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<document id="9271FD6C862E088EFCEF8D3BE96A837B" ID-CLB-Dataset="4184" ID-DOI="10.5852/ejt.2021.768.1501" ID-GBIF-Dataset="1361fbca-2ed1-4f60-85cc-1a3f62f73d79" ID-ISSN="2118-9773" ID-Zenodo-Dep="5535911" ID-ZooBank="58117DB6-B3F3-41BF-81B3-8AF089708922" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632918358292" checkinUser="felipe" docAuthor="Annisaqois, Manikmayang &amp; Wägele, J. Wolfgang" docDate="2021" docId="1A6987B821021F1BFD0167C8FCB5C27D" docLanguage="en" docName="ejt-768_1-52.pdf" docOrigin="European Journal of Taxonomy 768" docSource="http://dx.doi.org/10.5852/ejt.2021.768.1501" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Hyssura reptans Annisaqois &amp; Wägele 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="E650FFC021001F12FF846470FFA2C33A" masterDocTitle="Morphology and Taxonomy of Isopoda Anthuroidea (Crustacea) from Sulawesi with description of six new species" masterLastPageNumber="52" masterPageNumber="1" pageNumber="3" updateTime="1699458108869" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EF66C987719EE3564936621C5DA3B4AC">Morphology and Taxonomy of Isopoda Anthuroidea (Crustacea) from Sulawesi with description of six new species</mods:title>
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<mods:namePart id="C2E86402FA76C68B722579E7AE27C0DB">Annisaqois, Manikmayang</mods:namePart>
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<mods:affiliation id="DCE2FD7C0049779C09BC7BFBE708124F">Faculty of Fisheries and Marine Sciences, Sam Ratulangi University, Jln Kampus UNSRAT - Bahu, Manado 95115, Indonesia.</mods:affiliation>
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<mods:namePart id="52BF04E9D9BDB3A390CECC784E9D41ED">Wägele, J. Wolfgang</mods:namePart>
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<mods:affiliation id="323C006B5889E1F223CCC1BFA302BA8A">Zoological Research Museum Alexander Koenig, Adenauerallee 160, 53113 Bonn, Germany.</mods:affiliation>
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<mods:date id="CDCF7B00289E67C86859B0597AD703BD">2021</mods:date>
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<subSubSection id="DADA652521021F10FD0167C8FC0DC0E9" box="[645,943,952,979]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="927F36AE21021F10FD0167C8FC0DC0E9" blockId="2.[362,1225,952,1050]" box="[645,943,952,979]" pageId="2" pageNumber="3">
<heading id="C93781C221021F10FD0167C8FC0DC0E9" box="[645,943,952,979]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<taxonomicName id="55C04D2D21021F10FD0167C8FCEFC0E8" ID-CoL="8QCDJ" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[645,845,952,978]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="reptans" status="sp. nov.">
<emphasis id="A0B4EABC21021F10FD0167C8FCEFC0E8" bold="true" box="[645,845,952,978]" italics="true" pageId="2" pageNumber="3">Hyssura reptans</emphasis>
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<taxonomicNameLabel id="BB8757C721021F10FCD067C9FC0DC0E9" box="[852,943,953,979]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act: 
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<figureCitation id="0AFB2A2B21021F10FD63678FFCEEC720" box="[743,844,1023,1050]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="5.[189,232,1996,2022]" captionStartId-1="6.[189,232,1963,1989]" captionStartId-2="7.[189,232,1958,1984]" captionStartId-3="8.[189,232,1996,2022]" captionTargetBox-0="[370,1387,279,1889]" captionTargetBox-1="[215,1365,276,1883]" captionTargetBox-2="[198,1392,275,1833]" captionTargetBox-3="[201,1361,896,1922]" captionTargetId-0="figure-12@5.[247,1402,261,1922]" captionTargetId-1="figure-15@6.[186,1401,261,1907]" captionTargetId-2="figure-12@7.[186,1401,262,1923]" captionTargetId-3="figure-15@8.[189,1401,870,1960]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="Fig. 2. Hyssura reptans sp. nov., ♂, holotype in dorsal and lateral view (2.1 mm) (MZB Iso 099)." captionText-1="Fig. 3. Hyssura reptan sp. nov., ♂, holotype (MZB Iso 099). Abbreviations: A1 = antenna 1; A2 = antenna 2; P1–P3 = pereopods 1–3." captionText-2="Fig. 4. Hyssura reptans sp. nov., ♂, holotype (MZB Iso 099). Abbreviations: P4–P7 = pereopods 4–7; Plp1–2 = pleopods 1–2." captionText-3="Fig. 5. Hyssura reptans sp. nov., ♂, holotype. Abbreviations: Urp = uropod; Tel = telson." figureDoi-0="http://doi.org/10.5281/zenodo.5535915" figureDoi-1="http://doi.org/10.5281/zenodo.5535917" figureDoi-2="http://doi.org/10.5281/zenodo.5535921" figureDoi-3="http://doi.org/10.5281/zenodo.5535923" httpUri-0="https://zenodo.org/record/5535915/files/figure.png" httpUri-1="https://zenodo.org/record/5535917/files/figure.png" httpUri-2="https://zenodo.org/record/5535921/files/figure.png" httpUri-3="https://zenodo.org/record/5535923/files/figure.png" pageId="2" pageNumber="3">Figs 2–5</figureCitation>
</paragraph>
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<subSubSection id="DADA652521021F10FF396030FAE4C517" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="927F36AE21021F10FF396030FE6BC760" blockId="2.[189,457,1088,1114]" box="[189,457,1088,1114]" pageId="2" pageNumber="3">
<heading id="C93781C221021F10FF396030FE6BC760" bold="true" box="[189,457,1088,1114]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="A0B4EABC21021F10FF396030FE6BC760" bold="true" box="[189,457,1088,1114]" pageId="2" pageNumber="3">Differential diagnosis</emphasis>
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<paragraph id="927F36AE21021F10FF39601FFC42C6B8" blockId="2.[188,1401,1135,1410]" pageId="2" pageNumber="3">
Blind species of 
<taxonomicName id="55C04D2D21021F10FE0F6000FE4CC7B3" authorityName="Norman &amp; Stebbing" authorityYear="1886" box="[395,494,1136,1161]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21021F10FE0F6000FE4CC7B3" box="[395,494,1136,1161]" italics="true" pageId="2" pageNumber="3">Hyssura</emphasis>
</taxonomicName>
(eyes also lacking in male), pereopod 1 propodus palm concave in male, a comb of 6 setae on mesial surface in addition to two stronger medial setae and distally 4 setae, pleopod rami rectangular, pleopod 1 exopod not larger than endopod; exopods of pleopods 1 and 2 distally with more, 10 resp. 9 swimming setae, endopods with 5, uropod exopod narrow lanceolate, outer lateral margin smooth, medial margin slightly serrated, apex with several long simple setae, endopod of uropod as wide as sympod but about 2.5 times longer, a bit shorter than exopod, proximal two thirds of margins nearly parallel, distal margins with simple setae of varying length. Telson tongue-like, lateral margins serrated, not parallel; apex rounded and with 5 pairs of simple setae.
</paragraph>
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<taxonomicName id="55C04D2D21021F10FF3961DAFD96C6F9" authority="Wagele, 1981" authorityName="Wagele" authorityYear="1981" box="[189,564,1449,1475]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ligurica">
<emphasis id="A0B4EABC21021F10FF3961DAFE24C6F9" box="[189,390,1449,1475]" italics="true" pageId="2" pageNumber="3">Hyssura ligurica</emphasis>
Wägele, 1981
</taxonomicName>
(see 
<bibRefCitation id="F6514B5F21021F10FDF661D9FCB9C6F9" author="Wagele J. W." box="[626,795,1449,1475]" pageId="2" pageNumber="3" pagination="1 - 2" refId="ref13936" refString="Wagele J. W. 1981 a. Study of the Hyssuridae (Crustacea: Isopoda: Anthuridea) from the Mediterranean and the Red Sea. Israel Journal of Zoology 30: 1 - 2." type="journal article" year="1981">Wägele 1981a</bibRefCitation>
) is most similar, with a tail fan that resembles that of 
<taxonomicName id="55C04D2D21021F10FF6561BDFEC3C6DC" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[225,353,1485,1510]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="reptans" status="sp. nov.">
<emphasis id="A0B4EABC21021F10FF6561BDFEC3C6DC" box="[225,353,1485,1510]" italics="true" pageId="2" pageNumber="3">H. reptans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721021F10FEE861BDFE69C6DD" box="[364,459,1485,1511]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="55C04D2D21021F10FE5261BDFD00C6DC" authorityName="Wagele" authorityYear="1981" box="[470,674,1484,1510]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ligurica">
<emphasis id="A0B4EABC21021F10FE5261BDFD00C6DC" box="[470,674,1484,1510]" italics="true" pageId="2" pageNumber="3">Hyssura ligurica</emphasis>
</taxonomicName>
also has rectangular pleopod rami, but in comparison with 
<taxonomicName id="55C04D2D21021F10FF396180FE99C533" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[189,315,1520,1545]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="reptans">
<emphasis id="A0B4EABC21021F10FF396180FE99C533" box="[189,315,1520,1545]" italics="true" pageId="2" pageNumber="3">H. reptans</emphasis>
</taxonomicName>
longer meri and carpi in P4–7 and only one sensory spine on the propodi. The female has strong propodi of P1 and P2; unfortunately, the females of 
<taxonomicName id="55C04D2D21021F10FCEB6264FC49C517" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[879,1003,1556,1581]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="reptans">
<emphasis id="A0B4EABC21021F10FCEB6264FC49C517" box="[879,1003,1556,1581]" italics="true" pageId="2" pageNumber="3">H. reptans</emphasis>
</taxonomicName>
still remain to be discovered.
</paragraph>
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<subSubSection id="DADA652521021F10FF396225FCBDC5A4" box="[189,799,1621,1694]" pageId="2" pageNumber="3" type="etymology">
<paragraph id="927F36AE21021F10FF396225FEE6C555" blockId="2.[189,324,1621,1647]" box="[189,324,1621,1647]" pageId="2" pageNumber="3">
<heading id="C93781C221021F10FF396225FEE6C555" bold="true" box="[189,324,1621,1647]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="A0B4EABC21021F10FF396225FEE6C555" bold="true" box="[189,324,1621,1647]" pageId="2" pageNumber="3">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="927F36AE21021F10FF3962F3FCBDC5A4" blockId="2.[189,799,1667,1694]" box="[189,799,1667,1694]" pageId="2" pageNumber="3">
The species epithet is derived from 
<emphasis id="A0B4EABC21021F10FDE462F4FD3FC5A7" box="[608,669,1668,1693]" italics="true" pageId="2" pageNumber="3">repto</emphasis>
(to crawl).
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<subSubSection id="DADA652521021F10FF3962B5FB0EC46F" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="927F36AE21021F10FF3962B5FE08C5E5" blockId="2.[189,426,1733,1759]" box="[189,426,1733,1759]" pageId="2" pageNumber="3">
<heading id="C93781C221021F10FF3962B5FE08C5E5" bold="true" box="[189,426,1733,1759]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="A0B4EABC21021F10FF3962B5FE08C5E5" bold="true" box="[189,426,1733,1759]" pageId="2" pageNumber="3">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="927F36AE21021F10FF6E6284FB0EC46F" blockId="2.[189,1399,1779,1877]" pageId="2" pageNumber="3">
<materialsCitation id="22A83CF321021F10FF6E6284FB0EC46F" ID-GBIF-Occurrence="3359244302" collectedFrom="in coral rubble" collectingDate="2019-09" collectionCode="MZB" collectorName="M. Annisaqois &amp; J. W. Wagele" country="Indonesia" elevation="5" latitude="1.7743139" location="Bangka Island" longLatPrecision="1" longitude="125.18876" municipality="Bangka Island" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="A0B4EABC21021F10FF6E6284FD90C437" bold="true" box="[234,562,1779,1806]" pageId="2" pageNumber="3">
<typeStatus id="4D7B880C21021F10FF6E6284FEFEC434" box="[234,348,1780,1806]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
and type locality
</emphasis>
<collectingCountry id="EAD7763E21021F10FF396367FEF9C40B" box="[189,347,1815,1841]" name="Indonesia" pageId="2" pageNumber="3">INDONESIA</collectingCountry>
• 
<specimenCount id="84C6FD2721021F10FEF16368FE29C408" box="[373,395,1816,1842]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
(
<quantity id="55389B4B21021F10FE1F6367FE5EC408" box="[411,508,1815,1842]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="3" unit="mm" value="2.1">2.1 mm</quantity>
); 
<collectingMunicipality id="721BACD421021F10FD946367FD1FC40B" box="[528,701,1815,1841]" country="Indonesia" latitude="1.7743139" longLatPrecision="1" longitude="125.18876" municipality="Areng Kambing" name="Bangka Island" pageId="2" pageNumber="3">Bangka Island</collectingMunicipality>
, 
<locationDeviation id="83CD2F8C21021F10FD4C6367FC10C40B" box="[712,946,1815,1841]" pageId="2" pageNumber="3">off Areng Kambing</locationDeviation>
; 
<geoCoordinate id="F7F4506921021F10FC3A6366FBC9C408" box="[958,1131,1814,1842]" degrees="1" direction="north" minutes="46" orientation="latitude" pageId="2" pageNumber="3" precision="1" seconds="27.53" value="1.7743139">1°46′27.53″ N</geoCoordinate>
, 
<geoCoordinate id="F7F4506921021F10FBF36366FA9EC408" box="[1143,1340,1814,1842]" degrees="125" direction="east" minutes="11" orientation="longitude" pageId="2" pageNumber="3" precision="1" seconds="19.53" value="125.18876">125°11′19.53″ E</geoCoordinate>
; 
<elevation id="19EDD19D21021F10FACD6367FEA0C46E" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="3" unit="m" value="5.0">
<quantity id="55389B4B21021F10FACD6367FAD5C408" box="[1353,1399,1815,1842]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="3" unit="m" value="5.0">5 m</quantity>
depth
</elevation>
; 
<collectedFrom id="21DC755321021F10FE8A634AFE1FC46E" box="[270,445,1850,1876]" pageId="2" pageNumber="3">in coral rubble</collectedFrom>
; 
<collectorName id="3F35537821021F10FE4C634BFDD1C46E" box="[456,627,1850,1877]" pageId="2" pageNumber="3">M. Annisaqois</collectorName>
and 
<collectorName id="3F35537821021F10FD28634BFCE2C46E" box="[684,832,1850,1877]" pageId="2" pageNumber="3">J.W. Wägele</collectorName>
leg.; 
<date id="E67E106E21021F10FC06634BFC58C46F" box="[898,1018,1851,1877]" pageId="2" pageNumber="3" value="2019-09">
<collectingDate id="F63AE98621021F10FC06634BFC58C46F" box="[898,1018,1851,1877]" pageId="2" pageNumber="3" value="2019-09">Sep. 2019</collectingDate>
</date>
; 
<collectionCode id="F4D1AE6B21021F10FB83634BFBE5C46F" box="[1031,1095,1851,1877]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="3" type="Museum">MZB</collectionCode>
Iso 099.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DADA652521021F1AFF39630CFBE1C28A" lastPageId="8" lastPageNumber="9" pageId="2" pageNumber="3" type="description">
<paragraph id="927F36AE21021F10FF39630CFD84C4AC" blockId="2.[189,550,1916,1942]" box="[189,550,1916,1942]" pageId="2" pageNumber="3">
<heading id="C93781C221021F10FF39630CFD84C4AC" bold="true" box="[189,550,1916,1942]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="A0B4EABC21021F10FF39630CFD84C4AC" bold="true" box="[189,550,1916,1942]" pageId="2" pageNumber="3">
Description of 
<typeStatus id="4D7B880C21021F10FEF1630CFE09C4AC" box="[373,427,1916,1942]" pageId="2" pageNumber="3">type</typeStatus>
specimen
</emphasis>
</heading>
</paragraph>
<paragraph id="927F36AE21021F11FF3963DBFD39C530" blockId="2.[189,1399,1963,2025]" lastBlockId="3.[189,1399,1395,1546]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">BODY. Without chromatophores, 15–16 times as long as wide (rostrum to tip of telson). Head 1.7 times as long as wide, with a lateral constriction at half of its length. Eyes lacking. A small rostral projection between antennae. Pereonites dorsally smooth, as long as wide, length ratio of pereonites: 1&lt;2&gt;3&lt;4&gt; 5&gt; 6&gt; 7, pereonite 3 being about as long as pereonite 7. Pleonites 1–5 free, as wide as pereonites, total length of pleonites 1–5 exceeds length of pereonites 6 + 7. Telson about as long as uropods and shorter than three pleonites.</paragraph>
<caption id="C6BF662621031F11FF396579FC37C219" ID-Table-UUID="C6BF662621031F11FF396579FC37C219" box="[189,917,265,291]" httpUri="http://table.plazi.org/id/C6BF662621031F11FF396579FC37C219" pageId="3" pageNumber="4" startId="3.[189,259,265,291]" targetBox="[200,1387,337,1314]" targetIsTable="true" targetPageId="3">
<paragraph id="927F36AE21031F11FF396579FC37C219" blockId="3.[189,917,265,291]" box="[189,917,265,291]" pageId="3" pageNumber="4">
<emphasis id="A0B4EABC21031F11FF396579FE83C219" bold="true" box="[189,289,265,291]" pageId="3" pageNumber="4">Table 1.</emphasis>
Anthuridea collected on the shores of Bangka Island.
