160 lines
18 KiB
XML
160 lines
18 KiB
XML
<document id="AED5614D065F18A083E7F1E1B470EE97" ID-DOI="10.5281/zenodo.5160708" ID-ISSN="1942-1354" ID-Zenodo-Dep="5160708" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628109944637" checkinUser="felipe" docAuthor="Warner, William B." docDate="2011" docId="03D587FC172CE02DFF66142C363E8E3D" docLanguage="en" docName="InsectaMundi.0172.1-17.pdf" docOrigin="Insecta Mundi 2011 (172)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Acoma quadrilaminata Warner 2011, new species" docType="treatment" docVersion="3" lastPageNumber="5" masterDocId="FFECFF841729E02BFFA6106F32548D66" masterDocTitle="Four new species of Acoma Casey, with a key to species in the genus (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="4" updateTime="1699034678431" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="CDB9007565A7CADD63068B8BADFC17B6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="920E0BA6A2D080DF071A71739DFA1EE2">
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<mods:title id="2020BB3DA324F12CC55B11294AE2257D">Four new species of Acoma Casey, with a key to species in the genus (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="03CDD2A7E2900BF3C5548443AEAE53F9" type="personal">
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<mods:role id="81B89168C81048164004F46C1041F8C0">
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<mods:roleTerm id="C6DF0E37F7EE69D9309CE890E0B9EEEE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F3DD50BA6B32809FBEC255B6FF874D4F">Warner, William B.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A05FCC9FE4DF7140FD904073AD993A5E">text</mods:typeOfResource>
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<mods:titleInfo id="95C70DB269318B7D93E9E6D240431724">
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<mods:title id="91F30D11E17B4D4E624C34EFF7CE6C18">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="06AD4186158846C5E1F01E31C5626F89">
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<mods:date id="0159555772AA40D6BAEF00506548DFEA">2011</mods:date>
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<mods:detail id="F7373BDD603427A737E6DA6B86AA19F5" type="pubDate">
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<mods:number id="C25F6057EF0CCDFC6F66584BF77C67E4">2011-04-22</mods:number>
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</mods:detail>
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<mods:detail id="20470D31D8D912CC1061A648BE3D1430" type="volume">
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<mods:number id="6F9856B28A9F40CB6201C6F9811D82D0">2011</mods:number>
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</mods:detail>
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<mods:detail id="9A557261D7EDE800EA9F66054F25401B" type="issue">
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<mods:number id="F0AF8A95BCF0A1BA3CAB6DCAFD7BE8D2">172</mods:number>
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</mods:detail>
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<mods:extent id="37FDCC01470927759BCCF7ACCFE00AF1" unit="page">
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<mods:start id="58B1A13654B9E5788789893610F9A34B">1</mods:start>
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<mods:end id="E548CE545F0BBA53BC37F2679406A810">17</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="2B8B6583E7C9E477E60D8BD9D71A24C2">journal article</mods:classification>
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<mods:identifier id="A0A15AE17914378A1584059FA1FAC83F" type="DOI">10.5281/zenodo.5160708</mods:identifier>
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<mods:identifier id="3FBFD4EC5C4D9342FF8071EF8B2269CF" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="E28334B54F4F3DE136CCB915C1B7F67D" type="Zenodo-Dep">5160708</mods:identifier>
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</mods:mods>
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<treatment id="03D587FC172CE02DFF66142C363E8E3D" LSID="urn:lsid:plazi:treatment:03D587FC172CE02DFF66142C363E8E3D" httpUri="http://treatment.plazi.org/id/03D587FC172CE02DFF66142C363E8E3D" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
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<subSubSection id="C3666561172CE02EFF66142C30FC893D" box="[192,680,1090,1115]" pageId="5" pageNumber="4" type="nomenclature">
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<paragraph id="8BC336EA172CE02EFF66142C30FC893D" blockId="5.[192,680,1090,1147]" box="[192,680,1090,1115]" pageId="5" pageNumber="4">
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<heading id="D08B8186172CE02EFF66142C30FC893D" bold="true" box="[192,680,1090,1115]" fontSize="10" level="2" pageId="5" pageNumber="4" reason="3">
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<emphasis id="B908EAF8172CE02EFF66142C30FC893D" bold="true" box="[192,680,1090,1115]" pageId="5" pageNumber="4">
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<taxonomicName id="4C7C4D69172CE02EFF66142C3055893D" authority="Warner, 2011" authorityName="Warner" authorityYear="2011" box="[192,513,1090,1115]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="quadrilaminata" status="sp. nov.">
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<emphasis id="B908EAF8172CE02EFF66142C3055893D" bold="true" box="[192,513,1090,1115]" italics="true" pageId="5" pageNumber="4">Acoma quadrilaminata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23B5783172CE02EFDA0142C30FC893D" box="[518,680,1091,1115]" pageId="5" pageNumber="4" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3666561172CE02EFF66140C3330891D" box="[192,356,1123,1147]" pageId="5" pageNumber="4" type="description">
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<paragraph id="8BC336EA172CE02EFF66140C3330891D" blockId="5.[192,680,1090,1147]" box="[192,356,1123,1147]" pageId="5" pageNumber="4">
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<figureCitation id="13472A6F172CE02EFF66140C3330891D" box="[192,356,1123,1147]" captionStart="Figure 10-13" captionStartId="13.[192,276,1651,1673]" captionTargetBox="[192,1440,192,1628]" captionTargetId="figure-14@13.[192,1440,192,1628]" captionTargetPageId="13" captionText="Figure 10-13. Acoma quadrilaminata Warner: 10) Dorsal habitus. 11) Antenna. 12) Paramera. 13) Dorsolateral view of head and pronotum." figureDoi="http://doi.org/10.5281/zenodo.5160718" httpUri="https://zenodo.org/record/5160718/files/figure.png" pageId="5" pageNumber="4">Figures 10-13</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3666561172CE02EFF6614CC311B887D" pageId="5" pageNumber="4" type="materials_examined">
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<paragraph id="8BC336EA172CE02EFF6614CC33E489BD" blockId="5.[192,1440,1187,1308]" pageId="5" pageNumber="4">
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<emphasis id="B908EAF8172CE02EFF6614CC33DF89DD" bold="true" box="[192,395,1187,1211]" pageId="5" pageNumber="4">Type Material.</emphasis>
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<materialsCitation id="3B143CB7172CE02EFE3714CC33E489BD" collectingDate="1973-07-07" collectionCode="UNAM" country="Mexico" location="Las Arrastras" pageId="5" pageNumber="4" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54C78848172CE02EFE3714CC305589DD" box="[401,513,1187,1211]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="ED6DAE2F172CE02EFDF414CC30E489DD" box="[594,688,1187,1211]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="5" pageNumber="4">UNAM</collectionCode>
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): “
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<collectingCountry id="F36B767A172CE02EFD6B14CC314589DD" box="[717,785,1187,1211]" name="Mexico" pageId="5" pageNumber="4">MEX</collectingCountry>
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: Baja Calif. Norte,
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<quantity id="4C849B0F172CE02EFC5F14CC366289DD" box="[1017,1078,1187,1211]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="5" pageNumber="4" unit="mi" value="3.0">3 mi.</quantity>
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N
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<location id="8EA36031172CE02EFBFE14CB375389DD" LSID="urn:lsid:plazi:treatment:03D587FC172CE02DFF66142C363E8E3D:8EA36031172CE02EFBFE14CB375389DD" box="[1112,1287,1187,1211]" country="Mexico" name="Las Arrastras" pageId="5" pageNumber="4">Las Arrastras</location>
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,
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<date id="FFC2102A172CE02EFAB414CC37C889DD" box="[1298,1436,1187,1211]" pageId="5" pageNumber="4" value="1973-07-07">
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<collectingDate id="EF86E9C2172CE02EFAB414CC37C889DD" box="[1298,1436,1187,1211]" pageId="5" pageNumber="4" value="1973-07-07">VII-7-1973</collectingDate>
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</date>
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, Fisher & Westcott.”
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</materialsCitation>
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</paragraph>
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<paragraph id="8BC336EA172CE02EFF56148B316B899D" blockId="5.[192,1440,1187,1308]" box="[240,831,1251,1276]" pageId="5" pageNumber="4">
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<typeStatus id="54C78848172CE02EFF56148B3324899A" box="[240,368,1252,1276]" pageId="5" pageNumber="4" type="paratype">Paratypes</typeStatus>
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, (
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<specimenCount id="9D7AFD63172CE02EFE25148C33B4899D" box="[387,480,1251,1275]" pageId="5" pageNumber="4" type="male">5 males</specimenCount>
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) with data same as
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<typeStatus id="54C78848172CE02EFD68148C316E899D" box="[718,826,1251,1275]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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<paragraph id="8BC336EA172CE02EFF56156B311B887D" blockId="5.[192,1440,1187,1308]" box="[240,847,1283,1308]" pageId="5" pageNumber="4">
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<typeStatus id="54C78848172CE02EFF56156B3338887A" box="[240,364,1284,1308]" pageId="5" pageNumber="4" type="paratype">Paratypes</typeStatus>
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are deposited at
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<collectionCode id="ED6DAE2F172CE02EFD91156C30C1887D" box="[567,661,1283,1307]" pageId="5" pageNumber="4">WBWC</collectionCode>
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,
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<collectionCode id="ED6DAE2F172CE02EFD38156B30A7887D" box="[670,755,1284,1307]" pageId="5" pageNumber="4">RLWE</collectionCode>
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,
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<collectionCode id="ED6DAE2F172CE02EFD5B156C311F887D" box="[765,843,1283,1307]" pageId="5" pageNumber="4">CASC</collectionCode>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3666561172CE02EFF66152C37F488FD" pageId="5" pageNumber="4" type="diagnosis">
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<paragraph id="8BC336EA172CE02EFF66152C37F488FD" blockId="5.[192,1440,1347,1435]" pageId="5" pageNumber="4">
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<emphasis id="B908EAF8172CE02EFF66152C331A883D" bold="true" box="[192,334,1347,1371]" pageId="5" pageNumber="4">Diagnosis.</emphasis>
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Medium brown, vestiture light colored and contrasting with cuticle color, pronotum glabrous except for marginal fimbriae; elytral disc hairy; clypeal disc nearly impunctate; antenna 9-segmented, with 4-segmented club, all club lamellae of equal size; anterior tibia tridentate; paramera as in
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<figureCitation id="13472A6F172CE02EFA8015EC37C888FD" box="[1318,1436,1411,1435]" captionStart="Figure 10-13" captionStartId="13.[192,276,1651,1673]" captionTargetBox="[192,1440,192,1628]" captionTargetId="figure-14@13.[192,1440,192,1628]" captionTargetPageId="13" captionText="Figure 10-13. Acoma quadrilaminata Warner: 10) Dorsal habitus. 11) Antenna. 12) Paramera. 13) Dorsolateral view of head and pronotum." figureDoi="http://doi.org/10.5281/zenodo.5160718" httpUri="https://zenodo.org/record/5160718/files/figure.png" pageId="5" pageNumber="4">Figure 12</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3666561172CE02DFF6615AC332C8F1D" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="description">
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||
<paragraph id="8BC336EA172CE02EFF6615AC371488BD" blockId="5.[192,1441,1475,1915]" box="[192,1344,1475,1499]" pageId="5" pageNumber="4">
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<emphasis id="B908EAF8172CE02EFF6615AC333888BD" bold="true" box="[192,364,1475,1499]" pageId="5" pageNumber="4">Description.</emphasis>
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<typeStatus id="54C78848172CE02EFED415AC33B688BD" box="[370,482,1475,1499]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
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male, length:
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<quantity id="4C849B0F172CE02EFD2915AC30B988BD" box="[655,749,1475,1499]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="5" pageNumber="4" unit="mm" value="6.9">6.9 mm</quantity>
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, width:
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<quantity id="4C849B0F172CE02EFCEA15AC31FF88BD" box="[844,939,1475,1499]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="4" unit="mm" value="3.1">3.1 mm</quantity>
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, widest at about middle of elytra.
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</paragraph>
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<paragraph id="8BC336EA172CE02DFF56158C33DC8CDD" blockId="5.[192,1441,1475,1915]" lastBlockId="6.[192,1440,195,443]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
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BODY medium brown, shiny, hairs lighter colored and contrasting with ground color. HEAD with front and clypeal base scabrously, subreticulately punctate-rugose, weakly convex, more noticeably so on frontoclypeal transition; clypeus broadly parabolic, apically weakly emarginate, obtusely subdentate at each lateral terminus of emargination, margins moderately reflexed, only little more strongly so apically, disc concave, deeply punctate, punctures mostly separated by less than their own diameters; antenna 9- segmented, club four segmented, segment 5 (basal to club) unusually elongate, in form of flattened cylin- der and about as long as segments 2-4 combined. PRONOTUM glabrous except for marginal bead, length about 0.4 times elytral length, widest at about middle; lateral margins straight in posterior half, converging in anterior half, slightly sinuating before anterior pronotal angles which are slightly acute and drawn forward next to eyes; disc glabrous, moderately densely and distinctly punctate, punctures mostly separated by about 1 to 2 times their own widths, interpunctural integument weakly shagreened with micropunctures, midline impunctate and weakly impressed. SCUTELLUM subtriangular, apex broadly rounded. ELYTRON with disc setigerous, setal length about 1/4 to nearly 1/2 of elytron width; strial channels somewhat irregular but distinct, strial punctures obsolescent, intervals very irregularly uniseriately punctate, punctures distinct, mostly separated by about 2-4 times their own diameters, about half as wide as intervals. PYGIDIUM longer than broad, subscutelliform, apex arcuate, disc shagreened, shallowly subcontiguously punctate, punctures setigerous. LEGS: Anterior tibia moderately robust, tridentate, basal tooth about half as large as strong middle tooth; metatibia with widest corbel diameter little less than half pygidial length, posterior tarsal claws over 1/3 length of segment from which they arise. GENI- TALIA: paramera as in
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<figureCitation id="13472A6F172FE02DFE4811EC30138CFD" box="[494,583,387,411]" captionStart="Figure 10-13" captionStartId="13.[192,276,1651,1673]" captionTargetBox="[192,1440,192,1628]" captionTargetId="figure-14@13.[192,1440,192,1628]" captionTargetPageId="13" captionText="Figure 10-13. Acoma quadrilaminata Warner: 10) Dorsal habitus. 11) Antenna. 12) Paramera. 13) Dorsolateral view of head and pronotum." figureDoi="http://doi.org/10.5281/zenodo.5160718" httpUri="https://zenodo.org/record/5160718/files/figure.png" pageId="6" pageNumber="5">Fig. 12</figureCitation>
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, with apex only weakly deflexed, ventrobasal emargination shallow, moderately long.
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</paragraph>
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<paragraph id="8BC336EA172FE02DFF66118C332C8F1D" blockId="6.[192,1441,483,635]" pageId="6" pageNumber="5">
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<emphasis id="B908EAF8172FE02DFF66118C33198C9D" bold="true" box="[192,333,483,507]" pageId="6" pageNumber="5">Variation.</emphasis>
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Length:
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<quantity id="4C849B0F172FE02DFE1F118C30018C9D" box="[441,597,483,507]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" metricValueMax="8.3" metricValueMin="6.5" pageId="6" pageNumber="5" unit="mm" value="7.4" valueMax="8.3" valueMin="6.5">6.5 – 8.3 mm</quantity>
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, width: 3.0 – 4.0 mm. Clypeal form varies moderately in punctuation, depth of discal concavity, breadth and strength of apical emargination, but shape (lateral margins weakly convex) is similar. Medial mesotibial fringe of denticles varies from 7-12 (usually 9). In
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<specimenCount id="9D7AFD63172FE02DFB77124B37278F5D" box="[1233,1395,547,571]" pageId="6" pageNumber="5" type="generic">one specimen</specimenCount>
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the anterior pronotal angles are nearly right. One
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<typeStatus id="54C78848172FE02DFD45122B311B8F3A" box="[739,847,580,604]" pageId="6" pageNumber="5" type="paratype">paratype</typeStatus>
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has the left antenna deformed, with 3-segmented, shortened club.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3666561172FE02DFF6612CC364C8FDD" box="[192,1048,675,699]" pageId="6" pageNumber="5" type="etymology">
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||
<paragraph id="8BC336EA172FE02DFF6612CC364C8FDD" blockId="6.[192,1048,675,699]" box="[192,1048,675,699]" pageId="6" pageNumber="5">
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<emphasis id="B908EAF8172FE02DFF6612CC33088FDD" bold="true" box="[192,348,675,699]" pageId="6" pageNumber="5">Etymology.</emphasis>
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This species is named for its 4-segmented antennal clubs.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3666561172FE02DFF66128C363E8E3D" pageId="6" pageNumber="5" type="discussion">
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<paragraph id="8BC336EA172FE02DFF66128C363E8E3D" blockId="6.[192,1440,739,860]" pageId="6" pageNumber="5">
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<emphasis id="B908EAF8172FE02DFF66128C33108F9D" bold="true" box="[192,324,739,763]" pageId="6" pageNumber="5">Remarks.</emphasis>
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This species keys to
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<taxonomicName id="4C7C4D69172FE02DFD9A128B31418F9D" authority="Saylor" authorityName="Saylor" authorityYear="1948" box="[572,789,739,763]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="dilemma">
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<emphasis id="B908EAF8172FE02DFD9A128B30958F9D" box="[572,705,739,763]" italics="true" pageId="6" pageNumber="5">A dilemma</emphasis>
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Saylor
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</taxonomicName>
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in
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<bibRefCitation id="EFED4B1B172FE02DFC9F128C318B8F9D" author="Cazier, M. A." box="[825,991,739,763]" pageId="6" pageNumber="5" pagination="1 - 13" refId="ref7057" refString="Cazier, M. A. 1953. A review of the scarab genus Acoma (Coleoptera: Scarabaeidae). American Museum Novitates, No. 1624: 1 - 13." type="book chapter" year="1953">Cazier (1953)</bibRefCitation>
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.
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<bibRefCitation id="EFED4B1B172FE02DFC4E128C36DE8F9D" author="Saylor, L. W." box="[1000,1162,739,763]" pageId="6" pageNumber="5" pagination="337 - 374" refId="ref7155" refString="Saylor, L. W. 1948. Contributions toward a knowledge of the insect fauna of Lower California. No. 10. Coleoptera: Scarabaeidae. Proceedings of the California Academy of Sciences, 4 th Series 24 (10): 337 - 374." type="journal article" year="1948">Saylor (1948)</bibRefCitation>
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diagnosed that species on the combination of the 4-segmented antennal clubs and hairy elytra; however, the basal club segment in specimens of
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<taxonomicName id="4C7C4D69172FE02DFE2D134B30488E5D" authorityName="Saylor" authorityYear="1948" box="[395,540,803,827]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="dilemma">
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<emphasis id="B908EAF8172FE02DFE2D134B30488E5D" box="[395,540,803,827]" italics="true" pageId="6" pageNumber="5">A. dilemma</emphasis>
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</taxonomicName>
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is narrower than and slightly less than half as long as the more apical segments. In
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<taxonomicName id="4C7C4D69172FE02DFEC4132B30138E3D" authorityName="Warner" authorityYear="2011" box="[354,583,835,859]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="quadrilaminata">
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<emphasis id="B908EAF8172FE02DFEC4132B30138E3D" box="[354,583,835,859]" italics="true" pageId="6" pageNumber="5">A. quadrilaminata</emphasis>
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</taxonomicName>
|
||
, all antennal club segments are of equal size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |