaginoides</emphasis>
</taxonomicName>
and treats
<taxonomicName id="7CACA994B735FFDDFE7BDFEED125FE70" authorityName="Standley" authorityYear="1917" box="[390,512,410,437]" class="Magnoliopsida" family="Amaranthaceae" genus="Gomphrena" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="89" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis id="89D80E05B735FFDDFE7BDFEED125FE70" box="[390,512,410,437]" italics="true" pageId="11" pageNumber="89">G. palmeri</emphasis>
</taxonomicName>
as a different species.
</paragraph>
</subSubSection>
</treatment>
</document>