143 lines
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143 lines
16 KiB
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<document id="6896066E83A40A3D837089B629C7C2FB" ID-CLB-Dataset="30082" ID-DOI="http://dx.doi.org/10.3897/zookeys.758.24169" ID-GBIF-Dataset="5ae148eb-ed49-4509-8505-cf8c44aa04c8" ID-PMC="PMC5962634" ID-Pensoft-Pub="1313-2970-758-19" ID-PubMed="29844711" ID-ZooBank="A6F00F77979240CFBBD9D52E933B9E57" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-758-19" ModsDocOrigin="ZooKeys 758" ModsDocTitle="Revision of genus Pericalus from China, with descriptions of four new species (Carabidae, Lebiini, Pericalina)" checkinTime="1526381266857" checkinUser="pensoft" docAuthor="Shi, Hongliang & Liang, Hongbin" docDate="2018" docId="3BB07C5874352049FCCB98ABEFC57EEE" docLanguage="en" docName="ZooKeys 758: 19-54" docOrigin="ZooKeys 758" docSource="http://dx.doi.org/10.3897/zookeys.758.24169" docTitle="Pericalus ornatus subsp. formosanus Dupuis 1913" docType="treatment" docVersion="7" lastPageNumber="36" masterDocId="FFEEFF94FFF68844FF8EFF85FFA4E14C" masterDocTitle="Revision of genus Pericalus from China, with descriptions of four new species (Carabidae, Lebiini, Pericalina)" masterLastPageNumber="54" masterPageNumber="19" pageNumber="36" updateTime="1701379077708" updateUser="ExternalLinkService">
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<mods:mods id="9306EE46F441BB11C24DB9024A2EA603" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DB18B40A5A7B9CA4FA7B9BC72D4EEC83">
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<mods:title id="61790707A97551D01E44655E606C66B3">Revision of genus Pericalus from China, with descriptions of four new species (Carabidae, Lebiini, Pericalina)</mods:title>
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<mods:name id="11CF19ABF98273873F60AB53EA7D2343" type="personal">
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<mods:namePart id="831133F07CBE249DA0E7AD2F6B6CEDAC">Shi, Hongliang</mods:namePart>
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<mods:roleTerm id="CA53F2C7F2EC2AF817485AB4F57F5565">Author</mods:roleTerm>
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<mods:namePart id="133BF84FBA7F699FAD341D78B766B2BA">Liang, Hongbin</mods:namePart>
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<mods:typeOfResource id="E74BB3994622A122517C74693153205F">text</mods:typeOfResource>
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<mods:title id="CC5EA8644B0D925843644A36D1999CB1">ZooKeys</mods:title>
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<mods:part id="BA95015C27E3BAE030A698D67CF861D1">
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<mods:date id="FAEDE9CF1C48C01CB84F601FC3FA86D6">2018</mods:date>
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<mods:detail id="B35BC8E7CEB1DE658FD2D9A2F4767D22" type="volume">
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<mods:number id="3B95345A2758A191E490B3F458E69722">758</mods:number>
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<mods:extent id="891F48921BF6A34276E69B4594CD4F08" unit="page">
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<mods:start id="593F8118DE9218A3FCA61AFE7AAAA47E">19</mods:start>
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<mods:end id="93F2716EEAE091E9874BB336CD167F3E">54</mods:end>
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<mods:location id="D766BE0C9BF47F372284B0322EEE09D6">
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<mods:url id="D7A25C24AA4D27BD1FD92D5960AF650B">http://dx.doi.org/10.3897/zookeys.758.24169</mods:url>
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<mods:classification id="42536E4D3079C12C8C83B049CB60E997">journal article</mods:classification>
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<mods:identifier id="22E70B579AD13EDB1ECEF3EF43B26B35" type="DOI">http://dx.doi.org/10.3897/zookeys.758.24169</mods:identifier>
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<mods:identifier id="5770DBD735154134808ECED1E2399548" type="Pensoft-Pub">1313-2970-758-19</mods:identifier>
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<mods:identifier id="8306C231DEE347423C7C9AFCC7D3867B" type="ZooBank">A6F00F77979240CFBBD9D52E933B9E57</mods:identifier>
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</mods:mods>
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<treatment id="3BB07C5874352049FCCB98ABEFC57EEE" ID-GBIF-Taxon="143970616" LSID="urn:lsid:plazi:treatment:3BB07C5874352049FCCB98ABEFC57EEE" httpUri="http://treatment.plazi.org/id/3BB07C5874352049FCCB98ABEFC57EEE" lastPageNumber="36" pageId="17" pageNumber="36">
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<subSubSection id="EAA0A32982ED4AC0CE085AA46103F92E" pageId="17" pageNumber="36" type="nomenclature">
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<paragraph id="E13E390E821954260FD89A537CC079D4" pageId="17" pageNumber="36">
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<taxonomicName id="D7D082BD63CE69C26EB121DB15413CFC" ID-CoL="76T3W" authority="Dupuis, 1913" authorityName="Dupuis" authorityYear="1913" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus ornatus" order="Coleoptera" pageId="17" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="ornatus" subSpecies="formosanus">Pericalus (s. str.) ornatus formosanus Dupuis, 1913</taxonomicName>
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Figs 12, 25, 26, 39, 46, 57, 59
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</paragraph>
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</subSubSection>
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<subSubSection id="7AE02DCFC99F9CF3C16B7C860BE4A041" pageId="17" pageNumber="36" type="reference_group">
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<paragraph id="CA4487A95CE3A70DBFC80B1F3004328B" pageId="17" pageNumber="36">
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<bibRefCitation id="8AF5F389405C6EA4C028346CA791ED41" author="Dupuis, P" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="24" pageNumber="43" pagination="81 - 87" title="H. Sauter's Formosa-Ausbeute. (3 e contribution). Carabidae." volume="57" year="1913">Dupuis 1913</bibRefCitation>
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: 83 (type locality: Taiwan [Moozan, Sokutsu]; syntypes in IRSN);
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<bibRefCitation id="1CBA919D6EBEEA1F403BF983EE23BBD9" author="Jedlicka, A" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="24" pageNumber="43" pagination="269 - 579" title="Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleoptera, Carabidae)." volume="28" year="1963">
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<normalizedToken id="C128AB09FAC1BF0A78C18FD4146936B2" originalValue="Jedlička">Jedlicka</normalizedToken>
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1963
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</bibRefCitation>
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: 379;
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<bibRefCitation id="22446E094ACE3B1C2432A24063A65537" pageId="17" pageNumber="36">Kabak 2003</bibRefCitation>
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: 437 (catalogue).
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</paragraph>
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</subSubSection>
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<subSubSection id="843D631B8961B37B65BA99BE6F6E5D1B" pageId="17" pageNumber="36" type="type localities">
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<paragraph id="987C3647DDC47D78C7A79A14CE2E051B" pageId="17" pageNumber="36">Type localities.</paragraph>
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<paragraph id="BE6128E3C982AFBDFF3D80568BD21619" pageId="17" pageNumber="36">
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<normalizedToken id="3DCEBCD16ECEAEB9534A3FE56248A4F1" originalValue="“Moozan”">"Moozan"</normalizedToken>
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is a misspelling of Hoozan (sometimes also Hozan), referring to Fengshan (
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<geoCoordinate id="E76981FF050F338CB8DB9606105585EE" direction="north" orientation="latitude" precision="555" value="22.61">N22.61</geoCoordinate>
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,
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<geoCoordinate id="BD596B1004F56C26BEED2D795658EA0B" direction="east" orientation="longitude" precision="555" value="120.35">E120.35</geoCoordinate>
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) in Kaohsiung county, south Taiwan. Sokutsu refers to Hsiaolin (
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<geoCoordinate id="26B874E3C2A51B4466C200BC782F2F2A" direction="north" orientation="latitude" precision="555" value="23.16">N23.16</geoCoordinate>
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,
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<geoCoordinate id="B71687F62E4A1C1CAEE9478644699CFD" direction="east" orientation="longitude" precision="555" value="120.64">E120.64</geoCoordinate>
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) in Kaohsiung county.
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</paragraph>
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</subSubSection>
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<subSubSection id="299D5A2F0C836FCC5620D5B0C9A2AECF" pageId="17" pageNumber="36" type="materials_examined">
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<paragraph id="A225D6AF7C7EF2AD5EB2B115C16AAB25" pageId="17" pageNumber="36">Material examined</paragraph>
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<paragraph id="5DE4FB52751FE00BE9BE1D411F61E824" pageId="17" pageNumber="36">(13 ex.). 1 male (CCCC), "Taiwan, Pingtung County, Chunri, Dahanshan, 2009.V.3 D". 1 male, 2 females (CCCC), "Taiwan, Kaohsiung County, Hsiaoguanshan, 1996.X.12, Chou Wen-I leg". 1 female (CCCC), "Taiwan, Kaohsiung, Tengjhih, 2008.V.24 N". 2 males, 2 females (CCCC), "Taiwan, Nantou county, Xitou, 1995.III.25". 1 male, 1 female (CCCC), "Taiwan, Miaoli county, Sanyi, Guandaoshan, 1995.VI.2". 1 female (CCCC), "Taiwan, Yilan county, Datong, Renze, 1998.IV.11". 1 female (CCCC), "Taiwan, Taichung county, Heping, Anmashan".</paragraph>
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</subSubSection>
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<subSubSection id="46773147D4D00DFD1818CA1F61A60C56" pageId="17" pageNumber="36" type="diagnosis">
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<paragraph id="7957304829A710238FFB720ACB7FFA56" pageId="17" pageNumber="36">Diagnosis.</paragraph>
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<paragraph id="F15431774054EFD8377173EFE253E26B" pageId="17" pageNumber="36">
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Medium size species, body length 9.5-10.5 mm; dorsal surface black, elytra usually with faint cyan hue; anterior patch transverse, a little zigzag (more transverse and zigzag than
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<taxonomicName id="BE31D6FF7F6BA16D9D6076AF5F8C2DA0" lsidName="P. o." pageId="17" pageNumber="36" rank="subSpecies" species="o." subSpecies="ornatus">P. o. ornatus</taxonomicName>
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), usually in the fourth to sixth intervals; posterior patch separate, composed of three small spots, in intervals 2-3, 4-5, and 7 respectively. Pronotum transverse, PW/PL 1.50-1.58; lateral margins sinuate before posterior angles, posterior angles nearly rectangular; disc with fine wrinkles. Elytra flat; apical truncation distinctly curved; outer apical angles usually obtuse (Fig. 25), occasionally rounded (Fig. 26) or acute; sutural angles blunt; third interval usually with two setigerous pores (occasionally three as other species), the middle one usually missing. Median lobe of aedeagus with apical orifice opened to left-dorsal side (Fig. 39).
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</paragraph>
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</subSubSection>
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<subSubSection id="60BFD47BDF8AF34E67442AD3E86177BE" pageId="17" pageNumber="36" type="comparisons">
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<paragraph id="3BF79DC8180927535C1C0EAADC1F18A2" pageId="17" pageNumber="36">Comparison.</paragraph>
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<paragraph id="DE61B6650E4D3685EEE02D47BFD694BD" pageId="17" pageNumber="36">
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This subspecies is different from the nominate subspecies in: (1) elytral anterior patch generally more or less transverse (width a little more than length), and somewhat zigzag in
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<taxonomicName id="154507E2FD6E3BD4C9E610935A2F1B1A" lsidName="P. o." pageId="17" pageNumber="36" rank="subSpecies" species="o." subSpecies="formosanus">P. o. formosanus</taxonomicName>
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(Fig. 12); nearly round (width subequal to the length) and not zigzag in
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<taxonomicName id="937AB73D0D05269157C8D969DB27BBE4" lsidName="P. o." pageId="17" pageNumber="36" rank="subSpecies" species="o." subSpecies="ornatus">P. o. ornatus</taxonomicName>
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(Figs 9-11). (2)
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<taxonomicName id="E0823F85F29F8FDEA7CDE6EFC3FBDA25" lsidName="P. o." pageId="17" pageNumber="36" rank="subSpecies" species="o." subSpecies="formosanus">P. o. formosanus</taxonomicName>
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generally with only two setigerous pores on the third elytral interval (occasionally three); in
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<taxonomicName id="AE018B7AD56A7C4096CABD8364C68350" lsidName="P. o." pageId="17" pageNumber="36" rank="subSpecies" species="o." subSpecies="ornatus">P. o. ornatus</taxonomicName>
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always with three setigerous pores; (3) in
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<taxonomicName id="DB3C6A449DD9CAE76CBA52B45CEE30CC" lsidName="P. o." pageId="17" pageNumber="36" rank="subSpecies" species="o." subSpecies="formosanus">P. o. formosanus</taxonomicName>
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, the elytral outer apical angles obtuse in most examined specimens (Fig. 25); in
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<taxonomicName id="DD55D1B5A1DF5BF10D01577F8F667634" lsidName="P. o." pageId="17" pageNumber="36" rank="subSpecies" species="o." subSpecies="ornatus">P. o. ornatus</taxonomicName>
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usually acute (Figs 22, 24). But for some individuals, the shape of elytral outer apical angles cannot differentiate these two subspecies. The male genitalia of these two subspecies have no significant differences (Figs 38, 39).
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</paragraph>
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<paragraph id="678C6086913276A566A6C29556E7EF95" pageId="17" pageNumber="36">
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<taxonomicName id="E64AAE30958447F1FC3D8B5D2443968D" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus o." order="Coleoptera" pageId="17" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="o." subSpecies="formosanus">Pericalus o. formosanus</taxonomicName>
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is very similar with
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<taxonomicName id="DD06A67AD1AC8A2F6F22610D9BAF8569" lsidName="P. acutidens" pageId="17" pageNumber="36" rank="species" species="acutidens">P. acutidens</taxonomicName>
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in elytra pattern, but can be distinguished by two pores on elytral third interval, different male genitalia, and distribution. This subspecies is endemic and the only
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<taxonomicName id="C9698939FAE68F909DEC004A60656E4A" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus" order="Coleoptera" pageId="17" pageNumber="36" phylum="Arthropoda" rank="genus">Pericalus</taxonomicName>
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species in Taiwan.
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</paragraph>
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</subSubSection>
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<subSubSection id="DA0688EC6743AC5D3434FEAAA2E40E16" pageId="17" pageNumber="36" type="distribution">
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<paragraph id="DF0EB8060700D36F0FAA977C7B33142F" pageId="17" pageNumber="36">Distribution.</paragraph>
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<paragraph id="A868C15563BD3876061D78F9050BB63F" pageId="17" pageNumber="36">Only known from Taiwan (Fig. 59).</paragraph>
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</subSubSection>
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<subSubSection id="78BB46BB02283CF042B3E850035BA01A" pageId="17" pageNumber="36" type="remarks">
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<paragraph id="609FFE241CAA06F748F3A03E1E15623A" pageId="17" pageNumber="36">Remarks.</paragraph>
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<paragraph id="49AC056291B0EE3828502FACCE059396" pageId="17" pageNumber="36">
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According to our examined specimens, the male genitalia of
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<taxonomicName id="2EB321CA2D514CE5DD4E78902C8D53F6" lsidName="P. formosanus" pageId="17" pageNumber="36" rank="species" species="formosanus">P. formosanus</taxonomicName>
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have no significant differences from those of
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<taxonomicName id="C4BCC16DBB927A1ED9B9111E21887564" lsidName="P. ornatus" pageId="17" pageNumber="36" rank="species" species="ornatus">P. ornatus</taxonomicName>
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from the Asian continent. Considering the external differences between
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<taxonomicName id="B6AB5995E5FE1582E92BE9D7E689F51C" lsidName="P. ornatus" pageId="17" pageNumber="36" rank="species" species="ornatus">P. ornatus</taxonomicName>
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as well as their allopatric distributions, it is preferable to assign
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<taxonomicName id="3843B583AA0D682EC0F3DD17C83B240E" lsidName="P. formosanus" pageId="17" pageNumber="36" rank="species" species="formosanus">P. formosanus</taxonomicName>
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as a subspecies of
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<taxonomicName id="7BA3499E7D91CE09B1227BB76D3A0C72" lsidName="P. ornatus" pageId="17" pageNumber="36" rank="species" species="ornatus">P. ornatus</taxonomicName>
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rather than synonymize them.
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<bibRefCitation id="342E3F6D0363C8B60427BBB2BCE1EBC9" author="Fedorenko, DN" journalOrPublisher="Russian Entomological Journal" pageId="24" pageNumber="43" pagination="303 - 312" title="A new species of Pericalus s. str. (Coleoptera: Carabidae, Lebiini) from Vietnam, with notes on some Oriental members of the subgenus." volume="26" year="2017">Fedorenko (2017)</bibRefCitation>
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recorded
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<taxonomicName id="7C7978E01A7C67989B58AAD220A02CC2" lsidName="P. formosanus" pageId="17" pageNumber="36" rank="species" species="formosanus">P. formosanus</taxonomicName>
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from S. Vietnam based on a misidentification of
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<taxonomicName id="94A5FB72F246E71763ECA23F1D3297F8" lsidName="P. acutidens" pageId="17" pageNumber="36" rank="species" species="acutidens">P. acutidens</taxonomicName>
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sp. n.
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</paragraph>
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<paragraph id="38BCB1761E51F0D526623A5473C75DFA" pageId="17" pageNumber="36">In most examined specimens have only two setigerous pores in the elytral third interval (the middle one missing), except for two specimens with three pores on right elytron only (Fig. 12). We consider the presence of the middle pore as individual aberration, and consider that the number of pores in third elytral interval is a good taxonomical character to distinguish between these two subspecies. For the elytral outer apical angles, most examined specimens have an obtuse angle (Fig. 25), but it is rounded in one specimen from Nantou (Fig. 26), and acute in one specimen from Kaohsiung and two of the five photographed syntypes (http://digitmuse.nmns.edu.tw).</paragraph>
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</subSubSection>
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</treatment>
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</document> |