304 lines
57 KiB
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304 lines
57 KiB
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<document id="90E18637D9E855D6BE5FA96B314D941F" ID-CLB-Dataset="53877" ID-DOI="10.1016/j.cretres.2023.105590" ID-GBIF-Dataset="c69ee18a-f90f-472d-ad6e-71cf4d2f84ee" ID-Zenodo-Dep="8250332" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1692117422342" checkinUser="felipe" docAuthor="Jacobs, Megan L., Pérez-García, Adán, Martín-Jiménez, Marcos, Mottram, Catherine M., Martill, David M., Gale, Andrew S., Mattsson, Oliver L. & Wood, Charles" docDate="2023" docId="566E87D4966AFFA4287BFE020A6FF6AA" docLanguage="en" docName="CretRes.150.105590.pdf.imf" docOrigin="Cretaceous Research 150 (9)" docSource="http://dx.doi.org/10.1016/j.cretres.2023.105590" docTitle="Eodortoka morellana Perez-Garcia, Gasulla and Ortega 2014" docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="AA57FFAC966DFFAF2B49FFCB0B4FF417" masterDocTitle="A well preserved pan-pleurodiran (Dortokidae) turtle from the English Lower Cretaceous and the first radiometric date for the Wessex Formation (Hauterivian-Barremian) of the Isle of Wight, United Kingdom" masterLastPageNumber="429" masterPageNumber="428" pageNumber="8" updateTime="1692118029474" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="DF17BB56FBBF8AE61DAD6BEF4F99201E">A well preserved pan-pleurodiran (Dortokidae) turtle from the English Lower Cretaceous and the first radiometric date for the Wessex Formation (Hauterivian-Barremian) of the Isle of Wight, United Kingdom</mods:title>
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<mods:namePart id="F8DD3B9B06870784A1F62E048DA93E31">Jacobs, Megan L.</mods:namePart>
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<mods:affiliation id="EC74C96ACCFB9E6E9D28F53AD068454A">School of the Environment, Geography and Geosciences, University of Portsmouth, Portsmouth, PO 1 3 QL, UK</mods:affiliation>
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<mods:nameIdentifier id="5B5EA62670C157D0DE327454F48B065F" type="email">megan.jacobs@port.ac.uk</mods:nameIdentifier>
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<mods:namePart id="4FC792BA9D57E662132D85BFCE8117A4">Pérez-García, Adán</mods:namePart>
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<mods:affiliation id="CC0B009E7D1A7915EF736F7944D8FA97">Grupo de Biología Evolutiva, Facultad de Ciencias, UNED, 28232, Las Rozas, Madrid, Spain</mods:affiliation>
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<mods:namePart id="DE90A579D20FF9F6500D4DE5828EA8FD">Martín-Jiménez, Marcos</mods:namePart>
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<mods:affiliation id="AE8AAD91D4723105FAF911155D9D98F5">Grupo de Biología Evolutiva, Facultad de Ciencias, UNED, 28232, Las Rozas, Madrid, Spain</mods:affiliation>
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<mods:namePart id="7DA48D33AB0633E9F9EFB9A7D4C55279">Mottram, Catherine M.</mods:namePart>
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<mods:affiliation id="E0450A959AE61F96F27649EBC91FC9C2">School of the Environment, Geography and Geosciences, University of Portsmouth, Portsmouth, PO 1 3 QL, UK</mods:affiliation>
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<mods:namePart id="67A10573D7EE181BCF73CADB5C1769FC">Martill, David M.</mods:namePart>
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<mods:affiliation id="A0250989B149FDA75AC212152B332B55">School of the Environment, Geography and Geosciences, University of Portsmouth, Portsmouth, PO 1 3 QL, UK</mods:affiliation>
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<mods:namePart id="3154405752E9BEEA4577E1B2DC539238">Gale, Andrew S.</mods:namePart>
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<mods:affiliation id="2F2572D47AE28042AF3324CBC08C8532">School of the Environment, Geography and Geosciences, University of Portsmouth, Portsmouth, PO 1 3 QL, UK</mods:affiliation>
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<mods:namePart id="ED5A5D149F46DD5E646AA3D869F171F3">Mattsson, Oliver L.</mods:namePart>
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<mods:affiliation id="688EB8A0EECF0398B997EC2328DD4A89">Dinosaur Expeditions C. I. C., Military Road, Brighstone, Isle of Wight, PO 30 4 PG, UK</mods:affiliation>
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<mods:namePart id="1999DA83163FCB5466D92715599FCD5E">Wood, Charles</mods:namePart>
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<mods:affiliation id="FCB5C29739108C3224ADD0C7A009E80D">Future Technology Centre, School of Mechanical Design and Engineering, University of Portsmouth, Portsmouth, PO 1 3 HE, UK</mods:affiliation>
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<mods:title id="D2636A2A3EDDDB244038606D68BED29D">Cretaceous Research</mods:title>
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<treatment id="566E87D4966AFFA4287BFE020A6FF6AA" ID-DOI="http://doi.org/10.5281/zenodo.8250336" ID-Zenodo-Dep="8250336" LSID="urn:lsid:plazi:treatment:566E87D4966AFFA4287BFE020A6FF6AA" httpUri="http://treatment.plazi.org/id/566E87D4966AFFA4287BFE020A6FF6AA" lastPageId="11" lastPageNumber="12" pageId="7" pageNumber="8">
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<subSubSection id="96DD6549966AFFA8287BFE020EF6F5CA" box="[818,1465,457,477]" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="DE7836C2966AFFA8287BFE020EF6F5CA" blockId="7.[818,1465,457,505]" box="[818,1465,457,477]" pageId="7" pageNumber="8">
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<heading id="853081AE966AFFA8287BFE020EF6F5CA" box="[818,1465,457,477]" level="2" pageId="7" pageNumber="8">
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<taxonomicName id="19C74D41966AFFA8287BFE020EF6F5CA" authority="Perez-Garcia, Gasulla and Ortega, 2014" authorityName="Perez-Garcia, Gasulla and Ortega" authorityYear="2014" box="[818,1465,457,477]" family="Dortokidae" genus="Eodortoka" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="morellana">
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<emphasis id="ECB3EAD0966AFFA8287BFE0208D1F5CA" bold="true" box="[818,926,457,477]" italics="true" pageId="7" pageNumber="8">Eodortoka</emphasis>
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cf.
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<emphasis id="ECB3EAD0966AFFA8288BFE020F62F5CA" bold="true" box="[962,1069,457,477]" italics="true" pageId="7" pageNumber="8">morellana</emphasis>
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<bibRefCitation id="BA564B33966AFFA82F7AFE010EF6F5CA" author="Perez-Garcia, Gasulla and Ortega" box="[1075,1465,458,477]" firstAuthor="Perez-Garcia" pageId="7" pageNumber="8" pagination="130 - 138" refId="ref8214" refString="Perez-Garcia, A., Gasulla, J. M., Ortega, F., 2014. Eodortoka morellana gen. et sp. nov., the first pan-pleurodiran turtle (Dortokidae) defined in the Early Cretaceous of Europe. Cretaceous Research 48, 130 - 138." type="journal article" year="2014">Perez-García, Gasulla and Ortega, 2014</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="96DD6549966AFFA8287BFE2D08D1F5EE" box="[818,926,486,505]" pageId="7" pageNumber="8" type="description">
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<paragraph id="DE7836C2966AFFA8287BFE2D08D1F5EE" blockId="7.[818,1465,457,505]" box="[818,926,486,505]" pageId="7" pageNumber="8">
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(
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<figureCitation id="46FC2A47966AFFA82873FE2D08D8F5EE" box="[826,919,486,505]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStartId-0="3.[100,125,1945,1959]" captionStartId-1="4.[138,163,1288,1302]" captionStartId-2="5.[100,125,1874,1888]" captionStartId-3="6.[100,125,1870,1884]" captionStartId-4="7.[100,125,1932,1946]" captionStartId-5="8.[100,125,1945,1959]" captionStartId-6="9.[100,125,1814,1828]" captionTargetBox-0="[120,1475,391,1914]" captionTargetBox-1="[209,1869,201,1259]" captionTargetBox-2="[110,1478,543,1844]" captionTargetBox-3="[199,1389,181,1832]" captionTargetBox-4="[110,1478,912,1903]" captionTargetBox-5="[110,1478,1144,1915]" captionTargetBox-6="[110,1478,179,1785]" captionTargetId-0="figure-151@3.[110,1478,381,1915]" captionTargetId-1="figure-0@4.[207,1870,197,1259]" captionTargetId-2="figure-229@5.[110,1478,543,1844]" captionTargetId-3="figure-1@6.[196,1392,178,1841]" captionTargetId-4="figure-363@7.[110,1478,912,1903]" captionTargetId-5="figure-679@8.[110,1478,1144,1915]" captionTargetId-6="figure-0@9.[110,1478,178,1785]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="Fig. 3. IWCMS 2018.44, partial skeleton of Eodortoka cf. morellana. A, dorsal view, B, ventral view, C, anterior view, D, posterior view, E, left lateral view, F, right lateral view. G-H, details of the ornamental pattern on the outer surface of the shell, at the level of the left first costal (G) and of the left hypoplastron (H)." captionText-1="Fig. 4. IWCMS 2018.44, partial skeleton of Eodortoka cf. morellana. A,C,E, virtual three-dimensional reconstruction of the left shell plates and of the bones preserved inside the specimen; B,D,F, virtual three-dimensional reconstruction of the bones virtually removing the shell, in dorsal (A,B), ventral (C,D), and left lateral (E,F) views.The anterior and posterior ends of the shell cannot be reconstructed because they were not accurately scanned (black in A, E). The plates of the shell and the bones were reconstructed independently (shell plates in yellow, vertebrae in green, appendicular bones in blue, and girdles in pink). Abbreviations: c, costal; cav, caudal vertebra; cev7, seventh cervical vertebra; cev8, eighth cervical vertebra; dv, dorsal vertebra; hyo, hyoplastron; hyp, hypoplastron; lcor, left coracoid; lhu, left humerus; lil, left illium; lis; left ischium; lsc, left scapula; me, mesoplastron; ne, neural; p, peripheral; rfe, right femur; rfi, right fibula; rhu; right humerus; rpu, right pubis; rsc, right scapula; rti, right tibia; sv, sacral vertebra; xi, xiphiplastron." captionText-2="Fig. 5. Cervical vertebrae of IWCMS 2018.44. A-F, neural arch of the axis (cervical vertebra 2); G-L, cervical vertebra 3; M-R, cervical vertebra 4; S-X, cervical vertebra 5; Y-AD, cervical vertebra 6; AE-AJ, cervical vertebra 7; AK-AP, cervical vertebra 8. All of them are shown in left lateral (A,G,M,S,Y,AE,AK), right lateral (B,H,N,T,Z,AF,AL), anterior (C,I,O,U,AA,AG,AM), posterior (D,J,P,V,AB,AH,AN), dorsal (E,K,Q,W,AC,AI,AO), and ventral (F,L,R,X,AD,AJ,AP) views." captionText-3="Fig. 6. Dorsal vertebrae of IWCMS 2018.44. A-F, complete dorsal vertebra with fused neural; G-L, dorsal vertebra centrum; M-R, dorsal vertebra centrum; S-X, complete dorsal vertebra; Y-AD,neural arch of a posterior dorsal vertebra. All of them are shown in left lateral (A,G,M,S,Y), right lateral (B,H,N,T,Z), anterior (C,I,O,U,AA), posterior (D,J,P,V,AB), dorsal (E,K,Q,W,AC), and ventral (F,L,R,X,AD) views." captionText-4="Fig. 7. Sacral and caudal vertebrae of IWCMS 2018.44. A-D, sacral vertebrae; E-J, anterior caudal vertebra; K-L, posterior caudal vertebra; M-P posterior caudal vertebra. These elements are shown in anterior (A,G,K,M), posterior (B,H,L,N), dorsal (C,I,O), ventral (D,J,P), left lateral (E), and right lateral (F) views." captionText-5="Fig. 8. Elements of the girdles of IWCMS 2018.44. A-F, right scapula; G-L, left scapula; M-R, proximal end of the right coracoid; S-X, left coracoid. These elements are shown in posterior (A,G,M,S), ventral (B,H,O,V), dorsal (C,I,P,T), anterior (D,J,N,U), proximal (E,K,Q,W), and distal (F,L,R,X) views." captionText-6="Fig. 9. Visceral view of the left hypoplastron and xiphiplastron with the left pelvis (A), and virtually removing the pelvis (B) to show the pubic scar of IWCMS 2018.44.C-N, Left (C- H) and right (I-N) pelvis in posterior (C,I), anterior (D,J), dorsal (E,K), ventral (F,L), medial (G,M), and lateral (H,N) views. Abbreviations: hyp, hypoplastron; il, illium; is, ischium; ob, obturator foramen; ps, pubic scar; pu, pubis; xi, xiphiplastron." figureDoi-0="http://doi.org/10.5281/zenodo.8250340" figureDoi-1="http://doi.org/10.5281/zenodo.8250342" figureDoi-2="http://doi.org/10.5281/zenodo.8250344" figureDoi-3="http://doi.org/10.5281/zenodo.8250348" figureDoi-4="http://doi.org/10.5281/zenodo.8250350" figureDoi-5="http://doi.org/10.5281/zenodo.8250352" figureDoi-6="http://doi.org/10.5281/zenodo.8250354" httpUri-0="https://zenodo.org/record/8250340/files/figure.png" httpUri-1="https://zenodo.org/record/8250342/files/figure.png" httpUri-2="https://zenodo.org/record/8250344/files/figure.png" httpUri-3="https://zenodo.org/record/8250348/files/figure.png" httpUri-4="https://zenodo.org/record/8250350/files/figure.png" httpUri-5="https://zenodo.org/record/8250352/files/figure.png" httpUri-6="https://zenodo.org/record/8250354/files/figure.png" pageId="7" pageNumber="8">
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Figs. 3
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<emphasis id="ECB3EAD0966AFFA82830FE2D08C6F5EE" bold="true" box="[889,905,486,505]" italics="true" pageId="7" pageNumber="8">-</emphasis>
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9
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</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="96DD6549966AFFA8287BFDC40E3DF741" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph id="DE7836C2966AFFA8287BFDC40F5DF72D" blockId="7.[818,1488,527,854]" pageId="7" pageNumber="8">
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<emphasis id="ECB3EAD0966AFFA8287BFDC408C5F634" box="[818,906,527,547]" italics="true" pageId="7" pageNumber="8">Material.</emphasis>
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<materialsCitation id="6EAF3C9F966AFFA828C7FDDB0F5DF72D" collectionCode="IWCMS" country="United Kingdom" location="Brook Bay" pageId="7" pageNumber="8" specimenCode="IWCMS 2018.44" specimenCount="1" specimenCount-juvenile="1" stateProvince="Isle of Wight">
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<specimenCode id="8E619EB9966AFFA828C7FDDB0F63F634" box="[910,1068,528,547]" pageId="7" pageNumber="8">IWCMS 2018.44</specimenCode>
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is a partial skeleton of a probably
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<specimenCount id="C8C1FD4B966AFFA82E37FDDB0E80F634" box="[1406,1487,528,547]" pageId="7" pageNumber="8" type="juvenile">juvenile</specimenCount>
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individual (
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<figureCitation id="46FC2A47966AFFA828FAFDE708B8F628" box="[947,1015,556,575]" captionStart="Fig" captionStartId="3.[100,125,1945,1959]" captionTargetBox="[120,1475,391,1914]" captionTargetId="figure-151@3.[110,1478,381,1915]" captionTargetPageId="3" captionText="Fig. 3. IWCMS 2018.44, partial skeleton of Eodortoka cf. morellana. A, dorsal view, B, ventral view, C, anterior view, D, posterior view, E, left lateral view, F, right lateral view. G-H, details of the ornamental pattern on the outer surface of the shell, at the level of the left first costal (G) and of the left hypoplastron (H)." figureDoi="http://doi.org/10.5281/zenodo.8250340" httpUri="https://zenodo.org/record/8250340/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
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), comprising a relatively complete shell (maximum preserved length
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and maximum preserved width
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<quantity id="193F9B27966AFFA8283BFDA80884F660" box="[882,971,611,631]" metricMagnitude="-2" metricUnit="m" metricValue="4.76" pageId="7" pageNumber="8" unit="mm" value="47.6">47.6 mm</quantity>
|
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;
|
|
<figureCitation id="46FC2A47966AFFA8289DFDA80F42F660" box="[980,1037,611,631]" captionStart="Fig" captionStartId="3.[100,125,1945,1959]" captionTargetBox="[120,1475,391,1914]" captionTargetId="figure-151@3.[110,1478,381,1915]" captionTargetPageId="3" captionText="Fig. 3. IWCMS 2018.44, partial skeleton of Eodortoka cf. morellana. A, dorsal view, B, ventral view, C, anterior view, D, posterior view, E, left lateral view, F, right lateral view. G-H, details of the ornamental pattern on the outer surface of the shell, at the level of the left first costal (G) and of the left hypoplastron (H)." figureDoi="http://doi.org/10.5281/zenodo.8250340" httpUri="https://zenodo.org/record/8250340/files/figure.png" pageId="7" pageNumber="8">Figs 3</figureCitation>
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,
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|
<figureCitation id="46FC2A47966AFFA82F5FFDAF0F6CF660" box="[1046,1059,612,631]" captionStart="Fig" captionStartId="4.[138,163,1288,1302]" captionTargetBox="[209,1869,201,1259]" captionTargetId="figure-0@4.[207,1870,197,1259]" captionTargetPageId="4" captionText="Fig. 4. IWCMS 2018.44, partial skeleton of Eodortoka cf. morellana. A,C,E, virtual three-dimensional reconstruction of the left shell plates and of the bones preserved inside the specimen; B,D,F, virtual three-dimensional reconstruction of the bones virtually removing the shell, in dorsal (A,B), ventral (C,D), and left lateral (E,F) views.The anterior and posterior ends of the shell cannot be reconstructed because they were not accurately scanned (black in A, E). The plates of the shell and the bones were reconstructed independently (shell plates in yellow, vertebrae in green, appendicular bones in blue, and girdles in pink). Abbreviations: c, costal; cav, caudal vertebra; cev7, seventh cervical vertebra; cev8, eighth cervical vertebra; dv, dorsal vertebra; hyo, hyoplastron; hyp, hypoplastron; lcor, left coracoid; lhu, left humerus; lil, left illium; lis; left ischium; lsc, left scapula; me, mesoplastron; ne, neural; p, peripheral; rfe, right femur; rfi, right fibula; rhu; right humerus; rpu, right pubis; rsc, right scapula; rti, right tibia; sv, sacral vertebra; xi, xiphiplastron." figureDoi="http://doi.org/10.5281/zenodo.8250342" httpUri="https://zenodo.org/record/8250342/files/figure.png" pageId="7" pageNumber="8">4</figureCitation>
|
|
), whose external surface has been partially eroded, obscuring most boundaries of the scutes; as well as several vertebrae (most cervicals,
|
|
<figureCitation id="46FC2A47966AFFA82F08FD500F36F6B9" box="[1089,1145,667,686]" captionStart="Fig" captionStartId="5.[100,125,1874,1888]" captionTargetBox="[110,1478,543,1844]" captionTargetId="figure-229@5.[110,1478,543,1844]" captionTargetPageId="5" captionText="Fig. 5. Cervical vertebrae of IWCMS 2018.44. A-F, neural arch of the axis (cervical vertebra 2); G-L, cervical vertebra 3; M-R, cervical vertebra 4; S-X, cervical vertebra 5; Y-AD, cervical vertebra 6; AE-AJ, cervical vertebra 7; AK-AP, cervical vertebra 8. All of them are shown in left lateral (A,G,M,S,Y,AE,AK), right lateral (B,H,N,T,Z,AF,AL), anterior (C,I,O,U,AA,AG,AM), posterior (D,J,P,V,AB,AH,AN), dorsal (E,K,Q,W,AC,AI,AO), and ventral (F,L,R,X,AD,AJ,AP) views." figureDoi="http://doi.org/10.5281/zenodo.8250344" httpUri="https://zenodo.org/record/8250344/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
|
|
; some dorsals,
|
|
<figureCitation id="46FC2A47966AFFA82E55FD500E1CF6B9" box="[1308,1363,667,686]" captionStart="Fig" captionStartId="6.[100,125,1870,1884]" captionTargetBox="[199,1389,181,1832]" captionTargetId="figure-1@6.[196,1392,178,1841]" captionTargetPageId="6" captionText="Fig. 6. Dorsal vertebrae of IWCMS 2018.44. A-F, complete dorsal vertebra with fused neural; G-L, dorsal vertebra centrum; M-R, dorsal vertebra centrum; S-X, complete dorsal vertebra; Y-AD,neural arch of a posterior dorsal vertebra. All of them are shown in left lateral (A,G,M,S,Y), right lateral (B,H,N,T,Z), anterior (C,I,O,U,AA), posterior (D,J,P,V,AB), dorsal (E,K,Q,W,AC), and ventral (F,L,R,X,AD) views." figureDoi="http://doi.org/10.5281/zenodo.8250348" httpUri="https://zenodo.org/record/8250348/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
|
|
; and scarce caudals,
|
|
<figureCitation id="46FC2A47966AFFA828C0FD7C088FF6DC" box="[905,960,695,715]" captionStart="Fig" captionStartId="7.[100,125,1932,1946]" captionTargetBox="[110,1478,912,1903]" captionTargetId="figure-363@7.[110,1478,912,1903]" captionTargetPageId="7" captionText="Fig. 7. Sacral and caudal vertebrae of IWCMS 2018.44. A-D, sacral vertebrae; E-J, anterior caudal vertebra; K-L, posterior caudal vertebra; M-P posterior caudal vertebra. These elements are shown in anterior (A,G,K,M), posterior (B,H,L,N), dorsal (C,I,O), ventral (D,J,P), left lateral (E), and right lateral (F) views." figureDoi="http://doi.org/10.5281/zenodo.8250350" httpUri="https://zenodo.org/record/8250350/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
|
|
), partial pectoral and pelvic girdles (
|
|
<figureCitation id="46FC2A47966AFFA82E72FD7C0E32F6DD" box="[1339,1405,695,714]" captionStart="Fig" captionStartId="4.[138,163,1288,1302]" captionTargetBox="[209,1869,201,1259]" captionTargetId="figure-0@4.[207,1870,197,1259]" captionTargetPageId="4" captionText="Fig. 4. IWCMS 2018.44, partial skeleton of Eodortoka cf. morellana. A,C,E, virtual three-dimensional reconstruction of the left shell plates and of the bones preserved inside the specimen; B,D,F, virtual three-dimensional reconstruction of the bones virtually removing the shell, in dorsal (A,B), ventral (C,D), and left lateral (E,F) views.The anterior and posterior ends of the shell cannot be reconstructed because they were not accurately scanned (black in A, E). The plates of the shell and the bones were reconstructed independently (shell plates in yellow, vertebrae in green, appendicular bones in blue, and girdles in pink). Abbreviations: c, costal; cav, caudal vertebra; cev7, seventh cervical vertebra; cev8, eighth cervical vertebra; dv, dorsal vertebra; hyo, hyoplastron; hyp, hypoplastron; lcor, left coracoid; lhu, left humerus; lil, left illium; lis; left ischium; lsc, left scapula; me, mesoplastron; ne, neural; p, peripheral; rfe, right femur; rfi, right fibula; rhu; right humerus; rpu, right pubis; rsc, right scapula; rti, right tibia; sv, sacral vertebra; xi, xiphiplastron." figureDoi="http://doi.org/10.5281/zenodo.8250342" httpUri="https://zenodo.org/record/8250342/files/figure.png" pageId="7" pageNumber="8">Figs. 4</figureCitation>
|
|
,
|
|
<figureCitation id="46FC2A47966AFFA82EC1FD7C0EDAF6DD" box="[1416,1429,695,714]" captionStart="Fig" captionStartId="8.[100,125,1945,1959]" captionTargetBox="[110,1478,1144,1915]" captionTargetId="figure-679@8.[110,1478,1144,1915]" captionTargetPageId="8" captionText="Fig. 8. Elements of the girdles of IWCMS 2018.44. A-F, right scapula; G-L, left scapula; M-R, proximal end of the right coracoid; S-X, left coracoid. These elements are shown in posterior (A,G,M,S), ventral (B,H,O,V), dorsal (C,I,P,T), anterior (D,J,N,U), proximal (E,K,Q,W), and distal (F,L,R,X) views." figureDoi="http://doi.org/10.5281/zenodo.8250352" httpUri="https://zenodo.org/record/8250352/files/figure.png" pageId="7" pageNumber="8">8</figureCitation>
|
|
), and several appendicular bones including both humeri, right femur, fibula and tibia (
|
|
<figureCitation id="46FC2A47966AFFA82890FD240F40F715" box="[985,1039,751,770]" captionStart="Fig" captionStartId="9.[100,125,1814,1828]" captionTargetBox="[110,1478,179,1785]" captionTargetId="figure-0@9.[110,1478,178,1785]" captionTargetPageId="9" captionText="Fig. 9. Visceral view of the left hypoplastron and xiphiplastron with the left pelvis (A), and virtually removing the pelvis (B) to show the pubic scar of IWCMS 2018.44.C-N, Left (C- H) and right (I-N) pelvis in posterior (C,I), anterior (D,J), dorsal (E,K), ventral (F,L), medial (G,M), and lateral (H,N) views. Abbreviations: hyp, hypoplastron; il, illium; is, ischium; ob, obturator foramen; ps, pubic scar; pu, pubis; xi, xiphiplastron." figureDoi="http://doi.org/10.5281/zenodo.8250354" httpUri="https://zenodo.org/record/8250354/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
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|
).
|
|
<emphasis id="ECB3EAD0966AFFA8287BFCC108CDF709" box="[818,898,778,798]" italics="true" pageId="7" pageNumber="8">Locality.</emphasis>
|
|
<location id="DB186019966AFFA828C0FCC008BEF709" LSID="urn:lsid:plazi:treatment:566E87D4966AFFA4287BFE020A6FF6AA:DB186019966AFFA828C0FCC008BEF709" box="[905,1009,779,798]" country="United Kingdom" name="Brook Bay" pageId="7" pageNumber="8" stateProvince="Isle of Wight">Brook Bay</location>
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,
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|
<collectingRegion id="1C03F820966AFFA828B4FCC00FCBF709" box="[1021,1156,779,798]" country="United Kingdom" name="Isle of Wight" pageId="7" pageNumber="8">Isle of Wight</collectingRegion>
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,
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|
<collectingCountry id="A6D07652966AFFA82FD9FCC00E76F709" box="[1168,1337,779,798]" name="United Kingdom" pageId="7" pageNumber="8">United Kingdom</collectingCountry>
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, National Grid Reference SZ 385 836.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="DE7836C2966AFFA8287BFC890E3DF741" blockId="7.[818,1488,527,854]" box="[818,1394,834,854]" pageId="7" pageNumber="8">
|
|
<emphasis id="ECB3EAD0966AFFA8287BFC890830F741" box="[818,895,834,854]" italics="true" pageId="7" pageNumber="8">Horizon</emphasis>
|
|
. Wessex Formation, Early Cretaceous, Barremian.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption id="8AB8664A966AFFA82B2DF8470F37F3A6" ID-DOI="http://doi.org/10.5281/zenodo.8250350" ID-Zenodo-Dep="8250350" httpUri="https://zenodo.org/record/8250350/files/figure.png" pageId="7" pageNumber="8" startId="7.[100,125,1932,1946]" targetBox="[110,1478,912,1903]" targetPageId="7" targetType="figure">
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<paragraph id="DE7836C2966AFFA82B2DF8470F37F3A6" blockId="7.[100,1487,1932,1970]" pageId="7" pageNumber="8">
|
|
<emphasis id="ECB3EAD0966AFFA82B2DF8470BDBF38C" bold="true" box="[100,148,1932,1947]" pageId="7" pageNumber="8">Fig. 7.</emphasis>
|
|
Sacral and caudal vertebrae of IWCMS 2018.44. A-D, sacral vertebrae; E-J, anterior caudal vertebra; K-L, posterior caudal vertebra; M-P posterior caudal vertebra. These elements are shown in anterior (A,G,K,M), posterior (B,H,L,N), dorsal (C,I,O), ventral (D,J,P), left lateral (E), and right lateral (F) views.
|
|
</paragraph>
|
|
</caption>
|
|
<subSubSection id="96DD65499665FFA42B2DFF790A6FF6AA" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9" type="description">
|
|
<paragraph id="DE7836C29665FFA72B2DFF790A4DF4D1" blockId="8.[100,258,178,198]" box="[100,258,178,198]" pageId="8" pageNumber="9">
|
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<heading id="853081AE9665FFA72B2DFF790A4DF4D1" box="[100,258,178,198]" fontSize="8" level="3" pageId="8" pageNumber="9" reason="8">
|
|
<emphasis id="ECB3EAD09665FFA72B2DFF790A4DF4D1" box="[100,258,178,198]" italics="true" pageId="8" pageNumber="9">4.2. Description</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="DE7836C29665FFA72B2DFF210B9FF4E9" blockId="8.[100,771,234,701]" box="[100,208,234,254]" pageId="8" pageNumber="9">
|
|
<heading id="853081AE9665FFA72B2DFF210B9FF4E9" box="[100,208,234,254]" fontSize="8" level="3" pageId="8" pageNumber="9" reason="8">
|
|
<emphasis id="ECB3EAD09665FFA72B2DFF210B9FF4E9" box="[100,208,234,254]" italics="true" pageId="8" pageNumber="9">4.2.1. Shell</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="DE7836C29665FFA72BCDFECD0922F5CA" blockId="8.[100,771,234,701]" pageId="8" pageNumber="9">
|
|
<emphasis id="ECB3EAD09665FFA72BCDFECD0B92F50D" box="[132,221,262,282]" italics="true" pageId="8" pageNumber="9">Carapace</emphasis>
|
|
. The shell of
|
|
<collectionCode id="B8D6AE079665FFA72A31FECC0A88F50D" box="[376,455,263,282]" pageId="8" pageNumber="9">IWCMS</collectionCode>
|
|
.2018.44 has a high degree of erosion, the nuchal plate, most elements of the neural series, most peripherals and the pygal plate are missing, as are some costal plates of the right side in the central third of the carapace (
|
|
<figureCitation id="46FC2A479665FFA729F4FE9109B2F579" box="[701,765,346,366]" captionStart="Fig" captionStartId="3.[100,125,1945,1959]" captionTargetBox="[120,1475,391,1914]" captionTargetId="figure-151@3.[110,1478,381,1915]" captionTargetPageId="3" captionText="Fig. 3. IWCMS 2018.44, partial skeleton of Eodortoka cf. morellana. A, dorsal view, B, ventral view, C, anterior view, D, posterior view, E, left lateral view, F, right lateral view. G-H, details of the ornamental pattern on the outer surface of the shell, at the level of the left first costal (G) and of the left hypoplastron (H)." figureDoi="http://doi.org/10.5281/zenodo.8250340" httpUri="https://zenodo.org/record/8250340/files/figure.png" pageId="8" pageNumber="9">Figs. 3</figureCitation>
|
|
,
|
|
<figureCitation id="46FC2A479665FFA72B2DFEBD0B3EF59E" box="[100,113,374,393]" captionStart="Fig" captionStartId="4.[138,163,1288,1302]" captionTargetBox="[209,1869,201,1259]" captionTargetId="figure-0@4.[207,1870,197,1259]" captionTargetPageId="4" captionText="Fig. 4. IWCMS 2018.44, partial skeleton of Eodortoka cf. morellana. A,C,E, virtual three-dimensional reconstruction of the left shell plates and of the bones preserved inside the specimen; B,D,F, virtual three-dimensional reconstruction of the bones virtually removing the shell, in dorsal (A,B), ventral (C,D), and left lateral (E,F) views.The anterior and posterior ends of the shell cannot be reconstructed because they were not accurately scanned (black in A, E). The plates of the shell and the bones were reconstructed independently (shell plates in yellow, vertebrae in green, appendicular bones in blue, and girdles in pink). Abbreviations: c, costal; cav, caudal vertebra; cev7, seventh cervical vertebra; cev8, eighth cervical vertebra; dv, dorsal vertebra; hyo, hyoplastron; hyp, hypoplastron; lcor, left coracoid; lhu, left humerus; lil, left illium; lis; left ischium; lsc, left scapula; me, mesoplastron; ne, neural; p, peripheral; rfe, right femur; rfi, right fibula; rhu; right humerus; rpu, right pubis; rsc, right scapula; rti, right tibia; sv, sacral vertebra; xi, xiphiplastron." figureDoi="http://doi.org/10.5281/zenodo.8250342" httpUri="https://zenodo.org/record/8250342/files/figure.png" pageId="8" pageNumber="9">4</figureCitation>
|
|
). It is relatively low, and oval. The maximum width is at the level of the mesoplastron. Seven of the left costal plates are identified (all of them except the posterior one). Only the left bridge peripherals (4
|
|
<emphasis id="ECB3EAD09665FFA72B30FE010BC6F5CA" bold="true" box="[121,137,458,477]" italics="true" pageId="8" pageNumber="9">-</emphasis>
|
|
7) are preserved. No fontanelles are recognized.
|
|
</paragraph>
|
|
<paragraph id="DE7836C29665FFA72BCDFE2E0A85F6AB" blockId="8.[100,771,234,701]" pageId="8" pageNumber="9">
|
|
<emphasis id="ECB3EAD09665FFA72BCDFE2E0B9CF5EE" box="[132,211,485,505]" italics="true" pageId="8" pageNumber="9">Plastron</emphasis>
|
|
. The plastron is slightly crushed, seemingly having been depressed into the body cavity. The left side of the plastron is better preserved than the right (
|
|
<figureCitation id="46FC2A479665FFA72A2CFDD50AECF626" box="[357,419,542,561]" captionStart="Fig" captionStartId="3.[100,125,1945,1959]" captionTargetBox="[120,1475,391,1914]" captionTargetId="figure-151@3.[110,1478,381,1915]" captionTargetPageId="3" captionText="Fig. 3. IWCMS 2018.44, partial skeleton of Eodortoka cf. morellana. A, dorsal view, B, ventral view, C, anterior view, D, posterior view, E, left lateral view, F, right lateral view. G-H, details of the ornamental pattern on the outer surface of the shell, at the level of the left first costal (G) and of the left hypoplastron (H)." figureDoi="http://doi.org/10.5281/zenodo.8250340" httpUri="https://zenodo.org/record/8250340/files/figure.png" pageId="8" pageNumber="9">Figs. 3</figureCitation>
|
|
,
|
|
<figureCitation id="46FC2A479665FFA72AE4FDD50AF5F626" box="[429,442,542,561]" captionStart="Fig" captionStartId="4.[138,163,1288,1302]" captionTargetBox="[209,1869,201,1259]" captionTargetId="figure-0@4.[207,1870,197,1259]" captionTargetPageId="4" captionText="Fig. 4. IWCMS 2018.44, partial skeleton of Eodortoka cf. morellana. A,C,E, virtual three-dimensional reconstruction of the left shell plates and of the bones preserved inside the specimen; B,D,F, virtual three-dimensional reconstruction of the bones virtually removing the shell, in dorsal (A,B), ventral (C,D), and left lateral (E,F) views.The anterior and posterior ends of the shell cannot be reconstructed because they were not accurately scanned (black in A, E). The plates of the shell and the bones were reconstructed independently (shell plates in yellow, vertebrae in green, appendicular bones in blue, and girdles in pink). Abbreviations: c, costal; cav, caudal vertebra; cev7, seventh cervical vertebra; cev8, eighth cervical vertebra; dv, dorsal vertebra; hyo, hyoplastron; hyp, hypoplastron; lcor, left coracoid; lhu, left humerus; lil, left illium; lis; left ischium; lsc, left scapula; me, mesoplastron; ne, neural; p, peripheral; rfe, right femur; rfi, right fibula; rhu; right humerus; rpu, right pubis; rsc, right scapula; rti, right tibia; sv, sacral vertebra; xi, xiphiplastron." figureDoi="http://doi.org/10.5281/zenodo.8250342" httpUri="https://zenodo.org/record/8250342/files/figure.png" pageId="8" pageNumber="9">4</figureCitation>
|
|
). No remains of the epiplastra or entoplastron are preserved. The hyoplastron is almost fully preserved, except for its anterior margin. The specimen displays a laterally located pair of mesoplastra, being slightly wider than long. The anterior end of the xiphiplastron is present. At its thickest point, the plastron is
|
|
<quantity id="193F9B279665FFA72A74FD620AC5F6AA" box="[317,394,681,701]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="9" unit="mm" value="2.3">2.3 mm</quantity>
|
|
thick.
|
|
</paragraph>
|
|
<paragraph id="DE7836C29665FFA72B2DFD270A03F717" blockId="8.[100,771,748,1104]" box="[100,332,748,768]" pageId="8" pageNumber="9">
|
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<heading id="853081AE9665FFA72B2DFD270A03F717" box="[100,332,748,768]" fontSize="8" level="3" pageId="8" pageNumber="9" reason="8">
|
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<emphasis id="ECB3EAD09665FFA72B2DFD270A03F717" box="[100,332,748,768]" italics="true" pageId="8" pageNumber="9">4.2.2. Vertebral column</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="DE7836C29665FFA72BCDFCC30FA0F4E9" blockId="8.[100,771,748,1104]" lastBlockId="8.[818,1487,179,784]" pageId="8" pageNumber="9">
|
|
<emphasis id="ECB3EAD09665FFA72BCDFCC30A66F70B" box="[132,297,776,796]" italics="true" pageId="8" pageNumber="9">Cervical vertebrae</emphasis>
|
|
. There are seven cervical vertebrae preserved inside the shell of
|
|
<collectionCode id="B8D6AE079665FFA72A5AFCEE0A14F72F" box="[275,347,805,824]" pageId="8" pageNumber="9">IWCMS</collectionCode>
|
|
2018.44. 6 with a fused neural arch and centrum and one isolated neural arch of the axis (
|
|
<figureCitation id="46FC2A479665FFA7297FFC8A0929F743" box="[566,614,833,852]" captionStart="Fig" captionStartId="5.[100,125,1874,1888]" captionTargetBox="[110,1478,543,1844]" captionTargetId="figure-229@5.[110,1478,543,1844]" captionTargetPageId="5" captionText="Fig. 5. Cervical vertebrae of IWCMS 2018.44. A-F, neural arch of the axis (cervical vertebra 2); G-L, cervical vertebra 3; M-R, cervical vertebra 4; S-X, cervical vertebra 5; Y-AD, cervical vertebra 6; AE-AJ, cervical vertebra 7; AK-AP, cervical vertebra 8. All of them are shown in left lateral (A,G,M,S,Y,AE,AK), right lateral (B,H,N,T,Z,AF,AL), anterior (C,I,O,U,AA,AG,AM), posterior (D,J,P,V,AB,AH,AN), dorsal (E,K,Q,W,AC,AI,AO), and ventral (F,L,R,X,AD,AJ,AP) views." figureDoi="http://doi.org/10.5281/zenodo.8250344" httpUri="https://zenodo.org/record/8250344/files/figure.png" pageId="8" pageNumber="9">Fig.5</figureCitation>
|
|
A-AP).These are not in articulation within the carapace. The centra are ventrally keeled with the keel becoming less prominent through the series, and almost non-existent in cervical 8 (
|
|
<figureCitation id="46FC2A479665FFA72A28FC5E0ADCF7BF" box="[353,403,917,936]" captionStart="Fig" captionStartId="5.[100,125,1874,1888]" captionTargetBox="[110,1478,543,1844]" captionTargetId="figure-229@5.[110,1478,543,1844]" captionTargetPageId="5" captionText="Fig. 5. Cervical vertebrae of IWCMS 2018.44. A-F, neural arch of the axis (cervical vertebra 2); G-L, cervical vertebra 3; M-R, cervical vertebra 4; S-X, cervical vertebra 5; Y-AD, cervical vertebra 6; AE-AJ, cervical vertebra 7; AK-AP, cervical vertebra 8. All of them are shown in left lateral (A,G,M,S,Y,AE,AK), right lateral (B,H,N,T,Z,AF,AL), anterior (C,I,O,U,AA,AG,AM), posterior (D,J,P,V,AB,AH,AN), dorsal (E,K,Q,W,AC,AI,AO), and ventral (F,L,R,X,AD,AJ,AP) views." figureDoi="http://doi.org/10.5281/zenodo.8250344" httpUri="https://zenodo.org/record/8250344/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
|
|
). Centra 3
|
|
<emphasis id="ECB3EAD09665FFA72ABAFC5E094CF7BF" bold="true" box="[499,515,917,936]" italics="true" pageId="8" pageNumber="9">-</emphasis>
|
|
5 are opisthocoelus, centra 6
|
|
<emphasis id="ECB3EAD09665FFA72B39FC7A0BCFF7D3" bold="true" box="[112,128,945,964]" italics="true" pageId="8" pageNumber="9">-</emphasis>
|
|
7 are amphicoelous and centra 8 is procoelus (
|
|
<figureCitation id="46FC2A479665FFA7290CFC7A0937F7D3" box="[581,632,945,964]" captionStart="Fig" captionStartId="5.[100,125,1874,1888]" captionTargetBox="[110,1478,543,1844]" captionTargetId="figure-229@5.[110,1478,543,1844]" captionTargetPageId="5" captionText="Fig. 5. Cervical vertebrae of IWCMS 2018.44. A-F, neural arch of the axis (cervical vertebra 2); G-L, cervical vertebra 3; M-R, cervical vertebra 4; S-X, cervical vertebra 5; Y-AD, cervical vertebra 6; AE-AJ, cervical vertebra 7; AK-AP, cervical vertebra 8. All of them are shown in left lateral (A,G,M,S,Y,AE,AK), right lateral (B,H,N,T,Z,AF,AL), anterior (C,I,O,U,AA,AG,AM), posterior (D,J,P,V,AB,AH,AN), dorsal (E,K,Q,W,AC,AI,AO), and ventral (F,L,R,X,AD,AJ,AP) views." figureDoi="http://doi.org/10.5281/zenodo.8250344" httpUri="https://zenodo.org/record/8250344/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
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|
). The anterior end of the centra have an oval cross section, becoming more triangular through the series. Posterior end of the centum have a square cross section to triangular through the series. All vertebrae have a large circular neural canal, increasing in diameter through the vertebral series. From vertebrae 5
|
|
<emphasis id="ECB3EAD09665FFA72A44FBF70A52F058" bold="true" box="[269,285,1084,1103]" italics="true" pageId="8" pageNumber="9">-</emphasis>
|
|
8, the neural canal is larger in diameter than the diameter of the centrum (
|
|
<figureCitation id="46FC2A479665FFA72F6FFF780F18F4D1" box="[1062,1111,179,198]" captionStart="Fig" captionStartId="5.[100,125,1874,1888]" captionTargetBox="[110,1478,543,1844]" captionTargetId="figure-229@5.[110,1478,543,1844]" captionTargetPageId="5" captionText="Fig. 5. Cervical vertebrae of IWCMS 2018.44. A-F, neural arch of the axis (cervical vertebra 2); G-L, cervical vertebra 3; M-R, cervical vertebra 4; S-X, cervical vertebra 5; Y-AD, cervical vertebra 6; AE-AJ, cervical vertebra 7; AK-AP, cervical vertebra 8. All of them are shown in left lateral (A,G,M,S,Y,AE,AK), right lateral (B,H,N,T,Z,AF,AL), anterior (C,I,O,U,AA,AG,AM), posterior (D,J,P,V,AB,AH,AN), dorsal (E,K,Q,W,AC,AI,AO), and ventral (F,L,R,X,AD,AJ,AP) views." figureDoi="http://doi.org/10.5281/zenodo.8250344" httpUri="https://zenodo.org/record/8250344/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
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). The diapophyses are anteriorly placed in vertebrae 2
|
|
<emphasis id="ECB3EAD09665FFA728FEFF040888F4F5" bold="true" box="[951,967,207,226]" italics="true" pageId="8" pageNumber="9">-</emphasis>
|
|
4 and become more posteriorly placed through 5 to 8. Vertebrae 8 has a small, rounded neural spine.
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</paragraph>
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<paragraph id="DE7836C29665FFA72818FECD0E1AF602" blockId="8.[818,1487,179,784]" pageId="8" pageNumber="9">
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<emphasis id="ECB3EAD09665FFA72818FECD0F43F50D" box="[849,1036,262,282]" italics="true" pageId="8" pageNumber="9">Thoracic vertebrae</emphasis>
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. Five thoracic vertebrae are preserved, two with a fused neural arch (a neural plate being also preserved as a fused element to one of them), two isolated centra, and an isolated neural arch (
|
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<figureCitation id="46FC2A479665FFA72F52FE910F18F57A" box="[1051,1111,346,365]" captionStart="Fig" captionStartId="6.[100,125,1870,1884]" captionTargetBox="[199,1389,181,1832]" captionTargetId="figure-1@6.[196,1392,178,1841]" captionTargetPageId="6" captionText="Fig. 6. Dorsal vertebrae of IWCMS 2018.44. A-F, complete dorsal vertebra with fused neural; G-L, dorsal vertebra centrum; M-R, dorsal vertebra centrum; S-X, complete dorsal vertebra; Y-AD,neural arch of a posterior dorsal vertebra. All of them are shown in left lateral (A,G,M,S,Y), right lateral (B,H,N,T,Z), anterior (C,I,O,U,AA), posterior (D,J,P,V,AB), dorsal (E,K,Q,W,AC), and ventral (F,L,R,X,AD) views." figureDoi="http://doi.org/10.5281/zenodo.8250348" httpUri="https://zenodo.org/record/8250348/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
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A-AD). The centra are shallow and are laterally pinched. This is more evident mid series, with the centra becoming wider posteriorly. They possess a large neural canal, at least twice the depth of the centrum. The neural spines are tall and elongate, extending over the posterior and anterior portion of the centrum anteriorly in the series, and only posteriorly towards the posterior portion of the series.
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</paragraph>
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<paragraph id="DE7836C29665FFA72818FDD60830F693" blockId="8.[818,1487,179,784]" pageId="8" pageNumber="9">
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<emphasis id="ECB3EAD09665FFA72818FDD608A4F626" box="[849,1003,541,561]" italics="true" pageId="8" pageNumber="9">Sacral vertebrae</emphasis>
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. A portion of sacrum is preserved, with two right lateral processes (
|
|
<figureCitation id="46FC2A479665FFA728A4FDF10F69F65A" box="[1005,1062,570,589]" captionStart="Fig" captionStartId="7.[100,125,1932,1946]" captionTargetBox="[110,1478,912,1903]" captionTargetId="figure-363@7.[110,1478,912,1903]" captionTargetPageId="7" captionText="Fig. 7. Sacral and caudal vertebrae of IWCMS 2018.44. A-D, sacral vertebrae; E-J, anterior caudal vertebra; K-L, posterior caudal vertebra; M-P posterior caudal vertebra. These elements are shown in anterior (A,G,K,M), posterior (B,H,L,N), dorsal (C,I,O), ventral (D,J,P), left lateral (E), and right lateral (F) views." figureDoi="http://doi.org/10.5281/zenodo.8250350" httpUri="https://zenodo.org/record/8250350/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
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A-D). A small portion of centrum is preserved posteriorly. The neural arch is wide, with a
|
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<collectionCode id="B8D6AE079665FFA72E7AFD9D0E70F67E" box="[1331,1343,598,617]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="9">T</collectionCode>
|
|
-shaped cross section.
|
|
</paragraph>
|
|
<paragraph id="DE7836C29665FFA72818FD470E25F707" blockId="8.[818,1487,179,784]" pageId="8" pageNumber="9">
|
|
<emphasis id="ECB3EAD09665FFA72818FD4708B5F6B7" box="[849,1018,652,672]" italics="true" pageId="8" pageNumber="9">Caudal vertebrae</emphasis>
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. Three caudal vertebrae are preserved, with fused neural arches (
|
|
<figureCitation id="46FC2A479665FFA72F58FD620F04F6AA" box="[1041,1099,681,701]" captionStart="Fig" captionStartId="7.[100,125,1932,1946]" captionTargetBox="[110,1478,912,1903]" captionTargetId="figure-363@7.[110,1478,912,1903]" captionTargetPageId="7" captionText="Fig. 7. Sacral and caudal vertebrae of IWCMS 2018.44. A-D, sacral vertebrae; E-J, anterior caudal vertebra; K-L, posterior caudal vertebra; M-P posterior caudal vertebra. These elements are shown in anterior (A,G,K,M), posterior (B,H,L,N), dorsal (C,I,O), ventral (D,J,P), left lateral (E), and right lateral (F) views." figureDoi="http://doi.org/10.5281/zenodo.8250350" httpUri="https://zenodo.org/record/8250350/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
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E-P). The most anterior caudal has a complete centrum and neural arch preserved. It lacks a neural spine but possesses large pre- and postzygopophyses. The centrum is oval in cross section. Posterior caudals have a short centrum.
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</paragraph>
|
|
<paragraph id="DE7836C29665FFA7287BFC8B0F06F743" blockId="8.[818,1488,832,1104]" box="[818,1097,832,852]" pageId="8" pageNumber="9">
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|
<heading id="853081AE9665FFA7287BFC8B0F06F743" box="[818,1097,832,852]" fontSize="8" level="3" pageId="8" pageNumber="9" reason="8">
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<emphasis id="ECB3EAD09665FFA7287BFC8B0F06F743" box="[818,1097,832,852]" italics="true" pageId="8" pageNumber="9">4.2.3. Appendicular skeleton</emphasis>
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</heading>
|
|
</paragraph>
|
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<paragraph id="DE7836C29665FFA72818FC970E8FF7F7" blockId="8.[818,1488,832,1104]" pageId="8" pageNumber="9">
|
|
<emphasis id="ECB3EAD09665FFA72818FC9708AFF767" box="[849,992,860,880]" italics="true" pageId="8" pageNumber="9">Shoulder girdle</emphasis>
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|
. Both scapulae are preserved (
|
|
<figureCitation id="46FC2A479665FFA72E44FC960E0EF767" box="[1293,1345,861,880]" captionStart="Fig" captionStartId="8.[100,125,1945,1959]" captionTargetBox="[110,1478,1144,1915]" captionTargetId="figure-679@8.[110,1478,1144,1915]" captionTargetPageId="8" captionText="Fig. 8. Elements of the girdles of IWCMS 2018.44. A-F, right scapula; G-L, left scapula; M-R, proximal end of the right coracoid; S-X, left coracoid. These elements are shown in posterior (A,G,M,S), ventral (B,H,O,V), dorsal (C,I,P,T), anterior (D,J,N,U), proximal (E,K,Q,W), and distal (F,L,R,X) views." figureDoi="http://doi.org/10.5281/zenodo.8250352" httpUri="https://zenodo.org/record/8250352/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
|
|
A-L). The right one is complete. The acromion process is slightly oval in cross section. Both coracoids are preserved (
|
|
<figureCitation id="46FC2A479665FFA72F2BFC5E0FD8F7BF" box="[1122,1175,917,936]" captionStart="Fig" captionStartId="8.[100,125,1945,1959]" captionTargetBox="[110,1478,1144,1915]" captionTargetId="figure-679@8.[110,1478,1144,1915]" captionTargetPageId="8" captionText="Fig. 8. Elements of the girdles of IWCMS 2018.44. A-F, right scapula; G-L, left scapula; M-R, proximal end of the right coracoid; S-X, left coracoid. These elements are shown in posterior (A,G,M,S), ventral (B,H,O,V), dorsal (C,I,P,T), anterior (D,J,N,U), proximal (E,K,Q,W), and distal (F,L,R,X) views." figureDoi="http://doi.org/10.5281/zenodo.8250352" httpUri="https://zenodo.org/record/8250352/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
|
|
M-X). The left coracoid is complete, the right coracoid is only presented by the proximal end. The coracoid is dorsoventrally thin and considerably expanded distally.
|
|
</paragraph>
|
|
<paragraph id="DE7836C29665FFA42818FC230A6AF50D" blockId="8.[818,1488,832,1104]" lastBlockId="11.[100,770,179,701]" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
|
|
<emphasis id="ECB3EAD09665FFA72818FC230888F7EB" box="[849,967,1000,1020]" italics="true" pageId="8" pageNumber="9">Pelvic girdle</emphasis>
|
|
. The pelvic girdle is almost complete, preserved in their natural anatomical position (
|
|
<figureCitation id="46FC2A479665FFA72FC5FBCF0F8DF00F" box="[1164,1218,1028,1048]" captionStart="Fig" captionStartId="9.[100,125,1814,1828]" captionTargetBox="[110,1478,179,1785]" captionTargetId="figure-0@9.[110,1478,178,1785]" captionTargetPageId="9" captionText="Fig. 9. Visceral view of the left hypoplastron and xiphiplastron with the left pelvis (A), and virtually removing the pelvis (B) to show the pubic scar of IWCMS 2018.44.C-N, Left (C- H) and right (I-N) pelvis in posterior (C,I), anterior (D,J), dorsal (E,K), ventral (F,L), medial (G,M), and lateral (H,N) views. Abbreviations: hyp, hypoplastron; il, illium; is, ischium; ob, obturator foramen; ps, pubic scar; pu, pubis; xi, xiphiplastron." figureDoi="http://doi.org/10.5281/zenodo.8250354" httpUri="https://zenodo.org/record/8250354/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
|
|
). The sutures between the ilium, ischium and pubis are visible, and the pubic scar is visible on the xiphiplastron. The ilium is fused to the carapace, and the ischium and pubis are fused to the plastron. The ilium is robust and mediolaterally narrow. The ischium is rod-like, with a distally expanded head where it contacts the ilium. The pubis has a large obturator foramen.
|
|
</paragraph>
|
|
<caption id="8AB8664A9665FFA72B2DF85208B3F3A9" ID-DOI="http://doi.org/10.5281/zenodo.8250352" ID-Zenodo-Dep="8250352" httpUri="https://zenodo.org/record/8250352/files/figure.png" pageId="8" pageNumber="9" startId="8.[100,125,1945,1959]" targetBox="[110,1478,1144,1915]" targetPageId="8" targetType="figure">
|
|
<paragraph id="DE7836C29665FFA72B2DF85208B3F3A9" blockId="8.[100,1487,1945,1982]" pageId="8" pageNumber="9">
|
|
<emphasis id="ECB3EAD09665FFA72B2DF8520BDAF3B0" bold="true" box="[100,149,1945,1959]" pageId="8" pageNumber="9">Fig. 8.</emphasis>
|
|
Elements of the girdles of IWCMS 2018.44. A-F, right scapula; G-L, left scapula; M-R, proximal end of the right coracoid; S-X, left coracoid. These elements are shown in posterior (A,G,M,S), ventral (B,H,O,V), dorsal (C,I,P,T), anterior (D,J,N,U), proximal (E,K,Q,W), and distal (F,L,R,X) views.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="8AB8664A9664FFA62B2DF8DD0900F345" ID-DOI="http://doi.org/10.5281/zenodo.8250354" ID-Zenodo-Dep="8250354" httpUri="https://zenodo.org/record/8250354/files/figure.png" pageId="9" pageNumber="10" startId="9.[100,125,1814,1828]" targetBox="[110,1478,179,1785]" targetPageId="9" targetType="figure">
|
|
<paragraph id="DE7836C29664FFA62B2DF8DD0900F345" blockId="9.[100,1488,1814,1875]" pageId="9" pageNumber="10">
|
|
<emphasis id="ECB3EAD09664FFA62B2DF8DD0BDCF332" bold="true" box="[100,147,1814,1829]" pageId="9" pageNumber="10">Fig. 9.</emphasis>
|
|
Visceral view of the left hypoplastron and xiphiplastron with the left pelvis (A), and virtually removing the pelvis (B) to show the pubic scar of IWCMS 2018.44. C-N, Left (C- H) and right (I-N) pelvis in posterior (C,I), anterior (D,J), dorsal (E,K), ventral (F,L), medial (G,M), and lateral (H,N) views. Abbreviations: hyp, hypoplastron; il, illium; is, ischium; ob, obturator foramen; ps, pubic scar; pu, pubis; xi, xiphiplastron.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="8AB8664A9667FFA52B2DF8850E7DF364" ID-DOI="http://doi.org/10.5281/zenodo.8250356" ID-Zenodo-Dep="8250356" httpUri="https://zenodo.org/record/8250356/files/figure.png" pageId="10" pageNumber="11" startId="10.[100,125,1870,1884]" targetBox="[216,1371,178,1841]" targetPageId="10" targetType="figure">
|
|
<paragraph id="DE7836C29667FFA52B2DF8850E7DF364" blockId="10.[100,1487,1870,1907]" pageId="10" pageNumber="11">
|
|
<emphasis id="ECB3EAD09667FFA52B2DF8850BD4F34B" bold="true" box="[100,155,1870,1884]" pageId="10" pageNumber="11">Fig. 10.</emphasis>
|
|
Appendicular elements of IWCMS 2018.44. A-F, Right humerus; G-L, proximal end of the left humerus; M-R, right femur; S-X, proximal end of the right fibula; Y-AD, distal end of the right tibia; in ventral (A,G,M,S,Y), posterior (B,H,N,T,Z), dorsal (C,I,O,U,AA), anterior (D,J,P,V,AB), medial (E,K,Q,W,AC), and distal (F,L,R,X,AD) views.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="DE7836C29666FFA42BCDFEE90BB3F5B1" blockId="11.[100,770,179,701]" pageId="11" pageNumber="12">
|
|
<emphasis id="ECB3EAD09666FFA42BCDFEE90BA8F521" box="[132,231,290,310]" italics="true" pageId="11" pageNumber="12">Forelimbs.</emphasis>
|
|
Only the complete right humerus and part of the left are preserved (
|
|
<figureCitation id="46FC2A479666FFA42BB4FEF50A73F545" box="[253,316,318,338]" captionStart="Fig" captionStartId="10.[100,125,1870,1884]" captionTargetBox="[216,1371,178,1841]" captionTargetId="figure-1@10.[216,1371,178,1841]" captionTargetPageId="10" captionText="Fig.10. Appendicular elements of IWCMS 2018.44. A-F, Right humerus; G-L, proximal end of the left humerus; M-R, right femur; S-X, proximal end of the right fibula; Y-AD, distal end of the right tibia; in ventral (A,G,M,S,Y), posterior (B,H,N,T,Z), dorsal (C,I,O,U,AA), anterior (D,J,P,V,AB), medial (E,K,Q,W,AC), and distal (F,L,R,X,AD) views." figureDoi="http://doi.org/10.5281/zenodo.8250356" httpUri="https://zenodo.org/record/8250356/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
|
|
A-R). The diaphysis has a mediolateral curve, with an oval articular head. The distal end is wide, with a deep intertubercular fossa between a large medial process and a large lateral process.
|
|
</paragraph>
|
|
<paragraph id="DE7836C29666FFA42BCDFE660A6FF6AA" blockId="11.[100,770,179,701]" pageId="11" pageNumber="12">
|
|
<emphasis id="ECB3EAD09666FFA42BCDFE660BBEF5D6" box="[132,241,429,449]" italics="true" pageId="11" pageNumber="12">Hindlimbs.</emphasis>
|
|
The partial right femur, tibia and fibula are preserved. The right femur has its proximal end missing (
|
|
<figureCitation id="46FC2A479666FFA429E8FE0109ACF5CA" box="[673,739,458,477]" captionStart="Fig" captionStartId="10.[100,125,1870,1884]" captionTargetBox="[216,1371,178,1841]" captionTargetId="figure-1@10.[216,1371,178,1841]" captionTargetPageId="10" captionText="Fig.10. Appendicular elements of IWCMS 2018.44. A-F, Right humerus; G-L, proximal end of the left humerus; M-R, right femur; S-X, proximal end of the right fibula; Y-AD, distal end of the right tibia; in ventral (A,G,M,S,Y), posterior (B,H,N,T,Z), dorsal (C,I,O,U,AA), anterior (D,J,P,V,AB), medial (E,K,Q,W,AC), and distal (F,L,R,X,AD) views." figureDoi="http://doi.org/10.5281/zenodo.8250356" httpUri="https://zenodo.org/record/8250356/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
|
|
<collectionCode id="B8D6AE079666FFA42B2DFE2D0B39F5EE" box="[100,118,486,505]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="12">M-R</collectionCode>
|
|
). The diaphysis is laterally curved and is distally expanded forming a large tibial condyle. Only distal portion of the right tibia is preserved (
|
|
<figureCitation id="46FC2A479666FFA42A70FDD50A30F626" box="[313,383,542,561]" captionStart="Fig" captionStartId="10.[100,125,1870,1884]" captionTargetBox="[216,1371,178,1841]" captionTargetId="figure-1@10.[216,1371,178,1841]" captionTargetPageId="10" captionText="Fig.10. Appendicular elements of IWCMS 2018.44. A-F, Right humerus; G-L, proximal end of the left humerus; M-R, right femur; S-X, proximal end of the right fibula; Y-AD, distal end of the right tibia; in ventral (A,G,M,S,Y), posterior (B,H,N,T,Z), dorsal (C,I,O,U,AA), anterior (D,J,P,V,AB), medial (E,K,Q,W,AC), and distal (F,L,R,X,AD) views." figureDoi="http://doi.org/10.5281/zenodo.8250356" httpUri="https://zenodo.org/record/8250356/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
|
|
Y-AD). The shaft has an oval cross section. The distal end is expanded with a broad articular surface. Only the proximal end of the right fibula is preserved (
|
|
<figureCitation id="46FC2A479666FFA42B24FDBA0BFDF692" box="[109,178,625,645]" captionStart="Fig" captionStartId="10.[100,125,1870,1884]" captionTargetBox="[216,1371,178,1841]" captionTargetId="figure-1@10.[216,1371,178,1841]" captionTargetPageId="10" captionText="Fig.10. Appendicular elements of IWCMS 2018.44. A-F, Right humerus; G-L, proximal end of the left humerus; M-R, right femur; S-X, proximal end of the right fibula; Y-AD, distal end of the right tibia; in ventral (A,G,M,S,Y), posterior (B,H,N,T,Z), dorsal (C,I,O,U,AA), anterior (D,J,P,V,AB), medial (E,K,Q,W,AC), and distal (F,L,R,X,AD) views." figureDoi="http://doi.org/10.5281/zenodo.8250356" httpUri="https://zenodo.org/record/8250356/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
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S-X). It has a proximally expanded head with an oval cross section. The medial surface is straight, and the lateral surface is convex.
|
|
</paragraph>
|
|
</subSubSection>
|
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</treatment>
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</document> |