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<document ID-DOI="10.24349/5n37-k5b9" ID-ISSN="2107-7207" ID-Zenodo-Dep="7160442" approvalRequired="22" approvalRequired_for_textStreams="22" checkinTime="1665196901732" checkinUser="felipe" docAuthor="Stekolnikov, Alexandr A." docDate="2022" docId="03D0E364FFFBFF9FF2C37B7FFCE94C58" docLanguage="en" docName="Acarologia.62.2.418-425.pdf.pdf" docOrigin="Acarologia 62 (2)" docSource="https://www1.montpellier.inrae.fr/CBGP/acarologia/article.php?id=4520" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.4:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="4" docTitle="Minteracarus mombasa Stekolnikov 2022, n. sp." docType="treatment" docVersion="2" lastPageNumber="423" masterDocId="FFE99B1CFFF8FF99F328791CFFEA4947" masterDocTitle="A new genus and species of bat chiggers (Acariformes: Trombiculidae) from Kenya" masterLastPageNumber="425" masterPageNumber="418" pageNumber="420" updateTime="1665198357152" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new genus and species of bat chiggers (Acariformes: Trombiculidae) from Kenya</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stekolnikov, Alexandr A.</mods:namePart>
<mods:affiliation>Laboratory of Parasitic Arthropods, Zoological Institute of the Russian Academy of Sciences,</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Acarologia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-04-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>62</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>418</mods:start>
<mods:end>425</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>https://www1.montpellier.inrae.fr/CBGP/acarologia/article.php?id=4520</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.24349/5n37-k5b9</mods:identifier>
<mods:identifier type="ISSN">2107-7207</mods:identifier>
<mods:identifier type="Zenodo-Dep">7160442</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03D0E364FFFBFF9FF2C37B7FFCE94C58" httpUri="http://treatment.plazi.org/id/03D0E364FFFBFF9FF2C37B7FFCE94C58" lastPageId="6" lastPageNumber="423" pageId="3" pageNumber="420">
<subSubSection pageId="3" pageNumber="420" type="nomenclature">
<paragraph blockId="3.[491,904,611,639]" box="[491,904,611,639]" pageId="3" pageNumber="420">
<heading bold="true" box="[491,904,611,639]" fontSize="12" level="2" pageId="3" pageNumber="420" reason="6">
<emphasis bold="true" box="[491,904,611,639]" pageId="3" pageNumber="420">
<taxonomicName authority="Stekolnikov, 2022" authorityName="Stekolnikov" authorityYear="2022" box="[491,822,611,639]" class="Arachnida" family="Trombiculidae" genus="Minteracarus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="420" phylum="Arthropoda" rank="species" species="mombasa" status="n. sp.">
<emphasis bold="true" box="[491,822,611,639]" italics="true" pageId="3" pageNumber="420">MINteracarus mombasa</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[830,904,614,639]" pageId="3" pageNumber="420" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="3.[491,1188,669,758]" box="[491,995,669,688]" pageId="3" pageNumber="420">
Zoobank:
<uuid box="[580,995,669,688]" pageId="3" pageNumber="420">969EC25B-FAA0-4F94-B996-7C0E8F97E9BF</uuid>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="420" type="description">
<paragraph blockId="3.[491,1188,669,758]" box="[491,640,700,724]" pageId="3" pageNumber="420">
(
<figureCitation box="[498,605,700,724]" captionStart="Figure 1" captionStartId="4.[119,190,1163,1184]" captionTargetBox="[164,1436,237,1098]" captionTargetId="figure-2@4.[156,1441,221,1109]" captionTargetPageId="4" captionText="Figure 1 Minteracarus mombasa n. sp., holotype: A arrangement of dorsal idiosomal setae; B arrangement of ventral idiosomal setae; C" figureDoi="http://doi.org/10.5281/zenodo.7160444" httpUri="https://zenodo.org/record/7160444/files/figure.png" pageId="3" pageNumber="420">Figures 1</figureCitation>
,
<figureCitation box="[618,632,700,724]" captionStart="Figure 2" captionStartId="5.[119,190,1731,1752]" captionTargetBox="[130,1485,274,1668]" captionTargetId="figure-2@5.[103,1494,263,1678]" captionTargetPageId="5" captionText="Figure 2 Minteracarus mombasa n. sp.: A scutum (holotype); B ventral aspect of gnathosoma (paratype L:7263/2); C dorsal aspect" figureDoi="http://doi.org/10.5281/zenodo.7160448" httpUri="https://zenodo.org/record/7160448/files/figure.png" pageId="3" pageNumber="420">2</figureCitation>
)
</paragraph>
<paragraph blockId="3.[491,1188,669,758]" box="[491,1188,734,758]" pageId="3" pageNumber="420">
<emphasis box="[491,768,734,758]" italics="true" pageId="3" pageNumber="420">
<taxonomicName authorityName="Kocak &amp; Kemal" authorityYear="2009" box="[491,619,734,758]" class="Arachnida" family="Trombiculidae" genus="Grandjeana" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="420" phylum="Arthropoda" rank="genus">Grandjeana</taxonomicName>
(?)
<taxonomicName authorityName="Stekolnikov" authorityYear="2022" box="[666,768,734,758]" class="Arachnida" family="Trombiculidae" genus="Minteracarus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="420" phylum="Arthropoda" rank="species" species="mombasa">mombasa</taxonomicName>
</emphasis>
(
<emphasis box="[784,933,734,758]" italics="true" pageId="3" pageNumber="420">nomen nudum</emphasis>
): Stekolnikov 2019: 36.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="420" type="diagnosis">
<paragraph blockId="3.[491,617,799,825]" box="[491,617,799,825]" pageId="3" pageNumber="420">
<emphasis bold="true" box="[491,617,799,825]" pageId="3" pageNumber="420">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="3.[489,1532,851,979]" box="[491,1474,851,875]" pageId="3" pageNumber="420">SIF = 5B1N-N-2-3111.0000; fCx = 1.1.1; fSt = 2.2; fPp = B/B/NBB; DS = 100 119, V = 63</paragraph>
<paragraph blockId="3.[489,1532,851,979]" box="[489,1470,886,910]" pageId="3" pageNumber="420"> 81, NDV = 166 194; Ip = 681 763; PL ≥ AM ≥ AL; eyes absent; scutum bell-shaped,</paragraph>
<paragraph blockId="3.[489,1532,851,979]" pageId="3" pageNumber="420">
with moderate puncta, with prominent anterolateral shoulders; PLs situated off scutum; sensilla flagelliform, with 3 4 long branches in distal half. Measurements given in
<tableCitation box="[1337,1427,955,979]" captionStart="Table 1" captionStartId="3.[119,175,1245,1266]" captionText="Table 1 Morphometric (AW TaIIIW, µm) and meristic (DS NDV) traits ofMinteracarus mombasa n. sp. (N = 7). Abbreviations: AW distance between anterolateral scutal setae; PW distance between posterolateral scutal setae; SB distance between sensillary (trichobothrial" pageId="3" pageNumber="420">Tables 1</tableCitation>
and
<tableCitation box="[1485,1501,955,979]" captionStart="Table 2" captionStartId="4.[515,571,1627,1648]" captionText="Table 2 Minteracarus mombasa n. sp., lengths of specialized leg setae, µm (N = 4)." pageId="3" pageNumber="420">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="423" pageId="3" pageNumber="420" type="description">
<paragraph blockId="3.[491,741,1019,1045]" box="[491,741,1019,1045]" pageId="3" pageNumber="420">
<emphasis bold="true" box="[491,741,1019,1045]" pageId="3" pageNumber="420">Description of larva</emphasis>
</paragraph>
<paragraph blockId="3.[491,1535,1072,1165]" pageId="3" pageNumber="420">
<emphasis bold="true" box="[491,596,1072,1096]" pageId="3" pageNumber="420">Idiosoma</emphasis>
— (
<figureCitation box="[645,741,1072,1096]" captionStart="Figure 1" captionStartId="4.[119,190,1163,1184]" captionTargetBox="[164,1436,237,1098]" captionTargetId="figure-2@4.[156,1441,221,1109]" captionTargetPageId="4" captionText="Figure 1 Minteracarus mombasa n. sp., holotype: A arrangement of dorsal idiosomal setae; B arrangement of ventral idiosomal setae; C" figureDoi="http://doi.org/10.5281/zenodo.7160444" httpUri="https://zenodo.org/record/7160444/files/figure.png" pageId="3" pageNumber="420">Figure 1</figureCitation>
) — Slightly contracted medially in engorged specimens; eyes absent; two pairs of humeral setae densely covered with long barbs
<superScript attach="left" box="[1151,1163,1102,1119]" fontSize="7" pageId="3" pageNumber="420">st</superScript>
; 1posthumeral row (C excluding humeral setae) triple, with 35 41 setae; 2
<superScript attach="left" box="[946,969,1137,1154]" fontSize="7" pageId="3" pageNumber="420">nd</superScript>
posthumeral row (D) double, sometimes triple in
</paragraph>
<caption pageId="3" pageNumber="420" startId="3.[119,175,1245,1266]">
<paragraph blockId="3.[119,1549,1244,1475]" pageId="3" pageNumber="420">
<emphasis bold="true" box="[119,194,1245,1266]" pageId="3" pageNumber="420">Table 1</emphasis>
Morphometric (AW TaIIIW, µm) and meristic (DS NDV) traits of
<taxonomicName authority="Stekolnikov, 2022" authorityName="Stekolnikov" authorityYear="2022" box="[885,1114,1244,1266]" class="Arachnida" family="Trombiculidae" genus="Minteracarus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="420" phylum="Arthropoda" rank="species" species="mombasa" status="n. sp.">
<emphasis box="[885,1114,1244,1266]" italics="true" pageId="3" pageNumber="420">Minteracarus mombasa</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1120,1175,1245,1266]" pageId="3" pageNumber="420">
<taxonomicNameLabel box="[1120,1175,1245,1266]" pageId="3" pageNumber="420" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(N = 7). Abbreviations: AW distance between anterolateral scutal setae; PW distance between posterolateral scutal setae; SB distance between sensillary (trichobothrial
</paragraph>
</caption>
<paragraph blockId="3.[119,1549,1244,1475]" box="[1504,1511,1267,1289]" pageId="3" pageNumber="420">)</paragraph>
<paragraph blockId="3.[119,1549,1244,1475]" pageId="3" pageNumber="420">
bases; ASB distance from the level of sensillary bases to extreme anterior margin of scutum; PSB distance from the level of sensillary bases to extreme posterior margin of scutum; SD length of scutum (ASB + PSB); AP distance between antero- and posterolateral scutal seta on one side;
<collectionCode box="[220,259,1334,1356]" country="Australia" name="Australian Museum" pageId="3" pageNumber="420" type="Museum">AM</collectionCode>
length of anteromedian scutal seta; AL length of anterolateral scutal setae;
<collectionCode box="[1066,1094,1335,1356]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="3" pageNumber="420" type="Museum">PL</collectionCode>
length of posterolateral scutal setae; S length of sensilla (trichobothria); H length of humeral setae
<subScript attach="both" box="[737,762,1368,1382]" fontSize="6" pageId="3" pageNumber="420">min</subScript>
; D length of the shortest dorsal idiosomal seta
<subScript attach="both" box="[1228,1253,1368,1382]" fontSize="6" pageId="3" pageNumber="420">max</subScript>
; D length of the longest dorsal idiosomal seta;
<collectionCode box="[343,350,1383,1404]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="420" type="Museum">V</collectionCode>
<subScript attach="left" box="[349,374,1394,1408]" fontSize="6" pageId="3" pageNumber="420">min</subScript>
length of the shortest ventral idiosomal seta;
<subScript attach="right" box="[841,866,1394,1408]" fontSize="6" pageId="3" pageNumber="420">max</subScript>
<collectionCode box="[850,867,1383,1404]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="420" type="Museum">V</collectionCode>
length of the longest ventral idiosomal seta; pa length of leg
</paragraph>
<paragraph blockId="3.[119,1549,1244,1475]" box="[1510,1522,1335,1356]" pageId="3" pageNumber="420"></paragraph>
<paragraph blockId="3.[119,1549,1244,1475]" pageId="3" pageNumber="420">
I (including coxa, excluding claws); pm length of leg II (including coxa, excluding claws); pp length of leg III (including coxa, excluding claws); Ip sum of leg lengths (pa + pm + pp); TaIIIL length of leg tarsus III; TaIIIW width of leg tarsus III; DS number of dorsal idiosomal setae (excluding scutal);
<collectionCode box="[470,487,1454,1475]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="420" type="Museum">V</collectionCode>
number of ventral idiosomal setae (excluding coxal and sternal); NDV = DS +
<collectionCode box="[1281,1298,1454,1475]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="420" type="Museum">V</collectionCode>
.
</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[469,1230,1545,1564]" pageId="3" pageNumber="420">
Range Mean
<typeStatus box="[689,768,1545,1564]" pageId="3" pageNumber="420">Holotype</typeStatus>
Range Mean
<typeStatus box="[1151,1230,1545,1564]" pageId="3" pageNumber="420">Holotype</typeStatus>
</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[351,1201,1579,1601]" pageId="3" pageNumber="420">
AW 40-46 42 41
<subScript attach="left" box="[814,850,1579,1601]" fontSize="5" pageId="3" pageNumber="420">Dmin</subScript>
21-24 22 23
</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[351,1201,1613,1635]" pageId="3" pageNumber="420">
PW 77-106 87 87
<subScript attach="left" box="[814,853,1613,1635]" fontSize="5" pageId="3" pageNumber="420">Dmax</subScript>
32-38 36 32
</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[351,1201,1647,1669]" pageId="3" pageNumber="420">
SB 16-20 18 18
<subScript attach="left" box="[814,850,1647,1669]" fontSize="5" pageId="3" pageNumber="420">Vmin</subScript>
18-24 22 23
</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[351,1201,1682,1704]" pageId="3" pageNumber="420">
ASB 24-28 26 28
<subScript attach="left" box="[814,853,1682,1704]" fontSize="5" pageId="3" pageNumber="420">Vmax</subScript>
26-31 29 29
</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[351,1207,1717,1736]" pageId="3" pageNumber="420">PSB 15-17 16 17 pa 246-278 268 277</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[351,1207,1752,1771]" pageId="3" pageNumber="420">SD 40-44 43 44 pm 211-237 225 237</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[351,1207,1786,1805]" pageId="3" pageNumber="420">AP 29-41 34 33 pp 223-251 242 249</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[351,1207,1821,1840]" pageId="3" pageNumber="420">AM 32-37 35 37 Ip 681-763 735 763</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[351,1201,1855,1874]" pageId="3" pageNumber="420">AL 31-39 35 31 TaIIIL 63-75 69 73</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[351,1201,1889,1908]" pageId="3" pageNumber="420">PL 34-37 36 37 TaIIIW 15-17 16 15</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[351,1207,1924,1943]" pageId="3" pageNumber="420">S 60-60 60 60 DS 100-119 109 113</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[351,1201,1958,1977]" pageId="3" pageNumber="420">H 41-43 42 43 V 63-81 71 66</paragraph>
<paragraph blockId="3.[351,1230,1545,2011]" box="[814,1207,1992,2011]" pageId="3" pageNumber="420">NDV 166-194 179 179</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7160444" ID-Zenodo-Dep="7160444" box="[119,1509,1162,1184]" httpUri="https://zenodo.org/record/7160444/files/figure.png" pageId="4" pageNumber="421" startId="4.[119,190,1163,1184]" targetBox="[164,1436,237,1098]" targetPageId="4">
<paragraph blockId="4.[117,1509,1162,1233]" box="[119,1509,1162,1184]" pageId="4" pageNumber="421">
<emphasis bold="true" box="[119,209,1163,1184]" pageId="4" pageNumber="421">Figure 1</emphasis>
<taxonomicName authority="Stekolnikov, 2022" authorityName="Stekolnikov" authorityYear="2022" box="[217,445,1162,1184]" class="Arachnida" family="Trombiculidae" genus="Minteracarus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="mombasa" status="n. sp.">
<emphasis box="[217,445,1162,1184]" italics="true" pageId="4" pageNumber="421">Minteracarus mombasa</emphasis>
</taxonomicName>
<emphasis bold="true" box="[451,506,1163,1184]" pageId="4" pageNumber="421">
<taxonomicNameLabel box="[451,506,1163,1184]" pageId="4" pageNumber="421" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype: A arrangement of dorsal idiosomal setae; B arrangement of ventral idiosomal setae; C
</paragraph>
</caption>
<paragraph blockId="4.[117,1509,1162,1233]" box="[117,1493,1185,1210]" pageId="4" pageNumber="421">
humeral seta; D dorsal idiosomal seta of
<superScript attach="right" box="[559,570,1185,1199]" fontSize="6" pageId="4" pageNumber="421">st</superScript>
1posthumeral row (C); E preanal seta. Abbreviations: H humeral setae; u uropore (anus);
</paragraph>
<paragraph blockId="4.[117,1509,1162,1233]" box="[118,1299,1211,1233]" pageId="4" pageNumber="421">V ventral idiosomal setae. Borders between rows showed by dotted lines. Scale bars: A, B 100 µm; C E 50 µm.</paragraph>
<paragraph blockId="4.[491,1532,1347,1548]" pageId="4" pageNumber="421">
medial part, with 23 30 setae; distribution of other 36 47 setae by rows unclear; 1
<superScript attach="left" box="[1396,1408,1347,1364]" fontSize="7" pageId="4" pageNumber="421">st</superScript>
and 2
<superScript attach="left" box="[1475,1498,1347,1364]" fontSize="7" pageId="4" pageNumber="421">nd</superScript>
rows clearly isolated from each other and from posterior setae; total 100 119 short, heavily barbed dorsal idiosomal setae; four sternal and 63 81 ventral idiosomal setae, NDV = 166 194.
</paragraph>
<paragraph blockId="4.[491,1532,1347,1548]" lastBlockId="5.[117,1539,1731,1942]" lastPageId="5" lastPageNumber="422" pageId="4" pageNumber="421">
<emphasis bold="true" box="[531,676,1489,1513]" pageId="4" pageNumber="421">Gnathosoma</emphasis>
— (
<figureCitation box="[726,875,1489,1513]" captionStart="Figure 2" captionStartId="5.[119,190,1731,1752]" captionTargetBox="[130,1485,274,1668]" captionTargetId="figure-2@5.[103,1494,263,1678]" captionTargetPageId="5" captionText="Figure 2 Minteracarus mombasa n. sp.: A scutum (holotype); B ventral aspect of gnathosoma (paratype L:7263/2); C dorsal aspect" figureDoi="http://doi.org/10.5281/zenodo.7160448" httpUri="https://zenodo.org/record/7160448/files/figure.png" pageId="4" pageNumber="421">Figure 2B, C</figureCitation>
) — Cheliceral base with sparse puncta; cheliceral blade with tricuspid cap; gnathobase (infracapitulum) with sparse puncta and one pair of branched,
</paragraph>
<caption box="[515,1342,1627,1649]" pageId="4" pageNumber="421" startId="4.[515,571,1627,1648]">
<paragraph blockId="4.[515,1342,1627,1649]" box="[515,1342,1627,1649]" pageId="4" pageNumber="421">
<emphasis bold="true" box="[515,590,1627,1648]" pageId="4" pageNumber="421">Table 2</emphasis>
<taxonomicName authority="Stekolnikov, 2022" authorityName="Stekolnikov" authorityYear="2022" box="[602,830,1627,1649]" class="Arachnida" family="Trombiculidae" genus="Minteracarus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="mombasa" status="n. sp.">
<emphasis box="[602,830,1627,1649]" italics="true" pageId="4" pageNumber="421">Minteracarus mombasa</emphasis>
</taxonomicName>
<emphasis bold="true" box="[835,890,1627,1648]" pageId="4" pageNumber="421">
<taxonomicNameLabel box="[835,890,1627,1648]" pageId="4" pageNumber="421" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, lengths of specialized leg setae, µm (N = 4).
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="421">
<table box="[737,1200,1722,2004]" gridcols="3" gridrows="8" pageId="4" pageNumber="421">
<tr box="[737,1200,1722,1743]" gridrow="0" pageId="4" pageNumber="421" rowspan-0="1">
<th box="[997,1056,1722,1743]" gridcol="1" gridrow="0" pageId="4" pageNumber="421">Range</th>
<th box="[1148,1200,1722,1743]" gridcol="2" gridrow="0" pageId="4" pageNumber="421">Mean</th>
</tr>
<tr box="[737,1200,1759,1780]" gridrow="1" pageId="4" pageNumber="421">
<th box="[737,937,1759,1780]" gridcol="0" gridrow="1" pageId="4" pageNumber="421">
Tarsala I (
<emphasis box="[831,847,1759,1780]" italics="true" pageId="4" pageNumber="421">ω</emphasis>
)
</th>
<td box="[997,1056,1759,1780]" gridcol="1" gridrow="1" pageId="4" pageNumber="421">20-22</td>
<td box="[1148,1200,1759,1780]" gridcol="2" gridrow="1" pageId="4" pageNumber="421">21</td>
</tr>
<tr box="[737,1200,1797,1818]" gridrow="2" pageId="4" pageNumber="421">
<th box="[737,937,1797,1818]" gridcol="0" gridrow="2" pageId="4" pageNumber="421">
Tarsala II (
<emphasis box="[839,855,1797,1818]" italics="true" pageId="4" pageNumber="421">ω</emphasis>
)
</th>
<td box="[997,1056,1797,1818]" gridcol="1" gridrow="2" pageId="4" pageNumber="421">18-18</td>
<td box="[1148,1200,1797,1818]" gridcol="2" gridrow="2" pageId="4" pageNumber="421">18</td>
</tr>
<tr box="[737,1200,1834,1855]" gridrow="3" pageId="4" pageNumber="421">
<th box="[737,937,1834,1855]" gridcol="0" gridrow="3" pageId="4" pageNumber="421">
Distal tibiala I (
<emphasis box="[883,896,1834,1855]" italics="true" pageId="4" pageNumber="421">φ</emphasis>
)
</th>
<td box="[997,1056,1834,1855]" gridcol="1" gridrow="3" pageId="4" pageNumber="421">23-27</td>
<td box="[1148,1200,1834,1855]" gridcol="2" gridrow="3" pageId="4" pageNumber="421">25.5</td>
</tr>
<tr box="[737,1200,1871,1892]" gridrow="4" pageId="4" pageNumber="421">
<th box="[737,937,1871,1892]" gridcol="0" gridrow="4" pageId="4" pageNumber="421">
Proximal tibiala I (
<emphasis box="[912,925,1871,1892]" italics="true" pageId="4" pageNumber="421">φ</emphasis>
)
</th>
<td box="[997,1056,1871,1892]" gridcol="1" gridrow="4" pageId="4" pageNumber="421">22-22</td>
<td box="[1148,1200,1871,1892]" gridcol="2" gridrow="4" pageId="4" pageNumber="421">22</td>
</tr>
<tr box="[737,1200,1908,1929]" gridrow="5" pageId="4" pageNumber="421">
<th box="[737,937,1908,1929]" gridcol="0" gridrow="5" pageId="4" pageNumber="421">
Microtibiala (
<emphasis box="[865,876,1908,1929]" italics="true" pageId="4" pageNumber="421">κ</emphasis>
)
</th>
<td box="[997,1056,1908,1929]" gridcol="1" gridrow="5" pageId="4" pageNumber="421">8-11</td>
<td box="[1148,1200,1908,1929]" gridcol="2" gridrow="5" pageId="4" pageNumber="421">9.5</td>
</tr>
<tr box="[737,1200,1946,1967]" gridrow="6" pageId="4" pageNumber="421">
<th box="[737,937,1946,1967]" gridcol="0" gridrow="6" pageId="4" pageNumber="421">
Microgenuala (
<emphasis box="[879,890,1946,1967]" italics="true" pageId="4" pageNumber="421">κ</emphasis>
)
</th>
<td box="[997,1056,1946,1967]" gridcol="1" gridrow="6" pageId="4" pageNumber="421">8-8</td>
<td box="[1148,1200,1946,1967]" gridcol="2" gridrow="6" pageId="4" pageNumber="421">8</td>
</tr>
<tr box="[737,1200,1983,2004]" gridrow="7" pageId="4" pageNumber="421">
<th box="[737,937,1983,2004]" gridcol="0" gridrow="7" pageId="4" pageNumber="421">
Lateral genuala I (
<emphasis box="[906,917,1983,2004]" italics="true" pageId="4" pageNumber="421">σ</emphasis>
)
</th>
<td box="[997,1056,1983,2004]" gridcol="1" gridrow="7" pageId="4" pageNumber="421">19-19</td>
<td box="[1148,1200,1983,2004]" gridcol="2" gridrow="7" pageId="4" pageNumber="421">19</td>
</tr>
</table>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7160448" ID-Zenodo-Dep="7160448" box="[119,1496,1731,1753]" httpUri="https://zenodo.org/record/7160448/files/figure.png" pageId="5" pageNumber="422" startId="5.[119,190,1731,1752]" targetBox="[130,1485,274,1668]" targetPageId="5">
<paragraph blockId="5.[117,1539,1731,1942]" box="[119,1496,1731,1753]" pageId="5" pageNumber="422">
<emphasis bold="true" box="[119,210,1731,1752]" pageId="5" pageNumber="422">Figure 2</emphasis>
<taxonomicName authority="Stekolnikov, 2022" authorityName="Stekolnikov" authorityYear="2022" box="[221,450,1731,1753]" class="Arachnida" family="Trombiculidae" genus="Minteracarus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="422" phylum="Arthropoda" rank="species" species="mombasa" status="n. sp.">
<emphasis box="[221,450,1731,1753]" italics="true" pageId="5" pageNumber="422">Minteracarus mombasa</emphasis>
</taxonomicName>
<emphasis bold="true" box="[458,513,1731,1752]" pageId="5" pageNumber="422">
<taxonomicNameLabel box="[458,513,1731,1752]" pageId="5" pageNumber="422" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A scutum (holotype); B ventral aspect of gnathosoma (paratype L:7263/2); C dorsal aspect
</paragraph>
</caption>
<paragraph blockId="5.[117,1539,1731,1942]" pageId="5" pageNumber="422">
of gnathosoma (
<typeStatus box="[285,369,1754,1776]" pageId="5" pageNumber="422">paratype</typeStatus>
L:7263/2); D genu, tibia, and tarsus of leg II (
<typeStatus box="[856,950,1754,1776]" pageId="5" pageNumber="422">holotype</typeStatus>
); E genu, tibia, and tarsus of leg I (
<typeStatus box="[1330,1424,1754,1776]" pageId="5" pageNumber="422">holotype</typeStatus>
); F genu tibia, and tarsus of leg III (
<typeStatus box="[385,479,1776,1798]" pageId="5" pageNumber="422">holotype</typeStatus>
). Abbreviations: AL anterolateral scutal seta;
<collectionCode box="[942,981,1776,1798]" country="Australia" name="Australian Museum" pageId="5" pageNumber="422" type="Museum">AM</collectionCode>
anteromedian scutal seta; Cb cheliceral base; Cs cheliceral blade;
<subScript attach="right" box="[289,297,1810,1824]" fontSize="6" pageId="5" pageNumber="422">1</subScript>
f famulus I (
<emphasis box="[420,429,1800,1821]" italics="true" pageId="5" pageNumber="422">ε</emphasis>
); f
<subScript attach="left" box="[458,466,1810,1824]" fontSize="6" pageId="5" pageNumber="422">2</subScript>
famulus II (
<emphasis box="[602,611,1800,1821]" italics="true" pageId="5" pageNumber="422">ε</emphasis>
); Ga galeal (deutorostral) seta; ga genuala I
<emphasis box="[1076,1088,1800,1821]" italics="true" pageId="5" pageNumber="422">σ</emphasis>
(); gm genuala II
<emphasis box="[1271,1283,1800,1821]" italics="true" pageId="5" pageNumber="422">σ</emphasis>
(); gns gnathocoxal (tritorostral) seta; gp genuala III
<emphasis box="[461,473,1822,1843]" italics="true" pageId="5" pageNumber="422">σ</emphasis>
(); ic infracapitulum (gnathobase, gnathocoxa); µga microgenuala
<emphasis box="[1165,1176,1822,1843]" italics="true" pageId="5" pageNumber="422">κ</emphasis>
I);(µta microtibiala I (
<emphasis box="[1399,1410,1822,1843]" italics="true" pageId="5" pageNumber="422">κ</emphasis>
); PC palpal claw (odontus); PF palpal femur; PG palpal genu;
<collectionCode box="[711,739,1847,1868]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="5" pageNumber="422" type="Museum">PL</collectionCode>
posterolateral scutal seta; pST parasubterminala
<emphasis box="[1242,1251,1847,1868]" italics="true" pageId="5" pageNumber="422">z</emphasis>
(); PT
<emphasis bold="true" box="[1300,1306,1845,1858]" pageId="5" pageNumber="422">
<superScript attach="left" box="[1300,1306,1845,1858]" fontSize="6" pageId="5" pageNumber="422"></superScript>
</emphasis>
pretarsala I (
<emphasis box="[1452,1462,1847,1869]" italics="true" pageId="5" pageNumber="422">ζ</emphasis>
);
</paragraph>
<paragraph blockId="5.[117,1539,1731,1942]" box="[1501,1513,1777,1798]" pageId="5" pageNumber="422"></paragraph>
<paragraph blockId="5.[117,1539,1731,1942]" box="[119,1476,1870,1897]" pageId="5" pageNumber="422">
PT
<emphasis bold="true" box="[147,157,1870,1883]" pageId="5" pageNumber="422">
<superScript attach="left" box="[147,157,1870,1883]" fontSize="6" pageId="5" pageNumber="422"></superScript>
</emphasis>
pretarsala II (
<emphasis box="[311,321,1872,1894]" italics="true" pageId="5" pageNumber="422">ζ</emphasis>
); PTa palpal tarsus; PTi palpal tibia; S sensillum (trichobothrium);
<subScript attach="both" box="[1034,1042,1883,1897]" fontSize="6" pageId="5" pageNumber="422">0</subScript>
S palpal tarsala (
<emphasis box="[1208,1225,1873,1894]" italics="true" pageId="5" pageNumber="422">ω</emphasis>
); S
<subScript attach="left" box="[1259,1267,1883,1897]" fontSize="6" pageId="5" pageNumber="422">1</subScript>
leg tarsala I (
<emphasis box="[1417,1434,1873,1894]" italics="true" pageId="5" pageNumber="422">ω</emphasis>
); S
<subScript attach="left" box="[1468,1476,1883,1897]" fontSize="6" pageId="5" pageNumber="422">2</subScript>
</paragraph>
<paragraph blockId="5.[117,1539,1731,1942]" box="[117,1479,1897,1919]" pageId="5" pageNumber="422">
leg tarsala II (
<emphasis box="[268,285,1898,1919]" italics="true" pageId="5" pageNumber="422">ω</emphasis>
); ST subterminala (
<emphasis box="[495,505,1897,1919]" italics="true" pageId="5" pageNumber="422">ζ</emphasis>
); ta tibiala I (
<emphasis box="[657,670,1898,1919]" italics="true" pageId="5" pageNumber="422">φ</emphasis>
); tm tibiala II (
<emphasis box="[836,849,1898,1919]" italics="true" pageId="5" pageNumber="422">φ</emphasis>
); tp tibiala III (
<emphasis box="[1016,1029,1898,1919]" italics="true" pageId="5" pageNumber="422">φ</emphasis>
). Scale bars: A 50 µm; B, C 20 µm; D F
</paragraph>
<paragraph blockId="5.[117,1539,1731,1942]" box="[117,204,1920,1942]" pageId="5" pageNumber="422"> 50 µm.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7160450" ID-Zenodo-Dep="7160450" box="[515,1269,490,512]" httpUri="https://zenodo.org/record/7160450/files/figure.png" pageId="6" pageNumber="423" startId="6.[515,586,491,512]" targetBox="[692,1297,221,437]" targetPageId="6">
<paragraph blockId="6.[515,1269,490,512]" box="[515,1269,490,512]" pageId="6" pageNumber="423">
<emphasis bold="true" box="[515,605,491,512]" pageId="6" pageNumber="423">Figure 3</emphasis>
Microscope slide with the holotype of
<taxonomicName authority="Stekolnikov, 2022" authorityName="Stekolnikov" authorityYear="2022" box="[982,1210,490,512]" class="Arachnida" family="Trombiculidae" genus="Minteracarus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="423" phylum="Arthropoda" rank="species" species="mombasa" status="n. sp.">
<emphasis box="[982,1210,490,512]" italics="true" pageId="6" pageNumber="423">Minteracarus mombasa</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1216,1269,491,512]" pageId="6" pageNumber="423" rank="species">
<emphasis bold="true" box="[1216,1264,491,512]" pageId="6" pageNumber="423">n. sp</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph blockId="6.[491,1564,630,1311]" pageId="6" pageNumber="423">
gnathocoxal (tritorostral) setae; galeal (deutorostral) seta thick and nude; palpal claw (odontus) slender, with two short subequal prongs; palpal femur with few puncta and strong branched seta; palpal genu with branched seta; dorsal palpal tibial seta nude, lateral and ventral palpal tibial setae branched; palpal tarsus with 6 non-specialized setae, including 1 very thick, heavily barbed dorsal seta and 5 thin setae (4 bearing few branches and 1 nude), and long curved basal tarsala (
<emphasis box="[576,595,804,827]" italics="true" pageId="6" pageNumber="423">ω</emphasis>
).
</paragraph>
<paragraph blockId="6.[491,1564,630,1311]" pageId="6" pageNumber="423">
<emphasis bold="true" box="[531,618,838,862]" pageId="6" pageNumber="423">Scutum</emphasis>
— (
<figureCitation box="[670,788,838,862]" captionStart="Figure 2" captionStartId="5.[119,190,1731,1752]" captionTargetBox="[130,1485,274,1668]" captionTargetId="figure-2@5.[103,1494,263,1678]" captionTargetPageId="5" captionText="Figure 2 Minteracarus mombasa n. sp.: A scutum (holotype); B ventral aspect of gnathosoma (paratype L:7263/2); C dorsal aspect" figureDoi="http://doi.org/10.5281/zenodo.7160448" httpUri="https://zenodo.org/record/7160448/files/figure.png" pageId="6" pageNumber="423">Figure 2A</figureCitation>
) — Small, bell-shaped, with moderate puncta, with prominent anterolateral shoulders; anterior scutal margin sinuous, lateral margins with superimposed cuticular striations, posterior margin rounded; AM at level of ALs; PLs situated off scutum; all scutal setae covered with strong pointed barbs; sensilla flagelliform, with 3 4 long branches in distal half; arched crests present anterior to sensillary bases.
</paragraph>
<paragraph blockId="6.[491,1564,630,1311]" pageId="6" pageNumber="423">
<emphasis bold="true" box="[531,589,1011,1035]" pageId="6" pageNumber="423">Legs</emphasis>
— (
<figureCitation box="[631,788,1010,1034]" captionStart="Figure 2" captionStartId="5.[119,190,1731,1752]" captionTargetBox="[130,1485,274,1668]" captionTargetId="figure-2@5.[103,1494,263,1678]" captionTargetPageId="5" captionText="Figure 2 Minteracarus mombasa n. sp.: A scutum (holotype); B ventral aspect of gnathosoma (paratype L:7263/2); C dorsal aspect" figureDoi="http://doi.org/10.5281/zenodo.7160448" httpUri="https://zenodo.org/record/7160448/files/figure.png" pageId="6" pageNumber="423">Figure 2D F</figureCitation>
) — All 7-segmented (with divided femur), with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched setae (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 3 genualae (
<emphasis box="[881,894,1080,1103]" italics="true" pageId="6" pageNumber="423">σ</emphasis>
), long setiform microgenuala
<emphasis box="[1229,1242,1080,1103]" italics="true" pageId="6" pageNumber="423">κ</emphasis>
(); tibia 8B, 2 very long, curved, striated tibialae
<emphasis box="[756,771,1115,1138]" italics="true" pageId="6" pageNumber="423">φ</emphasis>
(), long setiform microtibiala (
<emphasis box="[1087,1100,1115,1138]" italics="true" pageId="6" pageNumber="423">κ</emphasis>
); tarsus 21B, tarsala (
<emphasis box="[1334,1353,1115,1138]" italics="true" pageId="6" pageNumber="423">ω</emphasis>
), famulus (
<emphasis box="[1477,1487,1115,1138]" italics="true" pageId="6" pageNumber="423">ε</emphasis>
) distal to tarsala, subterminala (
<emphasis box="[821,832,1148,1172]" italics="true" pageId="6" pageNumber="423">ζ</emphasis>
), nude parasubterminala (
<emphasis box="[1117,1127,1149,1172]" italics="true" pageId="6" pageNumber="423">z</emphasis>
), pretarsala (
<emphasis box="[1270,1281,1148,1172]" italics="true" pageId="6" pageNumber="423">ζ</emphasis>
). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala
<emphasis box="[1174,1187,1184,1207]" italics="true" pageId="6" pageNumber="423">σ</emphasis>
(); tibia 6B, 2 tibialae (
<emphasis box="[1428,1443,1184,1207]" italics="true" pageId="6" pageNumber="423">φ</emphasis>
) in tandem; tarsus 16B, tarsala II (
<emphasis box="[821,840,1219,1242]" italics="true" pageId="6" pageNumber="423">ω</emphasis>
), famulus (
<emphasis box="[964,974,1219,1242]" italics="true" pageId="6" pageNumber="423">ε</emphasis>
) near and slightly proximal to tarsala, pretarsala
</paragraph>
<paragraph blockId="6.[491,1564,630,1311]" box="[491,1493,1252,1277]" pageId="6" pageNumber="423">
(
<emphasis box="[499,510,1252,1276]" italics="true" pageId="6" pageNumber="423">ζ</emphasis>
). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala
<emphasis box="[1407,1420,1253,1276]" italics="true" pageId="6" pageNumber="423">σ</emphasis>
(); tibia
</paragraph>
<paragraph blockId="6.[491,1564,630,1311]" box="[491,771,1287,1311]" pageId="6" pageNumber="423">
6B, tibiala (
<emphasis box="[618,633,1288,1311]" italics="true" pageId="6" pageNumber="423">φ</emphasis>
); tarsus 14B.
</paragraph>
</subSubSection>
</treatment>
</document>