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<document id="123651B90FAD549809C6C3BAD2381166" ID-DOI="10.1206/593.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="10111354" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630275288352" checkinUser="felipe" docAuthor="Robert M. Hunt, Jr." docDate="2009" docId="03FF87A3FFFDF34FA3827D3887CAFD72" docLanguage="en" docName="B318.pdf" docOrigin="Bulletin of the American Museum of Natural History 2009 (318)" docStyle="DocumentStyle:2B34016C1EBFE58B07368F9517618743.3:BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleId="2B34016C1EBFE58B07368F9517618743" docStyleName="BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleVersion="3" docTitle="Daphoenodon (Borocyon) niobrarensis Loomis 1936" docType="treatment" docVersion="3" lastPageNumber="23" masterDocId="FFC6FFDBFFE9F359A1687914840BF831" masterDocTitle="Long-Legged Pursuit Carnivorans (Amphicyonidae, Daphoeninae) From The Early Miocene Of North America" masterLastPageNumber="95" masterPageNumber="1" pageNumber="21" updateTime="1699685760276" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="121CFB42A8288DFFBD555FAD0D6E515A">Long-Legged Pursuit Carnivorans (Amphicyonidae, Daphoeninae) From The Early Miocene Of North America</mods:title>
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<treatment id="03FF87A3FFFDF34FA3827D3887CAFD72" LSID="urn:lsid:plazi:treatment:03FF87A3FFFDF34FA3827D3887CAFD72" httpUri="http://treatment.plazi.org/id/03FF87A3FFFDF34FA3827D3887CAFD72" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
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<paragraph id="8BE936B5FFFDF34DA3827D388048FC4F" blockId="20.[746,1183,1068,1150]" pageId="20" pageNumber="21">
<heading id="D0A181D9FFFDF34DA3827D388048FC4F" centered="true" fontSize="9" level="2" pageId="20" pageNumber="21" reason="2">
<taxonomicName id="4C564D36FFFDF34DA3827D38801EFC51" authority="Loomis, 1936" authorityName="Loomis" authorityYear="1936" class="Mammalia" family="Amphicyonidae" genus="Daphoenodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="niobrarensis" subGenus="Borocyon">
<emphasis id="B922EAA7FFFDF34DA3827D38878DFC72" box="[746,902,1068,1091]" italics="true" pageId="20" pageNumber="21">Daphoenodon</emphasis>
(
<emphasis id="B922EAA7FFFDF34DA2F07D39800EFC73" box="[920,1029,1069,1090]" italics="true" pageId="20" pageNumber="21">Borocyon</emphasis>
)
<emphasis id="B922EAA7FFFDF34DA57E7D388094FC72" box="[1046,1183,1068,1091]" italics="true" pageId="20" pageNumber="21">niobrarensis</emphasis>
<bibRefCitation id="EFC74B44FFFDF34DA21D7D5E801EFC51" author="Loomis, F. B." box="[885,1045,1098,1120]" pageId="20" pageNumber="21" pagination="44 - 52" refId="ref47507" refString="Loomis, F. B. 1936. Three new Miocene dogs and their phylogeny. Journal of Paleontology 10 (1): 44 - 52." type="journal article" year="1936">Loomis, 1936</bibRefCitation>
</taxonomicName>
<figureCitation id="136D2A30FFFDF34DA22D7D7C87C8FC4F" box="[837,963,1128,1150]" captionStart="Fig" captionStartId="22.[150,188,718,737]" captionTargetBox="[148,1206,212,685]" captionTargetId="figure-392@22.[148,1208,209,687]" captionTargetPageId="22" captionText="Fig. 10. Holotype right mandible of Borocyon niobrarensis Loomis with c, p1p4, and m1m3 from Aletomeryx Quarry, lower Runningwater Fm., Cherry Co., Nebraska (ACM 3452)." figureDoi="http://doi.org/10.5281/zenodo.10111372" httpUri="https://zenodo.org/record/10111372/files/figure.png" pageId="20" pageNumber="21">Figures 10</figureCitation>
,
<figureCitation id="136D2A30FFFDF34DA2B97D7D87E4FC4F" box="[977,1007,1129,1150]" captionStart="Fig" captionStartId="23.[116,155,1552,1571]" captionTargetBox="[152,1133,210,1521]" captionTargetId="figure-64@23.[152,1135,209,1521]" captionTargetPageId="23" captionText="Fig. 11. Associated holotype right forelimb of Borocyon niobrarensis Loomis (ACM 3452), Aletomeryx Quarry, Cherry Co., Nebraska." figureDoi="http://doi.org/10.5281/zenodo.10111374" httpUri="https://zenodo.org/record/10111374/files/figure.png" pageId="20" pageNumber="21">11</figureCitation>
,
<figureCitation id="136D2A30FFFDF34DA2957D7D8010FC4F" box="[1021,1051,1129,1150]" captionStart="Fig" captionStartId="41.[116,155,1677,1696]" captionTargetBox="[156,1129,211,1647]" captionTargetId="figure-1@41.[156,1130,211,1647]" captionTargetPageId="41" captionText="Fig. 20. Comparison of the scapulae of (A) Borocyon niobrarensis (ACM 3452); (B) Ursus americanus" figureDoi="http://doi.org/10.5281/zenodo.10111396" httpUri="https://zenodo.org/record/10111396/files/figure.png" pageId="20" pageNumber="21">20</figureCitation>
,
<figureCitation id="136D2A30FFFDF34DA5407D7D8048FC4F" box="[1064,1091,1129,1150]" captionStart="Fig" captionStartId="45.[116,155,1347,1366]" captionTargetBox="[185,1100,210,1317]" captionTargetId="figure-131@45.[185,1100,210,1317]" captionTargetPageId="45" captionText="Fig. 23. Comparison of the ulnae of Amphicyon galushai (left) and Borocyon niobrarensis (center and right), showing the more elongate, slender, curved ulna of the latter species. No complete ulna of B. robustum is known; however, eight radii of this species demonstrate the existence of an even more slender, elongate ulna." figureDoi="http://doi.org/10.5281/zenodo.10111402" httpUri="https://zenodo.org/record/10111402/files/figure.png" pageId="20" pageNumber="21">23</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C653EFFFDF34CA3B27D818569FD4B" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BE936B5FFFDF34DA3B27D8180C7FD86" blockId="20.[701,1228,1173,1759]" pageId="20" pageNumber="21">
<materialsCitation id="3B3E3CE8FFFDF34DA3B27D8180C7FD86" collectionCode="ACM" pageId="20" pageNumber="21" specimenCode="ACM 3452, ACM 34-52" specimenCount="3" typeStatus="holotype">
<typeStatus id="54ED8817FFFDF34DA3B27D818751FC9A" box="[730,858,1173,1197]" pageId="20" pageNumber="21" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCode id="DBF09ECEFFFDF34DA2077D8387FCFC9C" box="[879,1015,1175,1197]" collectionCode="ACM" country="Australia" httpUri="http://grbio.org/cool/12ic-3ajm" name="Australian Collection of Microorganisms" pageId="20" pageNumber="21">ACM 3452</specimenCode>
(field no. 34-52), left mandible with c, p2m3, right mandible with c, p1m3, and isolated right M1, associated with nearly complete forelimbs;
<typeStatus id="54ED8817FFFDF34DA3D57C198728FD12" box="[701,803,1293,1315]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C564D36FFFDF34DA23D7C188752FD0E" authority="Loomis, 1936: 47" authorityName="Loomis" authorityPageNumber="47" authorityYear="1936" class="Mammalia" family="Amphicyonidae" genus="Daphoenodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="niobrarensis">
<emphasis id="B922EAA7FFFDF34DA23D7C18808DFD12" box="[853,1158,1292,1315]" italics="true" pageId="20" pageNumber="21">Daphoenodon niobrarensis</emphasis>
<bibRefCitation id="EFC74B44FFFDF34DA5FA7C198752FD0E" author="Loomis, F. B." pageId="20" pageNumber="21" pagination="44 - 52" refId="ref47507" refString="Loomis, F. B. 1936. Three new Miocene dogs and their phylogeny. Journal of Paleontology 10 (1): 44 - 52." type="journal article" year="1936">Loomis, 1936: 47</bibRefCitation>
</taxonomicName>
, figs. 2, 4. There is no doubt as to the identification of the
<typeStatus id="54ED8817FFFDF34DA5607C5C8065FD6F" box="[1032,1134,1352,1374]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
mandibles and forelimbs, which Loomis published as
<specimenCode id="DBF09ECEFFFDF34DA3B47C978761FDA8" box="[732,874,1411,1433]" collectionCode="ACM" country="Australia" httpUri="http://grbio.org/cool/12ic-3ajm" name="Australian Collection of Microorganisms" pageId="20" pageNumber="21">ACM 34-52</specimenCode>
, although the field number 34- 58 appears on many of the individual bones.
</materialsCitation>
</paragraph>
<paragraph id="8BE936B5FFFDF34CA3B27CA98569FD4B" blockId="20.[701,1228,1173,1759]" lastBlockId="21.[93,620,1291,1759]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<typeStatus id="54ED8817FFFDF34DA3B27CA98750FDE5" box="[730,859,1469,1493]" pageId="20" pageNumber="21" type="holotype">HOLOTYPE</typeStatus>
HORIZON
<collectionCode id="ED47AE70FFFDF34DA2967CD78039FDE5" box="[1022,1074,1475,1492]" pageId="20" pageNumber="21">AND</collectionCode>
LOCALITY: From the lower Runningwater Fm., Cherry Co.,
<collectingRegion id="4992F857FFFDF34DA3917CEE8765FE21" box="[761,878,1530,1552]" country="United States of America" name="Nebraska" pageId="20" pageNumber="21">Nebraska</collectingRegion>
.
<bibRefCitation id="EFC74B44FFFDF34DA2137CEE802DFE21" author="Loomis, F. B." box="[891,1062,1530,1552]" pageId="20" pageNumber="21" pagination="44 - 52" refId="ref47507" refString="Loomis, F. B. 1936. Three new Miocene dogs and their phylogeny. Journal of Paleontology 10 (1): 44 - 52." type="journal article" year="1936">Loomis (1936)</bibRefCitation>
reported that the
<typeStatus id="54ED8817FFFDF34DA3847F0C8759FE1F" box="[748,850,1560,1582]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
was Found in
<quantity id="4CAE9B50FFFDF34DA5737F0C8079FE1F" box="[1051,1138,1560,1582]" metricMagnitude="1" metricUnit="m" metricValue="4.91236" pageId="20" pageNumber="21" unit="in" value="1934.0">1934 in</quantity>
middle Miocene beds equivalent to the upper Harrison formation, near mouth of Antelope Creek,
<quantity id="4CAE9B50FFFDF34DA2717F648772FEB7" box="[793,889,1648,1670]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="20" pageNumber="21" unit="mi" value="12.0">12 miles</quantity>
southeast of Gordon,
<collectingRegion id="4992F857FFFDF34DA5C87F658704FE95" country="United States of America" name="Nebraska" pageId="20" pageNumber="21">Nebraska</collectingRegion>
. Associated with
<taxonomicName id="4C564D36FFFDF34DA2857F998078FE95" box="[1005,1139,1677,1700]" class="Mammalia" family="Palaeomerycidae" genus="Aletomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B922EAA7FFFDF34DA2857F998078FE95" box="[1005,1139,1677,1700]" italics="true" pageId="20" pageNumber="21">Aletomeryx</emphasis>
</taxonomicName>
[
<collectionCode id="ED47AE70FFFDF34DA5EF7F9A80C7FE92" box="[1159,1228,1678,1699]" country="Australia" httpUri="http://grbio.org/cool/12ic-3ajm" name="Australian Collection of Microorganisms" pageId="20" pageNumber="21">ACM</collectionCode>
1915]. In fact the
<typeStatus id="54ED8817FFFDF34DA2FF7FB887F6FEF3" box="[919,1021,1708,1730]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
was found with a large sample of
<taxonomicName id="4C564D36FFFDF34DA2E17FDC8004FEEE" box="[905,1039,1736,1759]" class="Mammalia" family="Palaeomerycidae" genus="Aletomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B922EAA7FFFDF34DA2E17FDC8004FEEE" box="[905,1039,1736,1759]" italics="true" pageId="20" pageNumber="21">Aletomeryx</emphasis>
</taxonomicName>
and the camel
<taxonomicName id="4C564D36FFFCF34CA1357C1F84C3FD13" authorityName="Frick &amp; Taylor" authorityYear="1971" box="[93,200,1291,1314]" class="Mammalia" family="Camelidae" genus="Michenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B922EAA7FFFCF34CA1357C1F84C3FD13" box="[93,200,1291,1314]" italics="true" pageId="21" pageNumber="22">Michenia</emphasis>
</taxonomicName>
[
<collectionCode id="ED47AE70FFFCF34CA1BC7C188512FD10" box="[212,281,1292,1313]" country="Australia" httpUri="http://grbio.org/cool/12ic-3ajm" name="Australian Collection of Microorganisms" pageId="21" pageNumber="22">ACM</collectionCode>
1844] in
<taxonomicName id="4C564D36FFFCF34CA0E87C18866DFD10" authority="Quarry" authorityName="Quarry" box="[384,614,1292,1314]" class="Mammalia" family="Palaeomerycidae" genus="Aletomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">Aletomeryx Quarry</taxonomicName>
, an early Miocene locality in the lower Runningwater Fm. worked by several museums during the 1930s.
</paragraph>
</subSubSection>
<caption id="DF29663DFFFCF34CA11C7D9F80A2FC88" ID-DOI="http://doi.org/10.5281/zenodo.10111370" ID-Zenodo-Dep="10111370" httpUri="https://zenodo.org/record/10111370/files/figure.png" pageId="21" pageNumber="22" startId="21.[116,155,1163,1182]" targetBox="[223,1063,210,1132]" targetPageId="21">
<paragraph id="8BE936B5FFFCF34CA11C7D9F80A2FCAF" blockId="21.[93,1193,1162,1209]" box="[116,1193,1162,1182]" pageId="21" pageNumber="22">
Fig. 9. Reconstructed palate of
<taxonomicName id="4C564D36FFFCF34CA0BA7D988638FCAF" authorityName="Peterson" authorityYear="1910" box="[466,563,1164,1182]" class="Mammalia" family="Amphicyonidae" genus="Borocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B922EAA7FFFCF34CA0BA7D988638FCAF" box="[466,563,1164,1182]" italics="true" pageId="21" pageNumber="22">Borocyon</emphasis>
</taxonomicName>
cf.
<taxonomicName id="4C564D36FFFCF34CA3347D9886D2FCAF" authorityName="Peterson" authorityYear="1910" box="[604,729,1162,1182]" class="Mammalia" family="Amphicyonidae" genus="Borocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="robustum">
<emphasis id="B922EAA7FFFCF34CA3347D9886D2FCAF" box="[604,729,1162,1182]" italics="true" pageId="21" pageNumber="22">B. robustum</emphasis>
</taxonomicName>
(cast, KU 114592) from sinkhole or fissure,
</paragraph>
<paragraph id="8BE936B5FFFCF34CA1357DB280A2FC88" blockId="21.[93,1193,1162,1209]" box="[93,1193,1189,1209]" pageId="21" pageNumber="22">Suwanee River, north Florida. Complete dentition includes I1I3, C, P1P4, M1M2. Note absence of M3.</paragraph>
</caption>
<subSubSection id="C34C653EFFFCF34CA1127C958660FEEE" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8BE936B5FFFCF34CA1127C9585DCFE92" blockId="21.[93,620,1291,1759]" pageId="21" pageNumber="22">
DIAGNOSIS:
<taxonomicName id="4C564D36FFFCF34CA0797C9085D2FDA8" box="[273,473,1411,1433]" class="Mammalia" family="Amphicyonidae" genus="Borocyon" kingdom="Animalia" order="Carnivora" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B922EAA7FFFCF34CA0797C908575FDA8" box="[273,382,1412,1433]" italics="true" pageId="21" pageNumber="22">Borocyon</emphasis>
species
</taxonomicName>
of intermediate size, with basilar skull length of,
<quantity id="4CAE9B50FFFCF34CA11C7CAA84DDFDE5" box="[116,214,1470,1492]" metricMagnitude="-1" metricUnit="m" metricValue="2.7" pageId="21" pageNumber="22" unit="mm" value="270.0">270 mm</quantity>
, and cm2 length of,
<quantity id="4CAE9B50FFFCF34CA35C7CAA84B4FDC0" metricMagnitude="-1" metricUnit="m" metricValue="1.09" metricValueMax="1.15" metricValueMin="1.03" pageId="21" pageNumber="22" unit="mm" value="109.0" valueMax="115.0" valueMin="103.0">103 115 mm</quantity>
, its skeletal and dental measurements (figs. 17, 18; tables 2, 3) generally falling between those of the large species
<taxonomicName id="4C564D36FFFCF34CA3397F0384CCFE7A" authorityName="Peterson" authorityYear="1910" class="Mammalia" family="Amphicyonidae" genus="Borocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="robustum">
<emphasis id="B922EAA7FFFCF34CA3397F0384CCFE7A" italics="true" pageId="21" pageNumber="22">B. robustum</emphasis>
</taxonomicName>
and small
<taxonomicName id="4C564D36FFFCF34CA0277F21861AFE7B" authorityName="Hunt" authorityYear="2009" box="[335,529,1589,1610]" class="Mammalia" family="Amphicyonidae" genus="Daphoenodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="neomexicanus">
<emphasis id="B922EAA7FFFCF34CA0277F21861AFE7B" box="[335,529,1589,1610]" italics="true" pageId="21" pageNumber="22">B. neomexicanus</emphasis>
</taxonomicName>
. Proximal tarsals (figs. 33, 34, astragalus, calcaneum) shorter and differently proportioned relative to those of
<taxonomicName id="4C564D36FFFCF34CA02C7F9A85DBFE92" authorityName="Peterson" authorityYear="1910" box="[324,464,1676,1699]" class="Mammalia" family="Amphicyonidae" genus="Borocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="robustum">
<emphasis id="B922EAA7FFFCF34CA02C7F9A85DBFE92" box="[324,464,1676,1699]" italics="true" pageId="21" pageNumber="22">B. robustum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE936B5FFFCF34CA1127FBE8660FEEE" blockId="21.[93,620,1291,1759]" pageId="21" pageNumber="22">
REFERRED SPECIMENS: Runningwater Formation, northwestern
<collectingRegion id="4992F857FFFCF34CA0E17FDD85F3FEEE" box="[393,504,1737,1759]" country="United States of America" name="Nebraska" pageId="21" pageNumber="22">Nebraska</collectingRegion>
<emphasis id="B922EAA7FFFCF34CA3737FDD8660FEEE" bold="true" box="[539,619,1737,1759]" pageId="21" pageNumber="22">Cherry</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34C653EFFFCF34FA3F37C1887CAFD72" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="8BE936B5FFFCF34CA3F37C1880A2FEEE" blockId="21.[667,1194,1292,1759]" pageId="21" pageNumber="22">
<emphasis id="B922EAA7FFFCF34CA3F37C188742FD10" bold="true" box="[667,841,1292,1314]" pageId="21" pageNumber="22">
Co.,
<collectingRegion id="4992F857FFFCF34CA3BB7C18874FFD10" box="[723,836,1292,1313]" country="United States of America" name="Nebraska" pageId="21" pageNumber="22">Nebraska</collectingRegion>
:
</emphasis>
(1) ACM 11796 (field no. 34- 58), left metacarpals 3 and 4,
<taxonomicName id="4C564D36FFFCF34CA5487C3E86F9FD6D" authority="Quarry" authorityName="Quarry" class="Mammalia" family="Palaeomerycidae" genus="Aletomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">Aletomeryx Quarry</taxonomicName>
; (2) F:AM 68269, right metatarsal 4, cuboid, navicular, phalanx,
<taxonomicName id="4C564D36FFFCF34CA5487C718781FDA9" authority="Quarry. Dawes Co." authorityName="Quarry. Dawes Co." class="Mammalia" family="Palaeomerycidae" genus="Aletomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
Aletomeryx Quarry.
<emphasis id="B922EAA7FFFCF34CA26D7C978781FDA9" bold="true" box="[773,906,1410,1432]" pageId="21" pageNumber="22">Dawes Co.</emphasis>
</taxonomicName>
<emphasis id="B922EAA7FFFCF34CA2E27C96801FFDA9" bold="true" box="[906,1044,1410,1432]" pageId="21" pageNumber="22">
,
<collectingRegion id="4992F857FFFCF34CA2F77C97801BFDA9" box="[927,1040,1411,1432]" country="United States of America" name="Nebraska" pageId="21" pageNumber="22">Nebraska</collectingRegion>
:
</emphasis>
(1) UNSM 25683 (field no. 1-11-9-36NP), left mandible, p4m2; (2) UNSM 25555 (field no. 1-13-8- 36NP), left humerus; (3) UNSM 44702 (field no. 9-12-8-36NP), right metacarpals 2, 3, and 4, possibly the same individual as UNSM 25555 and 25683. All from Red Horse Quarry (UNSM Dw-103).
<emphasis id="B922EAA7FFFCF34CA2967F46871AFEB4" bold="true" pageId="21" pageNumber="22">
Sheridan Co.,
<collectingRegion id="4992F857FFFCF34CA3F37F648707FEB4" box="[667,780,1648,1669]" country="United States of America" name="Nebraska" pageId="21" pageNumber="22">Nebraska</collectingRegion>
:
</emphasis>
(1) UNSM 26418 (no field no.), right m2, UNSM Loc. Sh-101B.
<emphasis id="B922EAA7FFFCF34CA5447F99871AFEF0" bold="true" pageId="21" pageNumber="22">
Sioux Co.,
<collectingRegion id="4992F857FFFCF34CA3F37FB88707FEF0" box="[667,780,1708,1729]" country="United States of America" name="Nebraska" pageId="21" pageNumber="22">Nebraska</collectingRegion>
:
</emphasis>
(1) F:AM 107601 (F:AM field nos. NEA 12, 8130; UNSM field no.
</paragraph>
<caption id="DF29663DFFFFF34FA1FE7BDA87F8FACA" ID-DOI="http://doi.org/10.5281/zenodo.10111372" ID-Zenodo-Dep="10111372" httpUri="https://zenodo.org/record/10111372/files/figure.png" pageId="22" pageNumber="23" startId="22.[150,188,718,737]" targetBox="[148,1206,212,685]" targetPageId="22">
<paragraph id="8BE936B5FFFFF34FA1FE7BDA80C0FAD0" blockId="22.[128,1227,717,764]" box="[150,1227,717,737]" pageId="22" pageNumber="23">
Fig. 10. Holotype right mandible of
<taxonomicName id="4C564D36FFFFF34FA3457BDA8761FAD0" authority="Loomis" authorityName="Loomis" box="[557,874,717,737]" class="Mammalia" family="Amphicyonidae" genus="Borocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="niobrarensis">
<emphasis id="B922EAA7FFFFF34FA3457BDA8719FAD0" box="[557,786,717,737]" italics="true" pageId="22" pageNumber="23">Borocyon niobrarensis</emphasis>
Loomis
</taxonomicName>
with c, p1p4, and m1m3 from
</paragraph>
<paragraph id="8BE936B5FFFFF34FA1E87BFC87F8FACA" blockId="22.[128,1227,717,764]" box="[128,1011,744,764]" pageId="22" pageNumber="23">
<taxonomicName id="4C564D36FFFFF34FA1E87BFC855BFACA" authority="Quarry" authorityName="Quarry" box="[128,336,744,763]" class="Mammalia" family="Palaeomerycidae" genus="Aletomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">Aletomeryx Quarry</taxonomicName>
, lower Runningwater Fm., Cherry Co., Nebraska (ACM 3452).
</paragraph>
</caption>
<paragraph id="8BE936B5FFFFF34FA1E87A2E84D8FDC2" blockId="22.[128,654,826,1759]" pageId="22" pageNumber="23">
RH104), right mandible with c, p1m3, partial right scapula, left ulna, left scapholunar, left metacarpal 2, right innominate, right calcaneum, astragalus, and cuboid, proximal metatarsal 5, proximal phalanx, intermediate phalanx, partial proximal phalanx. F:AM 107601 is an associated partial skeleton from a massive pinkish silty sandstone in SW1/4, SW1/4, sec. 1, T29N, R55W, northeast of the post office of Agate,
<collectingRegion id="4992F857FFFFF34FA0DC7D568622FC69" box="[436,553,1090,1112]" country="United States of America" name="Nebraska" pageId="22" pageNumber="23">Nebraska</collectingRegion>
. F:AM specimens were collected by Ron Brown in
<date id="FFE81075FFFFF34FA1E87D69851EFCA3" box="[128,277,1149,1170]" pageId="22" pageNumber="23" value="1970-08">August 1970</date>
; UNSM material (metacarpal 2, ulna, calcaneum, partial scapula) was collect- ed in
<date id="FFE81075FFFFF34FA1AF7DAC8533FCFC" box="[199,312,1208,1230]" pageId="22" pageNumber="23" value="1971-07">July 1971</date>
by R.M. Hunt, T. Hussain, and L. Hunt and donated to the American Museum. Associated with
<taxonomicName id="4C564D36FFFFF34FA0D27DE68663FD38" box="[442,616,1266,1289]" class="Mammalia" family="Merycoidodontidae" genus="Merycochoerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B922EAA7FFFFF34FA0D27DE68663FD38" box="[442,616,1266,1289]" italics="true" pageId="22" pageNumber="23">Merycochoerus</emphasis>
</taxonomicName>
cf.
<taxonomicName id="4C564D36FFFFF34FA1E87C05850FFD17" authorityName="Loomis" authorityYear="1924" box="[128,260,1297,1318]" class="Mammalia" family="Merycoidodontidae" genus="Merycochoerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B922EAA7FFFFF34FA1E87C05850FFD17" box="[128,260,1297,1318]" italics="true" pageId="22" pageNumber="23">M. magnus</emphasis>
</taxonomicName>
. (2) UNSM 44827, nearly complete skull and mandibles but lacking much of the rostrum; teeth extremely worn; right P3M2, left M1M3, partial P4; damaged right p4m3 and left m1m3; from massive pale reddish brown sandstone, Skavdahl Ranch, NW1/4, NW1/4, sec. 29, T29N, R
<specimenCount id="9D50FD3CFFFFF34FA1FC7CC984D8FDC2" box="[148,211,1501,1523]" pageId="22" pageNumber="23" type="generic">53W.</specimenCount>
</paragraph>
<paragraph id="8BE936B5FFFFF34FA1F47CEE8711FC44" blockId="22.[128,654,826,1759]" lastBlockId="22.[701,1228,826,1348]" pageId="22" pageNumber="23">
REFERRED SPECIMENS: Late early Miocene beds east of the Laramie Mountains, southeastern
<collectingRegion id="4992F857FFFFF34FA0717F228580FE7D" box="[281,395,1590,1612]" country="United States of America" name="Wyoming" pageId="22" pageNumber="23">Wyoming</collectingRegion>
(
<bibRefCitation id="EFC74B44FFFFF34FA0F27F228490FE5B" author="Cassiliano, M." pageId="22" pageNumber="23" pagination="25 - 68" refId="ref46044" refString="Cassiliano, M. 1980. Stratigraphy and vertebrate paleontology of the Horse Creek-Trail Creek area, Laramie County, Wyoming. Contributions to Geology University of Wyoming 19: 25 - 68." type="journal article" year="1980">Cassiliano, 1980: 30 39</bibRefCitation>
)—
<emphasis id="B922EAA7FFFFF34FA1AF7F4785D0FE58" bold="true" box="[199,475,1619,1641]" pageId="22" pageNumber="23">
Laramie Co.,
<collectingRegion id="4992F857FFFFF34FA00D7F4785DDFE58" box="[357,470,1619,1641]" country="United States of America" name="Wyoming" pageId="22" pageNumber="23">Wyoming</collectingRegion>
:
</emphasis>
(1) UW 10004, left mandible with p4m2 (a right mandible was retained by the field collector and is now lost), left M3, left P3, left canine, left scapula, humerus, ulna, and radius, right humerus and ulna, two right radii, right magnum, both pisiforms, left metacarpals 4 and 5, right innominate, left femur, tibia, and fibula, right astragalus and calcaneum, left navicular, right metatarsal 2, left metatarsals 3 and 4, four proximal phalanges, axis, two cervical vertebrae, three thoracic vertebrae, two lumbar vertebrae, one caudal vertebra, partial ribs. From the Middle Miocene Formation of
<bibRefCitation id="EFC74B44FFFFF34FA2477D5687F4FC69" author="Cassiliano, M." box="[815,1023,1090,1112]" pageId="22" pageNumber="23" pagination="25 - 68" refId="ref46044" refString="Cassiliano, M. 1980. Stratigraphy and vertebrate paleontology of the Horse Creek-Trail Creek area, Laramie County, Wyoming. Contributions to Geology University of Wyoming 19: 25 - 68." type="journal article" year="1980">Cassiliano (1980)</bibRefCitation>
at Horse Creek Quarry.
</paragraph>
<paragraph id="8BE936B5FFFFF34FA3B27D6887CAFD72" blockId="22.[701,1228,826,1348]" pageId="22" pageNumber="23">
REFERRED SPECIMEN: Carpenter Ranch Formation, southeastern
<collectingRegion id="4992F857FFFFF34FA2917D8F8060FC80" box="[1017,1131,1179,1201]" country="United States of America" name="Wyoming" pageId="22" pageNumber="23">Wyoming</collectingRegion>
(
<bibRefCitation id="EFC74B44FFFFF34FA5EE7D8F86FFFCFF" author="Hunt, R. M., Jr." pageId="22" pageNumber="23" pagination="1 - 45" refId="ref47153" refString="Hunt, R. M., Jr. 2005. An early Miocene domeskulled chalicothere from the '' Arikaree' ' conglomerates of Darton: calibrating the ages of High Plains paleovalleys against Rocky Mountain tectonism. American Museum Novitates 3486: 1 - 45." type="journal article" year="2005">Hunt, 2005</bibRefCitation>
)—
<emphasis id="B922EAA7FFFFF34FA24E7DAC8031FCFF" bold="true" box="[806,1082,1208,1230]" pageId="22" pageNumber="23">
Goshen Co.,
<collectingRegion id="4992F857FFFFF34FA2AD7DAC803DFCFF" box="[965,1078,1208,1230]" country="United States of America" name="Wyoming" pageId="22" pageNumber="23">Wyoming</collectingRegion>
:
</emphasis>
(1) UNSM 44815, partial left mandible with m1m3, partial p4, described in
<bibRefCitation id="EFC74B44FFFFF34FA2B67DE0809DFD39" author="Hunt, R. M., Jr." box="[990,1174,1267,1289]" pageId="22" pageNumber="23" pagination="1 - 45" refId="ref47153" refString="Hunt, R. M., Jr. 2005. An early Miocene domeskulled chalicothere from the '' Arikaree' ' conglomerates of Darton: calibrating the ages of High Plains paleovalleys against Rocky Mountain tectonism. American Museum Novitates 3486: 1 - 45." type="journal article" year="2005">Hunt (2005: 32</bibRefCitation>
, fig. 22). From indurated sandstone caprock at
<taxonomicName id="4C564D36FFFFF34FA3D57C3A87CAFD72" authority="Butte." authorityName="Butte." box="[701,961,1326,1348]" class="Mammalia" family="Merycoidodontidae" genus="Merycochoerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">Merycochoerus Butte.</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>