</paragraph>
</caption>
<paragraph id="927F36AE21031F11FF4C6521FA9FC7DE" pageId="3" pageNumber="4">
<table id="E0C0C40E2103E0EDFF4C6521FAC9C618" box="[200,1387,337,1314]" gridcols="6" gridrows="10" pageId="3" pageNumber="4">
<tr id="2CF034EC2103E0EDFF4C6521FAC9C2B3" box="[200,1387,337,393]" gridrow="0" pageId="3" pageNumber="4">
<th id="6F215D902103E0EDFF4C6521FE0FC2B3" box="[200,429,337,393]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="A0B4EABC21031F11FF4C6521FEC3C250" bold="true" box="[200,353,337,362]" pageId="3" pageNumber="4">Species name</emphasis>
</th>
<th id="6F215D902103E0EDFE4D6521FCE4C2B3" box="[457,838,337,393]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="A0B4EABC21031F11FE4D6521FDDBC253" bold="true" box="[457,633,337,361]" pageId="3" pageNumber="4">Sample locality</emphasis>
</th>
<th id="6F215D902103E0EDFCE56521FC49C2B3" box="[865,1003,337,393]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="A0B4EABC21031F11FCE56521FC72C253" bold="true" box="[865,976,337,361]" pageId="3" pageNumber="4">Substrate</emphasis>
</th>
<th id="6F215D902103E0EDFB846521FBE4C2B3" box="[1024,1094,337,393]" gridcol="3" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="A0B4EABC21031F11FB846522FBE4C250" bold="true" box="[1024,1094,338,362]" pageId="3" pageNumber="4">Depth</emphasis>
</th>
<th id="6F215D902103E0EDFBD86521FB4BC2B3" box="[1116,1257,337,393]" gridcol="4" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="A0B4EABC21031F11FBD86522FB30C250" bold="true" box="[1116,1170,338,362]" pageId="3" pageNumber="4">Date</emphasis>
</th>
<th id="6F215D902103E0EDFB7A6521FAC9C2B3" box="[1278,1387,337,393]" gridcol="5" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="A0B4EABC21031F11FB7A6521FACAC2B3" bold="true" box="[1278,1387,337,393]" pageId="3" pageNumber="4">Collected specimen</emphasis>
</th>
</tr>
<tr id="2CF034EC2103E0EDFF4C65E9FAC9C2EB" box="[200,1387,409,465]" gridrow="1" pageId="3" pageNumber="4">
<th id="6F215D902103E0EDFF4C65E9FE0FC2EB" box="[200,429,409,465]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">
<taxonomicName id="55C04D2D21031F11FF4C65EAFEDAC28B" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[200,376,410,433]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="reptans" status="sp. nov.">
<emphasis id="A0B4EABC21031F11FF4C65EAFEDAC28B" box="[200,376,410,433]" italics="true" pageId="3" pageNumber="4">Hyssura reptans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721031F11FEFB65EAFF57C2EB" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="6F215D902103E0EDFE4D65E9FCE4C2EB" box="[457,838,409,465]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">Bangka Island, off Areng Kambing, 1°46′27.53″N, 125°11′19.53”E</td>
<td id="6F215D902103E0EDFCE565E9FC49C2EB" box="[865,1003,409,465]" gridcol="2" gridrow="1" pageId="3" pageNumber="4">Coral rubble</td>
<td id="6F215D902103E0EDFB8465E9FBE4C2EB" box="[1024,1094,409,465]" gridcol="3" gridrow="1" pageId="3" pageNumber="4">5 m</td>
<td id="6F215D902103E0EDFBD865E9FB4BC2EB" box="[1116,1257,409,465]" gridcol="4" gridrow="1" pageId="3" pageNumber="4">24 Sept. 2019</td>
<td id="6F215D902103E0EDFB7A65E9FAC9C2EB" box="[1278,1387,409,465]" gridcol="5" gridrow="1" pageId="3" pageNumber="4">1</td>
</tr>
<tr id="2CF034EC2103E0EDFF4C6598FAC9C17A" box="[200,1387,488,576]" gridrow="2" pageId="3" pageNumber="4">
<th id="6F215D902103E0EDFF4C6598FE0FC17A" box="[200,429,488,576]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">
<taxonomicName id="55C04D2D21031F11FF4C6598FE9FC11A" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[200,336,488,544]" class="Malacostraca" family="Hyssuridae" genus="Kupellonura" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="indonesica" status="sp. nov.">
<emphasis id="A0B4EABC21031F11FF4C6598FE9FC11A" box="[200,336,488,544]" italics="true" pageId="3" pageNumber="4">Kupellonura indonesica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721031F11FEC76679FE34C11A" box="[323,406,521,544]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="6F215D902103E0EDFE4D6598FCE4C17A" box="[457,838,488,576]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">Bangka Island, off jetty of Coral Eye resort, 1°44′22.04″ N, 125°8′41.19″ E</td>
<td id="6F215D902103E0EDFCE56598FC49C17A" box="[865,1003,488,576]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">Coral rubble</td>
<td id="6F215D902103E0EDFB846598FBE4C17A" box="[1024,1094,488,576]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">2 m</td>
<td id="6F215D902103E0EDFBD86598FB4BC17A" box="[1116,1257,488,576]" gridcol="4" gridrow="2" pageId="3" pageNumber="4">17 Sept. 2019</td>
<td id="6F215D902103E0EDFB7A6598FAC9C17A" box="[1278,1387,488,576]" gridcol="5" gridrow="2" pageId="3" pageNumber="4">8</td>
</tr>
<tr id="2CF034EC2103E0EDFF4C6622FAC9C193" box="[200,1387,594,681]" gridrow="3" pageId="3" pageNumber="4">
<th id="6F215D902103E0EDFF4C6622FE0FC193" box="[200,429,594,681]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">
<taxonomicName id="55C04D2D21031F11FF4C6622FE9BC1B3" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[200,335,594,649]" class="Malacostraca" family="Hyssuridae" genus="Kupellonura" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="maccaroni" status="sp. nov.">
<emphasis id="A0B4EABC21031F11FF4C6622FE9BC1B3" box="[200,335,594,649]" italics="true" pageId="3" pageNumber="4">Kupellonura maccaroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721031F11FEBB6602FE30C1B3" box="[319,402,626,649]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="6F215D902103E0EDFE4D6622FCE4C193" box="[457,838,594,681]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">Bangka Island, off jetty of Coral Eye resort, 1°44′22.04″ N, 125°8′41.19″ E</td>
<td id="6F215D902103E0EDFCE56622FC49C193" box="[865,1003,594,681]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">Coral rubble</td>
<td id="6F215D902103E0EDFB846622FBE4C193" box="[1024,1094,594,681]" gridcol="3" gridrow="3" pageId="3" pageNumber="4">2 m</td>
<td id="6F215D902103E0EDFBD86622FB4BC193" box="[1116,1257,594,681]" gridcol="4" gridrow="3" pageId="3" pageNumber="4">17 Sept. 2019</td>
<td id="6F215D902103E0EDFB7A6622FAC9C193" box="[1278,1387,594,681]" gridcol="5" gridrow="3" pageId="3" pageNumber="4">2</td>
</tr>
<tr id="2CF034EC2103E0EDFF4C66CEFAC9C02C" box="[200,1387,702,790]" gridrow="4" pageId="3" pageNumber="4">
<th id="6F215D902103E0EDFF4C66CEFE0FC02C" box="[200,429,702,790]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">
<taxonomicName id="55C04D2D21031F11FF4C66CEFEEFC1CC" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[200,340,702,758]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bangkaensis" status="sp. nov.">
<emphasis id="A0B4EABC21031F11FF4C66CEFEEFC1CC" box="[200,340,702,758]" italics="true" pageId="3" pageNumber="4">Pendanthura bangkaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721031F11FED066AFFE04C1CC" box="[340,422,735,758]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="6F215D902103E0EDFE4D66CEFCE4C02C" box="[457,838,702,790]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">Bangka Island, off Coral Eye Resort, 1°44′22.04″ N, 125°8′41.19″ E</td>
<td id="6F215D902103E0EDFCE566CEFC49C02C" box="[865,1003,702,790]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">Coral rubble</td>
<td id="6F215D902103E0EDFB8466CEFBE4C02C" box="[1024,1094,702,790]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">1–2 m</td>
<td id="6F215D902103E0EDFBD866CEFB4BC02C" box="[1116,1257,702,790]" gridcol="4" gridrow="4" pageId="3" pageNumber="4">21 Sept. 2019</td>
<td id="6F215D902103E0EDFB7A66CEFAC9C02C" box="[1278,1387,702,790]" gridcol="5" gridrow="4" pageId="3" pageNumber="4">3</td>
</tr>
<tr id="2CF034EC2103E0EDFF4C6756FAC9C044" box="[200,1387,806,894]" gridrow="5" pageId="3" pageNumber="4">
<th id="6F215D902103E0EDFF4C6756FE0FC044" box="[200,429,806,894]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">
E 
<emphasis id="A0B4EABC21031F11FF5C6756FEF6C004" box="[216,340,806,830]" italics="true" pageId="3" pageNumber="4">xpanathura</emphasis>
<taxonomicName id="55C04D2D21031F11FEDF6756FED0C064" authority="(Kensley, 1979)" baseAuthorityName="Kensley" baseAuthorityYear="1979" class="Malacostraca" family="Expanathuridae" genus="Expanathura" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="collaris">
<emphasis id="A0B4EABC21031F11FEDF6756FE0FC004" box="[347,429,806,830]" italics="true" pageId="3" pageNumber="4">collaris</emphasis>
(
<bibRefCitation id="F6514B5F21031F11FF4B6736FEC9C064" author="Kensley B." box="[207,363,838,862]" pageId="3" pageNumber="4" pagination="814 - 836" refId="ref13186" refString="Kensley B. 1979. New species of anthurideans from the Cook and Fiji Islands (Crustacea: Isopoda: Anthuridea). Proceedings of the Biological Society of Washington 92 (4): 814 - 836." type="journal article" year="1979">Kensley, 1979</bibRefCitation>
)
</taxonomicName>
</th>
<td id="6F215D902103E0EDFE4D6756FCE4C044" box="[457,838,806,894]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">Bangka Island, in front of Coral Eye Resort near jetti, 1°44′20.22″ N, 125°8′46.07″ E</td>
<td id="6F215D902103E0EDFCE56756FC49C044" box="[865,1003,806,894]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">Coral rubble</td>
<td id="6F215D902103E0EDFB846756FBE4C044" box="[1024,1094,806,894]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">1–2 m</td>
<td id="6F215D902103E0EDFBD86756FB4BC044" box="[1116,1257,806,894]" gridcol="4" gridrow="5" pageId="3" pageNumber="4">26 Sept. 2019</td>
<td id="6F215D902103E0EDFB7A6756FAC9C044" box="[1278,1387,806,894]" gridcol="5" gridrow="5" pageId="3" pageNumber="4">5</td>
</tr>
<tr id="2CF034EC2103E0EDFF4C67E0FAC9C0D2" box="[200,1387,912,1000]" gridrow="6" pageId="3" pageNumber="4">
<th id="6F215D902103E0EDFF4C67E0FE0FC0D2" box="[200,429,912,1000]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">
<taxonomicName id="55C04D2D21031F11FF4C67E0FE04C092" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[200,422,912,936]" class="Malacostraca" family="Expanathuridae" genus="Expanathura" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="marcoi" status="sp. nov.">
<emphasis id="A0B4EABC21031F11FF4C67E0FE04C092" box="[200,422,912,936]" italics="true" pageId="3" pageNumber="4">Expanathura marcoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721031F11FF4C67C1FEB9C0F2" box="[200,283,945,968]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="6F215D902103E0EDFE4D67E0FCE4C0D2" box="[457,838,912,1000]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">Bangka Island, in front of Coral Eye Resort, 1°44′20.22″ N, 125°8′46.07″ E</td>
<td id="6F215D902103E0EDFCE567E0FC49C0D2" box="[865,1003,912,1000]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">Coral rubble</td>
<td id="6F215D902103E0EDFB8467E0FBE4C0D2" box="[1024,1094,912,1000]" gridcol="3" gridrow="6" pageId="3" pageNumber="4">1–2 m</td>
<td id="6F215D902103E0EDFBD867E0FB4BC0D2" box="[1116,1257,912,1000]" gridcol="4" gridrow="6" pageId="3" pageNumber="4">26 Sept. 2019</td>
<td id="6F215D902103E0EDFB7A67E0FAC9C0D2" box="[1278,1387,912,1000]" gridcol="5" gridrow="6" pageId="3" pageNumber="4">1</td>
</tr>
<tr id="2CF034EC2103E0EDFF4C678AFAC9C768" box="[200,1387,1018,1106]" gridrow="7" pageId="3" pageNumber="4">
<th id="6F215D902103E0EDFF4C678AFE0FC768" box="[200,429,1018,1106]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">
<taxonomicName id="55C04D2D21031F11FF4C678AFE3DC728" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[200,415,1018,1042]" class="Malacostraca" family="Paranthuridae" genus="Colanthura" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="gerungi" status="sp. nov.">
<emphasis id="A0B4EABC21031F11FF4C678AFE3DC728" box="[200,415,1018,1042]" italics="true" pageId="3" pageNumber="4">Colanthura gerungi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721031F11FF4C606BFEB9C708" box="[200,283,1051,1074]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="6F215D902103E0EDFE4D678AFCE4C768" box="[457,838,1018,1106]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">Bangka Island, west of Coral Eye Resort, 1°45′20.77″ N, 125°7′57.28″ E</td>
<td id="6F215D902103E0EDFCE5678AFC49C768" box="[865,1003,1018,1106]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">Seagrass bed</td>
<td id="6F215D902103E0EDFB84678AFBE4C768" box="[1024,1094,1018,1106]" gridcol="3" gridrow="7" pageId="3" pageNumber="4">1–2 m</td>
<td id="6F215D902103E0EDFBD8678AFB4BC768" box="[1116,1257,1018,1106]" gridcol="4" gridrow="7" pageId="3" pageNumber="4">18 Sept. 2019</td>
<td id="6F215D902103E0EDFB7A678AFAC9C768" box="[1278,1387,1018,1106]" gridcol="5" gridrow="7" pageId="3" pageNumber="4">14</td>
</tr>
<tr id="2CF034EC2103E0EDFF4C6012FAC9C780" box="[200,1387,1122,1210]" gridrow="8" pageId="3" pageNumber="4" rowspan-0="1">
<td id="6F215D902103E0EDFE4D6012FCE4C780" box="[457,838,1122,1210]" gridcol="1" gridrow="8" pageId="3" pageNumber="4">Bangka Island, west of Coral Eye Resort, 1°45′19.92″ N, 125°7′58.96″E</td>
<td id="6F215D902103E0EDFCE56012FC49C780" box="[865,1003,1122,1210]" gridcol="2" gridrow="8" pageId="3" pageNumber="4">M a n g r o v e roots</td>
<td id="6F215D902103E0EDFB846012FBE4C780" box="[1024,1094,1122,1210]" gridcol="3" gridrow="8" pageId="3" pageNumber="4">1–2 m</td>
<td id="6F215D902103E0EDFBD86012FB4BC780" box="[1116,1257,1122,1210]" gridcol="4" gridrow="8" pageId="3" pageNumber="4">21.Sept. 2019</td>
<td id="6F215D902103E0EDFB7A6012FAC9C780" box="[1278,1387,1122,1210]" gridcol="5" gridrow="8" pageId="3" pageNumber="4">2</td>
</tr>
<tr id="2CF034EC2103E0EDFF4C60BAFAC9C618" box="[200,1387,1226,1314]" gridrow="9" pageId="3" pageNumber="4" rowspan-0="1">
<td id="6F215D902103E0EDFE4D60BAFCE4C618" box="[457,838,1226,1314]" gridcol="1" gridrow="9" pageId="3" pageNumber="4">Bangka Island, off Coral Eye Resort, 1°44′22.04″ N, 125°8′41.19″ E</td>
<td id="6F215D902103E0EDFCE560BAFC49C618" box="[865,1003,1226,1314]" gridcol="2" gridrow="9" pageId="3" pageNumber="4">Coral rubble</td>
<td id="6F215D902103E0EDFB8460BAFBE4C618" box="[1024,1094,1226,1314]" gridcol="3" gridrow="9" pageId="3" pageNumber="4">1–2m</td>
<td id="6F215D902103E0EDFBD860BAFB4BC618" box="[1116,1257,1226,1314]" gridcol="4" gridrow="9" pageId="3" pageNumber="4">26 Sept. 2019</td>
<td id="6F215D902103E0EDFB7A60BAFAC9C618" box="[1278,1387,1226,1314]" gridcol="5" gridrow="9" pageId="3" pageNumber="4">1</td>
</tr>
</table>
</paragraph>
<paragraph id="927F36AE21031F11FF396248FDBDC477" blockId="3.[189,1399,1592,1869]" pageId="3" pageNumber="4">ANTENNAE. Antenna 1 peduncle of three articles, first 1.5 times as long as wide, articles 2 and 3 progressively shorter; flagellum with typical male modifications: nearly twice as long as peduncle and composed of 9 articles; distal articles narrower, last article tiny; all flagellum articles except the first with long and thin aesthetascs, arranged in a single transverse row per article, number of aesthetascs distally decreasing, last article with only one aesthetasc. Antenna 2 peduncle with 5 articles, second article longest and widest; flagellum of 8 articles, not as long as peduncle articles 4 + 5, with several simple setae, longest on distal article.</paragraph>
<paragraph id="927F36AE21031F11FF39630BFC3DC4D2" blockId="3.[189,1399,1915,2025]" pageId="3" pageNumber="4">MOUTHPARTS. Were lost during dissection; however, in situ a maxilliped palp with five articles was visible, not covering mouthparts but erected nearly perpendicularly, and a mandible with an atrophied endite. Presumably the mouthparts of males are not functional.</paragraph>
<paragraph id="927F36AE21041F16FF396579FB25C120" blockId="4.[188,1400,265,538]" pageId="4" pageNumber="5">PEREOPODS. Pereopod 1 weakly subchelate, propodus palm concave, a comb of 6 setae on mesial surface in addition to two stronger medial setae and distally 4 setae; carpus triangular with short, not prominent ventrodistal projection bearing a simple seta; more proximally another seta; merus dorsal portion nearly triangular. Pereopod 2 propodus less curved than in P1, elongated, about three times as long as wide, palm distally with 3 sensory spines and basally slightly concave; carpus ventrodistally only little projecting, as in pereopod 1. Pereopod 3 similar to pereopod 2. Pereopods 4–7 similar. Propodi distally with two short sensory spines on distoventral margin, with a tooth-like projection in between, carpus trapezoidal, with shorter dorsal margin, merus subtriangular and as long as carpus.</paragraph>
<paragraph id="927F36AE21041F16FF396631FC70C1FE" blockId="4.[188,1399,577,709]" pageId="4" pageNumber="5">
PLEOPODS. All of similar shape, rami nearly rectangular. Pleopod 1 exopod not larger than endopod; exopods of pleopods 1 and 2 distally with 10 resp. 9 swimming setae, endopods with 5. Exopod of Plp 2 composed of two articles. Appendix masculina on Plp 
<quantity id="55389B4B21041F16FCBE66F7FCC5C19B" box="[826,871,647,673]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="5" unit="in" value="2.0">2 in</quantity>
form of a cylindrical rod about 30% as long as endopod, distally rounded, without further conspicuous features.
</paragraph>
<caption id="C6BF662621041F16FF3963BEFBA8C4D2" ID-DOI="http://doi.org/10.5281/zenodo.5535913" ID-Zenodo-Dep="5535913" box="[189,1034,1998,2024]" httpUri="https://zenodo.org/record/5535913/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1998,2024]" targetBox="[291,1333,784,1914]" targetPageId="4">
<paragraph id="927F36AE21041F16FF3963BEFBA8C4D2" blockId="4.[189,1034,1998,2024]" box="[189,1034,1998,2024]" pageId="4" pageNumber="5">
<emphasis id="A0B4EABC21041F16FF3963BEFEA9C4D2" bold="true" box="[189,267,1998,2024]" pageId="4" pageNumber="5">Fig. 1.</emphasis>
Map of Northern Sulawesi with sampling sites on Bangka Island
</paragraph>
</caption>
<caption id="C6BF662621051F17FF3963BCFA94C4DC" ID-DOI="http://doi.org/10.5281/zenodo.5535915" ID-Zenodo-Dep="5535915" box="[189,1334,1996,2022]" httpUri="https://zenodo.org/record/5535915/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1996,2022]" targetBox="[370,1387,279,1889]" targetPageId="5">
<paragraph id="927F36AE21051F17FF3963BCFA94C4DC" blockId="5.[189,1334,1996,2022]" box="[189,1334,1996,2022]" pageId="5" pageNumber="6">
<emphasis id="A0B4EABC21051F17FF3963BCFEA9C4DC" bold="true" box="[189,267,1996,2022]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
<taxonomicName id="55C04D2D21051F17FE9763BDFE77C4DC" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[275,469,1997,2022]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="reptans" status="sp. nov.">
<emphasis id="A0B4EABC21051F17FE9763BDFE77C4DC" box="[275,469,1997,2022]" italics="true" pageId="5" pageNumber="6">Hyssura reptans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721051F17FE5963BCFD98C4DC" box="[477,570,1996,2022]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype in dorsal and lateral view (2.1 mm) (MZB Iso 099).
</paragraph>
</caption>
<caption id="C6BF662621061F14FF3963DBFDFEC4D2" ID-DOI="http://doi.org/10.5281/zenodo.5535917" ID-Zenodo-Dep="5535917" httpUri="https://zenodo.org/record/5535917/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1963,1989]" targetBox="[215,1365,276,1883]" targetPageId="6">
<paragraph id="927F36AE21061F14FF3963DBFDFEC4D2" blockId="6.[189,1399,1963,2025]" pageId="6" pageNumber="7">
<emphasis id="A0B4EABC21061F14FF3963DBFEADC4FF" bold="true" box="[189,271,1963,1989]" pageId="6" pageNumber="7">Fig. 3.</emphasis>
<taxonomicName id="55C04D2D21061F14FE9D63DCFE71C4FF" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[281,467,1964,1989]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="reptan" status="sp. nov.">
<emphasis id="A0B4EABC21061F14FE9D63DCFE71C4FF" box="[281,467,1964,1989]" italics="true" pageId="6" pageNumber="7">Hyssura reptan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721061F14FE5A63DCFD9CC4FC" box="[478,574,1964,1990]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (MZB Iso 099). Abbreviations: A1 = antenna 1; A2 = antenna 2; P1–P3 = pereopods 1–3.
</paragraph>
</caption>
<caption id="C6BF662621071F15FF3963D6FE76C4DE" ID-DOI="http://doi.org/10.5281/zenodo.5535921" ID-Zenodo-Dep="5535921" httpUri="https://zenodo.org/record/5535921/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1958,1984]" targetBox="[198,1392,275,1833]" targetPageId="7">
<paragraph id="927F36AE21071F15FF3963D6FE76C4DE" blockId="7.[189,1399,1958,2020]" pageId="7" pageNumber="8">
<emphasis id="A0B4EABC21071F15FF3963D6FEAEC4FA" bold="true" box="[189,268,1958,1984]" pageId="7" pageNumber="8">Fig. 4.</emphasis>
<taxonomicName id="55C04D2D21071F15FE9063D7FE75C4FA" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[276,471,1959,1984]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="reptans" status="sp. nov.">
<emphasis id="A0B4EABC21071F15FE9063D7FE75C4FA" box="[276,471,1959,1984]" italics="true" pageId="7" pageNumber="8">Hyssura reptans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721071F15FE5B63D7FD9FC4FB" box="[479,573,1959,1985]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (MZB Iso 099). Abbreviations: P4–P7 = pereopods 4–7; Plp1–2 = pleopods 1–2.
</paragraph>
</caption>
<paragraph id="927F36AE21081F1AFF396579FBE1C28A" blockId="8.[188,1399,265,433]" pageId="8" pageNumber="9">UROPOD. Exopod narrow lanceolate, length about 7.6 times greatest width, outer lateral margin smooth, medial margin slightly serrated with some single setae in notches; apex with several long simple setae. Endopod as wide as sympod but about 2.5 times longer, a bit shorter than exopod, proximal two thirds of margins nearly parallel, distal margins with simple setae of varying length. Telson tongue-like, lateral margins serrated, not parallel; apex rounded and with 5 pairs of simple seate.</paragraph>
</subSubSection>
<subSubSection id="DADA652521081F1BFF3965A7FCB5C27D" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
<paragraph id="927F36AE21081F1AFF3965A7FE8DC2CB" blockId="8.[189,303,471,497]" box="[189,303,471,497]" pageId="8" pageNumber="9">
<heading id="C93781C221081F1AFF3965A7FE8DC2CB" bold="true" box="[189,303,471,497]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="A0B4EABC21081F1AFF3965A7FE8DC2CB" bold="true" box="[189,303,471,497]" pageId="8" pageNumber="9">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="927F36AE21081F1AFF396676FB7BC02C" blockId="8.[188,1400,518,791]" pageId="8" pageNumber="9">
We place this species in the genus 
<taxonomicName id="55C04D2D21081F1AFDCB6677FC4DC11A" authority="Norman &amp; Stebbing, 1884" authorityName="Norman &amp; Stebbing" authorityYear="1884" box="[591,1007,518,544]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21081F1AFDCB6677FD10C11A" box="[591,690,519,544]" italics="true" pageId="8" pageNumber="9">Hyssura</emphasis>
Norman &amp; Stebbing, 1884
</taxonomicName>
because autapomorphies of other genera are missing. Species of 
<taxonomicName id="55C04D2D21081F1AFDB0665AFD35C179" authorityName="Norman &amp; Stebbing" authorityYear="1884" box="[564,663,554,579]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21081F1AFDB0665AFD35C179" box="[564,663,554,579]" italics="true" pageId="8" pageNumber="9">Hyssura</emphasis>
</taxonomicName>
are blind, the P1–3 carpus is barely projecting ventrodistally. A reduction of male mouthparts has been reported for 
<taxonomicName id="55C04D2D21081F1AFCBD663DFBDDC15C" authority="(Kensley, 1978 a)" baseAuthorityName="Kensley" baseAuthorityYear="1978" box="[825,1151,588,614]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vimsae">
<emphasis id="A0B4EABC21081F1AFCBD663DFC0DC15C" box="[825,943,589,614]" italics="true" pageId="8" pageNumber="9">H. vimsae</emphasis>
(
<bibRefCitation id="F6514B5F21081F1AFC3B663CFBC8C15C" author="Kensley B." box="[959,1130,588,614]" pageId="8" pageNumber="9" pagination="1 - 25" refId="ref13152" refString="Kensley B. 1978. The South African Museum's Meiring Naude Cruises. Part 8. Isopoda Anthuridea. Annals of the South African Museum 77 (1): 1 - 25." type="journal article" year="1978">Kensley, 1978</bibRefCitation>
a)
</taxonomicName>
(see 
<bibRefCitation id="F6514B5F21081F1AFB39663CFAC7C15C" author="Wagele J. W." box="[1213,1381,588,614]" pageId="8" pageNumber="9" pagination="1 - 2" refId="ref13936" refString="Wagele J. W. 1981 a. Study of the Hyssuridae (Crustacea: Isopoda: Anthuridea) from the Mediterranean and the Red Sea. Israel Journal of Zoology 30: 1 - 2." type="journal article" year="1981">Wägele 1981a</bibRefCitation>
), other males have not been described. The male of 
<taxonomicName id="55C04D2D21081F1AFC946600FC24C1B3" baseAuthorityName="Kensley" baseAuthorityYear="1978" box="[784,902,624,649]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vimsae">
<emphasis id="A0B4EABC21081F1AFC946600FC24C1B3" box="[784,902,624,649]" italics="true" pageId="8" pageNumber="9">H. vimsae</emphasis>
</taxonomicName>
also has a comb of setae on P1 propodus; similar combs have also been described for some species of 
<taxonomicName id="55C04D2D21081F1AFC1A66E2FB96C196" authorityName="Barnard" authorityYear="1925" box="[926,1076,658,684]" class="Malacostraca" family="Hyssuridae" genus="Kupellonura" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21081F1AFC1A66E2FB96C196" box="[926,1076,658,684]" italics="true" pageId="8" pageNumber="9">Kupellonura</emphasis>
</taxonomicName>
(
<emphasis id="A0B4EABC21081F1AFBCC66E3FAD4C196" box="[1096,1398,658,684]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="55C04D2D21081F1AFBCC66E3FB1CC196" authorityName="Poore &amp; Lew Ton" authorityYear="1988" box="[1096,1214,658,684]" class="Malacostraca" family="Hyssuridae" genus="Kupellonura" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="biriwa">K. biriwa</taxonomicName>
, 
<taxonomicName id="55C04D2D21081F1AFB4966E3FAD0C196" authorityName="Poore &amp; Lew Ton" authorityYear="1988" box="[1229,1394,659,684]" class="Malacostraca" family="Hyssuridae" genus="Kupellonura" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="marrongie">K. marrongie</taxonomicName>
,
</emphasis>
see 
<bibRefCitation id="F6514B5F21081F1AFF6F66C6FDABC1EA" author="Poore G. C. B. &amp; Lew Ton H. M." box="[235,521,694,720]" pageId="8" pageNumber="9" pagination="169 - 193" refId="ref13777" refString="Poore G. C. B. &amp; Lew Ton H. M. 1988. A generic review of the Hyssuridae (Crustacea: Isopoda) with a new genus and new species from Australia. Memoirs of the Museum of Victoria 49 (1): 169 - 193." type="journal article" year="1988">Poore &amp; Lew Ton 1988</bibRefCitation>
). It is remarkable that also the male is blind, while in other 
<taxonomicName id="55C04D2D21081F1AFB5166C6FAFBC1EA" authorityName="Wagele" authorityYear="1981" box="[1237,1369,694,720]" class="Malacostraca" family="Hyssuridae" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">Hyssuridae</taxonomicName>
at least males have prominent eyes. In the present species the carpus of P4–7 is shorter than in previously described species. Other characters are plesiomorphic and occur also in other 
<taxonomicName id="55C04D2D21081F1AFBC9668CFB77C02C" authorityName="Wagele" authorityYear="1981" box="[1101,1237,764,790]" class="Malacostraca" family="Hyssuridae" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">Hyssuridae</taxonomicName>
.
</paragraph>
<caption id="C6BF662621081F1AFF3963BCFB74C4DC" ID-DOI="http://doi.org/10.5281/zenodo.5535923" ID-Zenodo-Dep="5535923" box="[189,1238,1996,2022]" httpUri="https://zenodo.org/record/5535923/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1996,2022]" targetBox="[201,1361,896,1922]" targetPageId="8">
<paragraph id="927F36AE21081F1AFF3963BCFB74C4DC" blockId="8.[189,1238,1996,2022]" box="[189,1238,1996,2022]" pageId="8" pageNumber="9">
<emphasis id="A0B4EABC21081F1AFF3963BCFEA9C4DC" bold="true" box="[189,267,1996,2022]" pageId="8" pageNumber="9">Fig. 5.</emphasis>
<taxonomicName id="55C04D2D21081F1AFE9763BDFE77C4DC" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[275,469,1997,2022]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="reptans" status="sp. nov.">
<emphasis id="A0B4EABC21081F1AFE9763BDFE77C4DC" box="[275,469,1997,2022]" italics="true" pageId="8" pageNumber="9">Hyssura reptans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721081F1AFE5963BCFD98C4DC" box="[477,570,1996,2022]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype. Abbreviations: Urp = uropod; Tel = telson.
</paragraph>
</caption>
<paragraph id="927F36AE21091F1BFF396579FCB5C27D" blockId="9.[189,1399,265,327]" pageId="9" pageNumber="10">
This is the first 
<taxonomicName id="55C04D2D21091F1BFEF0657AFE75C219" authorityName="Norman &amp; Stebbing" authorityYear="1884" box="[372,471,266,291]" class="Malacostraca" family="Hyssuridae" genus="Hyssura" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21091F1BFEF0657AFE75C219" box="[372,471,266,291]" italics="true" pageId="9" pageNumber="10">Hyssura</emphasis>
</taxonomicName>
described from the Indo-Pacific. The other species occur in the Atlantic Ocean or in the Mediterranean and often in bathyal waters
</paragraph>
</subSubSection>
</treatment>
</document>