treatments-xml/data/03/D5/87/03D58784817DFFBFFDB8FD54FAD0FB33.xml

<document id="2AC7DAFBD3B24F34024AC413BE95DBF3" ID-CLB-Dataset="22393" ID-DOI="10.5852/ejt.2018.438" ID-GBIF-Dataset="34ad2255-e191-4870-83e8-74819ca0fe06" ID-Zenodo-Dep="3785478" ID-ZooBank="00B57E96-225C-4646-9CB8-B3514C4EC7E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588622038275" checkinUser="valdenar" docAuthor="Jia, Fenglong &amp; Tang, Yu-dan" docDate="2018" docId="03D58784817DFFBFFDB8FD54FAD0FB33" docLanguage="en" docName="ejt-438_jia_tang.pdf.imf" docOrigin="European Journal of Taxonomy 438" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Helochares songi Jia &amp; Tang 2018, sp. nov." docType="treatment" docUuid="2B1792C1-5D59-46B2-AE98-3440BA2FED57" docUuidSource="ZooBank" docVersion="6" lastPageNumber="6" masterDocId="FFECFFFC817FFFBAFFC7FF81FFF9FFFF" masterDocTitle="A revision of the Chinese Helochares (s. str.) Mulsant, 1844 (Coleoptera, Hydrophilidae)" masterLastPageNumber="27" masterPageNumber="1" pageNumber="3" updateTime="1698838436565" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D8313D8195B9679A8135C10A4E1FFB9F">A revision of the Chinese Helochares (s. str.) Mulsant, 1844 (Coleoptera, Hydrophilidae)</mods:title>
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<mods:namePart id="4E3F92270969FF45FFBBA1163D0360F3">Jia, Fenglong</mods:namePart>
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<mods:affiliation id="37539F0518E6D2E4F4B98C4299C6124B">Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China.</mods:affiliation>
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<mods:namePart id="F96D8A40B9FFE65C4DB93E52F4BAA3D7">Tang, Yu-dan</mods:namePart>
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<mods:affiliation id="0B4F3DDDC98D3005BF022C672AF959D4">Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China.</mods:affiliation>
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<treatment id="03D58784817DFFBFFDB8FD54FAD0FB33" ID-DOI="http://doi.org/10.5281/zenodo.3794615" ID-GBIF-Taxon="163943952" ID-Zenodo-Dep="3794615" LSID="urn:lsid:zoobank.org:act:2B1792C1-5D59-46B2-AE98-3440BA2FED57" httpUri="http://treatment.plazi.org/id/03D58784817DFFBFFDB8FD54FAD0FB33" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
<subSubSection id="C3666519817DFFB8FDB8FD54FB3FFCEB" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BC33692817DFFB8FDB8FD54FC4DFD0F" blockId="2.[366,1222,725,822]" box="[639,948,725,752]" pageId="2" pageNumber="3">
<heading id="D08B81FE817DFFB8FDB8FD54FC4DFD0F" box="[639,948,725,752]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<taxonomicName id="4C7C4D11817DFFB8FDB8FD54FCABFD0F" ID-CoL="8Y49F" authority="Jia &amp; Tang, 2018" authorityName="Jia &amp; Tang" authorityYear="2018" box="[639,850,725,752]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="songi" status="sp. nov.">
<emphasis id="B908EA80817DFFB8FDB8FD54FCABFD0F" bold="true" box="[639,850,725,752]" italics="true" pageId="2" pageNumber="3">Helochares songi</emphasis>
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<taxonomicNameLabel id="A23B57FB817DFFB8FC9DFD57FC4DFD0F" box="[858,948,726,752]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act: 
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<subSubSection id="C3666519817DFFB8FD7CFC9DFC81FCC9" box="[699,888,796,822]" pageId="2" pageNumber="3" type="description">
<paragraph id="8BC33692817DFFB8FD7CFC9DFC81FCC9" blockId="2.[366,1222,725,822]" box="[699,888,796,822]" pageId="2" pageNumber="3">
<figureCitation id="13472A17817DFFB8FD7CFC9DFCDBFCC9" box="[699,802,796,822]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="2" pageNumber="3">Figs 1–8</figureCitation>
, 
<figureCitation id="13472A17817DFFB8FCE8FC9DFC81FCC9" box="[815,888,796,822]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="2" pageNumber="3">48–49</figureCitation>
</paragraph>
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<subSubSection id="C3666519817DFFB8FF7AFCDCFAF4FAB2" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BC33692817DFFB8FF7AFCDCFECFFC88" blockId="2.[189,310,861,887]" box="[189,310,861,887]" pageId="2" pageNumber="3">
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<emphasis id="B908EA80817DFFB8FF7AFCDCFECFFC88" bold="true" box="[189,310,861,887]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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<paragraph id="8BC33692817DFFB8FF7AFC0DFAF4FAB2" blockId="2.[189,1399,908,1357]" pageId="2" pageNumber="3">
Length 
<quantity id="4C849B77817DFFB8FEDEFC0DFE52FC58" box="[281,427,908,935]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.3" metricValueMin="4.8" pageId="2" pageNumber="3" unit="mm" value="5.05" valueMax="5.3" valueMin="4.8">4.8–5.3 mm</quantity>
, width 
<quantity id="4C849B77817DFFB8FDC3FC0DFD6FFC58" box="[516,662,908,935]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="2" pageNumber="3" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
. Dark brown or black (
<figureCitation id="13472A17817DFFB8FC6AFC0DFC0BFC59" box="[941,1010,908,934]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
), clypeus uniformly dark brown to black (
<figureCitation id="13472A17817DFFB8FEE8FC2EFE81FC35" box="[303,376,943,970]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
), anterior half of lateral margin of pronotum yellowish brown (
<figureCitation id="13472A17817DFFB8FBB6FC2EFAE5FC35" box="[1137,1308,943,970]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="2" pageNumber="3">Figs 1, 3–4, 6</figureCitation>
), elytra uniformly dark brown to black (
<figureCitation id="13472A17817DFFB8FD8DFC52FD6CFC12" box="[586,661,979,1005]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
). Dorsum with uniform dense and coarse ground punctures. Clypeus moderately expanded laterally, with somewhat rounded lateral margin, clearly impressed in front of eyes (
<figureCitation id="13472A17817DFFB8FEACFB98FE4FFBCC" box="[363,438,1049,1075]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
). Maxillary palps ca 1.2× as long as width of head, dark brown to black, each palpomere with light colour apically (
<figureCitation id="13472A17817DFFB8FD4EFBBDFCE1FBA9" box="[649,792,1084,1110]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="2" pageNumber="3">Figs 1–2, 4</figureCitation>
). Pedicel subequal in length to antennomeres 3–6 combined. Submentum with a small glabrous area medially and with a few coarse punctures laterally (
<figureCitation id="13472A17817DFFB8FF01FB02FEF7FB62" box="[198,270,1154,1181]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
). Elytra almost parallel from base to posterior third, somewhat explanate laterally, with distinct systematic punctures (
<figureCitation id="13472A17817DFFB8FE06FB27FDF1FB3F" box="[449,520,1190,1216]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
). Mesoventrite with a low transverse ridge medially and a low longitudinal carina (
<figureCitation id="13472A17817DFFB8FED9FB48FE97FB1B" box="[286,366,1225,1252]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
). Metaventrite without glabrous area medially. Femora densely pubescent except for apical eighth (
<figureCitation id="13472A17817DFFB8FE52FB6DFDFDFAF9" box="[405,516,1260,1286]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="2" pageNumber="3">Figs 2, 8</figureCitation>
). Aedeagus slender, length of paramere ca 10 × as long as width of apex (
<figureCitation id="13472A17817DFFB8FF01FA8EFEDBFAD5" box="[198,290,1295,1322]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="2" pageNumber="3">Fig. 48</figureCitation>
), subparallel, apex with a small tooth inwards (
<figureCitation id="13472A17817DFFB8FCB7FA8EFC07FAD5" box="[880,1022,1295,1322]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="2" pageNumber="3">Figs 48–49</figureCitation>
); apex of median lobe slightly overlapping parameres; basal strut of median lobe long, ca ¼ × as long as parameres (
<figureCitation id="13472A17817DFFB8FB6FFAB2FAF9FAB2" box="[1192,1280,1331,1357]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="2" pageNumber="3">Fig. 48</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3666519817DFFB8FF7AFAF5FB29FA42" box="[189,1232,1396,1469]" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8BC33692817DFFB8FF7AFAF5FEBDFA71" blockId="2.[189,324,1396,1422]" box="[189,324,1396,1422]" pageId="2" pageNumber="3">
<heading id="D08B81FE817DFFB8FF7AFAF5FEBDFA71" bold="true" box="[189,324,1396,1422]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B908EA80817DFFB8FF7AFAF5FEBDFA71" bold="true" box="[189,324,1396,1422]" pageId="2" pageNumber="3">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BC33692817DFFB8FF7AFA22FB29FA42" blockId="2.[189,1232,1443,1469]" box="[189,1232,1443,1469]" pageId="2" pageNumber="3">
The specific name is given in honor of Keqing Song, the collector of the 
<typeStatus id="54C78830817DFFB8FBD3FA22FBBEFA42" box="[1044,1095,1443,1469]" pageId="2" pageNumber="3">type</typeStatus>
specimens.
</paragraph>
</subSubSection>
<subSubSection id="C3666519817DFFB8FF7AFA65FE1DF854" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BC33692817DFFB8FF7AFA65FE53FA01" blockId="2.[189,426,1508,1534]" box="[189,426,1508,1534]" pageId="2" pageNumber="3">
<heading id="D08B81FE817DFFB8FF7AFA65FE53FA01" bold="true" box="[189,426,1508,1534]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B908EA80817DFFB8FF7AFA65FE53FA01" bold="true" box="[189,426,1508,1534]" pageId="2" pageNumber="3">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BC33692817DFFB8FF2DF992FEA5F9D2" blockId="2.[189,1399,1555,1651]" box="[234,348,1555,1581]" pageId="2" pageNumber="3">
<emphasis id="B908EA80817DFFB8FF2DF992FEA5F9D2" bold="true" box="[234,348,1555,1581]" pageId="2" pageNumber="3">
<typeStatus id="54C78830817DFFB8FF2DF992FEA5F9D2" box="[234,348,1555,1581]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC33692817DFFB8FF7AF9B7FD23F98C" blockId="2.[189,1399,1555,1651]" pageId="2" pageNumber="3">
<materialsCitation id="3B143CCF817DFFB8FF7AF9B7FD23F98C" ID-GBIF-Occurrence="2610058766" collectingDate="2011-07-21" collectionCode="SYSU" collectorName="Keqing Song" country="China" elevation="316" location="Nalin River" municipality="Shiwandashan" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<collectingCountry id="F36B7602817DFFB8FF7AF9B7FEE6F9AF" box="[189,287,1590,1616]" name="China" pageId="2" pageNumber="3">CHINA</collectingCountry>
: 1 ♂, 
<collectingRegion id="49B8F870817DFFB8FEACF9B7FDBCF9AF" box="[363,581,1590,1616]" country="China" name="Guangxi" pageId="2" pageNumber="3">Guangxi Province</collectingRegion>
, 
<collectingMunicipality id="6BA7ACE8817DFFB8FD94F9B7FD06F9AF" box="[595,767,1590,1616]" pageId="2" pageNumber="3">Shiwandashan</collectingMunicipality>
, 
<location id="8EA36049817DFFB8FCCAF9B7FC62F9AF" LSID="urn:lsid:plazi:treatment:03D58784817DFFBFFDB8FD54FAD0FB33:8EA36049817DFFB8FCCAF9B7FC62F9AF" box="[781,923,1590,1616]" country="China" municipality="Shiwandashan" name="Nalin River" pageId="2" pageNumber="3" stateProvince="Guangxi">Nalin River</location>
, 
<elevation id="0051D1A1817DFFB8FC6EF9B7FBCEF9AF" box="[937,1079,1590,1616]" metricMagnitude="2" metricUnit="m" metricValue="3.16" pageId="2" pageNumber="3" unit="m" value="316.0">
<quantity id="4C849B77817DFFB8FC6EF9B7FC0FF9AF" box="[937,1014,1590,1616]" metricMagnitude="2" metricUnit="m" metricValue="3.16" pageId="2" pageNumber="3" unit="m" value="316.0">316 m</quantity>
a.s.l.
</elevation>
, in water at edge of upper river, 
<date id="FFC21052817DFFB8FEC5F9D8FE6AF98C" box="[258,403,1625,1651]" pageId="2" pageNumber="3" value="2011-07-21">
<collectingDate id="EF86E9BA817DFFB8FEC5F9D8FE6AF98C" box="[258,403,1625,1651]" pageId="2" pageNumber="3" value="2011-07-21">21 Jul. 2011</collectingDate>
</date>
, 
<collectorName id="26895344817DFFB8FE67F9D8FDC0F98C" box="[416,569,1625,1651]" pageId="2" pageNumber="3">Keqing Song</collectorName>
leg. (
<collectionCode id="ED6DAE57817DFFB8FDBBF9D8FD34F98C" box="[636,717,1625,1651]" pageId="2" pageNumber="3">SYSU</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BC33692817DFFB8FF2DF91AFC94F94A" blockId="2.[189,1399,1691,1963]" box="[234,877,1691,1717]" pageId="2" pageNumber="3">
<emphasis id="B908EA80817DFFB8FF2DF91AFE9EF94A" bold="true" box="[234,359,1691,1717]" pageId="2" pageNumber="3">
<typeStatus id="54C78830817DFFB8FF2DF91AFE9EF94A" box="[234,359,1691,1717]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D7AFD1B817DFFB8FEBFF91AFE25F94A" box="[376,476,1691,1717]" pageId="2" pageNumber="3" type="generic">77 spec.</specimenCount>
, 
<collectionCode id="ED6DAE57817DFFB8FE2FF91AFDBAF94A" box="[488,579,1691,1717]" pageId="2" pageNumber="3">BMNH</collectionCode>
, 
<collectionCode id="ED6DAE57817DFFB8FD89F91AFD52F94A" box="[590,683,1691,1717]" pageId="2" pageNumber="3">KMNH</collectionCode>
, 
<collectionCode id="ED6DAE57817DFFB8FD70F91AFCF4F94A" box="[695,781,1691,1717]" country="Czech Republic" name="National Museum Prague" pageId="2" pageNumber="3">NMPC</collectionCode>
, 
<collectionCode id="ED6DAE57817DFFB8FCDFF91AFC9AF94A" box="[792,867,1691,1717]" pageId="2" pageNumber="3">SYSU</collectionCode>
)
</paragraph>
<paragraph id="8BC33692817DFFB8FF7AF93FFE1DF854" blockId="2.[189,1399,1691,1963]" pageId="2" pageNumber="3">
<collectingCountry id="F36B7602817DFFB8FF7AF93FFEE6F927" box="[189,287,1726,1752]" name="China" pageId="2" pageNumber="3">CHINA</collectingCountry>
: 
<emphasis id="B908EA80817DFFB8FEEEF93CFE6CF928" bold="true" box="[297,405,1725,1751]" pageId="2" pageNumber="3">
<collectingRegion id="49B8F870817DFFB8FEEEF93CFE6CF928" box="[297,405,1725,1751]" country="China" name="Guangxi" pageId="2" pageNumber="3">Guangxi</collectingRegion>
</emphasis>
: 2 ƋƋ, 
<specimenCount id="9D7AFD1B817DFFB8FE33F93FFDCFF927" box="[500,566,1726,1752]" pageId="2" pageNumber="3" type="female">4 ♀♀</specimenCount>
, 
<specimenCount id="9D7AFD1B817DFFB8FD83F93CFD5FF927" box="[580,678,1725,1752]" pageId="2" pageNumber="3" type="generic">15 spec.</specimenCount>
, same collection data as for 
<typeStatus id="54C78830817DFFB8FC37F93FFBA3F927" box="[1008,1114,1726,1752]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
; 
<specimenCount id="9D7AFD1B817DFFB8FBA1F93FFB43F927" box="[1126,1210,1726,1752]" pageId="2" pageNumber="3" type="generic">3 spec.</specimenCount>
, Shiwandashan, Forest Park, 
<quantity id="4C849B77817DFFB8FE94F960FE67F904" box="[339,414,1761,1787]" metricMagnitude="2" metricUnit="m" metricValue="2.3899999999999997" pageId="2" pageNumber="3" unit="m" value="239.0">239 m</quantity>
a.s.l., edge of upper river, 
<date id="FFC21052817DFFB8FD1DF960FC94F904" box="[730,877,1761,1787]" pageId="2" pageNumber="3" value="2011-07-19">19 Jul. 2011</date>
, Keqing Song leg.; 
<specimenCount id="9D7AFD1B817DFFB8FB90F960FB40F904" box="[1111,1209,1761,1787]" pageId="2" pageNumber="3" type="generic">11 spec.</specimenCount>
, Shiwandashan, Forest Park, edge of river, 
<date id="FFC21052817DFFB8FDD6F885FD45F8E1" box="[529,700,1796,1822]" pageId="2" pageNumber="3" value="2011-08-17">17 Aug. 2011</date>
, Keqing Song leg.; 
<specimenCount id="9D7AFD1B817DFFB8FC7DF885FBEDF8E0" box="[954,1044,1796,1823]" pageId="2" pageNumber="3" type="generic">3 spec.</specimenCount>
, Shiwandashan, Forest Park, 
<quantity id="4C849B77817DFFB8FF7AF8A6FEF6F8BD" box="[189,271,1831,1858]" metricMagnitude="2" metricUnit="m" metricValue="2.21" pageId="2" pageNumber="3" unit="m" value="221.0">221 m</quantity>
a.s.l., in water at edge of lower stone river, 
<date id="FFC21052817DFFB8FC98F8A6FBF9F8BE" box="[863,1024,1831,1857]" pageId="2" pageNumber="3" value="2011-07-12">12 Jul. 2011</date>
, Keqing Song leg.; 
<specimenCount id="9D7AFD1B817DFFB8FAC0F8A6FA8BF8BD" box="[1287,1394,1831,1858]" pageId="2" pageNumber="3" type="generic">20 spec.</specimenCount>
, Shiwandashan, 
<quantity id="4C849B77817DFFB8FEB2F8CAFE46F89A" box="[373,447,1867,1893]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="3" unit="m" value="430.0">430 m</quantity>
a.s.l., edge of Nalinhe River (in water), 
<date id="FFC21052817DFFB8FC50F8CAFBD1F89A" box="[919,1064,1866,1893]" pageId="2" pageNumber="3" value="2011-07-21">21 Jul. 2011</date>
, Keqing Song leg.; 
<specimenCount id="9D7AFD1B817DFFB8FAC8F8CAFA8BF89A" box="[1295,1394,1867,1893]" pageId="2" pageNumber="3" type="generic">19 spec.</specimenCount>
, Shiwandashan, 
<quantity id="4C849B77817DFFB8FEB2F8ECFE39F877" box="[373,448,1901,1928]" metricMagnitude="2" metricUnit="m" metricValue="2.69" pageId="2" pageNumber="3" unit="m" value="269.0">269 m</quantity>
a.s.l., edge of river outside Forest Park, 
<date id="FFC21052817DFFB8FC5CF8EFFBD4F877" box="[923,1069,1902,1928]" pageId="2" pageNumber="3" value="2011-07-20">20 Jul. 2011</date>
, Keqing Song leg. (
<collectionCode id="ED6DAE57817DFFB8FAD2F8EFFA8BF877" box="[1301,1394,1902,1928]" pageId="2" pageNumber="3">BMNH</collectionCode>
, 
<collectionCode id="ED6DAE57817DFFB8FF7AF810FEE3F854" box="[189,282,1937,1963]" pageId="2" pageNumber="3">KMNH</collectionCode>
, 
<collectionCode id="ED6DAE57817DFFB8FEE2F810FE82F854" box="[293,379,1937,1963]" country="Czech Republic" name="National Museum Prague" pageId="2" pageNumber="3">NMPC</collectionCode>
, 
<collectionCode id="ED6DAE57817DFFB8FE40F810FE2EF854" box="[391,471,1937,1963]" pageId="2" pageNumber="3">SYSU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3666519817CFFB9FF7AFE8AFC1BF8D6" pageId="3" pageNumber="4" type="description">
<paragraph id="8BC33692817CFFB9FF7AFE8AFEB7FEDA" blockId="3.[189,334,267,293]" box="[189,334,267,293]" pageId="3" pageNumber="4">
<heading id="D08B81FE817CFFB9FF7AFE8AFEB7FEDA" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B908EA80817CFFB9FF7AFE8AFEB7FEDA" bold="true" box="[189,334,267,293]" pageId="3" pageNumber="4">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BC33692817CFFB9FF7AFEBBFA8FFDFB" blockId="3.[189,1399,313,516]" pageId="3" pageNumber="4">
FORM AND COLOUR. Body oval, length 
<quantity id="4C849B77817CFFB9FDB9FEB8FCE9FEAB" box="[638,784,313,340]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.3" metricValueMin="4.8" pageId="3" pageNumber="4" unit="mm" value="5.05" valueMax="5.3" valueMin="4.8">4.8–5.3 mm</quantity>
, width 
<quantity id="4C849B77817CFFB9FCAEFEB8FC05FEAB" box="[873,1020,313,340]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="3" pageNumber="4" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
, moderately convex. Head dark brown to black. Pronotum dark brown to black, anterior half with narrow yellowish brown lateral margins (
<figureCitation id="13472A17817CFFB9FF02FE01FE90FE64" box="[197,361,384,411]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Figs 1, 3–4, 6</figureCitation>
). Elytra uniformly dark brown to black (
<figureCitation id="13472A17817CFFB9FC82FE01FC54FE65" box="[837,941,384,410]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Figs 1, 3</figureCitation>
). Maxillary palps dark brown to black, each palpomere light yellowish brown apically (
<figureCitation id="13472A17817CFFB9FCC6FE22FC74FE42" box="[769,909,419,445]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Figs 1–2, 4</figureCitation>
). Labial palpomeres yellow to yellowish brown. Antennae yellowish brown, club with dense yellow setae. Ventral side dark brown or black (
<figureCitation id="13472A17817CFFB9FF01FE68FEF4FDFB" box="[198,269,489,516]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
), legs with same colour as ventral side, but with reddish to yellowish brown tarsomeres (
<figureCitation id="13472A17817CFFB9FAE7FE68FA91FDFB" box="[1312,1384,489,516]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BC33692817CFFB9FF7AFDAAFE07FBF0" blockId="3.[189,1399,555,1039]" pageId="3" pageNumber="4">
HEAD. Labrum with fine and dense punctures, smooth between punctures, anterior margin broadly emarginate. Clypeus broad, anterior margin clearly broadly emarginate, moderately expanded in front of eyes, with somewhat rounded lateral margin (
<figureCitation id="13472A17817CFFB9FCCBFDF0FCAEFD74" box="[780,855,625,651]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
), clearly impressed in front of eyes (
<figureCitation id="13472A17817CFFB9FADBFDF0FA90FD74" box="[1308,1385,625,651]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
). Ground punctures on clypeus coarser and sparser than those on labrum, surface smooth between punctures. Systematic punctures on clypeus almost same size as ground punctures. Eyes of moderate size, clearly emarginate anteriorly (
<figureCitation id="13472A17817CFFB9FDA1FD5AFD2CFD0A" box="[614,725,731,757]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Figs 1, 4</figureCitation>
), separated by ca 4.0–4.1 × the width of one eye. Frons with slightly coarser and stronger punctures than on clypeus, with clear systematic punctures. Maxillary palps long and slender, ca 1.1–1.2 × as wide as head, second and third palpomeres curved inwards, apical palpomere almost as long as penultimate, asymmetrical (
<figureCitation id="13472A17817CFFB9FC9CFCC5FC5CFCA0" box="[859,933,836,863]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
). Antennae with 9 antennomeres, scape ca 1.5× as long as pedicel, pedicel subequal in length to antennomeres 3–6 combined, club loosely segmented, with dense pubescence. 
<collectingRegion id="49B8F870817CFFB9FDBEFC0AFD25FC5A" box="[633,732,907,933]" country="Myanmar" name="Chin" pageId="3" pageNumber="4">Mentum</collectingRegion>
ca as wide as long, subquadrate, with deep v-shaped impression anteromedially, with coarse punctures and wrinkles posteriorly (
<figureCitation id="13472A17817CFFB9FBFEFC2FFB7BFC36" box="[1081,1154,942,969]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
). Submentum with a small glabrous area medially and with a few coarse punctures laterally. Maxilla with coarse punctures, smooth between punctures.
</paragraph>
<paragraph id="8BC33692817CFFB9FF7AFBB7FE85F9C2" blockId="3.[189,1399,1078,1597]" pageId="3" pageNumber="4">
THORAX. Pronotum ca 2× as wide as long, widest posteriorly, posterolateral margins broadly round, anterior margin smooth, with a very fine transverse groove, lateral margin with stronger groove, posterior margin without such groove; posterior margin almost straight. Ground punctures slightly coarser than those on head, systematic punctures on pronotum distinct, almost of same size as ground punctures (
<figureCitation id="13472A17817CFFB9FF02FB42FECEFB21" box="[197,311,1219,1246]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Figs 4, 6</figureCitation>
). Prosternum bulging in middle, not carinate, pubescent, with transverse groove (
<figureCitation id="13472A17817CFFB9FADBFB42FA9EFB22" box="[1308,1383,1218,1245]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
), protruding anteriorly. Mesoventrite with a low transverse ridge medially, which does not bear an elevated tooth or projection but with long setae (
<figureCitation id="13472A17817CFFB9FCC1FA88FCA9FADB" box="[774,848,1289,1316]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
); behind ridge with a low longitudinal carina. Metaventrite pubescent, with somewhat convex middle portion, without glabrous area. Metepisterna ca 3.5× as long as wide, subparallel. Scutellum triangular, with similar punctures to elytra. Elytra almost parallel from base to posterior third, somewhat explanate laterally, ground punctures similar to those on pronotum, with 3 distinct rows of systematic punctures (
<figureCitation id="13472A17817CFFB9FC55FA17FC25FA4F" box="[914,988,1430,1456]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
), without sutural stria; epipleuron very broad anteriorly, reaching posterior margin of first visible abdominal sternite, with sparse, coarse punctures. Femora densely pubescent, with apical eighth glabrous (
<figureCitation id="13472A17817CFFB9FC37FA5DFB98FA09" box="[1008,1121,1500,1526]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Figs 2, 8</figureCitation>
). Metatarsomeres with dense white hairs ventrally and a few long swimming hairs dorsally. Claws of moderate size, rather strongly curved.
</paragraph>
<paragraph id="8BC33692817CFFB9FF7AF9E5FA97F981" blockId="3.[189,1390,1636,1663]" box="[189,1390,1636,1663]" pageId="3" pageNumber="4">
ABDOMEN. All visible abdominal ventrites with dense pubescence; first ventrite not carinate (
<figureCitation id="13472A17817CFFB9FB32F9E5FAA7F981" box="[1269,1374,1636,1662]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Figs 2, 8</figureCitation>
).
</paragraph>
<paragraph id="8BC33692817CFFB9FF7AF927FC1BF8D6" blockId="3.[189,1398,1701,1834]" pageId="3" pageNumber="4">
AEDEAGUS. Slender, length of parameres ca 10 × as long as width of apex, subparallel, a little expanded subapically and round apically (
<figureCitation id="13472A17817CFFB9FDEAF948FD7AF91C" box="[557,643,1737,1763]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="3" pageNumber="4">Fig. 48</figureCitation>
), apex with a small, inwardly directed tooth (
<figureCitation id="13472A17817CFFB9FB41F948FAF7F91C" box="[1158,1294,1737,1763]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="3" pageNumber="4">Figs 48–49</figureCitation>
); median lobe slightly protruding beyond parameres, gradually narrowed apically, apex ca 0.5× as wide as apex of parameres; basal strut long, ca ¼ × as long as parameres (
<figureCitation id="13472A17817CFFB9FCBAF88EFC2CF8D6" box="[893,981,1807,1833]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="3" pageNumber="4">Fig. 48</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3666519817CFFBFFF7AF8D1FD6DFCC9" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BC33692817CFFB9FF7AF8D1FE30F895" blockId="3.[189,457,1872,1898]" box="[189,457,1872,1898]" pageId="3" pageNumber="4">
<heading id="D08B81FE817CFFB9FF7AF8D1FE30F895" bold="true" box="[189,457,1872,1898]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B908EA80817CFFB9FF7AF8D1FE30F895" bold="true" box="[189,457,1872,1898]" pageId="3" pageNumber="4">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BC33692817CFFBFFF7AF8FEFD6DFCC9" blockId="3.[189,1398,1919,2016]" lastBlockId="5.[189,1399,267,822]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
This species is very similar to 
<taxonomicName id="4C7C4D11817CFFB9FDE5F801FC41F866" authority="d'Orchymont, 1932" authorityName="d'Orchymont" authorityYear="1932" box="[546,952,1919,1945]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B908EA80817CFFB9FDE5F801FD3EF866" box="[546,711,1919,1945]" italics="true" pageId="3" pageNumber="4">H. fuliginosus</emphasis>
<bibRefCitation id="EFED4B63817CFFB9FD09F8FEFC41F866" author="d'Orchymont A." box="[718,952,1919,1945]" pageId="3" pageNumber="4" pagination="623 - 714" refId="ref16315" refString="d'Orchymont A. 1932. Zur Kenntnis der Kolbenwasserkafer (Palpicornia) von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement IX (Tropische Binnengewasser II): 623 - 714." type="book chapter" year="1932">d’Orchymont, 1932</bibRefCitation>
</taxonomicName>
, but may easily be distinguished from the latter by its larger size (
<quantity id="4C849B77817CFFB9FDCBF823FD67F842" box="[524,670,1954,1981]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.3" metricValueMin="3.6" pageId="3" pageNumber="4" unit="mm" value="3.95" valueMax="4.3" valueMin="3.6">3.6–4.3 mm</quantity>
in 
<taxonomicName id="4C7C4D11817CFFB9FD0FF822FC89F843" authorityName="d'Orchymont" authorityYear="1932" box="[712,880,1954,1980]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B908EA80817CFFB9FD0FF822FC89F843" box="[712,880,1954,1980]" italics="true" pageId="3" pageNumber="4">H. fuliginosus</emphasis>
</taxonomicName>
), lateral margin of clypeus and elytra with same colour as disc (
<figureCitation id="13472A17817CFFB9FE78F844FDF3F81F" box="[447,522,1989,2016]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
), pronotum and elytra more coarsely punctate, and maxillary palps dark brown or black, each palpomere with light colour apically (
<figureCitation id="13472A17817AFFBFFCBAFE8AFC10FED9" box="[893,1001,267,294]" captionStart="Figs 1–8" captionStartId="4.[189,241,1962,1988]" captionTargetBox="[193,1410,264,1930]" captionTargetId="figure@4.[137,1447,253,1936]" captionTargetPageId="4" captionText="Figs 1–8. Helochares songi sp. nov. 1–3. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 4. Head and pronotum. 5. Head and prosternum. 6. Pronotum, lateral. 7. Mesoventrite. 8. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="5" pageNumber="6">Figs 4–5</figureCitation>
). Aedeagus slender, parameres ca 10 × as long as width of apex (
<figureCitation id="13472A17817AFFBFFDE6FEAFFD80FEB6" box="[545,633,302,329]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="5" pageNumber="6">Fig. 48</figureCitation>
), apex with a small inwardly directed tooth (
<figureCitation id="13472A17817AFFBFFB45FEAFFAF5FEB6" box="[1154,1292,302,329]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="5" pageNumber="6">Figs 48–49</figureCitation>
); median lobe slightly projecting beyond parameres apically (
<figureCitation id="13472A17817AFFBFFCDAFED3FC8CFE93" box="[797,885,338,364]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="5" pageNumber="6">Fig. 48</figureCitation>
). It can be distinguished from 
<taxonomicName id="4C7C4D11817AFFBFFB1FFED2FE7AFE70" authority="Regimbart, 1903" authorityName="Regimbart" authorityYear="1903" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="atropiceus">
<emphasis id="B908EA80817AFFBFFB1FFED2FA8EFE93" box="[1240,1399,339,364]" italics="true" pageId="5" pageNumber="6">H. atropiceus</emphasis>
<bibRefCitation id="EFED4B63817AFFBFFF7AFEF4FE7AFE70" author="Regimbart M." box="[189,387,373,399]" pageId="5" pageNumber="6" pagination="52 - 64" refId="ref16408" refString="Regimbart M. 1903. Contribution a la faune Indo-Chinoise. 19 e memoire. Hydrophilidae. Annales de la Societe entomologique de France 72: 52 - 64. Available from https: // biodiversitylibrary. org / page / 8247940 [accessed 6 Apr. 2018]." type="journal article" year="1903">Régimbart, 1903</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7C4D11817AFFBFFE4AFEF7FD3CFE70" authority="Hebauer, 1999" authorityName="Hebauer" authorityYear="1999" box="[397,709,373,399]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ciniensis">
<emphasis id="B908EA80817AFFBFFE4AFEF7FDEFFE70" box="[397,534,374,399]" italics="true" pageId="5" pageNumber="6">H. ciniensis</emphasis>
Hebauer, 1999
</taxonomicName>
and 
<taxonomicName id="4C7C4D11817AFFBFFD3DFEF7FBB0FE70" authority="Sharp, 1890" authorityName="Sharp" authorityYear="1890" box="[762,1097,373,399]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="taprobanicus">
<emphasis id="B908EA80817AFFBFFD3DFEF7FC4FFE70" box="[762,950,373,399]" italics="true" pageId="5" pageNumber="6">H. taprobanicus</emphasis>
Sharp, 1890
</taxonomicName>
, occurring in the Oriental, by its slender aedeagus, parameres ca 10× as long as width of apex, without branches, not concave outside, apex with a small inwardly directed tooth (
<figureCitation id="13472A17817AFFBFFCDDFE3AFC5DFE2A" box="[794,932,443,469]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="5" pageNumber="6">Figs 48–49</figureCitation>
). 
<bibRefCitation id="EFED4B63817AFFBFFC70FE3AFB88FE29" author="Hebauer F." box="[951,1137,443,470]" pageId="5" pageNumber="6" pagination="3 - 14" refId="ref15291" refString="Hebauer F. 1995. Neues zu den Acidocerina Hansen (Helocharae d'Orchymont) der indomalaiischen Region (Coleoptera, Hydrophilidae). Acta Coleopterologica 11 (3): 3 - 14." type="journal article" year="1995">Hebauer (1995)</bibRefCitation>
described 
<taxonomicName id="4C7C4D11817AFFBFFB37FE3AFED4FE06" authorityName="Hebauer" authorityYear="1995" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fulgurans">
<emphasis id="B908EA80817AFFBFFB37FE3AFED4FE06" italics="true" pageId="5" pageNumber="6">Helochares fulgurans</emphasis>
</taxonomicName>
based on a unique female from 
<collectingCountry id="F36B7602817AFFBFFD07FE5FFCD5FE07" box="[704,812,478,504]" name="Thailand" pageId="5" pageNumber="6">Thailand</collectingCountry>
. 
<taxonomicName id="4C7C4D11817AFFBFFCFCFE5EFBF5FE07" authority="Jia &amp; Tang, 2018" authorityName="Jia &amp; Tang" authorityYear="2018" box="[827,1036,479,505]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="songi" status="sp. nov.">
<emphasis id="B908EA80817AFFBFFCFCFE5EFBF5FE07" box="[827,1036,479,505]" italics="true" pageId="5" pageNumber="6">Helochares songi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57FB817AFFBFFBD0FE5EFB8CFE06" box="[1047,1141,479,505]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from 
<taxonomicName id="4C7C4D11817AFFBFFF38FD82FE6DFDE3" authorityName="Hebauer" authorityYear="1995" box="[255,404,514,540]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fulgurans">
<emphasis id="B908EA80817AFFBFFF38FD82FE6DFDE3" box="[255,404,514,540]" italics="true" pageId="5" pageNumber="6">H. fulgurans</emphasis>
</taxonomicName>
by its larger size (
<quantity id="4C849B77817AFFBFFDB5FD83FD36FDE3" box="[626,719,514,541]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="6" unit="mm" value="3.5">3.5 mm</quantity>
in 
<taxonomicName id="4C7C4D11817AFFBFFD30FD82FC75FDE3" authorityName="Hebauer" authorityYear="1995" box="[759,908,514,540]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fulgurans">
<emphasis id="B908EA80817AFFBFFD30FD82FC75FDE3" box="[759,908,514,540]" italics="true" pageId="5" pageNumber="6">H. fulgurans</emphasis>
</taxonomicName>
), dark colour (bright yellow brown, only dark between eyes, with two dark basal spots on pronotum in 
<taxonomicName id="4C7C4D11817AFFBFFC62FDA7FBC2FDC0" authorityName="Hebauer" authorityYear="1995" box="[933,1083,549,575]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fulgurans">
<emphasis id="B908EA80817AFFBFFC62FDA7FBC2FDC0" box="[933,1083,549,575]" italics="true" pageId="5" pageNumber="6">H. fulgurans</emphasis>
</taxonomicName>
) and maxillary palps dark brown or black, each palpomere with light colour apically (yellow in 
<taxonomicName id="4C7C4D11817AFFBFFBC1FDC8FB65FD9D" authorityName="Hebauer" authorityYear="1995" box="[1030,1180,584,610]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fulgurans">
<emphasis id="B908EA80817AFFBFFBC1FDC8FB65FD9D" box="[1030,1180,584,610]" italics="true" pageId="5" pageNumber="6">H. fulgurans</emphasis>
</taxonomicName>
). From 
<taxonomicName id="4C7C4D11817AFFBFFAC7FDC8FE5FFD7A" authority="d'Orchymont, 1919" authorityName="d'Orchymont" authorityYear="1919" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B908EA80817AFFBFFAC7FDC8FA8EFD9D" box="[1280,1399,584,610]" italics="true" pageId="5" pageNumber="6">H. vitalisi</emphasis>
d’Orchymont, 1919
</taxonomicName>
(
<figureCitation id="13472A17817AFFBFFE71FDEAFDF4FD79" box="[438,525,619,646]" captionStart="Figs 41–46. – 41–42" captionStartId="20.[189,241,1930,1956]" captionTargetBox="[235,1344,268,1888]" captionTargetId="figure@20.[220,1367,254,1900]" captionTargetPageId="20" captionText="Figs 41–46. – 41–42. Helochares pallens (MacLeay, 1825). Elytra. 41. Dorsal. 42. Ventral. 43, 45. Paratype of H. vitalisi d’Orchymont, 1919. 43. Data of labels. 45. Habitus, dorsal. 44, 46. Paratype of H. fuliginosus d’Orchymont, 1932. 44. Data of labels. 46. Habitus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3785492" httpUri="https://zenodo.org/record/3785492/files/figure.png" pageId="5" pageNumber="6">Fig. 45</figureCitation>
) it differs in having the head, pronotum and elytra with coarser punctures (ground punctures very fine in 
<taxonomicName id="4C7C4D11817AFFBFFDE1FD0EFD63FD56" authorityName="d'Orchymont" authorityYear="1919" box="[550,666,655,681]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B908EA80817AFFBFFDE1FD0EFD63FD56" box="[550,666,655,681]" italics="true" pageId="5" pageNumber="6">H. vitalisi</emphasis>
</taxonomicName>
), each maxillary palpomere with light colour apically, clypeus moderately expanded laterally, with somewhat rounded lateral margin, clearly impressed in front of eyes (clypeus not expanded laterally and not impressed in front of eyes in 
<taxonomicName id="4C7C4D11817AFFBFFBE0FD57FB62FD10" authorityName="d'Orchymont" authorityYear="1919" box="[1063,1179,725,751]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B908EA80817AFFBFFBE0FD57FB62FD10" box="[1063,1179,725,751]" italics="true" pageId="5" pageNumber="6">H. vitalisi</emphasis>
</taxonomicName>
) and elytra almost parallel from base to posterior third (widest at middle in 
<taxonomicName id="4C7C4D11817AFFBFFC9CFD78FC36FCED" authorityName="d'Orchymont" authorityYear="1919" box="[859,975,760,786]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B908EA80817AFFBFFC9CFD78FC36FCED" box="[859,975,760,786]" italics="true" pageId="5" pageNumber="6">H. vitalisi</emphasis>
</taxonomicName>
), somewhat explanate laterally (not explanate in 
<taxonomicName id="4C7C4D11817AFFBFFE94FC9DFE3EFCCA" authorityName="d'Orchymont" authorityYear="1919" box="[339,455,795,821]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="B908EA80817AFFBFFE94FC9DFE3EFCCA" box="[339,455,795,821]" italics="true" pageId="5" pageNumber="6">H. vitalisi</emphasis>
</taxonomicName>
, as in 
<taxonomicName id="4C7C4D11817AFFBFFDD4FC9AFD7AFCCA" authorityName="C.G. Thomson" authorityYear="1859" box="[531,643,795,821]" class="Insecta" family="Hydrophilidae" genus="Enochrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B908EA80817AFFBFFDD4FC9AFD7AFCCA" box="[531,643,795,821]" italics="true" pageId="5" pageNumber="6">Enochrus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF03661A817BFFBEFF7AF82BFBC2F817" ID-DOI="http://doi.org/10.5281/zenodo.3785482" ID-Zenodo-Dep="3785482" httpUri="https://zenodo.org/record/3785482/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,241,1962,1988]" targetBox="[193,1410,264,1930]" targetPageId="4">
<paragraph id="8BC33692817BFFBEFF7AF82BFBC2F817" blockId="4.[189,1399,1962,2024]" pageId="4" pageNumber="5">
<emphasis id="B908EA80817BFFBEFF7AF82BFED3F83B" bold="true" box="[189,298,1962,1988]" pageId="4" pageNumber="5">Figs 1–8.</emphasis>
<taxonomicName id="4C7C4D11817BFFBEFEF7F82BFE02F83B" authority="Jia &amp; Tang, 2018" authorityName="Jia &amp; Tang" authorityYear="2018" box="[304,507,1962,1988]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="songi" status="sp. nov.">
<emphasis id="B908EA80817BFFBEFEF7F82BFE02F83B" box="[304,507,1962,1988]" italics="true" pageId="4" pageNumber="5">Helochares songi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57FB817BFFBEFDC7F82AFDA0F83A" box="[512,601,1963,1989]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B908EA80817BFFBEFD99F82BFD73F83B" bold="true" box="[606,650,1962,1988]" pageId="4" pageNumber="5">1–3</emphasis>
. Habitus. 1. Dorsal. 2. Ventral. 3. Lateral. 
<emphasis id="B908EA80817BFFBEFBA9F82BFB84F83B" bold="true" box="[1134,1149,1962,1988]" pageId="4" pageNumber="5">4</emphasis>
. Head and pronotum. 
<emphasis id="B908EA80817BFFBEFF7AF84FFF35F817" bold="true" box="[189,204,1998,2024]" pageId="4" pageNumber="5">5</emphasis>
. Head and prosternum. 
<emphasis id="B908EA80817BFFBEFE22F84FFE0DF817" bold="true" box="[485,500,1998,2024]" pageId="4" pageNumber="5">6</emphasis>
. Pronotum, lateral. 
<emphasis id="B908EA80817BFFBEFD1AF84FFD15F817" bold="true" box="[733,748,1998,2024]" pageId="4" pageNumber="5">7</emphasis>
. Mesoventrite. 
<emphasis id="B908EA80817BFFBEFC63F84FFC4AF817" bold="true" box="[932,947,1998,2024]" pageId="4" pageNumber="5">8</emphasis>
. Abdomen.
</paragraph>
</caption>
<subSubSection id="C3666519817AFFBFFF7AFCDCFDDBFC37" pageId="5" pageNumber="6" type="biology_ecology">
<paragraph id="8BC33692817AFFBFFF7AFCDCFEE5FC88" blockId="5.[189,284,861,887]" box="[189,284,861,887]" pageId="5" pageNumber="6">
<heading id="D08B81FE817AFFBFFF7AFCDCFEE5FC88" bold="true" box="[189,284,861,887]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B908EA80817AFFBFFF7AFCDCFEE5FC88" bold="true" box="[189,284,861,887]" pageId="5" pageNumber="6">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BC33692817AFFBFFF7AFC0AFDDBFC37" blockId="5.[189,1398,907,968]" pageId="5" pageNumber="6">Lives along the sides of rivers with a sandy bottom and stagnant water. The female carries the egg case under the abdominal ventrites.</paragraph>
</subSubSection>
<subSubSection id="C3666519817AFFBFFF7AFC71FBF8FBA3" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BC33692817AFFBFFF7AFC71FED6FBF5" blockId="5.[189,303,1008,1034]" box="[189,303,1008,1034]" pageId="5" pageNumber="6">
<heading id="D08B81FE817AFFBFFF7AFC71FED6FBF5" bold="true" box="[189,303,1008,1034]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B908EA80817AFFBFFF7AFC71FED6FBF5" bold="true" box="[189,303,1008,1034]" pageId="5" pageNumber="6">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BC33692817AFFBFFF7AFB9FFBF8FBA3" blockId="5.[189,1399,1054,1116]" pageId="5" pageNumber="6">
The density is very high at the 
<typeStatus id="54C78830817AFFBFFDE4FB9EFDAFFBC6" box="[547,598,1055,1081]" pageId="5" pageNumber="6">type</typeStatus>
locality, but no specimen was collected at light in July. It is probable that this species can be collected at light during its reproductive period.
</paragraph>
</subSubSection>
<subSubSection id="C3666519817AFFBFFF7AFB02FAD0FB33" box="[189,1321,1155,1229]" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8BC33692817AFFBFFF7AFB02FEAFFB62" blockId="5.[189,342,1155,1181]" box="[189,342,1155,1181]" pageId="5" pageNumber="6">
<heading id="D08B81FE817AFFBFFF7AFB02FEAFFB62" bold="true" box="[189,342,1155,1181]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B908EA80817AFFBFFF7AFB02FEAFFB62" bold="true" box="[189,342,1155,1181]" pageId="5" pageNumber="6">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC33692817AFFBFFF7AFB33FAD0FB33" blockId="5.[189,1321,1202,1229]" box="[189,1321,1202,1229]" pageId="5" pageNumber="6">
<collectingCountry id="F36B7602817AFFBFFF7AFB33FEFAFB33" box="[189,259,1202,1228]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
(
<collectingRegion id="49B8F870817AFFBFFED4FB33FE87FB32" box="[275,382,1202,1229]" country="China" name="Guangxi" pageId="5" pageNumber="6">Guangxi</collectingRegion>
), so far only known from the Shiwandashan National Park and its surroundings.
</paragraph>
</subSubSection>
</treatment>
</document>