354 lines
48 KiB
XML
354 lines
48 KiB
XML
<document id="8957E4F5E3FA138E6A136F1CD313CC78" ID="10.11646/zootaxa.4161.2.9" ID-DOI="10.11646/zootaxa.4161.2.9" ID-GBIF-Dataset="b529abbf-bbda-4810-b959-3f537bffd427" ID-ISSN="1175-5326" ID-Zenodo-Dep="254867" ID-ZooBank="09043B98-9518-48FF-9EFC-EAE7C9690E03" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1473058318798" checkinUser="plazi" docAuthor="Zhou, De-Yao & Li, Li-Zhen" docDate="2016" docId="FB0487902B253C3439F7EB87FBABFB38" docLanguage="en" docName="zootaxa.4161.2.9.pdf" docOrigin="Zootaxa 4161 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Horaeomorphus bicornis" docType="treatment" docVersion="9" lastPageNumber="276" masterDocId="073DFFE82B243C313960EE23FFBCFF98" masterDocTitle="New data on Horaeomorphus Schaufuss (Coleoptera: Staphylinidae: Scydmaeninae) from the Oriental region, with description of a bizarre new species from Yunnan, Southeast China" masterLastPageNumber="281" masterPageNumber="271" pageNumber="272" updateTime="1698384064788" updateUser="plazi">
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<mods:mods id="ACDD52B9E62EB08B7C77E51378F176BD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CDC0E172F16256703BC959A280F7FA81">
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<mods:title id="1C7B806BDE27F11A2CB8E89126586668">New data on Horaeomorphus Schaufuss (Coleoptera: Staphylinidae: Scydmaeninae) from the Oriental region, with description of a bizarre new species from Yunnan, Southeast China</mods:title>
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<mods:name id="E7BE36E7E5382EB9A4BE945B7DAEB2FF" type="personal">
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<mods:namePart id="9A0631A81B7EBC257D68F9935D0DDBBC">Zhou, De-Yao</mods:namePart>
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<mods:roleTerm id="5BA43DD583AA128B471600D03959EC2F">Author</mods:roleTerm>
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<mods:namePart id="09BEE33B0A13D476996BF132D8056328">Li, Li-Zhen</mods:namePart>
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<mods:typeOfResource id="E892C499F9CD6CFB654B990152D5B164">text</mods:typeOfResource>
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<mods:title id="2A510AD520A1CDF8ED2C1C7127A8ED6C">Zootaxa</mods:title>
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<mods:part id="EFB4BB51C15914FAE877321972136844">
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<mods:date id="BA72415C3C2397AC8B2CE336F3319CF3">2016</mods:date>
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<mods:number id="2CF91AA1EBF6403E20C4477DBD8F986F">4161</mods:number>
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<mods:identifier id="B39602D0D8DEE65AFDBE486670AC7314" type="DOI">10.11646/zootaxa.4161.2.9</mods:identifier>
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<mods:identifier id="F4A28B30AA4012A3D676915A901A61ED" type="GBIF-Dataset">b529abbf-bbda-4810-b959-3f537bffd427</mods:identifier>
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<treatment id="FB0487902B253C3439F7EB87FBABFB38" ID-DOI="http://doi.org/10.5281/zenodo.6091238" ID-GBIF-Taxon="124409936" ID-Zenodo-Dep="6091238" LSID="urn:lsid:plazi:treatment:FB0487902B253C3439F7EB87FBABFB38" httpUri="http://treatment.plazi.org/id/FB0487902B253C3439F7EB87FBABFB38" lastPageId="5" lastPageNumber="276" pageId="1" pageNumber="272">
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<paragraph id="731236862B253C3039F7EB87FC3CFA26" blockId="1.[151,896,1444,1503]" box="[151,896,1444,1470]" pageId="1" pageNumber="272">
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<heading id="285A81EA2B253C3039F7EB87FC3CFA26" bold="true" box="[151,896,1444,1470]" fontSize="11" level="1" pageId="1" pageNumber="272" reason="1">
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<emphasis id="41D9EA942B253C3039F7EB87FC3CFA26" bold="true" box="[151,896,1444,1470]" pageId="1" pageNumber="272">
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<taxonomicName id="B4AD4D052B253C3039F7EB87FE7BFA26" ID-CoL="3MNJJ" box="[151,455,1444,1470]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="bicornis">
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<emphasis id="41D9EA942B253C3039F7EB87FE7BFA26" bold="true" box="[151,455,1444,1470]" italics="true" pageId="1" pageNumber="272">Horaeomorphus bicornis</emphasis>
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</taxonomicName>
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D.-Y. Zhou & L.-Z. Li, new species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="731236862B253C3039F7EBE5FEF0FA47" blockId="1.[151,896,1444,1503]" box="[151,332,1478,1503]" pageId="1" pageNumber="272">
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<figureCitation id="EB962A032B253C3039F7EBE5FF60FA47" box="[151,220,1478,1503]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1882,1905]" captionTargetBox="[163,1425,526,1854]" captionTargetId="figure@2.[151,1435,517,1862]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Dorsal habitus of Horaeomorphus species. H. bicornis, holotype male (1); H. punctatissimus, male (2)." httpUri="https://zenodo.org/record/254868/files/figure.png" pageId="1" pageNumber="272">Figs 1</figureCitation>
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,
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<figureCitation id="EB962A032B253C30398AEBE4FE99FA47" box="[234,293,1479,1503]" captionStart-0="FIGURES 3 – 7" captionStart-1="FIGURES 8 – 12" captionStartId-0="3.[151,264,1973,1996]" captionStartId-1="4.[151,264,1894,1917]" captionTargetBox-0="[151,1430,661,1952]" captionTargetBox-1="[153,1436,211,1859]" captionTargetId-0="figure@3.[151,1435,661,1953]" captionTargetId-1="figure@4.[151,1436,193,1873]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 3 – 7. Diagnostic characters of Horaeomorphus bicornis. Head in dorsal view (3), in ventral view (4), and in lateral view (5); left metatibia in dorsal view (6), and in lateral view (7). Abbreviation: mn, mentum." captionText-1="FIGURES 8 – 12. Diagnostic characters of Horaeomorphus bicornis. Aedeagus in ventral view (8), in lateral view (9), and in dorsal view; endophallus, enlarged, in ventral view (11); and left antenna, in dorsal view (12); photographed (a) and schematic (b)." httpUri-0="https://zenodo.org/record/254869/files/figure.png" httpUri-1="https://zenodo.org/record/254870/files/figure.png" pageId="1" pageNumber="272">3–12</figureCitation>
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,
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<figureCitation id="EB962A032B253C303850EBE4FEF0FA47" box="[304,332,1479,1503]" captionStart="FIGURES 21 – 23" captionStartId="8.[151,264,1570,1593]" captionTargetBox="[151,1435,193,1548]" captionTargetId="figure@8.[151,1435,193,1549]" captionTargetPageId="8" captionText="FIGURES 21 – 23. Habitat of H. bicornis in Yinjiang Hsien (21). Living male adult of H. punctatissimus on a rotten log in Mt. T" httpUri="https://zenodo.org/record/254873/files/figure.png" pageId="1" pageNumber="272">21</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="3BB7650D2B253C3039F7E831FB0AF9EF" pageId="1" pageNumber="272" type="materials_examined">
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<paragraph id="731236862B253C3039F7E831FB0AF9EF" blockId="1.[151,1436,1553,2015]" pageId="1" pageNumber="272">
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<materialsCitation id="C3C53CDB2B253C3039F7E831FDDDF9EF" ID-GBIF-Occurrence="1303591652" collectingDate="2016-02-17" collectorName="Jian-Yue Qiu & Hao Xu" country="China" county="China" elevation="1000" latitude="24.62988" location="Tongbiguan Country" longLatPrecision="1" longitude="97.58305" municipality="Yinjiang Hsien" pageId="1" pageNumber="272" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
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<emphasis id="41D9EA942B253C3039F7E831FE82F9B3" bold="true" box="[151,318,1554,1579]" pageId="1" pageNumber="272">
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<typeStatus id="AC1688242B253C3039F7E831FF6EF9B3" box="[151,210,1554,1579]" pageId="1" pageNumber="272">Type</typeStatus>
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material
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</emphasis>
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(2 ƋƋ).
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<emphasis id="41D9EA942B253C3038C7E831FD37F9B3" bold="true" box="[423,651,1554,1579]" pageId="1" pageNumber="272">
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<typeStatus id="AC1688242B253C3038C7E831FDA4F9B3" box="[423,536,1554,1579]" pageId="1" pageNumber="272" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="0BBA76162B253C303B45E831FD3BF9B3" box="[549,647,1554,1579]" name="China" pageId="1" pageNumber="272">CHINA</collectingCountry>
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:
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</emphasis>
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Ƌ, labeled ‘
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<collectingCounty id="9A734E0A2B253C303A74E831FCDCF9B3" box="[788,864,1554,1579]" pageId="1" pageNumber="272">China</collectingCounty>
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:
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<collectingRegion id="B169F8642B253C303A0BE830FC78F9B2" box="[875,964,1555,1578]" country="China" name="Yunnan" pageId="1" pageNumber="272">Yunnan</collectingRegion>
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,
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<collectingMunicipality id="9376ACFC2B253C303AAFE831FBC5F9B3" box="[975,1145,1554,1579]" pageId="1" pageNumber="272">Yinjiang Hsien</collectingMunicipality>
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(盈江县),
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<location id="7672605D2B253C303C76E831FF4FF9CA" LSID="urn:lsid:plazi:treatment:FB0487902B253C3439F7EB87FBABFB38:7672605D2B253C303C76E831FF4FF9CA" country="China" county="China" latitude="24.62988" longLatPrecision="1" longitude="97.58305" municipality="Yinjiang Hsien" name="Tongbiguan Country" pageId="1" pageNumber="272" stateProvince="Yunnan">Tongbiguan Country</location>
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(铜壁关乡) to
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<location id="7672605D2B253C3038A4E819FDDEF9CA" LSID="urn:lsid:plazi:treatment:FB0487902B253C3439F7EB87FBABFB38:7672605D2B253C3038A4E819FDDEF9CA" box="[452,610,1594,1619]" country="China" county="China" latitude="24.62988" longLatPrecision="1" longitude="97.58305" municipality="Yinjiang Hsien" name="Nabang Town" pageId="1" pageNumber="272" stateProvince="Yunnan">Nabang Town</location>
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(那邦镇),
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<geoCoordinate id="169950412B253C303B9FE819FC14F9CB" box="[767,936,1594,1619]" direction="north" orientation="latitude" pageId="1" pageNumber="272" precision="1" value="24.62988">24°37'47.57''N</geoCoordinate>
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,
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<geoCoordinate id="169950412B253C303AD3E819FBE5F9CB" box="[947,1113,1594,1619]" direction="east" orientation="longitude" pageId="1" pageNumber="272" precision="1" value="97.58305">97°34'59.01''E</geoCoordinate>
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, rotten wood, sifted,
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<quantity id="B4559B632B253C303C27E819FA2BF9CA" box="[1351,1431,1594,1618]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="272" unit="m" value="1000.0">
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<elevation id="F880D1B52B253C303C27E819FA2BF9CA" box="[1351,1431,1594,1618]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="272" unit="m" value="1000.0">1000m</elevation>
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</quantity>
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,
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<date id="071310462B253C3039F7E87DFEB7F9EF" box="[151,267,1630,1655]" pageId="1" pageNumber="272" value="2016-02-17">
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<collectingDate id="1757E9AE2B253C3039F7E87DFEB7F9EF" box="[151,267,1630,1655]" pageId="1" pageNumber="272" value="2016-02-17">17.ii.2016</collectingDate>
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</date>
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,
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<collectorName id="DE5853502B253C303876E87DFE15F9EF" box="[278,425,1630,1655]" pageId="1" pageNumber="272">Jian-Yue Qiu</collectorName>
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&
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<collectorName id="DE5853502B253C3038ADE87CFD99F9EE" box="[461,549,1631,1654]" pageId="1" pageNumber="272">Hao Xu</collectorName>
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leg.’
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</materialsCitation>
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.
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<materialsCitation id="C3C53CDB2B253C303B0CE87DFB0EF9EF" ID-GBIF-Occurrence="1303591653" box="[620,1202,1630,1655]" country="China" location="CHINA" pageId="1" pageNumber="272" specimenCount="2" typeStatus="paratype">
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<emphasis id="41D9EA942B253C303B0CE87DFCECF9EF" bold="true" box="[620,848,1630,1655]" pageId="1" pageNumber="272">
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<typeStatus id="AC1688242B253C303B0CE87DFD61F9EF" box="[620,733,1630,1655]" pageId="1" pageNumber="272" type="paratype">Paratype</typeStatus>
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:
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<collectingCountry id="0BBA76162B253C303B8AE87DFCF0F9EF" box="[746,844,1630,1655]" name="China" pageId="1" pageNumber="272">CHINA</collectingCountry>
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:
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</emphasis>
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Ƌ, same label data as
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||
<typeStatus id="AC1688242B253C303D2CE87DFB0EF9EF" box="[1100,1202,1630,1655]" pageId="1" pageNumber="272" type="holotype">holotype</typeStatus>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="3BB7650D2B253C3039A7E8A1FB7DF8B3" pageId="1" pageNumber="272" type="diagnosis">
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<paragraph id="731236862B253C3039A7E8A1FB7DF8B3" blockId="1.[151,1436,1553,2015]" pageId="1" pageNumber="272">
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<emphasis id="41D9EA942B253C3039A7E8A1FEFDF903" bold="true" box="[199,321,1666,1691]" pageId="1" pageNumber="272">Diagnosis.</emphasis>
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The labral projection gradually broadening anteriorly together with the projection on the clypeus allow for an unambiguous identification of the males of
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<taxonomicName id="B4AD4D052B253C303A6EE884FC30F926" box="[782,908,1702,1726]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="41D9EA942B253C303A6EE884FC30F926" box="[782,908,1702,1726]" italics="true" pageId="1" pageNumber="272">H. bicornis</emphasis>
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||
</taxonomicName>
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||
. The mentum with a pair of large lateral lobes; each metatibiae strongly bent at anterior third with the posterior 2/3 broadened and flattened; the coarsely punctate elytra each with a longitudinal, glossy, sparsely and finely punctured area located along the elytral suture; and the aedeagus with setiferous, emarginate apex of ventral wall are also unique among all congeners.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="3BB7650D2B253C3439A7E915FE22FEFD" lastPageId="5" lastPageNumber="276" pageId="1" pageNumber="272" type="description">
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<paragraph id="731236862B253C3039A7E915FE6CF8EB" blockId="1.[151,1436,1553,2015]" pageId="1" pageNumber="272">
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<emphasis id="41D9EA942B253C3039A7E915FEE5F8D7" bold="true" box="[199,345,1846,1871]" pageId="1" pageNumber="272">Description.</emphasis>
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Male (
|
||
<figureCitation id="EB962A032B253C3038D4E915FE47F8D7" box="[436,507,1846,1871]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1882,1905]" captionTargetBox="[163,1425,526,1854]" captionTargetId="figure@2.[151,1435,517,1862]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Dorsal habitus of Horaeomorphus species. H. bicornis, holotype male (1); H. punctatissimus, male (2)." httpUri="https://zenodo.org/record/254868/files/figure.png" pageId="1" pageNumber="272">Fig. 1</figureCitation>
|
||
). BL
|
||
<quantity id="B4559B632B253C303B2AE915FD4AF8D6" box="[586,758,1846,1871]" metricMagnitude="-3" metricUnit="m" metricValue="1.815" metricValueMax="1.87" metricValueMin="1.76" pageId="1" pageNumber="272" unit="mm" value="1.815" valueMax="1.87" valueMin="1.76">1.76–1.87 mm</quantity>
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; flattened, reddish brown, legs and palpi slightly lighter, vestiture of setae yellowish.
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||
</paragraph>
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||
<paragraph id="731236862B253C3339A7E95DFB5CFE84" blockId="1.[151,1436,1553,2015]" lastBlockId="2.[151,1436,151,464]" lastPageId="2" lastPageNumber="273" pageId="1" pageNumber="272">
|
||
Head (
|
||
<figureCitation id="EB962A032B253C303872E95DFECBF80F" box="[274,375,1918,1943]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1973,1996]" captionTargetBox="[151,1430,661,1952]" captionTargetId="figure@3.[151,1435,661,1953]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Diagnostic characters of Horaeomorphus bicornis. Head in dorsal view (3), in ventral view (4), and in lateral view (5); left metatibia in dorsal view (6), and in lateral view (7). Abbreviation: mn, mentum." httpUri="https://zenodo.org/record/254869/files/figure.png" pageId="1" pageNumber="272">Figs 3–5</figureCitation>
|
||
) broadest at moderately large and moderately strongly convex eyes, HL
|
||
<quantity id="B4559B632B253C303DDAE95CFADCF80E" box="[1210,1376,1919,1943]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="1" pageNumber="272" unit="mm" value="0.28" valueMax="0.29" valueMin="0.27">0.27–0.29 mm</quantity>
|
||
, HW
|
||
<quantity id="B4559B632B253C3039F7E981FE83F822" box="[151,319,1954,1978]" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.0" metricValueMin="3.9" pageId="1" pageNumber="272" unit="mm" value="0.395" valueMax="0.4" valueMin="0.39">0.39–0.40 mm</quantity>
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||
; tempora rounded and about as long as eye in dorsal view; vertex strongly transverse and weakly convex, with pair of shallow but distinct pits located near posterior margins of supraantennal tubercles; frons strongly convex, median region of clypeus strongly projecting (
|
||
<figureCitation id="EB962A032B263C333A3DEEB4FC7EFF28" box="[861,962,151,176]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1973,1996]" captionTargetBox="[151,1430,661,1952]" captionTargetId="figure@3.[151,1435,661,1953]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Diagnostic characters of Horaeomorphus bicornis. Head in dorsal view (3), in ventral view (4), and in lateral view (5); left metatibia in dorsal view (6), and in lateral view (7). Abbreviation: mn, mentum." httpUri="https://zenodo.org/record/254869/files/figure.png" pageId="2" pageNumber="273">Figs 3, 5</figureCitation>
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||
); supra-antennal tubercles strongly raised, well delimited from frons but confluent with vertex. Punctures on vertex and frons fine and inconspicuous; setae moderately long, sparse, suberect to erect. Antennae (
|
||
<figureCitation id="EB962A032B263C333B92EEFCFCF8FF60" box="[754,836,223,248]" captionStart="FIGURES 8 – 12" captionStartId="4.[151,264,1894,1917]" captionTargetBox="[153,1436,211,1859]" captionTargetId="figure@4.[151,1436,193,1873]" captionTargetPageId="4" captionText="FIGURES 8 – 12. Diagnostic characters of Horaeomorphus bicornis. Aedeagus in ventral view (8), in lateral view (9), and in dorsal view; endophallus, enlarged, in ventral view (11); and left antenna, in dorsal view (12); photographed (a) and schematic (b)." httpUri="https://zenodo.org/record/254870/files/figure.png" pageId="2" pageNumber="273">Fig. 12</figureCitation>
|
||
) slender, shorter than half length of body, AnL
|
||
<quantity id="B4559B632B263C333C3DEEFCFF42FE84" metricMagnitude="-4" metricUnit="m" metricValue="7.8" metricValueMax="8.1" metricValueMin="7.5" pageId="2" pageNumber="273" unit="mm" value="0.78" valueMax="0.81" valueMin="0.75">0.75– 0.81 mm</quantity>
|
||
; relative lengths of antennomeres: 2.0: 1.6: 1.9: 1.8: 1.6: 1.6: 1.4: 1.4:1.8: 1.8: 2.8.
|
||
</paragraph>
|
||
<paragraph id="731236862B263C3339A7EF04FCE9FE48" blockId="2.[151,1436,151,464]" pageId="2" pageNumber="273">
|
||
Labrum transverse, anterior margin rounded, posterior margin strongly projecting at middle, forming a slightly reflexed, flattened horn (see in
|
||
<figureCitation id="EB962A032B263C333892EF6FFD88FEFD" box="[498,564,332,357]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1973,1996]" captionTargetBox="[151,1430,661,1952]" captionTargetId="figure@3.[151,1435,661,1953]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Diagnostic characters of Horaeomorphus bicornis. Head in dorsal view (3), in ventral view (4), and in lateral view (5); left metatibia in dorsal view (6), and in lateral view (7). Abbreviation: mn, mentum." httpUri="https://zenodo.org/record/254869/files/figure.png" pageId="2" pageNumber="273">Fig. 5</figureCitation>
|
||
), which in dorsal view (
|
||
<figureCitation id="EB962A032B263C333A21EF6FFC3FFEFC" box="[833,899,332,357]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1973,1996]" captionTargetBox="[151,1430,661,1952]" captionTargetId="figure@3.[151,1435,661,1953]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Diagnostic characters of Horaeomorphus bicornis. Head in dorsal view (3), in ventral view (4), and in lateral view (5); left metatibia in dorsal view (6), and in lateral view (7). Abbreviation: mn, mentum." httpUri="https://zenodo.org/record/254869/files/figure.png" pageId="2" pageNumber="273">Fig. 3</figureCitation>
|
||
) is gradually broadening apically and forming a transverse, rounded anterior margin. Left mandible with two subapical teeth, right mandible with only one subapical tooth (in dorsal view,
|
||
<figureCitation id="EB962A032B263C333B66EFB7FDF7FE34" box="[518,587,404,429]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1973,1996]" captionTargetBox="[151,1430,661,1952]" captionTargetId="figure@3.[151,1435,661,1953]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Diagnostic characters of Horaeomorphus bicornis. Head in dorsal view (3), in ventral view (4), and in lateral view (5); left metatibia in dorsal view (6), and in lateral view (7). Abbreviation: mn, mentum." httpUri="https://zenodo.org/record/254869/files/figure.png" pageId="2" pageNumber="273">Fig. 3</figureCitation>
|
||
). Mentum with each lateral margin strongly bent ventrally, in lateral view each forming a large lateral lobe with rounded margin (
|
||
<figureCitation id="EB962A032B263C333A62EF94FCFBFE48" box="[770,839,439,464]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1973,1996]" captionTargetBox="[151,1430,661,1952]" captionTargetId="figure@3.[151,1435,661,1953]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Diagnostic characters of Horaeomorphus bicornis. Head in dorsal view (3), in ventral view (4), and in lateral view (5); left metatibia in dorsal view (6), and in lateral view (7). Abbreviation: mn, mentum." httpUri="https://zenodo.org/record/254869/files/figure.png" pageId="2" pageNumber="273">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="27D2660E2B263C3339F7E979FAE8F8E9" box="[151,1364,1882,1905]" httpUri="https://zenodo.org/record/254868/files/figure.png" pageId="2" pageNumber="273" targetBox="[163,1425,526,1854]" targetPageId="2">
|
||
<paragraph id="731236862B263C3339F7E979FAE8F8E9" blockId="2.[151,1364,1882,1905]" box="[151,1364,1882,1905]" pageId="2" pageNumber="273">
|
||
<emphasis id="41D9EA942B263C3339F7E979FE85F8E8" bold="true" box="[151,313,1882,1905]" pageId="2" pageNumber="273">FIGURES 1–2.</emphasis>
|
||
Dorsal habitus of
|
||
<taxonomicName id="B4AD4D052B263C333895E978FD27F8E9" box="[501,667,1883,1905]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="41D9EA942B263C333895E978FD27F8E9" box="[501,667,1883,1905]" italics="true" pageId="2" pageNumber="273">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<taxonomicName id="B4AD4D052B263C333B97E97FFCD4F8E9" box="[759,872,1883,1905]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="41D9EA942B263C333B97E97FFCD4F8E9" box="[759,872,1883,1905]" italics="true" pageId="2" pageNumber="273">H. bicornis</emphasis>
|
||
</taxonomicName>
|
||
, holotype male (1);
|
||
<taxonomicName id="B4AD4D052B263C333D57E97FFB50F8E9" box="[1079,1260,1884,1905]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="species" species="punctatissimus">
|
||
<emphasis id="41D9EA942B263C333D57E97FFB50F8E9" box="[1079,1260,1884,1905]" italics="true" pageId="2" pageNumber="273">H. punctatissimus</emphasis>
|
||
</taxonomicName>
|
||
, male (2).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="731236862B263C3239A7E9BAFD26FE84" blockId="2.[151,1436,1944,2005]" lastBlockId="3.[151,1436,151,608]" lastPageId="3" lastPageNumber="274" pageId="2" pageNumber="273">
|
||
Pronotum subtrapezoidal, elongate, broadest near anterior third, PL
|
||
<quantity id="B4559B632B263C333ADEE9BBFBDEF828" box="[958,1122,1944,1969]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" metricValueMax="5.4" metricValueMin="5.0" pageId="2" pageNumber="273" unit="mm" value="0.52" valueMax="0.54" valueMin="0.5">0.50–0.54 mm</quantity>
|
||
, PWb
|
||
<quantity id="B4559B632B263C333DCCE9BBFAEDF828" box="[1196,1361,1944,1969]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.7" metricValueMin="3.5" pageId="2" pageNumber="273" unit="mm" value="0.36" valueMax="0.37" valueMin="0.35">0.35–0.37 mm</quantity>
|
||
, PWm
|
||
<quantity id="B4559B632B263C3339F7E99FFEFCF84C" box="[151,320,1980,2005]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.1" metricValueMin="4.9" pageId="2" pageNumber="273" unit="mm" value="0.5" valueMax="0.51" valueMin="0.49">0.49–0.51 mm</quantity>
|
||
; hind angles obtuse and blunt; posterior margin rounded; posterior collar delimited from disc by broad, but shallow transverse groove, which is slightly convex posteriorly and connects two lateral pits, median pit large and deep, located anteriorly to the antebasal groove, which is anteriorly connected to a short and narrow longitudinal groove. Punctures on disc much coarser and denser than those on frons and vertex; vestiture composed of sparse, moderately long and suberect setae.
|
||
</paragraph>
|
||
<paragraph id="731236862B273C3239A7EF04FCA8FD80" blockId="3.[151,1436,151,608]" pageId="3" pageNumber="274">
|
||
Elytra strongly elongate, oval, slightly more convex than pronotum, broadest between middle and anterior third, EL
|
||
<quantity id="B4559B632B273C32386EEF6FFE05FEFC" box="[270,441,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.4849999999999999" metricValueMax="1.5" metricValueMin="1.47" pageId="3" pageNumber="274" unit="mm" value="1.4849999999999999" valueMax="1.5" valueMin="1.47">1.47–1.50 mm</quantity>
|
||
, EW
|
||
<quantity id="B4559B632B273C323B62EF6FFD11FEFC" box="[514,685,332,357]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="6.8999999999999995" metricValueMin="6.6" pageId="3" pageNumber="274" unit="mm" value="0.675" valueMax="0.69" valueMin="0.66">0.66–0.69 mm</quantity>
|
||
, elytral index 1.4 7–1.50. Each elytron with moderately distinct humerus. Punctures deep and large, with relatively sharp margins, in anterior half dense, distances between punctures are smaller than or equal to puncture diameter, in posterior half of elytra punctures gradually reducing in diameter and becoming shallower, less distinct, each elytra with a longitudinal, sparsely and finely punctured area located along elytral suture, setation moderately dense and long, composed of setae slightly thicker than those on pronotum, suberect. Metathoracic wings well developed.
|
||
</paragraph>
|
||
<paragraph id="731236862B273C3239A7EC07FC30FDF8" blockId="3.[151,1436,151,608]" pageId="3" pageNumber="274">
|
||
Hind legs strongly modified, with each metatibia (
|
||
<figureCitation id="EB962A032B273C323B99EC07FCE1FDA5" box="[761,861,548,573]" captionStart="FIGURES 3 – 7" captionStartId="3.[151,264,1973,1996]" captionTargetBox="[151,1430,661,1952]" captionTargetId="figure@3.[151,1435,661,1953]" captionTargetPageId="3" captionText="FIGURES 3 – 7. Diagnostic characters of Horaeomorphus bicornis. Head in dorsal view (3), in ventral view (4), and in lateral view (5); left metatibia in dorsal view (6), and in lateral view (7). Abbreviation: mn, mentum." httpUri="https://zenodo.org/record/254869/files/figure.png" pageId="3" pageNumber="274">Figs 6–7</figureCitation>
|
||
) strongly bent at anterior third with the posterior 2/ 3 flattened, forming sharp outer maginal projecting at bending side.
|
||
</paragraph>
|
||
<caption id="27D2660E2B273C3239F7E996FBF0F873" httpUri="https://zenodo.org/record/254869/files/figure.png" pageId="3" pageNumber="274" targetBox="[151,1430,661,1952]" targetPageId="3">
|
||
<paragraph id="731236862B273C3239F7E996FBF0F873" blockId="3.[151,1436,1973,2027]" pageId="3" pageNumber="274">
|
||
<emphasis id="41D9EA942B273C3239F7E996FE86F854" bold="true" box="[151,314,1973,1996]" pageId="3" pageNumber="274">FIGURES 3–7.</emphasis>
|
||
Diagnostic characters of
|
||
<taxonomicName id="B4AD4D052B273C323B20E995FCFCF854" box="[576,832,1974,1996]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="41D9EA942B273C323B20E995FCFCF854" box="[576,832,1974,1996]" italics="true" pageId="3" pageNumber="274">Horaeomorphus bicornis</emphasis>
|
||
</taxonomicName>
|
||
. Head in dorsal view (3), in ventral view (4), and in lateral view (5); left metatibia in dorsal view (6), and in lateral view (7). Abbreviation: mn, mentum.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="27D2660E2B203C3539F7E945FF07F822" httpUri="https://zenodo.org/record/254870/files/figure.png" pageId="4" pageNumber="275" targetBox="[153,1436,211,1859]" targetPageId="4">
|
||
<paragraph id="731236862B203C3539F7E945FF07F822" blockId="4.[151,1436,1894,1978]" pageId="4" pageNumber="275">
|
||
<emphasis id="41D9EA942B203C3539F7E945FEF4F8E4" bold="true" box="[151,328,1894,1917]" pageId="4" pageNumber="275">FIGURES 8–12.</emphasis>
|
||
Diagnostic characters of
|
||
<taxonomicName id="B4AD4D052B203C353B2FE945FCECF8E4" box="[591,848,1894,1916]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="275" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="41D9EA942B203C353B2FE945FCECF8E4" box="[591,848,1894,1916]" italics="true" pageId="4" pageNumber="275">Horaeomorphus bicornis</emphasis>
|
||
</taxonomicName>
|
||
. Aedeagus in ventral view (8), in lateral view (9), and in dorsal view; endophallus, enlarged, in ventral view (11); and left antenna, in dorsal view (12); photographed (a) and schematic (b).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="731236862B213C3439A7EEB4FCE9FEA7" blockId="5.[151,1436,151,1184]" pageId="5" pageNumber="276">
|
||
Aedeagus (
|
||
<figureCitation id="EB962A032B213C343828EEB4FE02FF28" box="[328,446,151,176]" captionStart="FIGURES 8 – 12" captionStartId="4.[151,264,1894,1917]" captionTargetBox="[153,1436,211,1859]" captionTargetId="figure@4.[151,1436,193,1873]" captionTargetPageId="4" captionText="FIGURES 8 – 12. Diagnostic characters of Horaeomorphus bicornis. Aedeagus in ventral view (8), in lateral view (9), and in dorsal view; endophallus, enlarged, in ventral view (11); and left antenna, in dorsal view (12); photographed (a) and schematic (b)." httpUri="https://zenodo.org/record/254870/files/figure.png" pageId="5" pageNumber="276">Figs 8–10</figureCitation>
|
||
) oval in ventral and dorsal view, AeL
|
||
<quantity id="B4559B632B213C343AEDEEB4FC4BFF37" box="[909,1015,151,176]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="5" pageNumber="276" unit="mm" value="0.36">0.36 mm</quantity>
|
||
, broadest near middle, with slightly emarginate apex, two lateral groups of three short setae each are located apically ondorsal wall; parameres slender, each with one apical and one subapical seta; endophallus (
|
||
<figureCitation id="EB962A032B213C343A2CEEFCFC1CFF60" box="[844,928,223,248]" captionStart="FIGURES 8 – 12" captionStartId="4.[151,264,1894,1917]" captionTargetBox="[153,1436,211,1859]" captionTargetId="figure@4.[151,1436,193,1873]" captionTargetPageId="4" captionText="FIGURES 8 – 12. Diagnostic characters of Horaeomorphus bicornis. Aedeagus in ventral view (8), in lateral view (9), and in dorsal view; endophallus, enlarged, in ventral view (11); and left antenna, in dorsal view (12); photographed (a) and schematic (b)." httpUri="https://zenodo.org/record/254870/files/figure.png" pageId="5" pageNumber="276">Fig. 11</figureCitation>
|
||
) complicated, in dorsal view with strongly developed central structure, which is inversely triangular and connected to a ligulate sclerite by a M-shaped structure, and relatively weakly developed lateral components.
|
||
</paragraph>
|
||
<paragraph id="731236862B213C3439A7EF6FFE22FEFD" blockId="5.[151,1436,151,1184]" box="[199,414,332,357]" pageId="5" pageNumber="276">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3BB7650D2B213C3439A7EF4CFF7FFBC0" pageId="5" pageNumber="276" type="discussion">
|
||
<paragraph id="731236862B213C3439A7EF4CFEE9FC8C" blockId="5.[151,1436,151,1184]" pageId="5" pageNumber="276">
|
||
<emphasis id="41D9EA942B213C3439A7EF4CFE87FE10" bold="true" box="[199,315,367,392]" pageId="5" pageNumber="276">Remarks.</emphasis>
|
||
<taxonomicName id="B4AD4D052B213C343821EF53FDE7FE10" box="[321,603,368,392]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="41D9EA942B213C343821EF53FDE7FE10" box="[321,603,368,392]" italics="true" pageId="5" pageNumber="276">Horaeomorphus bicornis</emphasis>
|
||
</taxonomicName>
|
||
is the first species of the genus with a strongly modified head bearing both median labral and clypeal projections. Such characters were also rarely reported in other
|
||
<taxonomicName id="B4AD4D052B213C343D1BEFB7FAA8FE35" box="[1147,1300,404,429]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="tribe" tribe="Glandulariini">Glandulariini</taxonomicName>
|
||
. In the huge genus
|
||
<taxonomicName id="B4AD4D052B213C343980EF9AFE7EFE48" authority="Thomson" authorityName="Thomson" box="[224,450,439,464]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="41D9EA942B213C343980EF9AFEF2FE48" box="[224,334,441,464]" italics="true" pageId="5" pageNumber="276">Euconnus</emphasis>
|
||
Thomson
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B4AD4D052B213C3438AEEF9AFCBDFE48" authority="Franz, 1980" authorityName="Franz" authorityYear="1980" box="[462,769,440,464]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="fronticornis">
|
||
<emphasis id="41D9EA942B213C3438AEEF9AFDCCFE48" box="[462,624,440,464]" italics="true" pageId="5" pageNumber="276">E. fronticornis</emphasis>
|
||
<bibRefCitation id="173C4B772B213C343B18EF9BFCBDFE48" author="Franz" box="[632,769,440,464]" pageId="5" pageNumber="276" refString="Franz, H. (1980) Monographie der sudamerikanischen Scydmaenidae mit Einschluss einiger mittelamerikanischer Arten (Coleoptera). Folia Entomologica Hungarica, 41, 39 - 264." type="journal article" year="1980">Franz, 1980</bibRefCitation>
|
||
</taxonomicName>
|
||
, described as having a horn on the anterior margin of frons (“Besitz eines stumpfen Hornes nahe dem Vorderrande der Stirn”,
|
||
<bibRefCitation id="173C4B772B213C343AEFEFFEFBA8FE6C" author="Franz" box="[911,1044,476,500]" pageId="5" pageNumber="276" refString="Franz, H. (1980) Monographie der sudamerikanischen Scydmaenidae mit Einschluss einiger mittelamerikanischer Arten (Coleoptera). Folia Entomologica Hungarica, 41, 39 - 264." type="journal article" year="1980">Franz 1980</bibRefCitation>
|
||
), a similar structure also occurs in the Peruvian monotypic genus
|
||
<taxonomicName id="B4AD4D052B213C343894EC23FCFBFD80" authority="Franz, 1989" authorityName="Franz" authorityYear="1989" box="[500,839,512,536]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="41D9EA942B213C343894EC23FD0AFD80" box="[500,694,512,536]" italics="true" pageId="5" pageNumber="276">Protandroconnus</emphasis>
|
||
<bibRefCitation id="173C4B772B213C343BDEEC23FCFBFD80" author="Franz" box="[702,839,512,536]" pageId="5" pageNumber="276" refString="Franz, H. (1989) Neue Scydmaeniden (Coleoptera) aus dem tropischen Sudamerika. Revue Suisse de Zoologie 96, 629 - 636. http: // dx. doi. org / 10.5962 / bhl. part. 82052" type="journal article" year="1989">Franz, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
. However, none of the foregoing species has a labral projection. The mentum in
|
||
<taxonomicName id="B4AD4D052B213C34388CEC07FD18FDA4" box="[492,676,548,572]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="41D9EA942B213C34388CEC07FD18FDA4" box="[492,676,548,572]" italics="true" pageId="5" pageNumber="276">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
shows some interspecifically variable, with its lateral margin sometimes elevated (Jałoszyński, unpublished observations), but no other species has such large lateral lobes as those found in
|
||
<taxonomicName id="B4AD4D052B213C343828EC4EFE76FD1C" box="[328,458,620,644]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="41D9EA942B213C343828EC4EFE76FD1C" box="[328,458,620,644]" italics="true" pageId="5" pageNumber="276">H. bicornis</emphasis>
|
||
</taxonomicName>
|
||
. Strongly bent metafemora are seen in
|
||
<taxonomicName id="B4AD4D052B213C343AF9EC4EFB44FD1C" authority="Franz, 1992" authorityName="Franz" authorityYear="1992" box="[921,1272,620,644]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="punctatissimus">
|
||
<emphasis id="41D9EA942B213C343AF9EC4EFBD8FD1C" box="[921,1124,621,644]" italics="true" pageId="5" pageNumber="276">H. punctatissimus</emphasis>
|
||
<bibRefCitation id="173C4B772B213C343D0FEC4EFB44FD1C" author="Franz" box="[1135,1272,620,644]" pageId="5" pageNumber="276" refString="Franz, H. (1992) Monographie der Scydmaenidae (Coleoptera) von Sabah (NO-Borneo), Revue suisse de Zoologie, 99 (4), 859 - 953. http: // dx. doi. org / 10.5962 / bhl. part. 79857" type="journal article" year="1992">Franz, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
(illustrated in
|
||
<bibRefCitation id="173C4B772B213C3439F7ECACFEDDFD30" author="Jaloszynski" box="[151,353,655,680]" pageId="5" pageNumber="276" refString="Jaloszynski, P. (2006) Revision of Horaeomorphus Schaufuss of East Malaysia, Singapore, and Sunda Islands (Coleoptera, Scydmaenidae). Genus 17 (1), 19 - 66." type="journal article" year="2006">Jałoszyński 2006</bibRefCitation>
|
||
), and
|
||
<taxonomicName id="B4AD4D052B213C3438D1ECB2FCFBFD30" authority="Jaloszynski, 2009" authorityName="Jaloszynski" authorityYear="2009" box="[433,839,655,680]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="samosirensis">
|
||
<emphasis id="41D9EA942B213C3438D1ECB2FDD4FD3F" box="[433,616,656,680]" italics="true" pageId="5" pageNumber="276">H. samosirensis</emphasis>
|
||
<bibRefCitation id="173C4B772B213C343B16ECACFCFBFD30" author="Jaloszynski" box="[630,839,655,680]" pageId="5" pageNumber="276" refString="Jaloszynski, P. (2009) Four new species of Horaeomorphus Schaufuss from Oriental Region (Coleoptera: Scydmaenidae). Raffles Bulletin of Zoology 57 (2), 297 - 303." type="journal article" year="2009">Jałoszyński, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
, but no species was known with each metafemur projecting laterally at the bending side. The presence of setae on the apex of aedeagal dorsal wall was reported in the genus
|
||
<taxonomicName id="B4AD4D052B213C343868ECFBFECAFD77" box="[264,374,728,751]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="280" phylum="Arthropoda" rank="genus" sensu="stricto">Euconnus</taxonomicName>
|
||
(e.g.,
|
||
<bibRefCitation id="173C4B772B213C3438DCECF4FD30FD68" author="Jaloszynski" box="[444,652,727,752]" pageId="5" pageNumber="276" refString="Jaloszynski, P. (2012 b) Taxonomy of ' Euconnus complex'. Part I. Morphology of Euconnus s. str. and revision of Euconnomorphus Franz and Venezolanoconnus Franz (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3555, 55 - 82." type="journal article" year="2012" yearSuffix="b">Jałoszyński 2012b</bibRefCitation>
|
||
, figs 43–44), but this character is for the first time reported to occur in
|
||
<taxonomicName id="B4AD4D052B213C3439F7ECDFFEF3FC8C" box="[151,335,764,788]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="41D9EA942B213C3439F7ECDFFEF3FC8C" box="[151,335,764,788]" italics="true" pageId="5" pageNumber="276">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="731236862B213C3439A7ED3CFA3FFC18" blockId="5.[151,1436,151,1184]" pageId="5" pageNumber="276">
|
||
Within the genus,
|
||
<taxonomicName id="B4AD4D052B213C3438F4ED02FDAFFCA0" box="[404,531,800,824]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="41D9EA942B213C3438F4ED02FDAFFCA0" box="[404,531,800,824]" italics="true" pageId="5" pageNumber="276">H. bicornis</emphasis>
|
||
</taxonomicName>
|
||
shares the small and flattened body, the elongate, subtrapezoidal pronotum with paired lateral antebasal pits connected by a shallow groove, and the median antebasal pit connected anteriorly to a narrow longitudinal groove, with its Malaysian congener
|
||
<taxonomicName id="B4AD4D052B213C343A7BED4AFBF3FC18" authority="Lhoste, 1939" authorityName="Lhoste" authorityYear="1939" box="[795,1103,871,896]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="wailimae">
|
||
<emphasis id="41D9EA942B213C343A7BED4AFC18FC18" box="[795,932,872,896]" italics="true" pageId="5" pageNumber="276">H. wailimae</emphasis>
|
||
(Lhoste, 1939)
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="173C4B772B213C343D3EED44FA9FFC18" author="Jaloszynski" box="[1118,1315,871,896]" pageId="5" pageNumber="276" refString="Jaloszynski, P. (2006) Revision of Horaeomorphus Schaufuss of East Malaysia, Singapore, and Sunda Islands (Coleoptera, Scydmaenidae). Genus 17 (1), 19 - 66." type="journal article" year="2006">Jałoszyński 2006</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="173C4B772B213C343C4EED4BFAC4FC18" author="Jaloszynski" box="[1326,1400,872,896]" pageId="5" pageNumber="276" refString="Jaloszynski, P. (2014 a) Oriental Horaeomorphus Schaufuss: a new species, synonymic notes and distributional synopsis (Coleoptera: Staphylinidae: Scydmaeninae). Raffles Bulletin of Zoology 62, 83 - 88." type="journal article" year="2014" yearSuffix="a">2014a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="731236862B213C3439A7EDAFFC6AFB88" blockId="5.[151,1436,151,1184]" pageId="5" pageNumber="276">
|
||
This is the third species described from a small area of Southwest
|
||
<collectingRegion id="B169F8642B213C343ACFEDAEFBB8FC3C" box="[943,1028,909,932]" country="China" name="Yunnan" pageId="5" pageNumber="276">Yunnan</collectingRegion>
|
||
Prov. (another two are
|
||
<taxonomicName id="B4AD4D052B213C343C6CEDAEFE30FC50" authority="Zhou & Zhang, 2016" authorityName="Zhou & Zhang" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="41D9EA942B213C343C6CEDAEFA20FC3C" box="[1292,1436,909,932]" italics="true" pageId="5" pageNumber="276">H. punctatus</emphasis>
|
||
<bibRefCitation id="173C4B772B213C3439F7ED8CFE30FC50" author="Zhou" box="[151,396,943,968]" pageId="5" pageNumber="276" refString="Zhou, D. Y., Zhang, S. J. & Li, L. Z. (2016) Contributions to the knowledge of the genus Horaeomorphus Schaufuss (Coleoptera, Staphylinidae, Scydmaeninae) in mainland China. Zookeys, 572, 51 - 70." type="journal article" year="2016">Zhou & Zhang, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
from Yingjiang Hsien, and
|
||
<taxonomicName id="B4AD4D052B213C343BB4ED92FBC6FC50" authority="Zhou & Zhang, 2016" authorityName="Zhou & Zhang" authorityYear="2016" box="[724,1146,943,968]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="pengzhongi">
|
||
<emphasis id="41D9EA942B213C343BB4ED92FCC6FC50" box="[724,890,944,968]" italics="true" pageId="5" pageNumber="276">H. pengzhongi</emphasis>
|
||
<bibRefCitation id="173C4B772B213C343AE4ED8CFBC6FC50" author="Zhou" box="[900,1146,943,968]" pageId="5" pageNumber="276" refString="Zhou, D. Y., Zhang, S. J. & Li, L. Z. (2016) Contributions to the knowledge of the genus Horaeomorphus Schaufuss (Coleoptera, Staphylinidae, Scydmaeninae) in mainland China. Zookeys, 572, 51 - 70." type="journal article" year="2016">Zhou & Zhang, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
from Tengchong Hsien). Together with an unidentifiable female mentioned below, there are already four species discovered in this area, which reveals a biodiversity hotspot of
|
||
<taxonomicName id="B4AD4D052B213C343B30EDDBFCB4FB88" box="[592,776,1016,1040]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="41D9EA942B213C343B30EDDBFCB4FB88" box="[592,776,1016,1040]" italics="true" pageId="5" pageNumber="276">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
out of Sundaland.
|
||
</paragraph>
|
||
<paragraph id="731236862B213C3439A7EA3FFF7FFBC0" blockId="5.[151,1436,151,1184]" pageId="5" pageNumber="276">
|
||
<emphasis id="41D9EA942B213C3439A7EA3FFEF6FBAD" bold="true" box="[199,330,1052,1077]" pageId="5" pageNumber="276">Bionomics.</emphasis>
|
||
<typeStatus id="AC1688242B213C343833EA3EFE37FBAD" box="[339,395,1053,1077]" pageId="5" pageNumber="276">Type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="B4AD4D052B213C343B4FEA3EFD13FBAC" box="[559,687,1052,1076]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="41D9EA942B213C343B4FEA3EFD13FBAC" box="[559,687,1052,1076]" italics="true" pageId="5" pageNumber="276">H. bicornis</emphasis>
|
||
</taxonomicName>
|
||
were collected by sifting chips of rotten log on the roadside (
|
||
<figureCitation id="EB962A032B213C343C0FEA3FFF0AFBC0" captionStart="FIGURES 21 – 23" captionStartId="8.[151,264,1570,1593]" captionTargetBox="[151,1435,193,1548]" captionTargetId="figure@8.[151,1435,193,1549]" captionTargetPageId="8" captionText="FIGURES 21 – 23. Habitat of H. bicornis in Yinjiang Hsien (21). Living male adult of H. punctatissimus on a rotten log in Mt. T" httpUri="https://zenodo.org/record/254873/files/figure.png" pageId="5" pageNumber="276">Fig. 21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3BB7650D2B213C3439A7EA47FD2EFBE4" box="[199,658,1124,1149]" pageId="5" pageNumber="276" type="distribution">
|
||
<paragraph id="731236862B213C3439A7EA47FD2EFBE4" blockId="5.[151,1436,151,1184]" box="[199,658,1124,1149]" pageId="5" pageNumber="276">
|
||
<emphasis id="41D9EA942B213C3439A7EA47FEDCFBE5" bold="true" box="[199,352,1124,1149]" pageId="5" pageNumber="276">Distribution.</emphasis>
|
||
Southwest
|
||
<collectingCountry id="0BBA76162B213C343883EA47FD97FBE5" box="[483,555,1124,1149]" name="China" pageId="5" pageNumber="276">China</collectingCountry>
|
||
:
|
||
<collectingRegion id="B169F8642B213C343B55EA46FD32FBE4" box="[565,654,1125,1148]" country="China" name="Yunnan" pageId="5" pageNumber="276">Yunnan</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3BB7650D2B213C3439A7EAA4FBABFB38" box="[199,1047,1159,1184]" pageId="5" pageNumber="276" type="etymology">
|
||
<paragraph id="731236862B213C3439A7EAA4FBABFB38" blockId="5.[151,1436,151,1184]" box="[199,1047,1159,1184]" pageId="5" pageNumber="276">
|
||
<emphasis id="41D9EA942B213C3439A7EAA4FEF2FB38" bold="true" box="[199,334,1159,1184]" pageId="5" pageNumber="276">Etymology.</emphasis>
|
||
The specific epithet refers to the labral and clypeal projections.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |