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505 lines
68 KiB
XML
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<mods:title id="74E5C062D652D64FACF08987D08D9CE7">A revision of the Chinese Helochares (s. str.) Mulsant, 1844 (Coleoptera, Hydrophilidae)</mods:title>
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<mods:namePart id="B9A204411EA249C239BADABAAFB38A73">Jia, Fenglong</mods:namePart>
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<mods:affiliation id="5F1E0E50B10F8439FFECF710798EB94E">Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China.</mods:affiliation>
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<mods:affiliation id="CC340A1D790E1DA2A875542636EF7A1A">Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China.</mods:affiliation>
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<heading id="D08B81FE8174FFB1FD86F9A7FC0BF9BE" box="[577,1010,1574,1601]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
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<taxonomicName id="4C7C4D118174FFB1FD86F9A7FC0BF9BE" ID-CoL="8XQKG" authority="(Muller, 1776)" authorityName="Fikacek et al." authorityYear="2015" baseAuthorityName="Muller" baseAuthorityYear="1776" box="[577,1010,1574,1601]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="obscurus">
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<emphasis id="B908EA808174FFB1FD86F9A7FCB9F9BE" bold="true" box="[577,832,1574,1601]" italics="true" pageId="11" pageNumber="12">Helochares obscurus</emphasis>
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(
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<bibRefCitation id="EFED4B638174FFB1FC88F9A7FC13F9BE" author="Muller O. F." box="[847,1002,1574,1601]" pageId="11" pageNumber="12" refId="ref16019" refString="Muller O. F. 1776. Zoologicae Danicae Prodromus, seu Animalium Daniae et Norvegiae indigenarum Characteres, Nomina et Synonyma imprimis Popularium. Hallager, Havniae [Copenhagen]. https: // doi. org / 10.5962 / bhl. title. 13268" type="book" year="1776">Müller, 1776</bibRefCitation>
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)
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</paragraph>
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<subSubSection id="C36665198174FFB1FD6BF9C8FC71F99B" box="[684,904,1609,1636]" pageId="11" pageNumber="12" type="description">
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<figureCitation id="13472A178174FFB1FD6BF9C8FCCBF99C" box="[684,818,1609,1635]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="11" pageNumber="12">Figs 26–31</figureCitation>
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,
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<figureCitation id="13472A178174FFB1FCF9F9C8FC71F99B" box="[830,904,1609,1636]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="11" pageNumber="12">53–55</figureCitation>
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<paragraph id="8BC336928174FFB1FF7AF90AFD6AF95B" blockId="11.[189,943,1674,1877]" box="[189,659,1674,1701]" pageId="11" pageNumber="12">
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<treatmentCitation id="0ADD10838174FFB1FF7AF90AFD69F95B" author="Muller O. F." box="[189,656,1674,1701]" page="69" pageId="11" pageNumber="12" year="1776">
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<taxonomicName id="4C7C4D118174FFB1FF7AF90AFD69F95B" ID-CoL="3N852" authority="Muller, 1776: 69" authorityName="Muller" authorityPageNumber="69" authorityYear="1776" box="[189,656,1674,1701]" class="Insecta" family="Hydrophilidae" genus="Hydrophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="obscurus">
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<emphasis id="B908EA808174FFB1FF7AF90AFE39F95A" box="[189,448,1675,1701]" italics="true" pageId="11" pageNumber="12">Hydrophilus obscurus</emphasis>
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<bibRefCitation id="EFED4B638174FFB1FE00F90BFD69F95B" author="Muller O. F." box="[455,656,1674,1700]" pageId="11" pageNumber="12" refId="ref16019" refString="Muller O. F. 1776. Zoologicae Danicae Prodromus, seu Animalium Daniae et Norvegiae indigenarum Characteres, Nomina et Synonyma imprimis Popularium. Hallager, Havniae [Copenhagen]. https: // doi. org / 10.5962 / bhl. title. 13268" type="book" year="1776">Müller, 1776: 69</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BC336928174FFB1FF7AF92FFCEDF936" blockId="11.[189,943,1674,1877]" box="[189,788,1710,1737]" pageId="11" pageNumber="12">
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<treatmentCitation id="0ADD10838174FFB1FF7AF92FFCE9F936" author="Fabricius J. C." box="[189,784,1710,1737]" page="185" pageId="11" pageNumber="12" year="1792">
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<taxonomicName id="4C7C4D118174FFB1FF7AF92FFCE9F936" ID-CoL="3N82J" authority="Fabricius, 1792: 185" authorityName="Fabricius" authorityPageNumber="185" authorityYear="1792" box="[189,784,1710,1737]" class="Insecta" family="Hydrophilidae" genus="Hydrophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="erythrocephalus">
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<emphasis id="B908EA808174FFB1FF7AF92FFDEDF937" box="[189,532,1710,1736]" italics="true" pageId="11" pageNumber="12">Hydrophilus erythrocephalus</emphasis>
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<bibRefCitation id="EFED4B638174FFB1FDDCF92FFCE9F936" author="Fabricius J. C." box="[539,784,1710,1737]" pageId="11" pageNumber="12" refId="ref14949" refString="Fabricius J. C. 1792. Entomologia Systematica emendata et aucta, secundum Classes, Ordines, Genera, Species, adjectis Synonymis, Locis, Observationibus, Descriptionibus. Vol. 1, part 1. C. G. Proft, Hafniae [Copenhagen]. Available from https: // biodiversitylibrary. org / page / 51985494 [accessed 11 Apr. 2018]." type="url" year="1792">Fabricius, 1792: 185</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BC336928174FFB1FF7AF950FC90F914" blockId="11.[189,943,1674,1877]" box="[189,873,1745,1772]" pageId="11" pageNumber="12">
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<treatmentCitation id="0ADD10838174FFB1FF7AF950FC9FF914" author="Herbst J. F. W." box="[189,870,1745,1772]" page="304" pageId="11" pageNumber="12" year="1797">
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<taxonomicName id="4C7C4D118174FFB1FF7AF950FC9FF914" ID-CoL="6MJRR" authority="var. variegatus Herbst, 1797: 304" authorityName="Herbst" authorityPageNumber="304" authorityYear="1797" box="[189,870,1745,1772]" class="Insecta" family="Hydrophilidae" genus="Hydrophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="variety" species="variegatus" variety="variegatus">
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<emphasis id="B908EA808174FFB1FF7AF950FE2BF914" box="[189,466,1745,1771]" italics="true" pageId="11" pageNumber="12">Hydrophilus variegatus</emphasis>
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var.
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<emphasis id="B908EA808174FFB1FDCBF953FD71F914" box="[524,648,1746,1771]" italics="true" pageId="11" pageNumber="12">variegatus</emphasis>
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<bibRefCitation id="EFED4B638174FFB1FD48F950FC9FF914" author="Herbst J. F. W." box="[655,870,1745,1771]" pageId="11" pageNumber="12" refId="ref15535" refString="Herbst J. F. W. 1797. Natursystem aller bekannten in- und auslandischen Insekten. Der Kafer 7. Pauli, Berlin." type="book" year="1797">Herbst, 1797: 304</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BC336928174FFB1FF7AF975FD54F8F1" blockId="11.[189,943,1674,1877]" box="[189,685,1780,1807]" pageId="11" pageNumber="12">
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<treatmentCitationGroup id="AB6C11BC8174FFB1FF7AF975FD54F8F1" box="[189,685,1780,1807]" pageId="11" pageNumber="12">
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<treatmentCitation id="0ADD10838174FFB1FF7AF975FD51F8F1" author="Rey C." box="[189,680,1780,1807]" page="14" pageId="11" pageNumber="12" year="1885">
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<taxonomicName id="4C7C4D118174FFB1FF7AF975FD51F8F1" ID-CoL="3KGX9" authority="Rey, 1885: 14" authorityName="Rey" authorityPageNumber="14" authorityYear="1885" box="[189,680,1780,1807]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="subcompressus">
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<emphasis id="B908EA808174FFB1FF7AF975FE02F8F1" box="[189,507,1780,1806]" italics="true" pageId="11" pageNumber="12">Helochares subcompressus</emphasis>
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<bibRefCitation id="EFED4B638174FFB1FDC5F974FD51F8F1" author="Rey C." box="[514,680,1780,1807]" pageId="11" pageNumber="12" pagination="13 - 32" refId="ref16643" refString="Rey C. 1885. Descriptions de coleopteres nouveaux ou peu connus de la tribu des Palpicornes. Annales de la Societe linneenne de Lyon 31: 13 - 32. Available from https: // biodiversitylibrary. org / page / 54600262 [accessed 11 Apr. 2018]." type="journal article" year="1885">Rey, 1885: 14</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</paragraph>
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<paragraph id="8BC336928174FFB1FF7AF896FC56F8CD" blockId="11.[189,943,1674,1877]" box="[189,943,1815,1842]" pageId="11" pageNumber="12">
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<treatmentCitationGroup id="AB6C11BC8174FFB1FF7AF896FC56F8CD" box="[189,943,1815,1842]" pageId="11" pageNumber="12">
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<treatmentCitation id="0ADD10838174FFB1FF7AF896FC52F8CD" author="Sahlberg J." box="[189,939,1815,1842]" page="20" pageId="11" pageNumber="12" year="1913">
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<taxonomicName id="4C7C4D118174FFB1FF7AF896FC52F8CD" ID-CoL="6LN8V" authority="Sahlberg, 1913: 20" authorityName="Sahlberg" authorityPageNumber="20" authorityYear="1913" box="[189,939,1815,1842]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="variety" species="erythrocephalus" variety="substriatus">
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<emphasis id="B908EA808174FFB1FF7AF896FDF1F8CE" box="[189,520,1815,1841]" italics="true" pageId="11" pageNumber="12">Helochares erythrocephalus</emphasis>
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||
var.
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||
<emphasis id="B908EA808174FFB1FD85F896FD3AF8CE" box="[578,707,1815,1841]" italics="true" pageId="11" pageNumber="12">substriatus</emphasis>
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<bibRefCitation id="EFED4B638174FFB1FD0DF896FC52F8CD" author="Sahlberg J." box="[714,939,1815,1842]" pageId="11" pageNumber="12" pagination="1 - 108" refId="ref16694" refString="Sahlberg J. 1913. Coleoptera balcanica quae mensibus Octobri et Decembri 1903 atque Martis et Aprili 1906 in peninsula balcanica collegerunt John Sahlberg et Unio Saalas. Ofversigt av Finska Vetenskaps- Societetens Forhandlingar 55 A (15): 1 - 108. Available from https: // biodiversitylibrary. org / page / 14567987 [accessed 11 Apr. 2018]." type="journal article" year="1913">Sahlberg, 1913: 20</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BC336928174FFB1FF7AF8BAFD04F8AB" blockId="11.[189,943,1674,1877]" box="[189,765,1850,1877]" pageId="11" pageNumber="12">
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<taxonomicName id="4C7C4D118174FFB1FF7AF8BAFE66F8AB" ID-CoL="6MK45" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[189,415,1851,1877]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="griseus">
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<emphasis id="B908EA808174FFB1FF7AF8BAFE66F8AB" box="[189,415,1851,1877]" italics="true" pageId="11" pageNumber="12">Helochares griseus</emphasis>
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</taxonomicName>
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a.
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||
<emphasis id="B908EA808174FFB1FE05F8BAFDE3F8AA" box="[450,538,1851,1877]" italics="true" pageId="11" pageNumber="12">Mülleri</emphasis>
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<treatmentCitation id="0ADD10838174FFB1FDE5F8BBFD00F8AB" author="Reitter E." box="[546,761,1850,1877]" page="364" pageId="11" pageNumber="12" year="1909">
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<bibRefCitation id="EFED4B638174FFB1FDE5F8BBFD00F8AB" author="Reitter E." box="[546,761,1850,1877]" pageId="11" pageNumber="12" refId="ref16617" refString="Reitter E. 1909. Fauna Germanica. Die Kafer des Deutschen Reiches. Vol. 2. K. G. Lutz, Stuttgart." type="book" year="1909">Reitter, 1909: 364</bibRefCitation>
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</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC336928174FFB1FF7AF8FDFC8FF869" blockId="11.[189,1072,1916,2013]" box="[189,886,1916,1943]" pageId="11" pageNumber="12">
|
||
<treatmentCitationGroup id="AB6C11BC8174FFB1FF7AF8FDFC8FF869" box="[189,886,1916,1943]" pageId="11" pageNumber="12">
|
||
<taxonomicName id="4C7C4D118174FFB1FF7AF8FDFD99F869" ID-CoL="6MJRR" box="[189,608,1916,1943]" class="Insecta" family="Hydrophilidae" genus="Hydrophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="variety" species="griseus" variety="variegatus">
|
||
<emphasis id="B908EA808174FFB1FF7AF8FDFE52F869" box="[189,427,1916,1942]" italics="true" pageId="11" pageNumber="12">Hydrophilus griseus</emphasis>
|
||
var.
|
||
<emphasis id="B908EA808174FFB1FE23F8FCFD99F869" box="[484,608,1917,1942]" italics="true" pageId="11" pageNumber="12">variegatus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADD10838174FFB1FDB9F8FDFC8BF869" author="Gyllenhal L." box="[638,882,1916,1942]" page="122" pageId="11" pageNumber="12" year="1808">
|
||
<bibRefCitation id="EFED4B638174FFB1FDB9F8FDFC8BF869" author="Gyllenhal L." box="[638,882,1916,1942]" pageId="11" pageNumber="12" refId="ref15158" refString="Gyllenhal L. 1808. Insecta Suecica. Vol. 1, part 1. F. J. Leverentz, Scaris [Skara, Sweden]. Available from https: // biodiversitylibrary. org / page / 9330983 [accessed 11 Apr. 2018]." type="url" year="1808">Gyllenhal 1808: 122</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC336928174FFB1FF7AF81EFC74F845" blockId="11.[189,1072,1916,2013]" box="[189,909,1951,1978]" pageId="11" pageNumber="12">
|
||
<treatmentCitationGroup id="AB6C11BC8174FFB1FF7AF81EFC74F845" box="[189,909,1951,1978]" pageId="11" pageNumber="12">
|
||
<taxonomicName id="4C7C4D118174FFB1FF7AF81EFD72F846" box="[189,651,1951,1978]" class="Insecta" family="Syrphidae" genus="Helophilus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="variety" species="lividus" variety="erythrocephalus">
|
||
<emphasis id="B908EA808174FFB1FF7AF81EFE6AF846" box="[189,403,1951,1977]" italics="true" pageId="11" pageNumber="12">Helophilus lividus</emphasis>
|
||
var.
|
||
<emphasis id="B908EA808174FFB1FE0AF81EFD72F846" box="[461,651,1951,1977]" italics="true" pageId="11" pageNumber="12">erythrocephalus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADD10838174FFB1FD6FF81EFC70F845" author="Mulsant E." box="[680,905,1951,1978]" page="135" pageId="11" pageNumber="12" year="1844">
|
||
<bibRefCitation id="EFED4B638174FFB1FD6FF81EFC70F845" author="Mulsant E." box="[680,905,1951,1978]" pageId="11" pageNumber="12" refId="ref16065" refString="Mulsant E. 1844. Histoire Naturelle des Coleopteres de France. Vol. 5. Palpicornes. L. Maison, Paris; Ch. Savy Jeune, Lyon. Available from https: // biodiversitylibrary. org / page / 9628710 [accessed 11 Apr. 2018]." type="url" year="1844">Mulsant 1844: 135</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC336928174FFB1FF7AF843FBC9F822" blockId="11.[189,1072,1916,2013]" box="[189,1072,1986,2013]" pageId="11" pageNumber="12">
|
||
<treatmentCitationGroup id="AB6C11BC8174FFB1FF7AF843FBC9F822" box="[189,1072,1986,2013]" pageId="11" pageNumber="12">
|
||
<taxonomicName id="4C7C4D118174FFB1FF7AF843FD72F823" box="[189,651,1986,2013]" class="Insecta" family="Hydrophilidae" genus="Philhydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="variety" species="lividus" variety="erythrocephalus">
|
||
<emphasis id="B908EA808174FFB1FF7AF843FE6AF823" box="[189,403,1986,2012]" italics="true" pageId="11" pageNumber="12">Philhydrus lividus</emphasis>
|
||
var.
|
||
<emphasis id="B908EA808174FFB1FE0AF843FD72F823" box="[461,651,1986,2012]" italics="true" pageId="11" pageNumber="12">erythrocephalus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADD10838174FFB1FD6FF843FBD4F822" author="Gemminger M. & Harold B." box="[680,1069,1986,2013]" page="481" pageId="11" pageNumber="12" year="1868">
|
||
<bibRefCitation id="EFED4B638174FFB1FD6FF843FBD4F822" author="Gemminger M. & Harold B." box="[680,1069,1986,2013]" pageId="11" pageNumber="12" pagination="425 - 752" refId="ref15102" refString="Gemminger M. & Harold B. 1868. Catalogus Coleopterorum hucusque Descriptorum Synonymicus et Systematicus. Vol. 2: 425 - 752. E. H. Gummi, Monachii [Munich]. Available from https: // biodiversitylibrary. org / page / 10174177 [accessed 11 Apr. 2018]." type="journal article" year="1868">Gemminger & Harold 1868: 481</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC336928173FFB6FF7AFE8AFC17FEDA" blockId="12.[189,1006,267,399]" box="[189,1006,267,294]" pageId="12" pageNumber="13">
|
||
<treatmentCitationGroup id="AB6C11BC8173FFB6FF7AFE8AFC17FEDA" box="[189,1006,267,294]" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C7C4D118173FFB6FF7AFE8AFDB0FEDA" box="[189,585,267,294]" class="Insecta" family="Hydrophilidae" genus="Philhydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="variety" species="lividus" variety="variegatus">
|
||
<emphasis id="B908EA808173FFB6FF7AFE8AFE6AFEDA" box="[189,403,267,293]" italics="true" pageId="12" pageNumber="13">Philhydrus lividus</emphasis>
|
||
var.
|
||
<emphasis id="B908EA808173FFB6FE0AFE8DFDB0FEDA" box="[461,585,268,293]" italics="true" pageId="12" pageNumber="13">variegatus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADD10838173FFB6FDA1FE8AFC12FEDA" author="Gemminger M. & Harold B." box="[614,1003,267,293]" page="481" pageId="12" pageNumber="13" year="1868">
|
||
<bibRefCitation id="EFED4B638173FFB6FDA1FE8AFC12FEDA" author="Gemminger M. & Harold B." box="[614,1003,267,293]" pageId="12" pageNumber="13" pagination="425 - 752" refId="ref15102" refString="Gemminger M. & Harold B. 1868. Catalogus Coleopterorum hucusque Descriptorum Synonymicus et Systematicus. Vol. 2: 425 - 752. E. H. Gummi, Monachii [Munich]. Available from https: // biodiversitylibrary. org / page / 10174177 [accessed 11 Apr. 2018]." type="journal article" year="1868">Gemminger & Harold 1868: 481</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC336928173FFB6FF7AFEAEFCBEFEB6" blockId="12.[189,1006,267,399]" box="[189,839,302,329]" pageId="12" pageNumber="13">
|
||
<treatmentCitationGroup id="AB6C11BC8173FFB6FF7AFEAEFCBEFEB6" box="[189,839,302,329]" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C7C4D118173FFB6FF7AFEAEFEBAFEB6" ID-CoL="8VY5S" authorityName="Mulsant" authorityYear="1844" box="[189,323,303,329]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B908EA808173FFB6FF7AFEAEFEBAFEB6" box="[189,323,303,329]" italics="true" pageId="12" pageNumber="13">Helochares</emphasis>
|
||
</taxonomicName>
|
||
(i. sp.)
|
||
<taxonomicName id="4C7C4D118173FFB6FE59FEAEFDA5FEB6" ID-CoL="3N82J" box="[414,604,303,329]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="erythrocephalus">
|
||
<emphasis id="B908EA808173FFB6FE59FEAEFDA5FEB6" box="[414,604,303,329]" italics="true" pageId="12" pageNumber="13">erythrocephalus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADD10838173FFB6FDBFFEAEFCBAFEB6" author="Kuwert A." box="[632,835,303,329]" page="37" pageId="12" pageNumber="13" year="1890">
|
||
<bibRefCitation id="EFED4B638173FFB6FDBFFEAEFCBAFEB6" author="Kuwert A." box="[632,835,303,329]" pageId="12" pageNumber="13" pagination="1 - 121" refId="ref15701" refString="Kuwert A. 1890. Bestimmungs-Tabellen der Hydrophiliden Europas, Westasiens und Nordafrikas. Verhandlungen des naturforschenden Vereins in Brunn 28: 1 - 121. Available from https: // biodiversitylibrary. org / page / 43763386 [accessed 11 Apr. 2018]." type="journal article" year="1890">Kuwert 1890: 37</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC336928173FFB6FF7AFED3FC55FE93" blockId="12.[189,1006,267,399]" box="[189,940,338,364]" pageId="12" pageNumber="13">
|
||
<treatmentCitationGroup id="AB6C11BC8173FFB6FF7AFED3FC55FE93" box="[189,940,338,364]" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C7C4D118173FFB6FF7AFED3FD4BFE93" ID-CoL="6LN8V" box="[189,690,338,364]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="variety" species="griseus" variety="substriatus">
|
||
<emphasis id="B908EA808173FFB6FF7AFED3FEBAFE93" box="[189,323,338,364]" italics="true" pageId="12" pageNumber="13">Helochares</emphasis>
|
||
(s. str.)
|
||
<emphasis id="B908EA808173FFB6FE65FED2FE0EFE93" box="[418,503,339,364]" italics="true" pageId="12" pageNumber="13">griseus</emphasis>
|
||
var.
|
||
<emphasis id="B908EA808173FFB6FDF6FED3FD4BFE93" box="[561,690,338,364]" italics="true" pageId="12" pageNumber="13">substriatus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADD10838173FFB6FD08FED3FC50FE93" author="Zaitzev F. A." box="[719,937,338,364]" page="382" pageId="12" pageNumber="13" year="1908">
|
||
<bibRefCitation id="EFED4B638173FFB6FD08FED3FC50FE93" author="Zaitzev F. A." box="[719,937,338,364]" pageId="12" pageNumber="13" pagination="283 - 420" refId="ref17043" refString="Zaitzev F. A. 1908. Catalogue des coleopteres aquatiques des familles Dryopidae, Georyssidae, Cyathoceridae, Heteroceridae et Hydrophilidae. Horae Societatis entomologicae rossicae 38: 283 - 420." type="journal article" year="1908">Zaitzev 1908: 382</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC336928173FFB6FF7AFEF4FCB8FE70" blockId="12.[189,1006,267,399]" box="[189,833,373,399]" pageId="12" pageNumber="13">
|
||
<treatmentCitationGroup id="AB6C11BC8173FFB6FF7AFEF4FCB8FE70" box="[189,833,373,399]" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C7C4D118173FFB6FF7AFEF4FEBAFE70" ID-CoL="8VY5S" authorityName="Mulsant" authorityYear="1844" box="[189,323,373,399]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B908EA808173FFB6FF7AFEF4FEBAFE70" box="[189,323,373,399]" italics="true" pageId="12" pageNumber="13">Helochares</emphasis>
|
||
</taxonomicName>
|
||
(s. str.)
|
||
<emphasis id="B908EA808173FFB6FE65FEF4FDF8FE70" box="[418,513,373,399]" italics="true" pageId="12" pageNumber="13">obscuus</emphasis>
|
||
–
|
||
<treatmentCitation id="0ADD10838173FFB6FDD9FEF4FCC7FE70" author="d'Orchymont A." box="[542,830,373,399]" page="306" pageId="12" pageNumber="13" year="1933">
|
||
<bibRefCitation id="EFED4B638173FFB6FDD9FEF4FCC7FE70" author="d'Orchymont A." box="[542,830,373,399]" pageId="12" pageNumber="13" pagination="271 - 314" refId="ref16350" refString="d'Orchymont A. 1933. Contribution a l'etude des Palpicornia VIII. Bulletin et Annales de la Societe entomologique de Belgique 73: 271 - 314." type="journal article" year="1933">d’Orchymont 1933: 306</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36665198173FFB6FF7AFE37FD50FC37" pageId="12" pageNumber="13" type="diagnosis">
|
||
<paragraph id="8BC336928173FFB6FF7AFE37FECFFE2F" blockId="12.[189,310,438,464]" box="[189,310,438,464]" pageId="12" pageNumber="13">
|
||
<heading id="D08B81FE8173FFB6FF7AFE37FECFFE2F" bold="true" box="[189,310,438,464]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis id="B908EA808173FFB6FF7AFE37FECFFE2F" bold="true" box="[189,310,438,464]" pageId="12" pageNumber="13">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336928173FFB6FF7AFE64FD50FC37" blockId="12.[189,1399,485,968]" pageId="12" pageNumber="13">
|
||
Length 5.0–
|
||
<quantity id="4C849B778173FFB6FE8AFE64FE52FE00" box="[333,427,485,512]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="12" pageNumber="13" unit="mm" value="6.4">6.4 mm</quantity>
|
||
. Yellowish brown to reddish brown with dark head (
|
||
<figureCitation id="13472A178173FFB6FBD8FE64FB80FDFF" box="[1055,1145,485,512]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 26</figureCitation>
|
||
). Dorsum with dense and coarse punctures. Systematic punctures on clypeus of the same size as ground punctures. Clypeus never impressed in front of eyes (
|
||
<figureCitation id="13472A178173FFB6FD8AFDAAFD5CFDBA" box="[589,677,555,581]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 28</figureCitation>
|
||
), lateral margin distinct narrowed anteriorly. Eyes somewhat emarginate anteriorly (
|
||
<figureCitation id="13472A178173FFB6FE17FDCFFDD2FD96" box="[464,555,590,617]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 28</figureCitation>
|
||
). Maxillary palps yellowish brown with infuscate apex (
|
||
<figureCitation id="13472A178173FFB6FB1AFDCFFA93FD96" box="[1245,1386,590,617]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Figs 28–29</figureCitation>
|
||
), ca 1.3× as long as width of head. Antennae with pedicel slightly shorter than antennomeres 3–6 combined. Submentum with small glabrous areas medially and with uneven, dense, coarse and strong punctures (
|
||
<figureCitation id="13472A178173FFB6FE85FD39FE64FD2D" box="[322,413,696,722]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 29</figureCitation>
|
||
). Elytra usually with some longitudinal rows of black spots (
|
||
<figureCitation id="13472A178173FFB6FB40FD39FB1AFD2C" box="[1159,1251,696,723]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 26</figureCitation>
|
||
), systematic punctures on pronotum and elytra indistinct (
|
||
<figureCitation id="13472A178173FFB6FD27FD5AFC96FD0A" box="[736,879,731,757]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Figs 26, 28</figureCitation>
|
||
). Mesoventrite with a low transverse ridge medially, which does not bear an elevated tooth or projection but with a tuft of long setae (
|
||
<figureCitation id="13472A178173FFB6FAC8FD7FFA90FCE6" box="[1295,1385,766,793]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 30</figureCitation>
|
||
). Metaventrite without glabrous area medially. Femora densely pubescent except apical eighth (
|
||
<figureCitation id="13472A178173FFB6FF02FCC4FEACFCA0" box="[197,341,837,863]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Figs 27, 31</figureCitation>
|
||
). Aedeagus (
|
||
<figureCitation id="13472A178173FFB6FE32FCC4FD86FC9F" box="[501,639,837,864]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="12" pageNumber="13">Figs 53–55</figureCitation>
|
||
) ca 2.4× as long as wide. Apex of paramere dilated inwards, rounded near apex, not distinctly angular (dorsal view). Membranous inner sac with several small dentiform bulges on each side, corresponding to a number of sclerotized spines of median lobe that are visible when the median lobe is extended.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36665198173FFB6FF7AFC71FD8EFBA3" pageId="12" pageNumber="13" type="materials_examined">
|
||
<paragraph id="8BC336928173FFB6FF7AFC71FE53FBF5" blockId="12.[189,426,1008,1034]" box="[189,426,1008,1034]" pageId="12" pageNumber="13">
|
||
<heading id="D08B81FE8173FFB6FF7AFC71FE53FBF5" bold="true" box="[189,426,1008,1034]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis id="B908EA808173FFB6FF7AFC71FE53FBF5" bold="true" box="[189,426,1008,1034]" pageId="12" pageNumber="13">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336928173FFB6FF7AFB9EFD8EFBA3" blockId="12.[189,1398,1053,1116]" pageId="12" pageNumber="13">
|
||
<materialsCitation id="3B143CCF8173FFB6FF7AFB9EFD8EFBA3" ID-GBIF-Occurrence="2610058764" pageId="12" pageNumber="13">
|
||
<collectingCountry id="F36B76028173FFB6FF7AFB9EFEE6FBC6" box="[189,287,1055,1081]" name="China" pageId="12" pageNumber="13">CHINA</collectingCountry>
|
||
:
|
||
<emphasis id="B908EA808173FFB6FEEDFB9FFE6EFBC7" bold="true" box="[298,407,1054,1080]" pageId="12" pageNumber="13">
|
||
<collectingRegion id="49B8F8708173FFB6FEEDFB9FFE6EFBC7" box="[298,407,1054,1080]" country="China" name="Xinjiang Uygur" pageId="12" pageNumber="13">Xinjiang</collectingRegion>
|
||
</emphasis>
|
||
: 1 ♂, Tacheng, Haerdun River dike,
|
||
<date id="FFC210528173FFB6FC88FB9EFC1CFBC6" box="[847,997,1054,1081]" pageId="12" pageNumber="13" value="2005-08-04">4 Aug. 2005</date>
|
||
, Ling Zhao leg.;
|
||
<specimenCount id="9D7AFD1B8173FFB6FB76FB9EFB27FBC6" box="[1201,1246,1055,1081]" pageId="12" pageNumber="13" type="female">1 ♀</specimenCount>
|
||
, Yining, Yili River,
|
||
<date id="FFC210528173FFB6FECBFBC0FE67FBA3" box="[268,414,1089,1116]" pageId="12" pageNumber="13" value="1984-08-05">5 Aug. 1984</date>
|
||
, Zhihe Huang leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36665198173FFB4FF7AFB02FDF2FCB0" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
|
||
<paragraph id="8BC336928173FFB6FF7AFB02FE92FB62" blockId="12.[189,363,1155,1181]" box="[189,363,1155,1181]" pageId="12" pageNumber="13">
|
||
<heading id="D08B81FE8173FFB6FF7AFB02FE92FB62" bold="true" box="[189,363,1155,1181]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis id="B908EA808173FFB6FF7AFB02FE92FB62" bold="true" box="[189,363,1155,1181]" pageId="12" pageNumber="13">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336928173FFB6FF7AFB33FAFEFA83" blockId="12.[189,1399,1201,1404]" pageId="12" pageNumber="13">
|
||
FORM AND COLOUR. Body length 5.0–
|
||
<quantity id="4C849B778173FFB6FD9EFB30FD4AFB33" box="[601,691,1201,1228]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="12" pageNumber="13" unit="mm" value="6.4">6.4 mm</quantity>
|
||
, width
|
||
<quantity id="4C849B778173FFB6FCC4FB33FC68FB33" box="[771,913,1202,1228]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.1" metricValueMin="2.4" pageId="12" pageNumber="13" unit="mm" value="2.75" valueMax="3.1" valueMin="2.4">2.4–3.1 mm</quantity>
|
||
, oval, moderately convex. Head yellowish to dark brown, labrum dark brown or black. Pronotum yellowish to reddish brown (
|
||
<figureCitation id="13472A178173FFB6FB56FB54FAE3FB10" box="[1169,1306,1237,1263]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Figs 26, 28</figureCitation>
|
||
). Elytra with same colour as pronotum but with some longitudinal rows of black spots (
|
||
<figureCitation id="13472A178173FFB6FB48FB79FB13FAEC" box="[1167,1258,1272,1299]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 26</figureCitation>
|
||
). Maxillary palps yellowish brown with last palpomere black apically (
|
||
<figureCitation id="13472A178173FFB6FCAFFA9AFC08FACA" box="[872,1009,1307,1333]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Figs 28–29</figureCitation>
|
||
). Antennae yellowish brown with somewhat dark club. Labial palps reddish or yellowish brown and dark apically (
|
||
<figureCitation id="13472A178173FFB6FB45FABFFAF7FAA6" box="[1154,1294,1342,1369]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Figs 27–28</figureCitation>
|
||
). Ventral side dark brown, legs with apices of femora, tibiae and tarsomeres yellowish brown (
|
||
<figureCitation id="13472A178173FFB6FB65FAE3FB03FA83" box="[1186,1274,1378,1404]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC336928173FFB6FF7AFA25FB31F8E2" blockId="12.[189,1399,1443,1822]" pageId="12" pageNumber="13">
|
||
HEAD. Labrum with fine punctures, systematic punctures detectable. Clypeus broad, anterior margin broadly emarginate, lightly expanded in front of eyes, not impressed in front of eyes (
|
||
<figureCitation id="13472A178173FFB6FB17FA47FAD3FA1F" box="[1232,1322,1478,1504]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 28</figureCitation>
|
||
), with moderately coarse and dense punctures, systematic punctures of same size as ground punctures (
|
||
<figureCitation id="13472A178173FFB6FAD4FA68FA90F9FB" box="[1299,1385,1513,1540]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 28</figureCitation>
|
||
). Eyes moderate in size, emarginate anteriorly; interval between eyes ca 4× as wide as one eye. Ground punctures on frons similar to those on clypeus, systematic punctures only slightly coarser than ground punctures. Maxillary palps long and slender, ca 1.3× as long as width of head, 1
|
||
<superScript id="7C099BDA8173FFB6FBB8F9D3FB73F99F" attach="left" box="[1151,1162,1618,1632]" fontSize="6" pageId="12" pageNumber="13">st</superScript>
|
||
and 2
|
||
<superScript id="7C099BDA8173FFB6FB12F9D0FB1FF9A0" attach="left" box="[1237,1254,1617,1631]" fontSize="6" pageId="12" pageNumber="13">nd</superScript>
|
||
palpomeres curved inwards, apical palpomere a little shorter than penultimate, slightly bending inwards. Antennae with 9 antennomeres, scape ca 1.5 × as long as pedicel, pedicel almost as long as antennomeres 3–6 combined, club loosely segmented, bearing dense pubescence (
|
||
<figureCitation id="13472A178173FFB6FC5CF93CFBDBF928" box="[923,1058,1725,1751]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Figs 27, 29</figureCitation>
|
||
).
|
||
<collectingRegion id="49B8F8708173FFB6FBF1F93CFB60F928" box="[1078,1177,1725,1751]" country="Myanmar" name="Chin" pageId="12" pageNumber="13">Mentum</collectingRegion>
|
||
ca 1.3 × as wide as long, subquadrate, moderately impressed anteromedially, with coarse punctures and wrinkles (
|
||
<figureCitation id="13472A178173FFB6FAD6F961FA90F905" box="[1297,1385,1760,1786]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 29</figureCitation>
|
||
). Submentum with uneven, coarse punctures and small, glabrous areas medially (
|
||
<figureCitation id="13472A178173FFB6FBA4F882FB42F8E2" box="[1123,1211,1795,1821]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 29</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC336928173FFB4FF7AF8C4FD80FDE3" blockId="12.[189,1398,1860,1993]" lastBlockId="14.[189,1399,267,540]" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
|
||
THORAX. Pronotum ca 2× as wide as long, widest medially; anterior margin and posterior margin almost straight; with punctures as on head, systematic punctures arranged as two transverse oblique rows, almost of same size as ground punctures (
|
||
<figureCitation id="13472A178173FFB6FD96F80AFD50F85A" box="[593,681,1931,1957]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="12" pageNumber="13">Fig. 28</figureCitation>
|
||
). Prosternum bulging in middle, not carinate, with transverse groove, protruding anteriorly. Mesoventrite with a low transverse ridge medially, which does not bear an elevated tooth or projection but with a tuft of long setae, behind the ridge, with a low, longitudinal carina between mesocoxae (
|
||
<figureCitation id="13472A178171FFB4FDC9FEAFFD9EFEB6" box="[526,615,302,329]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="14" pageNumber="15">Fig. 30</figureCitation>
|
||
). Metaventrite pubescent, with somewhat convex middle portion, without glabrous area; metepisterna ca 3.5× as long as wide, subparallel. Scutellum triangular, with similar punctures to those on elytra. Ground punctures on elytra similar to those on pronotum, with 2 or 3 rows of indistinct systematic punctures (
|
||
<figureCitation id="13472A178171FFB4FD6CFE19FCFAFE4C" box="[683,771,408,435]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="14" pageNumber="15">Fig. 26</figureCitation>
|
||
). Epipleura very wide anteriorly, gradually narrowed, reaching level of first visible abdominal sternite. Femora densely pubescent with apical eighth glabrous. Metatarsomeres with dense white hairs ventrally and a few long swimming hairs dorsally. Claws of moderate size, rather strongly curved.
|
||
</paragraph>
|
||
<caption id="DF03661A8172FFB7FF7AF82CFC09F815" ID-DOI="http://doi.org/10.5281/zenodo.3785488" ID-Zenodo-Dep="3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,241,1965,1991]" targetBox="[285,1305,265,1912]" targetPageId="13">
|
||
<paragraph id="8BC336928172FFB7FF7AF82CFC09F815" blockId="13.[189,1398,1965,2027]" pageId="13" pageNumber="14">
|
||
<emphasis id="B908EA808172FFB7FF7AF82CFEB1F838" bold="true" box="[189,328,1965,1991]" pageId="13" pageNumber="14">Figs 26–31.</emphasis>
|
||
<taxonomicName id="4C7C4D118172FFB7FE88F82CFD05F837" authority="(Muller, 1776)" authorityName="Fikacek et al." authorityYear="2015" baseAuthorityName="Muller" baseAuthorityYear="1776" box="[335,764,1965,1992]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="B908EA808172FFB7FE88F82CFDBFF838" box="[335,582,1965,1991]" italics="true" pageId="13" pageNumber="14">Helochares obscurus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFED4B638172FFB7FD92F82CFD0BF837" author="Muller O. F." box="[597,754,1965,1992]" pageId="13" pageNumber="14" refId="ref16019" refString="Muller O. F. 1776. Zoologicae Danicae Prodromus, seu Animalium Daniae et Norvegiae indigenarum Characteres, Nomina et Synonyma imprimis Popularium. Hallager, Havniae [Copenhagen]. https: // doi. org / 10.5962 / bhl. title. 13268" type="book" year="1776">Müller, 1776</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B908EA808172FFB7FCC1F82CFCB6F838" bold="true" box="[774,847,1965,1991]" pageId="13" pageNumber="14">26–27</emphasis>
|
||
. Habitus. 26. Dorsal. 27. Ventral.
|
||
<emphasis id="B908EA808172FFB7FB1BF82CFB00F838" bold="true" box="[1244,1273,1965,1991]" pageId="13" pageNumber="14">28</emphasis>
|
||
. Head and pronotum.
|
||
<emphasis id="B908EA808172FFB7FEF9F851FEA2F815" bold="true" box="[318,347,2000,2026]" pageId="13" pageNumber="14">29</emphasis>
|
||
. Head and prosternum.
|
||
<emphasis id="B908EA808172FFB7FDB2F851FD6BF815" bold="true" box="[629,658,2000,2026]" pageId="13" pageNumber="14">30</emphasis>
|
||
. Mesoventrite.
|
||
<emphasis id="B908EA808172FFB7FC8DF851FC9EF815" bold="true" box="[842,871,2000,2026]" pageId="13" pageNumber="14">31</emphasis>
|
||
. Abdomen.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC336928171FFB4FF7AFDC2FAA2FDA2" blockId="14.[189,1371,579,606]" box="[189,1371,579,606]" pageId="14" pageNumber="15">
|
||
ABDOMEN. All visible abdominal ventrites with dense pubescence; first ventrite not carinate (
|
||
<figureCitation id="13472A178171FFB4FB31FDC2FAB4FDA2" box="[1270,1357,579,605]" captionStart="Figs 26–31" captionStartId="13.[189,241,1965,1991]" captionTargetBox="[285,1305,265,1912]" captionTargetId="figure@13.[263,1325,248,1922]" captionTargetPageId="13" captionText="Figs 26–31. Helochares obscurus (Müller, 1776). 26–27. Habitus. 26. Dorsal. 27. Ventral. 28. Head and pronotum. 29. Head and prosternum. 30. Mesoventrite. 31. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785488" httpUri="https://zenodo.org/record/3785488/files/figure.png" pageId="14" pageNumber="15">Fig. 31</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC336928171FFB4FF7AFD04FDF2FCB0" blockId="14.[189,1399,645,847]" pageId="14" pageNumber="15">
|
||
AEDEAGUS. Ca 2.4× as long as wide. Apex of parameres dilated inwards, rounded near apex, not distinctly angular (dorsal view). Membranous inner sac with several small dentiform bulges on each side, corresponding to a number of sclerotized spines of median lobe, visible when median lobe is extended. Median lobe with spine-formed apex (
|
||
<figureCitation id="13472A178171FFB4FCE7FD6FFC48FCF6" box="[800,945,750,777]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="14" pageNumber="15">Figs 53–55</figureCitation>
|
||
). Apex of median lobe of aedeagus (
|
||
<figureCitation id="13472A178171FFB4FF01FC90FEE3FCD3" box="[198,282,785,812]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="14" pageNumber="15">Fig. 53</figureCitation>
|
||
) damaged during our dissection of the unique male. We used the figures of
|
||
<bibRefCitation id="EFED4B638171FFB4FB48FC93FAC4FCD3" author="Hansen M." box="[1167,1341,786,812]" pageId="14" pageNumber="15" pagination="1 - 254" refId="ref15206" refString="Hansen M. 1987. The Hydrophiloidea (Coleoptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica 18: 1 - 254." type="journal article" year="1987">Hansen (1987)</bibRefCitation>
|
||
here for illustration (
|
||
<figureCitation id="13472A178171FFB4FEB2FCB4FE06FCB0" box="[373,511,820,847]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="14" pageNumber="15">Figs 54–55</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36665198171FFB4FF7AFCF7FCA5FB90" pageId="14" pageNumber="15" type="diagnosis">
|
||
<paragraph id="8BC336928171FFB4FF7AFCF7FE30FC6F" blockId="14.[189,457,886,912]" box="[189,457,886,912]" pageId="14" pageNumber="15">
|
||
<heading id="D08B81FE8171FFB4FF7AFCF7FE30FC6F" bold="true" box="[189,457,886,912]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="B908EA808171FFB4FF7AFCF7FE30FC6F" bold="true" box="[189,457,886,912]" pageId="14" pageNumber="15">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336928171FFB4FF7AFC25FCA5FB90" blockId="14.[189,1399,932,1135]" pageId="14" pageNumber="15">
|
||
This species can easily be distinguished from other species occurring in
|
||
<collectingCountry id="F36B76028171FFB4FC30FC25FBC4FC41" box="[1015,1085,932,958]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
|
||
by its having the pronotum and elytra yellow brown, head only dark between the eyes, elytra with some longitudinal rows of black spots, aedeagus with a membranous inner sac, median lobe with a number of sclerotized spines and with a spine-formed apex. For the differential characters between it and
|
||
<taxonomicName id="4C7C4D118171FFB4FC1BFB8EFAE5FBD6" authority="(McLeay, 1825)" authorityName="d'Orchymont" authorityYear="1935" baseAuthorityName="MacLeay" baseAuthorityYear="1825" box="[988,1308,1038,1065]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pallens">
|
||
<emphasis id="B908EA808171FFB4FC1BFB8EFBAFFBD7" box="[988,1110,1038,1064]" italics="true" pageId="14" pageNumber="15">H. pallens</emphasis>
|
||
(McLeay, 1825)
|
||
</taxonomicName>
|
||
see the Differential diagnosis of that species. This species is close to the Palaearctic
|
||
<taxonomicName id="4C7C4D118171FFB4FB84FBB3FA8BFBB4" authority="Sharp, 1869" authorityName="Sharp" authorityYear="1869" box="[1091,1394,1073,1099]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="B908EA808171FFB4FB84FBB3FB20FBB4" box="[1091,1241,1074,1099]" italics="true" pageId="14" pageNumber="15">H. punctatus</emphasis>
|
||
Sharp, 1869
|
||
</taxonomicName>
|
||
. The differential characters were given by
|
||
<bibRefCitation id="EFED4B638171FFB4FD60FBD4FCA1FB90" author="Hansen M." box="[679,856,1109,1135]" pageId="14" pageNumber="15" pagination="1 - 254" refId="ref15206" refString="Hansen M. 1987. The Hydrophiloidea (Coleoptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica 18: 1 - 254." type="journal article" year="1987">Hansen (1987)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36665198171FFB4FF7AFB17FE22FAFD" pageId="14" pageNumber="15" type="discussion">
|
||
<paragraph id="8BC336928171FFB4FF7AFB17FED6FB4F" blockId="14.[189,303,1174,1200]" box="[189,303,1174,1200]" pageId="14" pageNumber="15">
|
||
<heading id="D08B81FE8171FFB4FF7AFB17FED6FB4F" bold="true" box="[189,303,1174,1200]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="B908EA808171FFB4FF7AFB17FED6FB4F" bold="true" box="[189,303,1174,1200]" pageId="14" pageNumber="15">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336928171FFB4FF7AFB45FE22FAFD" blockId="14.[189,1398,1220,1282]" pageId="14" pageNumber="15">This species seems to live in water at the edge of dams and the margins of rivers. Other biological characters are unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36665198171FFB4FF7AFAA8FB5DFA24" pageId="14" pageNumber="15" type="distribution">
|
||
<paragraph id="8BC336928171FFB4FF7AFAA8FEAFFABC" blockId="14.[189,342,1321,1347]" box="[189,342,1321,1347]" pageId="14" pageNumber="15">
|
||
<heading id="D08B81FE8171FFB4FF7AFAA8FEAFFABC" bold="true" box="[189,342,1321,1347]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="B908EA808171FFB4FF7AFAA8FEAFFABC" bold="true" box="[189,342,1321,1347]" pageId="14" pageNumber="15">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC336928171FFB4FF7AFAD9FB5DFA24" blockId="14.[189,1398,1368,1500]" pageId="14" pageNumber="15">
|
||
<collectingCountry id="F36B76028171FFB4FF7AFAD9FEFAFA8D" box="[189,259,1368,1394]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="49B8F8708171FFB4FED5FAD9FE79FA8C" box="[274,384,1368,1395]" country="China" name="Xinjiang Uygur" pageId="14" pageNumber="15">Xinjiang</collectingRegion>
|
||
); Palearctic:
|
||
<collectingCountry id="F36B76028171FFB4FDD1FAD9FD96FA8D" box="[534,623,1368,1394]" name="Austria" pageId="14" pageNumber="15">Austria</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FDBEFAD9FD04FA8D" box="[633,765,1368,1394]" name="Azerbaijan" pageId="14" pageNumber="15">Azerbaijan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FCC0FAD9FC9DFA8D" box="[775,868,1368,1394]" name="Belarus" pageId="14" pageNumber="15">Belarus</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FCA8FAD9FC31FA8D" box="[879,968,1368,1394]" name="Croatia" pageId="14" pageNumber="15">Croatia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FC13FAD9FB68FA8D" box="[980,1169,1368,1394]" name="Czech Republic" pageId="14" pageNumber="15">Czech Republic</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FB5BFAD9FAF5FA8D" box="[1180,1292,1368,1394]" name="Denmark" pageId="14" pageNumber="15">Denmark</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FAD0FAD9FA8BFA8D" box="[1303,1394,1368,1394]" name="Estonia" pageId="14" pageNumber="15">Estonia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FF7AFAFAFEE3FA6A" box="[189,282,1403,1429]" name="Finland" pageId="14" pageNumber="15">Finland</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FEECFAFAFE87FA6A" box="[299,382,1403,1429]" name="France" pageId="14" pageNumber="15">France</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FE57FAFAFE07FA6A" box="[400,510,1403,1429]" name="Germany" pageId="14" pageNumber="15">Germany</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FDC8FAFAFD4BFA6A" box="[527,690,1403,1429]" name="United Kingdom" pageId="14" pageNumber="15">Great Britain</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FD03FAFAFCE3FA6A" box="[708,794,1403,1429]" name="Greece" pageId="14" pageNumber="15">Greece</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FCECFAFAFC6AFA6A" box="[811,915,1403,1429]" name="Hungary" pageId="14" pageNumber="15">Hungary</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FC62FAFAFC2EFA6A" box="[933,983,1403,1429]" name="Iran" pageId="14" pageNumber="15">Iran</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FC2EFAFAFBD4FA6A" box="[1001,1069,1403,1429]" name="Israel" pageId="14" pageNumber="15">Israel</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FBF9FAFAFB8CFA6A" box="[1086,1141,1403,1429]" name="Italy" pageId="14" pageNumber="15">Italy</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FB40FAFAFAEAFA6A" box="[1159,1299,1403,1429]" name="Kazakhstan" pageId="14" pageNumber="15">Kazakhstan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FAE3FAFAFA8BFA6A" box="[1316,1394,1403,1429]" name="Latvia" pageId="14" pageNumber="15">Latvia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FF7AFA1FFEC8FA47" box="[189,305,1438,1464]" name="Lithuania" pageId="14" pageNumber="15">Lithuania</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FEFCFA1FFE3FFA47" box="[315,454,1438,1464]" name="Luxembourg" pageId="14" pageNumber="15">Luxemburg</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FE17FA1EFD9BFA46" box="[464,610,1439,1465]" name="Serbia" pageId="14" pageNumber="15">Montenegro</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FDAAFA1FFD07FA47" box="[621,766,1438,1464]" name="Netherlands" pageId="14" pageNumber="15">Netherlands</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FCCFFA1EFC91FA46" box="[776,872,1439,1465]" name="Norway" pageId="14" pageNumber="15">Norway</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FCB5FA1FFC3FFA47" box="[882,966,1438,1464]" name="Poland" pageId="14" pageNumber="15">Poland</collectingCountry>
|
||
, Russian Fed. (
|
||
<collectingCountry id="F36B76028171FFB4FBB1FA1FFB30FA47" box="[1142,1225,1438,1464]" name="Russia" pageId="14" pageNumber="15">Russia</collectingCountry>
|
||
, West Siberia),
|
||
<collectingCountry id="F36B76028171FFB4FF7AFA40FEDEFA24" box="[189,295,1473,1499]" name="Slovakia" pageId="14" pageNumber="15">Slovakia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FEF5FA40FE6BFA24" box="[306,402,1473,1499]" name="Sweden" pageId="14" pageNumber="15">Sweden</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FE59FA40FDD4FA24" box="[414,557,1473,1499]" name="Switzerland" pageId="14" pageNumber="15">Switzerland</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FDFFFA40FD74FA24" box="[568,653,1473,1499]" name="Turkey" pageId="14" pageNumber="15">Turkey</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36B76028171FFB4FD5EFA40FD0EFA24" box="[665,759,1473,1499]" name="Ukraine" pageId="14" pageNumber="15">Ukraine</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFED4B638171FFB4FCC1FA40FC15FA23" author="Fikacek M. & Angus R. B. & Gentili E. & Jia F. & Minoshima Y. N. & Prokin A. & Przevozny M. & Ryndevich S. K." box="[774,1004,1473,1500]" pageId="14" pageNumber="15" pagination="1" refId="ref15018" refString="Fikacek M., Angus R. B., Gentili E., Jia F., Minoshima Y. N., Prokin A., Przevozny M. & Ryndevich S. K. 2015. Helophoridae, Hydrochidae, Hydrophilidae. In: Lobl I. & Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1 (revised and updated edition), Hydrophiloidea - Staphylinoidea. Brill, Leiden, Boston." type="book chapter" year="2015">
|
||
Fikáček
|
||
<emphasis id="B908EA808171FFB4FCABFA43FC5EFA24" box="[876,935,1473,1499]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFED4B638171FFB4FC3FFA43FB6EFA24" author="Hansen M." box="[1016,1175,1473,1500]" pageId="14" pageNumber="15" refId="ref15262" refString="Hansen M. 1999. World Catalogue of Insects 2: Hydrophiloidea (s. str.) (Coleoptera). Apollo Books, Stenstrup, Denmark." type="book" year="1999">Hansen 1999</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |