treatments-xml/data/3B/43/E5/3B43E574AA6CA44A2212C635381D18A0.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.806.21916" ID-GBIF-Dataset="c93c0643-4af1-435a-bad7-15ec4672211c" ID-PMC="PMC6302070" ID-Pensoft-Pub="1313-2970-806-121" ID-PubMed="30588159" ID-ZBK="996E3193C180461CB3E582BB5E0014A1" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-806-121" ModsDocOrigin="ZooKeys 806" ModsDocTitle="Nine new species of Clada from Madagascar (Coleoptera, Ptinidae)" checkinTime="1545315492460" checkinUser="pensoft" docAuthor="Zahradnik, Petr &amp; ryzna, Milos" docDate="2018" docId="3B43E574AA6CA44A2212C635381D18A0" docLanguage="en" docName="ZooKeys 806: 121-140" docOrigin="ZooKeys 806" docSource="http://dx.doi.org/10.3897/zookeys.806.21916" docTitle="Clada (Clada) njakai Zahradnik &amp; ryzna, 2018, sp. n." docType="treatment" docUuid="D3CD1585-C840-41A0-9290-E8FFCD375206" docUuidSource="ZooBank" docVersion="5" lastPageNumber="121" masterDocId="FFB8BD0CFFDC4671FFE0FFC5FFB45748" masterDocTitle="Nine new species of Clada from Madagascar (Coleoptera, Ptinidae)" masterLastPageNumber="140" masterPageNumber="121" pageNumber="121" updateTime="1668166581846" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Nine new species of Clada from Madagascar (Coleoptera, Ptinidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zahradnik, Petr</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>ryzna, Milos</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>806</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>121</mods:start>
<mods:end>140</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.806.21916</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.806.21916</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-806-121</mods:identifier>
<mods:identifier type="ZBK">996E3193C180461CB3E582BB5E0014A1</mods:identifier>
<mods:identifier type="ZooBank">996E3193C180461CB3E582BB5E0014A1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="154127124" LSID="urn:lsid:zoobank.org:act:D3CD1585-C840-41A0-9290-E8FFCD375206" httpUri="http://treatment.plazi.org/id/3B43E574AA6CA44A2212C635381D18A0" lastPageNumber="121" pageId="0" pageNumber="121">
<subSubSection pageId="0" pageNumber="121" type="nomenclature">
<paragraph pageId="0" pageNumber="121">
<taxonomicName LSID="http://zoobank.org/D3CD1585-C840-41A0-9290-E8FFCD375206" class="Insecta" family="Ptinidae" genus="Clada" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clada (Clada) njakai" order="Coleoptera" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="njakai" subGenus="Clada">Clada (Clada) njakai</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="121">sp. n.</taxonomicNameLabel>
Figs 7, 16, 25, 34
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="type material">
<paragraph pageId="0" pageNumber="121">Type material.</paragraph>
<paragraph pageId="0" pageNumber="121">
Holotype male: Madagascar, Toliara prov., Tsimanampetsotsa N. P., Mitoho camp,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-24.0483">24°02.898'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="43.7523">43°45.138'E</geoCoordinate>
, 10 m a. s. l., 12.-13.i.2014, M.
<normalizedToken originalValue="Trýzna">Tryzna</normalizedToken>
leg. Paratypes (15): 1 male: the same data as holotype; 13 males: Madagascar, Toliara prov., Tsimanampetsotsa N. P., Andranovao camp,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-24.025084">24°01.505'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="43.738434">43°44.306'E</geoCoordinate>
, 15 m a. s. l., 14.-15.i.2014, M.
<normalizedToken originalValue="Trýzna">Tryzna</normalizedToken>
leg. (FGMRI 7 ex., LBVC 2 ex., MTDC 2 ex., NHMUK 2 ex.).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="differential diagnosis">
<paragraph pageId="0" pageNumber="121">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="121">
This species is similar to
<taxonomicName lsidName="C. (C.) humeralis" pageId="0" pageNumber="121" rank="species" species="humeralis" subGenus="C.">C. (C.) humeralis</taxonomicName>
Pic, 1926, but differs by the lighter colour of the elytra and missing lighter humeri. Fully differs by shape of the aedeagus. For differences from other Madagascan species, see key.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="description">
<paragraph pageId="0" pageNumber="121">Description.</paragraph>
<paragraph pageId="0" pageNumber="121">Male (holotype). Elongate-elliptical, transversally convex. Body length 5.3 mm, maximum width 2.1 mm (Figure 7). Ratio length:width of elytra 1.8. Whole body dark brown, only antennae, maxillary and labial palpi and humeri on elytra moderately lighter. Pubescence yellowish-white.</paragraph>
<paragraph pageId="0" pageNumber="121">Head matt-shiny, with double punctation - one coarse, dense, umbilicate, distance between punctures approximately the same as their diameter; other one very fine, punctures almost touching. Pubescence recumbent or semi-erect, short, inclined backwards; on sides of head semi-erect and long, inclined forwards. Clypeus with shallow transverse depression. Eyes large, globular with short erect sparse pubescence. Frons twice as wide as diameters of eye, from dorsal view. Antennae consisting of eleven antennomeres, 4th to 10th pectinate (Figure 25). 1stantennomere robust, twice as long as wide; 2nd smallest, only one-half length of 1st, as long as wide, slightly narrower than 1st. 3rd serrate, as long as wide; 4th and 5th 1.8 times wider than long; 6th and 7th 1.5 times wider than long; the 8th and 9th 1.3 times wider as long; and 10th 1.1 times as wide as long. Apical antennomere is longest, oblong oval, 4.3 times as long as wide. All antennomeres on margin with short erect dense setae. Apical maxillary palpomere long, spindle shaped.</paragraph>
<paragraph pageId="0" pageNumber="121">Pronotum convex, matt-shiny, rounded, transverse (ratio length:width of pronotum 0.7); widest at 2/3 posteriorly. Base of pronotum finely bordered. Middle of pronotum at base with blunt small swlling, posteriorly slightly sharpened. Surface of pronotum with double punctation, one coarse, dense, umbilicate, distance between punctures approximately the same as their diameter; other one is very fine, punctures almost touching. Pubescence long, sparse, recumbent, inclined more or less to middle of pronotum.</paragraph>
<paragraph pageId="0" pageNumber="121">Scutellum triangular, narrow, 1.3 times as long as wide, very densely recumbent pubescence, inclined backwards, surface is not visible.</paragraph>
<paragraph pageId="0" pageNumber="121">Elytra oval, transversally convex, shining, with distinct humeri. Each elytron with five fine costae, almost invisible, but apex more distinct. Surface of elytra with double punctation, one coarse, dense, umbilicate, distance between punctures approximately the same as their diameter; other one is very fine, punctures almost touching. Pubescence relatively sparse, recumbent or semi-erect, inclined backwards. Posterior margin of each elytron with approximately 25 small teeth.</paragraph>
<paragraph pageId="0" pageNumber="121">Legs stout, with short and dense recumbent pubescence. All tarsi robust, same length as tibia. 1st metatarsomere as long as 2nd and 3rd together, same width, slightly emarginate on top, 4th slightly shorter than previous, more emarginate, almost to 2/3 of their length. 5th is same length as 3rd and 4th together, rectangular, wider on the top, with two large claws, without teeth.</paragraph>
<paragraph pageId="0" pageNumber="121">For aedeagus see Figure 34.</paragraph>
<paragraph pageId="0" pageNumber="121">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="variability">
<paragraph pageId="0" pageNumber="121">Variability.</paragraph>
<paragraph pageId="0" pageNumber="121">Body length 4.7-7.1 mm, maximum width 1.8-2.8 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="name derivation">
<paragraph pageId="0" pageNumber="121">Name derivation.</paragraph>
<paragraph pageId="0" pageNumber="121">Patronym, dedicated to Adolphe Randrianjaka (University of Antananarivo, Faculty of Sciences, Department of Entomology), whom we called Njaka.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="biology_ecology">
<paragraph pageId="0" pageNumber="121">Biology.</paragraph>
<paragraph pageId="0" pageNumber="121">Unknown. All specimens were collected at light.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="distribution">
<paragraph pageId="0" pageNumber="121">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="121">This species is found in the southwestern part of Madagascar (Figure 16).</paragraph>
</subSubSection>
</treatment>
</document>