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<document id="B54C9FDC62305C52603FB272D08AFDCF" ID-CLB-Dataset="27005" ID-DOI="10.11646/zootaxa.4586.3.2" ID-GBIF-Dataset="915dc71e-0d15-4a61-a2e7-f24bc6d647e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="2647202" ID-ZooBank="DAFD77FB-77E6-4C1C-BCB0-09B4420977ED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555663597398" checkinUser="plazi" docAuthor="Marceniuk, Alexandre Pires, Ingenito, Leonardo Ferreira Da Silva, Lima, Flávio César Thadeo, Gasparini, João Luiz Rosetti &amp; Oliveira, Claudio" docDate="2019" docId="03D0CE16C02A034D988DFC57BD9CFAAA" docLanguage="en" docName="zootaxa.4586.3.2.pdf" docOrigin="Zootaxa 4586 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paragenidens grandoculis Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira 2019, new combination" docType="treatment" docVersion="12" lastPageNumber="434" masterDocId="FFE9B66EC02F0344981AFFD8BC2CFFD2" masterDocTitle="Systematics, biogeography and conservation of Paragenidens grandoculis n. gen. and n. comb. (Siluriformes; Ariidae), a critically endangered species from southeastern Brazil" masterLastPageNumber="444" masterPageNumber="425" pageNumber="430" updateTime="1698723098970" updateUser="ExternalLinkService">
<mods:mods id="EE135D5B6093D5C7E007630885752115" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1BFD8FDDCA4A55DAA472A43858CE5832">Systematics, biogeography and conservation of Paragenidens grandoculis n. gen. and n. comb. (Siluriformes; Ariidae), a critically endangered species from southeastern Brazil</mods:title>
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<mods:name id="7F03199159E197D03B9ABF1EC4B090DA" type="personal">
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<mods:namePart id="438755DF434D1634EA389090C28580E5">Marceniuk, Alexandre Pires</mods:namePart>
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<mods:namePart id="EAC0EF9DCE3CEE128DC494F4AA1610FE">Ingenito, Leonardo Ferreira Da Silva</mods:namePart>
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<mods:name id="A2B3A7194CEE30405109E12D73B56582" type="personal">
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<mods:namePart id="618FF181795CF24A679CCAE636A5D482">Lima, Flávio César Thadeo</mods:namePart>
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<mods:namePart id="32DAADA6D404471D634154C5A09A4B5B">Gasparini, João Luiz Rosetti</mods:namePart>
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<mods:namePart id="6629E5328153B56AF175834EF5F51FA8">Oliveira, Claudio</mods:namePart>
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<mods:title id="DE55D2BFAFEEC0DF678EE41C13AC2D12">Zootaxa</mods:title>
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<mods:date id="7980E5BB1E6151247B330334CCF127F4">2019</mods:date>
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<mods:number id="55300DB7F6B8BF2CCBB6B92C54069919">2019-04-18</mods:number>
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<mods:number id="AE6209A8F1019C163E059FE7E4388B24">4586</mods:number>
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<treatment id="03D0CE16C02A034D988DFC57BD9CFAAA" ID-DOI="http://doi.org/10.5281/zenodo.3510438" ID-GBIF-Taxon="156827773" ID-Zenodo-Dep="3510438" LSID="urn:lsid:plazi:treatment:03D0CE16C02A034D988DFC57BD9CFAAA" httpUri="http://treatment.plazi.org/id/03D0CE16C02A034D988DFC57BD9CFAAA" lastPageId="9" lastPageNumber="434" pageId="5" pageNumber="430">
<subSubSection id="C3632C8BC02A0341988DFC57BF96FC7B" box="[151,954,911,937]" pageId="5" pageNumber="430" type="nomenclature">
<paragraph id="8BC67F00C02A0341988DFC57BF96FC7B" blockId="5.[151,954,911,971]" box="[151,954,911,937]" pageId="5" pageNumber="430">
<heading id="D08EC86CC02A0341988DFC57BF96FC7B" bold="true" box="[151,954,911,937]" fontSize="11" level="1" pageId="5" pageNumber="430" reason="1">
<emphasis id="B90DA312C02A0341988DFC57BF96FC7B" bold="true" box="[151,954,911,937]" pageId="5" pageNumber="430">
<taxonomicName id="4C790483C02A0341988DFC57BEFAFC7B" authority="(Steindachner 1877)" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[151,726,911,937]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="grandoculis" status="comb. nov.">
<emphasis id="B90DA312C02A0341988DFC57BDFFFC7B" bold="true" box="[151,467,911,937]" italics="true" pageId="5" pageNumber="430">Paragenidens grandoculis</emphasis>
(Steindachner 1877)
</taxonomicName>
,
<taxonomicNameLabel id="A23E1E69C02A03419AF9FC57BF96FC7B" box="[739,954,911,937]" pageId="5" pageNumber="430" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3632C8BC02A0341988DFC6ABD94FC19" box="[151,440,946,971]" pageId="5" pageNumber="430" type="description">
<paragraph id="8BC67F00C02A0341988DFC6ABD94FC19" blockId="5.[151,954,911,971]" box="[151,440,946,971]" pageId="5" pageNumber="430">
(
<figureCitation id="13426385C02A03419884FC6ABD27FC19" box="[158,267,946,971]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1757]" captionTargetBox="[151,1435,193,1714]" captionTargetId="figure@7.[151,1435,193,1714]" captionTargetPageId="7" captionText="FIGURE 2. Body and head in lateral and dorsal view of all Paragenidens gradoculis specimens in zoological collections. (A). NMW 48272, rio Doce, Espírito Santo state. (B). MCZ 7681 (Museum of Comparative Zoology, Harvard University), rio Doce, Espírito Santo state, scale 50 mm. (C). MZUSP 2295, lagoa Feia, Rio de Janeiro, 251 mm SL. (D). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo. (E). MZUSP 1010414, 170 mm SL, lagoa Juparanã, Espírito Santo state. (F). LBP 24068, 240 mm SL, lagoa Nova, Espírito Santo state." httpUri="https://zenodo.org/record/2647206/files/figure.png" pageId="5" pageNumber="430">Figures 2</figureCitation>
,
<figureCitation id="13426385C02A0341990DFC6BBD09FC19" box="[279,293,947,971]" captionStart="FIGURE 3" captionStartId="8.[151,250,1718,1740]" captionTargetBox="[151,1435,841,1697]" captionTargetId="figure@8.[151,1435,841,1697]" captionTargetPageId="8" captionText="FIGURE 3. Accessory tooth plates and mesial face of the first and second branchial arches. (A). holotype NMW 48272, rio Doce, Espírito Santo. (B). MZUSP 1010414, 156 mm SL, lagoa Juparanã, Espírito Santo state. (C). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo state. (D). MNRJ 2295, 251 mm SL, lagoa Feia, São João da Barra, Rio de Janeiro state." httpUri="https://zenodo.org/record/2647208/files/figure.png" pageId="5" pageNumber="430">3</figureCitation>
and
<tableCitation id="C6FB4ABBC02A03419946FC6ABD80FC19" box="[348,428,946,971]" captionStart="TABLE 1" captionStartId="6.[151,239,152,175]" captionTargetPageId="6" captionText="TABLE 1. Morphometric data for Paragenidens grandoculis. Standard length is expressed in millimeters and all other measurements are expressed in percentages of standard length. A. Type-specimens (MCZ 7680 and MCZ 7681). B. Non type-specimens from lagoa Juparanã and lagoa Nova, Espirito Santo State. C. Non type-specimens from lagoa Feia Rio de Janeiro State." httpUri="http://table.plazi.org/id/DF062F88C0290342988DFF40BD6AFEDD" pageId="5" pageNumber="430" tableUuid="DF062F88C0290342988DFF40BD6AFEDD">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3632C8BC02A0341988DFC23B8CAFBBE" pageId="5" pageNumber="430" type="reference_group">
<paragraph id="8BC67F00C02A0341988DFC23B889FBC2" blockId="5.[151,1436,1018,1132]" box="[151,1189,1018,1040]" pageId="5" pageNumber="430">
<taxonomicName id="4C790483C02A0341988DFC23BE68FBC2" ID-CoL="GQV6" authority="Steindachner 1877: 644" authorityName="Steindachner" authorityPageNumber="644" authorityYear="1877" box="[151,580,1018,1040]" class="Actinopterygii" family="Ariidae" genus="Arius" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C02A0341988DFC23BD61FBC2" box="[151,333,1018,1040]" italics="true" pageId="5" pageNumber="430">Arius grandoculis</emphasis>
Steindachner 1877: 644
</taxonomicName>
646, pl. 11 (type-locality: &quot;Rio Doce&quot; southeastern
<collectingCountry id="F36E3F90C02A03419C4DFC22B8B5FBC2" box="[1111,1177,1018,1040]" name="Brazil" pageId="5" pageNumber="430">Brazil</collectingCountry>
).
</paragraph>
<paragraph id="8BC67F00C02A0341988DFBC2B8CAFBBE" blockId="5.[151,1436,1018,1132]" pageId="5" pageNumber="430">
<treatmentCitationGroup id="AB69582EC02A0341988DFBC2B8CAFBBE" pageId="5" pageNumber="430">
<taxonomicName id="4C790483C02A0341988DFBC2BDBEFBFD" ID-CoL="6W7KH" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[151,402,1049,1071]" class="Actinopterygii" family="Ariidae" genus="Potamarius" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C02A0341988DFBC2BDBEFBFD" box="[151,402,1049,1071]" italics="true" pageId="5" pageNumber="430">Potamarius grandoculis</emphasis>
</taxonomicName>
.-Marceniuk &amp; Ferraris 2003:452 (only name).—Marceniuk &amp; Menezes 2003:43 (only name).—
<treatmentCitation id="0AD85911C02A034198DDFBEFBDB8FB9F" author="Marceniuk, A. P." box="[199,404,1079,1101]" page="98" pageId="5" pageNumber="430" year="2005">
<bibRefCitation id="EFE802F1C02A034198DDFBEFBD5BFB9F" author="Marceniuk, A. P." box="[199,375,1079,1101]" pageId="5" pageNumber="430" pagination="89 - 101" refId="ref11913" refString="Marceniuk, A. P. (2005) Chave para identificacao das especies de bagres marinhos (Siluriformes, Ariidae) da costa Brasileira. Boletim do Instituto do Pesca, 31, 89 - 101." type="journal article" year="2005">Marceniuk 2005</bibRefCitation>
:98
</treatmentCitation>
(identification key).-
<treatmentCitation id="0AD85911C02A03419A69FBEFBFE0FB9F" ID-DOI="http://doi.org/10.5281/zenodo.3509301" ID-Zenodo-Dep="3509301" author="Marceniuk, A. P. &amp; Menezes, N. A." box="[627,972,1079,1101]" httpUri="http://treatment.plazi.org/id/84FBD122474C0161716BEAC83456A60B" page="98" pageId="5" pageNumber="430" year="2007">
<bibRefCitation id="EFE802F1C02A03419A69FBEFBF83FB9F" author="Marceniuk, A. P. &amp; Menezes, N. A." box="[627,943,1079,1101]" pageId="5" pageNumber="430" pagination="1 - 126" refId="ref12009" refString="Marceniuk, A. P. &amp; Menezes, N. A. (2007) Systematics of the family Ariidae (Ostariophysi, Siluriformes), with a redefinition of the genera. Zootaxa, 1416 (1), 1 - 126. https: // doi. org / 10.11646 / zootaxa. 1416.1.1" type="journal article" year="2007">Marceniuk &amp; Menezes 2007</bibRefCitation>
:98
</treatmentCitation>
(taxonomy),
<treatmentCitation id="0AD85911C02A03419C7BFBEFB9B0FB9F" author="Vieira, F. &amp; Gasparini, J. L." box="[1121,1436,1079,1101]" page="90" pageId="5" pageNumber="430" year="2007">
<bibRefCitation id="EFE802F1C02A03419C7BFBEFB95EFB9F" author="Vieira, F. &amp; Gasparini, J. L." box="[1121,1394,1079,1101]" pageId="5" pageNumber="430" pagination="87 - 104" refId="ref13922" refString="Vieira, F. &amp; Gasparini, J. L. (2007) Os Peixes Ameacados de Extincao no Estado do Espirito Santo. 87 - 104. In: Passamani, M. &amp; Mendes, S. L. (Orgs.). Especies da Fauna Ameacada de Extincao no Estado do Espirito Santo. Vitoria: Instituto de Pesquisas da Mata Atlantica, 140 pp." type="book chapter" year="2007">Vieira &amp; Gasparini 2007</bibRefCitation>
: 90
</treatmentCitation>
(conservation).-Ferraris 2007:53 (only name).—
<treatmentCitation id="0AD85911C02A03419AB7FB8EBF91FBBE" author="Marceniuk, A. P. &amp; Menezes, N. A. &amp; Britto, M. R." box="[685,957,1110,1132]" page="668" pageId="5" pageNumber="430" year="2012">
<bibRefCitation id="EFE802F1C02A03419AB7FB8EBFB8FBBE" author="Marceniuk, A. P. &amp; Menezes, N. A. &amp; Britto, M. R." box="[685,916,1110,1132]" pageId="5" pageNumber="430" pagination="534 - 669" refId="ref12122" refString="Marceniuk, A. P., Menezes, N. A. &amp; Britto, M. R. (2012) Phylogenetic analysis of the family Ariidae (Ostariophysi: Siluriformes), with an hypothesis on the monophyly and relationships of the genera. Zoological Journal of the Linnean Society, 165, 534 - 669. https: // doi. org / 10.1111 / j. 1096 - 3642.2012.00822. x" type="journal article" year="2012">
Marceniuk
<emphasis id="B90DA312C02A03419B3EFB8FBF7DFBBE" box="[804,849,1110,1132]" italics="true" pageId="5" pageNumber="430">et al</emphasis>
. 2012
</bibRefCitation>
:668
</treatmentCitation>
(phylogenetic relationships).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3632C8BC02A0341988DFB41BF09F9CB" pageId="5" pageNumber="430" type="diagnosis">
<paragraph id="8BC67F00C02A0341988DFB41BF09F9CB" blockId="5.[151,1437,1177,1994]" pageId="5" pageNumber="430">
<emphasis id="B90DA312C02A0341988DFB41BD26FB60" bold="true" box="[151,266,1177,1202]" pageId="5" pageNumber="430">Diagnosis</emphasis>
.
<taxonomicName id="4C790483C02A03419902FB41BE6DFB63" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[280,577,1177,1201]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C02A03419902FB41BE6DFB63" box="[280,577,1177,1201]" italics="true" pageId="5" pageNumber="430">Paragenidens grandoculis</emphasis>
</taxonomicName>
is distinguished from other Neotropical ariids by having vestigial accessory tooth plates [
<emphasis id="B90DA312C02A03419935FB66BD6AFB07" box="[303,326,1214,1237]" italics="true" pageId="5" pageNumber="430">vs</emphasis>
. very conspicuous accessory tooth plates in all other ariid representatives including
<taxonomicName id="4C790483C02A03419D03FB66BD1CFB2B" class="Actinopterygii" family="Ariidae" genus="Potamarius" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="ussumacintae">
<emphasis id="B90DA312C02A03419D03FB66BD1CFB2B" italics="true" pageId="5" pageNumber="430">Potamarius ussumacintae</emphasis>
</taxonomicName>
and excluding
<emphasis id="B90DA312C02A034199C4FB39BF48FB2B" box="[478,868,1249,1273]" italics="true" pageId="5" pageNumber="430">
<taxonomicName id="4C790483C02A034199C4FB39BEC9FB2B" authorityName="Boulenger" authorityYear="1898" box="[478,741,1249,1273]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="labiata">Chinchaysuyoa labiata</taxonomicName>
,
<taxonomicName id="4C790483C02A03419AEBFB39BF48FB2B" box="[753,868,1249,1273]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="ortegai">C. ortegai</taxonomicName>
</emphasis>
and mouth-brooders males of
<taxonomicName id="4C790483C02A03419CDAFB39B9B8FB2B" authority="genidens" authorityName="genidens (Cuvier" authorityYear="1829" box="[1216,1428,1249,1273]" class="Actinopterygii" family="Ariidae" genus="Genidens" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="genidens">
<emphasis id="B90DA312C02A03419CDAFB39B9B8FB2B" box="[1216,1428,1249,1273]" italics="true" pageId="5" pageNumber="430">Genidens genidens</emphasis>
</taxonomicName>
, or absent in
<taxonomicName id="4C790483C02A03419930FADEBDEAFACF" authorityName="Hubbs &amp; Miller" authorityYear="1960" box="[298,454,1285,1309]" class="Actinopterygii" family="Ariidae" genus="Potamarius" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="izabalensis">
<emphasis id="B90DA312C02A03419930FADEBDEAFACF" box="[298,454,1285,1309]" italics="true" pageId="5" pageNumber="430">P. izabalensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C790483C02A03419A18FADEBE5EFACF" baseAuthorityName="Evermann &amp; Goldsborough" baseAuthorityYear="1902" box="[514,626,1285,1309]" class="Actinopterygii" family="Ariidae" genus="Potamarius" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="nelsoni">
<emphasis id="B90DA312C02A03419A18FADEBE5EFACF" box="[514,626,1285,1309]" italics="true" pageId="5" pageNumber="430">P. nelsoni</emphasis>
</taxonomicName>
,
<figureCitation id="13426385C02A03419A99FADDBEC8FACF" box="[643,740,1285,1310]" captionStart="FIGURE 3" captionStartId="8.[151,250,1718,1740]" captionTargetBox="[151,1435,841,1697]" captionTargetId="figure@8.[151,1435,841,1697]" captionTargetPageId="8" captionText="FIGURE 3. Accessory tooth plates and mesial face of the first and second branchial arches. (A). holotype NMW 48272, rio Doce, Espírito Santo. (B). MZUSP 1010414, 156 mm SL, lagoa Juparanã, Espírito Santo state. (C). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo state. (D). MNRJ 2295, 251 mm SL, lagoa Feia, São João da Barra, Rio de Janeiro state." httpUri="https://zenodo.org/record/2647208/files/figure.png" pageId="5" pageNumber="430">Figure 3</figureCitation>
(a), (b)].
<taxonomicName id="4C790483C02A03419B4AFADDB850FACF" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[848,1148,1285,1309]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C02A03419B4AFADDB850FACF" box="[848,1148,1285,1309]" italics="true" pageId="5" pageNumber="430">Paragenidens grandoculis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C790483C02A03419D03FADEBD38FA93" authorityName="Hubbs &amp; Miller" authorityYear="1960" class="Actinopterygii" family="Ariidae" genus="Potamarius" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="izabalensis">
<emphasis id="B90DA312C02A03419D03FADEBD38FA93" italics="true" pageId="5" pageNumber="430">Potamarius izabalensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C790483C02A03419955FAF2BD92FA93" baseAuthorityName="Evermann &amp; Goldsborough" baseAuthorityYear="1902" box="[335,446,1321,1345]" class="Actinopterygii" family="Ariidae" genus="Potamarius" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="nelsoni">
<emphasis id="B90DA312C02A03419955FAF2BD92FA93" box="[335,446,1321,1345]" italics="true" pageId="5" pageNumber="430">P. nelsoni</emphasis>
</taxonomicName>
from Central America, and the sympatric
<taxonomicName id="4C790483C02A03419BB6FAF1B8AEFA93" authority="genidens" authorityName="genidens (Cuvier" authorityYear="1829" box="[940,1154,1321,1345]" class="Actinopterygii" family="Ariidae" genus="Genidens" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="genidens">
<emphasis id="B90DA312C02A03419BB6FAF1B8AEFA93" box="[940,1154,1321,1345]" italics="true" pageId="5" pageNumber="430">Genidens genidens</emphasis>
</taxonomicName>
, by having snout length 1.01.5 times in cephalic shield width at frontal area [
<emphasis id="B90DA312C02A03419B0BFA96BF03FAB7" box="[785,815,1358,1381]" italics="true" pageId="5" pageNumber="430">vs.</emphasis>
1.61.7 times in
<taxonomicName id="4C790483C02A03419BE5FA95B8A9FAB7" authorityName="genidens (Cuvier" authorityYear="1829" box="[1023,1157,1357,1381]" class="Actinopterygii" family="Ariidae" genus="Genidens" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="genidens">
<emphasis id="B90DA312C02A03419BE5FA95B8A9FAB7" box="[1023,1157,1357,1381]" italics="true" pageId="5" pageNumber="430">G. genidens</emphasis>
</taxonomicName>
and 0.50.8 times in
<taxonomicName id="4C790483C02A03419D9DFA96BD38FA5B" authorityName="Hubbs &amp; Miller" authorityYear="1960" class="Actinopterygii" family="Ariidae" genus="Potamarius" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="izabalensis">
<emphasis id="B90DA312C02A03419D9DFA96BD38FA5B" italics="true" pageId="5" pageNumber="430">P. izabalensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C790483C02A0341994BFAAABD91FA5B" baseAuthorityName="Evermann &amp; Goldsborough" baseAuthorityYear="1902" box="[337,445,1393,1417]" class="Actinopterygii" family="Ariidae" genus="Potamarius" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="nelsoni">
<emphasis id="B90DA312C02A0341994BFAAABD91FA5B" box="[337,445,1393,1417]" italics="true" pageId="5" pageNumber="430">P. nelsoni</emphasis>
</taxonomicName>
,
<figureCitation id="13426385C02A034199C8FAA9BE19FA58" box="[466,565,1393,1418]" captionStart="FIGURE 4" captionStartId="12.[151,250,1816,1838]" captionTargetBox="[410,1166,206,1795]" captionTargetId="figure@12.[397,1186,193,1814]" captionTargetPageId="12" captionText="FIGURE 4. (A). Principal component analysis. Type-specimens of Paragenidens grandoculis (black), specimens from Espirito Santo state (light blue), from Rio de Janeiro state (dark blue), specimens of Genidens genidens from Espírito Santo, São Paulo, Paraná and Rio Grande do Sul states (light green), from the rio Doce channel (dark green), Potamarius izabalensis (orange), Potamarius nelsoni (red). (B). Plot of width of cephalic shield at frontal area versus snout length. (C). Body width versus snout length. Paragenidens grandoculis type-specimens (filled square), non type-specimens from Espito Santo state (opened square), non-type specimens from Rio de Janeiro state (asterisk), Genidens genidens from Espírito Santo, São Paulo and Rio Grande do Sul states (filled triangle), P. izabalensis (circle filled), and P. nelsoni (trapeze filled)." httpUri="https://zenodo.org/record/2647210/files/figure.png" pageId="5" pageNumber="430">Figure 4</figureCitation>
(b)], and snout length 2.22.7 times in body width [
<emphasis id="B90DA312C02A03419C8FFAAAB880FA5B" box="[1173,1196,1394,1417]" italics="true" pageId="5" pageNumber="430">vs</emphasis>
. 2.94.3 times in
<taxonomicName id="4C790483C02A03419D9EFAA9BCD7FA7F" authorityName="genidens (Cuvier" authorityYear="1829" class="Actinopterygii" family="Ariidae" genus="Genidens" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="genidens">
<emphasis id="B90DA312C02A03419D9EFAA9BCD7FA7F" italics="true" pageId="5" pageNumber="430">G. genidens</emphasis>
</taxonomicName>
, 1.61.8 times in
<taxonomicName id="4C790483C02A034199CEFA4EBE5DFA7F" authorityName="Hubbs &amp; Miller" authorityYear="1960" box="[468,625,1429,1453]" class="Actinopterygii" family="Ariidae" genus="Potamarius" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="izabalensis">
<emphasis id="B90DA312C02A034199CEFA4EBE5DFA7F" box="[468,625,1429,1453]" italics="true" pageId="5" pageNumber="430">P. izabalensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C790483C02A03419AB5FA4EBF33FA7F" baseAuthorityName="Evermann &amp; Goldsborough" baseAuthorityYear="1902" box="[687,799,1429,1453]" class="Actinopterygii" family="Ariidae" genus="Potamarius" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="nelsoni">
<emphasis id="B90DA312C02A03419AB5FA4EBF33FA7F" box="[687,799,1429,1453]" italics="true" pageId="5" pageNumber="430">P. nelsoni</emphasis>
</taxonomicName>
,
<figureCitation id="13426385C02A03419B28FA4DBFBAFA7F" box="[818,918,1429,1454]" captionStart="FIGURE 4" captionStartId="12.[151,250,1816,1838]" captionTargetBox="[410,1166,206,1795]" captionTargetId="figure@12.[397,1186,193,1814]" captionTargetPageId="12" captionText="FIGURE 4. (A). Principal component analysis. Type-specimens of Paragenidens grandoculis (black), specimens from Espirito Santo state (light blue), from Rio de Janeiro state (dark blue), specimens of Genidens genidens from Espírito Santo, São Paulo, Paraná and Rio Grande do Sul states (light green), from the rio Doce channel (dark green), Potamarius izabalensis (orange), Potamarius nelsoni (red). (B). Plot of width of cephalic shield at frontal area versus snout length. (C). Body width versus snout length. Paragenidens grandoculis type-specimens (filled square), non type-specimens from Espito Santo state (opened square), non-type specimens from Rio de Janeiro state (asterisk), Genidens genidens from Espírito Santo, São Paulo and Rio Grande do Sul states (filled triangle), P. izabalensis (circle filled), and P. nelsoni (trapeze filled)." httpUri="https://zenodo.org/record/2647210/files/figure.png" pageId="5" pageNumber="430">Figure 4</figureCitation>
(c)] and from
<taxonomicName id="4C790483C02A03419C26FA4DB96FFA7F" authorityName="Boulenger" authorityYear="1898" box="[1084,1347,1429,1453]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="labiata">
<emphasis id="B90DA312C02A03419C26FA4DB96FFA7F" box="[1084,1347,1429,1453]" italics="true" pageId="5" pageNumber="430">Chinchaysuyoa labiata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C790483C02A03419D98FA4DBCC4FA03" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="ortegai">
<emphasis id="B90DA312C02A03419D98FA4DBCC4FA03" italics="true" pageId="5" pageNumber="430">C. ortegai</emphasis>
</taxonomicName>
from
<collectingCountry id="F36E3F90C02A0341992AFA61BDA1FA00" box="[304,397,1465,1490]" name="Ecuador" pageId="5" pageNumber="430">Ecuador</collectingCountry>
and
<collectingCountry id="F36E3F90C02A034199D2FA62BDD2FA03" box="[456,510,1466,1489]" name="Peru" pageId="5" pageNumber="430">Peru</collectingCountry>
, in the eastern Pacific, by longer head (27.029.4% SL
<emphasis id="B90DA312C02A03419C98FA62B8B5FA03" box="[1154,1177,1466,1489]" italics="true" pageId="5" pageNumber="430">vs</emphasis>
. 17.919.7% SL in
<taxonomicName id="4C790483C02A03419D98FA61BCCAFA27" authorityName="Boulenger" authorityYear="1898" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="labiata">
<emphasis id="B90DA312C02A03419D98FA61BCCAFA27" italics="true" pageId="5" pageNumber="430">C. labiata</emphasis>
</taxonomicName>
and 19.921.7% SL in
<taxonomicName id="4C790483C02A034199E1FA05BE43FA27" box="[507,623,1501,1525]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="ortegai">
<emphasis id="B90DA312C02A034199E1FA05BE43FA27" box="[507,623,1501,1525]" italics="true" pageId="5" pageNumber="430">C. ortegai</emphasis>
</taxonomicName>
) and narrow cephalic shield at lateral ethmoid area (9.711.5% SL
<emphasis id="B90DA312C02A03419D67FA06B9B8FA27" box="[1405,1428,1502,1525]" italics="true" pageId="5" pageNumber="430">vs</emphasis>
. 11.712.0% SL in
<taxonomicName id="4C790483C02A03419972F9D9BDF4F9CB" authorityName="Boulenger" authorityYear="1898" box="[360,472,1537,1561]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="labiata">
<emphasis id="B90DA312C02A03419972F9D9BDF4F9CB" box="[360,472,1537,1561]" italics="true" pageId="5" pageNumber="430">C. labiata</emphasis>
</taxonomicName>
and
<quantity id="4C81D2E5C02A03419A15F9D9BEB0F9C8" box="[527,668,1537,1562]" metricMagnitude="-1" metricUnit="m" metricValue="3.2639000000000005" metricValueMax="3.3782000000000005" metricValueMin="3.1496000000000004" pageId="5" pageNumber="430" unit="in" value="12.85" valueMax="13.3" valueMin="12.4">12.413.3 in</quantity>
<taxonomicName id="4C790483C02A03419AB9F9D9BF39F9CB" box="[675,789,1537,1561]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="430" phylum="Chordata" rank="species" species="ortegai">
<emphasis id="B90DA312C02A03419AB9F9D9BF39F9CB" box="[675,789,1537,1561]" italics="true" pageId="5" pageNumber="430">C. ortegai</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3632C8BC02A034C98DDF9FDBDABFCC7" lastPageId="8" lastPageNumber="433" pageId="5" pageNumber="430" type="description">
<paragraph id="8BC67F00C02A034198DDF9FDBF8AF818" blockId="5.[151,1437,1177,1994]" pageId="5" pageNumber="430">
<emphasis id="B90DA312C02A034198DDF9FDBD7DF9EC" bold="true" box="[199,337,1573,1598]" pageId="5" pageNumber="430">Description</emphasis>
. Morphometrics data summarized in
<tableCitation id="C6FB4ABBC02A03419B1CF9FDBF71F9EF" box="[774,861,1573,1598]" captionStart="TABLE 1" captionStartId="6.[151,239,152,175]" captionTargetPageId="6" captionText="TABLE 1. Morphometric data for Paragenidens grandoculis. Standard length is expressed in millimeters and all other measurements are expressed in percentages of standard length. A. Type-specimens (MCZ 7680 and MCZ 7681). B. Non type-specimens from lagoa Juparanã and lagoa Nova, Espirito Santo State. C. Non type-specimens from lagoa Feia Rio de Janeiro State." httpUri="http://table.plazi.org/id/DF062F88C0290342988DFF40BD6AFEDD" pageId="5" pageNumber="430" tableUuid="DF062F88C0290342988DFF40BD6AFEDD">Table 1</tableCitation>
. Head long, very narrow and depressed at lateral ethmoid and frontal area, with profile elevated posteriorly, straight from mesethmoid to parieto-supraoccipital process. Snout long and narrow. Anterior nostril round, with fleshy edge, posterior nostril covered by flap of skin; nostrils quite close to one another and distant to orbit, not connected by fleshy furrow. Eye lateral and very large; relatively close to each other. Three pairs of long teretiform barbels; maxillary barbel surpassing base of pectoralfin spine, lateral mental barbel surpassing margin of gill membrane, and mesial mental barbel not reaching margin of gill membrane. Mouth subterminal to terminal, relatively small and lower jaw arched. Lips thin, lower lip thinner than upper lip. Vomerian tooth plates absent. Accessory tooth plates vestigial. Premaxilla relatively long and narrow, with sharp teeth. Dentary separated at midline, with sharp teeth. Gill membranes fused, attached to isthmus. Fifteen to 17 acicular gill rakers on first arch, 1618 spike-shaped gill rakers on second arch. Mesial surfaces of first and second gill arches with or without developed gill rakers, lateral and mesial surfaces of first and second gill arches lacking fleshy papillae intercalated with gill rakers.
</paragraph>
<caption id="DF062F88C0290342988DFF40BD6AFEDD" ID-Table-UUID="DF062F88C0290342988DFF40BD6AFEDD" httpUri="http://table.plazi.org/id/DF062F88C0290342988DFF40BD6AFEDD" pageId="6" pageNumber="431" targetBox="[159,1399,298,1974]" targetIsTable="true" targetPageId="6">
<paragraph id="8BC67F00C0290342988DFF40BD6AFEDD" blockId="6.[151,1436,151,271]" pageId="6" pageNumber="431">
<emphasis id="B90DA312C0290342988DFF40BD27FF7D" bold="true" box="[151,267,152,175]" pageId="6" pageNumber="431">TABLE 1.</emphasis>
Morphometric data for
<taxonomicName id="4C790483C02903429A0AFF4FBF07FF7D" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[528,811,151,175]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="431" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C02903429A0AFF4FBF07FF7D" box="[528,811,151,175]" italics="true" pageId="6" pageNumber="431">Paragenidens grandoculis</emphasis>
</taxonomicName>
. Standard length is expressed in millimeters and all other measurements are expressed in percentages of standard length.
<emphasis id="B90DA312C02903429B26FF60BF63FF1D" bold="true" box="[828,847,184,207]" pageId="6" pageNumber="431">A</emphasis>
. Type-specimens (MCZ 7680 and MCZ 7681).
<emphasis id="B90DA312C02903429D55FF60B94DFF1D" bold="true" box="[1359,1377,184,207]" pageId="6" pageNumber="431">B</emphasis>
. Non type-specimens from lagoa Juparanã and lagoa Nova, Espirito Santo State.
<emphasis id="B90DA312C02903429BDAFF00BFFFFF3D" bold="true" box="[960,979,216,239]" pageId="6" pageNumber="431">C</emphasis>
. Non type-specimens from lagoa Feia Rio de Janeiro State.
</paragraph>
</caption>
<paragraph id="8BC67F00C02903429AFFFEF3B880F864" pageId="6" pageNumber="431">
<table id="F9798DA0C029FCBB9885FEF2B95BF864" box="[159,1399,298,1974]" gridcols="5" gridrows="41" pageId="6" pageNumber="431">
<tr id="35497D42C029FCBB9885FEF2B95BFE92" box="[159,1399,298,320]" gridrow="0" pageId="6" pageNumber="431" rowspan-0="1" rowspan-2="1">
<th id="7698143EC029FCBB9AFFFEF2BF66FE92" box="[741,842,298,320]" gridcol="1" gridrow="0" pageId="6" pageNumber="431">A</th>
<th id="7698143EC029FCBB9C1EFEF2B881FE92" box="[1028,1197,298,320]" gridcol="3" gridrow="0" pageId="6" pageNumber="431">B</th>
<th id="7698143EC029FCBB9D51FEF2B95BFE92" box="[1355,1399,298,320]" gridcol="4" gridrow="0" pageId="6" pageNumber="431">C</th>
</tr>
<tr id="35497D42C029FCBB9885FE8CB95BFEB8" box="[159,1399,340,362]" gridrow="1" pageId="6" pageNumber="431">
<th id="7698143EC029FCBB9885FE8CBE52FEB8" box="[159,638,340,362]" gridcol="0" gridrow="1" pageId="6" pageNumber="431">Standard length (mm)</th>
<td id="7698143EC029FCBB9AFFFE8CBF66FEB8" box="[741,842,340,362]" gridcol="1" gridrow="1" pageId="6" pageNumber="431">193196</td>
<td id="7698143EC029FCBB9BB4FE8CBFE4FEB8" box="[942,968,340,362]" gridcol="2" gridrow="1" pageId="6" pageNumber="431">10</td>
<td id="7698143EC029FCBB9C1EFE8CB881FEB8" box="[1028,1197,340,362]" gridcol="3" gridrow="1" pageId="6" pageNumber="431">156251</td>
<td id="7698143EC029FCBB9D51FE8CB95BFEB8" box="[1355,1399,340,362]" gridcol="4" gridrow="1" pageId="6" pageNumber="431">251</td>
</tr>
<tr id="35497D42C029FCBB9885FEA5B95BFE41" box="[159,1399,381,403]" gridrow="2" pageId="6" pageNumber="431" rowspan-1="1">
<th id="7698143EC029FCBB9885FEA5BE52FE41" box="[159,638,381,403]" gridcol="0" gridrow="2" pageId="6" pageNumber="431">Head length</th>
<td id="7698143EC029FCBB9BB4FEA5BFE4FE41" box="[942,968,381,403]" gridcol="2" gridrow="2" pageId="6" pageNumber="431">10</td>
<td id="7698143EC029FCBB9C1EFEA5B881FE41" box="[1028,1197,381,403]" gridcol="3" gridrow="2" pageId="6" pageNumber="431">27.0 (25.129.4)</td>
<td id="7698143EC029FCBB9D51FEA5B95BFE41" box="[1355,1399,381,403]" gridcol="4" gridrow="2" pageId="6" pageNumber="431">27.4</td>
</tr>
<tr id="35497D42C029FCBB9885FE7EB95BFE6E" box="[159,1399,422,444]" gridrow="3" pageId="6" pageNumber="431">
<th id="7698143EC029FCBB9885FE7EBE52FE6E" box="[159,638,422,444]" gridcol="0" gridrow="3" pageId="6" pageNumber="431">Snout length</th>
<td id="7698143EC029FCBB9AFFFE7EBF66FE6E" box="[741,842,422,444]" gridcol="1" gridrow="3" pageId="6" pageNumber="431">9.19.1</td>
<td id="7698143EC029FCBB9BB4FE7EBFE4FE6E" box="[942,968,422,444]" gridcol="2" gridrow="3" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFE7EB881FE6E" box="[1028,1197,422,444]" gridcol="3" gridrow="3" pageId="6" pageNumber="431">7.7 (7.09.0)</td>
<td id="7698143EC029FCBB9D51FE7EB95BFE6E" box="[1355,1399,422,444]" gridcol="4" gridrow="3" pageId="6" pageNumber="431">7.0</td>
</tr>
<tr id="35497D42C029FCBB9885FE08B95BFE34" box="[159,1399,464,486]" gridrow="4" pageId="6" pageNumber="431" rowspan-4="1">
<th id="7698143EC029FCBB9885FE08BE52FE34" box="[159,638,464,486]" gridcol="0" gridrow="4" pageId="6" pageNumber="431">Distance between anterior nostrils</th>
<td id="7698143EC029FCBB9AFFFE08BF66FE34" box="[741,842,464,486]" gridcol="1" gridrow="4" pageId="6" pageNumber="431">4.74.9</td>
<td id="7698143EC029FCBB9BB4FE08BFE4FE34" box="[942,968,464,486]" gridcol="2" gridrow="4" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EFE08B881FE34" box="[1028,1197,464,486]" gridcol="3" gridrow="4" pageId="6" pageNumber="431">4,7 (4,35.4)</td>
</tr>
<tr id="35497D42C029FCBB9885FE21B95BFDDD" box="[159,1399,505,527]" gridrow="5" pageId="6" pageNumber="431" rowspan-1="1">
<th id="7698143EC029FCBB9885FE21BE52FDDD" box="[159,638,505,527]" gridcol="0" gridrow="5" pageId="6" pageNumber="431">Distance between posterior nostrils</th>
<td id="7698143EC029FCBB9BB4FE21BFE4FDDD" box="[942,968,505,527]" gridcol="2" gridrow="5" pageId="6" pageNumber="431">10</td>
<td id="7698143EC029FCBB9C1EFE21B881FDDD" box="[1028,1197,505,527]" gridcol="3" gridrow="5" pageId="6" pageNumber="431">5.0 (4.65.4)</td>
<td id="7698143EC029FCBB9D51FE21B95BFDDD" box="[1355,1399,505,527]" gridcol="4" gridrow="5" pageId="6" pageNumber="431">5.1</td>
</tr>
<tr id="35497D42C029FCBB9885FDFAB95BFDEA" box="[159,1399,546,568]" gridrow="6" pageId="6" pageNumber="431" rowspan-4="1">
<th id="7698143EC029FCBB9885FDFABE52FDEA" box="[159,638,546,568]" gridcol="0" gridrow="6" pageId="6" pageNumber="431">Distance between anterior nostril and orbit</th>
<td id="7698143EC029FCBB9AFFFDFABF66FDEA" box="[741,842,546,568]" gridcol="1" gridrow="6" pageId="6" pageNumber="431">6.06.2</td>
<td id="7698143EC029FCBB9BB4FDFABFE4FDEA" box="[942,968,546,568]" gridcol="2" gridrow="6" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EFDFAB881FDEA" box="[1028,1197,546,568]" gridcol="3" gridrow="6" pageId="6" pageNumber="431">7.3 (7.07.7)</td>
</tr>
<tr id="35497D42C029FCBB9885FD94B95BFDB0" box="[159,1399,588,610]" gridrow="7" pageId="6" pageNumber="431" rowspan-4="1">
<th id="7698143EC029FCBB9885FD94BE52FDB0" box="[159,638,588,610]" gridcol="0" gridrow="7" pageId="6" pageNumber="431">Distance between posterior nostril and orbit</th>
<td id="7698143EC029FCBB9AFFFD94BF66FDB0" box="[741,842,588,610]" gridcol="1" gridrow="7" pageId="6" pageNumber="431">4.54.5</td>
<td id="7698143EC029FCBB9BB4FD94BFE4FDB0" box="[942,968,588,610]" gridcol="2" gridrow="7" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFD94B881FDB0" box="[1028,1197,588,610]" gridcol="3" gridrow="7" pageId="6" pageNumber="431">5.5 (5.16.2)</td>
</tr>
<tr id="35497D42C029FCBB9885FDADB95BFD59" box="[159,1399,629,651]" gridrow="8" pageId="6" pageNumber="431">
<th id="7698143EC029FCBB9885FDADBE52FD59" box="[159,638,629,651]" gridcol="0" gridrow="8" pageId="6" pageNumber="431">Orbital diameter</th>
<td id="7698143EC029FCBB9AFFFDADBF66FD59" box="[741,842,629,651]" gridcol="1" gridrow="8" pageId="6" pageNumber="431">5.35.5</td>
<td id="7698143EC029FCBB9BB4FDADBFE4FD59" box="[942,968,629,651]" gridcol="2" gridrow="8" pageId="6" pageNumber="431">10</td>
<td id="7698143EC029FCBB9C1EFDADB881FD59" box="[1028,1197,629,651]" gridcol="3" gridrow="8" pageId="6" pageNumber="431">5.8 (4.76.8)</td>
<td id="7698143EC029FCBB9D51FDADB95BFD59" box="[1355,1399,629,651]" gridcol="4" gridrow="8" pageId="6" pageNumber="431">5.0</td>
</tr>
<tr id="35497D42C029FCBB9885FD46B95BFD66" box="[159,1399,670,692]" gridrow="9" pageId="6" pageNumber="431" rowspan-1="1">
<th id="7698143EC029FCBB9885FD46BE52FD66" box="[159,638,670,692]" gridcol="0" gridrow="9" pageId="6" pageNumber="431">Interorbital distance</th>
<td id="7698143EC029FCBB9BB4FD46BFE4FD66" box="[942,968,670,692]" gridcol="2" gridrow="9" pageId="6" pageNumber="431">10</td>
<td id="7698143EC029FCBB9C1EFD46B881FD66" box="[1028,1197,670,692]" gridcol="3" gridrow="9" pageId="6" pageNumber="431">10.0 (9.311.8)</td>
<td id="7698143EC029FCBB9D51FD46B95BFD66" box="[1355,1399,670,692]" gridcol="4" gridrow="9" pageId="6" pageNumber="431">11.0</td>
</tr>
<tr id="35497D42C029FCBB9885FD10B95BFD0C" box="[159,1399,712,734]" gridrow="10" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885FD10BE52FD0C" box="[159,638,712,734]" gridcol="0" gridrow="10" pageId="6" pageNumber="431">Maxillary barbel length</th>
<td id="7698143EC029FCBB9BB4FD10BFE4FD0C" box="[942,968,712,734]" gridcol="2" gridrow="10" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFD10B881FD0C" box="[1028,1197,712,734]" gridcol="3" gridrow="10" pageId="6" pageNumber="431">24.9 (17.528.7)</td>
</tr>
<tr id="35497D42C029FCBB9885FD29B95BFCD5" box="[159,1399,753,775]" gridrow="11" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885FD29BE52FCD5" box="[159,638,753,775]" gridcol="0" gridrow="11" pageId="6" pageNumber="431">Lateral mental barbel length</th>
<td id="7698143EC029FCBB9BB4FD29BFE4FCD5" box="[942,968,753,775]" gridcol="2" gridrow="11" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFD29B881FCD5" box="[1028,1197,753,775]" gridcol="3" gridrow="11" pageId="6" pageNumber="431">16.6 (11.918.9)</td>
</tr>
<tr id="35497D42C029FCBB9885FCC2B95BFCE2" box="[159,1399,794,816]" gridrow="12" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885FCC2BE52FCE2" box="[159,638,794,816]" gridcol="0" gridrow="12" pageId="6" pageNumber="431">Mesial mental barbel length</th>
<td id="7698143EC029FCBB9BB4FCC2BFE4FCE2" box="[942,968,794,816]" gridcol="2" gridrow="12" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EFCC2B881FCE2" box="[1028,1197,794,816]" gridcol="3" gridrow="12" pageId="6" pageNumber="431">10.8 (7.412.6)</td>
</tr>
<tr id="35497D42C029FCBB9885FC9CB95BFC88" box="[159,1399,836,858]" gridrow="13" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885FC9CBE52FC88" box="[159,638,836,858]" gridcol="0" gridrow="13" pageId="6" pageNumber="431">Mouth width</th>
<td id="7698143EC029FCBB9BB4FC9CBFE4FC88" box="[942,968,836,858]" gridcol="2" gridrow="13" pageId="6" pageNumber="431">3</td>
<td id="7698143EC029FCBB9C1EFC9CB881FC88" box="[1028,1197,836,858]" gridcol="3" gridrow="13" pageId="6" pageNumber="431">8.6 (8.19.1)</td>
</tr>
<tr id="35497D42C029FCBB9885FCB5B95BFC51" box="[159,1399,877,899]" gridrow="14" pageId="6" pageNumber="431" rowspan-4="1">
<th id="7698143EC029FCBB9885FCB5BE52FC51" box="[159,638,877,899]" gridcol="0" gridrow="14" pageId="6" pageNumber="431">Width of cephalic shield at lateral ethmoid area</th>
<td id="7698143EC029FCBB9AFFFCB5BF66FC51" box="[741,842,877,899]" gridcol="1" gridrow="14" pageId="6" pageNumber="431">10.710.7</td>
<td id="7698143EC029FCBB9BB4FCB5BFE4FC51" box="[942,968,877,899]" gridcol="2" gridrow="14" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFCB5B881FC51" box="[1028,1197,877,899]" gridcol="3" gridrow="14" pageId="6" pageNumber="431">10.7 (9.711.5)</td>
</tr>
<tr id="35497D42C029FCBB9885FC4EB95BFC7E" box="[159,1399,918,940]" gridrow="15" pageId="6" pageNumber="431">
<th id="7698143EC029FCBB9885FC4EBE52FC7E" box="[159,638,918,940]" gridcol="0" gridrow="15" pageId="6" pageNumber="431">Width of cephalic shield at frontals area</th>
<td id="7698143EC029FCBB9AFFFC4EBF66FC7E" box="[741,842,918,940]" gridcol="1" gridrow="15" pageId="6" pageNumber="431">9.09.6</td>
<td id="7698143EC029FCBB9BB4FC4EBFE4FC7E" box="[942,968,918,940]" gridcol="2" gridrow="15" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFC4EB881FC7E" box="[1028,1197,918,940]" gridcol="3" gridrow="15" pageId="6" pageNumber="431">6.2 (5.36.9)</td>
<td id="7698143EC029FCBB9D51FC4EB95BFC7E" box="[1355,1399,918,940]" gridcol="4" gridrow="15" pageId="6" pageNumber="431">7.2</td>
</tr>
<tr id="35497D42C029FCBB9885FC18B95BFC04" box="[159,1399,960,982]" gridrow="16" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885FC18BE52FC04" box="[159,638,960,982]" gridcol="0" gridrow="16" pageId="6" pageNumber="431">Width of cephalic shield at epioccipital area</th>
<td id="7698143EC029FCBB9BB4FC18BFE4FC04" box="[942,968,960,982]" gridcol="2" gridrow="16" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EFC18B881FC04" box="[1028,1197,960,982]" gridcol="3" gridrow="16" pageId="6" pageNumber="431">9.9 (8.011.5)</td>
</tr>
<tr id="35497D42C029FCBB9885FC31B95BFC2D" box="[159,1399,1001,1023]" gridrow="17" pageId="6" pageNumber="431" rowspan-1="1">
<th id="7698143EC029FCBB9885FC31BE52FC2D" box="[159,638,1001,1023]" gridcol="0" gridrow="17" pageId="6" pageNumber="431">Width of cephalic shield at supracleithrum area</th>
<td id="7698143EC029FCBB9BB4FC31BFE4FC2D" box="[942,968,1001,1023]" gridcol="2" gridrow="17" pageId="6" pageNumber="431">10</td>
<td id="7698143EC029FCBB9C1EFC31B881FC2D" box="[1028,1197,1001,1023]" gridcol="3" gridrow="17" pageId="6" pageNumber="431">16.1 (15.417.4)</td>
<td id="7698143EC029FCBB9D51FC31B95BFC2D" box="[1355,1399,1001,1023]" gridcol="4" gridrow="17" pageId="6" pageNumber="431">15.4</td>
</tr>
<tr id="35497D42C029FCBB9885FBCAB95BFBFA" box="[159,1399,1042,1064]" gridrow="18" pageId="6" pageNumber="431" rowspan-1="1">
<th id="7698143EC029FCBB9885FBCABE52FBFA" box="[159,638,1042,1064]" gridcol="0" gridrow="18" pageId="6" pageNumber="431">delsc</th>
<td id="7698143EC029FCBB9BB4FBCABFE4FBFA" box="[942,968,1042,1064]" gridcol="2" gridrow="18" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFBCAB881FBFA" box="[1028,1197,1042,1064]" gridcol="3" gridrow="18" pageId="6" pageNumber="431">22.2 (20.523.7)</td>
<td id="7698143EC029FCBB9D51FBCAB95BFBFA" box="[1355,1399,1042,1064]" gridcol="4" gridrow="18" pageId="6" pageNumber="431">22.7</td>
</tr>
<tr id="35497D42C029FCBB9885FBE4B95BFB80" box="[159,1399,1084,1106]" gridrow="19" pageId="6" pageNumber="431" rowspan-1="1">
<th id="7698143EC029FCBB9885FBE4BE52FB80" box="[159,638,1084,1106]" gridcol="0" gridrow="19" pageId="6" pageNumber="431">Supraoccipital process length</th>
<td id="7698143EC029FCBB9BB4FBE4BFE4FB80" box="[942,968,1084,1106]" gridcol="2" gridrow="19" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFBE4B881FB80" box="[1028,1197,1084,1106]" gridcol="3" gridrow="19" pageId="6" pageNumber="431">9.4 (8.51.2)</td>
<td id="7698143EC029FCBB9D51FBE4B95BFB80" box="[1355,1399,1084,1106]" gridcol="4" gridrow="19" pageId="6" pageNumber="431">8.1</td>
</tr>
<tr id="35497D42C029FCBB9885FBBDB95BFBA9" box="[159,1399,1125,1147]" gridrow="20" pageId="6" pageNumber="431" rowspan-1="1">
<th id="7698143EC029FCBB9885FBBDBE52FBA9" box="[159,638,1125,1147]" gridcol="0" gridrow="20" pageId="6" pageNumber="431">Supraoccipital process width</th>
<td id="7698143EC029FCBB9BB4FBBDBFE4FBA9" box="[942,968,1125,1147]" gridcol="2" gridrow="20" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFBBDB881FBA9" box="[1028,1197,1125,1147]" gridcol="3" gridrow="20" pageId="6" pageNumber="431">2.3 (1.92.8)</td>
<td id="7698143EC029FCBB9D51FBBDB95BFBA9" box="[1355,1399,1125,1147]" gridcol="4" gridrow="20" pageId="6" pageNumber="431">2.2</td>
</tr>
<tr id="35497D42C029FCBB9885FB56B95BFB76" box="[159,1399,1166,1188]" gridrow="21" pageId="6" pageNumber="431" rowspan-1="1">
<th id="7698143EC029FCBB9885FB56BE52FB76" box="[159,638,1166,1188]" gridcol="0" gridrow="21" pageId="6" pageNumber="431">Nuchalplate length</th>
<td id="7698143EC029FCBB9BB4FB56BFE4FB76" box="[942,968,1166,1188]" gridcol="2" gridrow="21" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFB56B881FB76" box="[1028,1197,1166,1188]" gridcol="3" gridrow="21" pageId="6" pageNumber="431">6.2 (5.96.6)</td>
<td id="7698143EC029FCBB9D51FB56B95BFB76" box="[1355,1399,1166,1188]" gridcol="4" gridrow="21" pageId="6" pageNumber="431">5.8</td>
</tr>
<tr id="35497D42C029FCBB9885FB60B95BFB1C" box="[159,1399,1208,1230]" gridrow="22" pageId="6" pageNumber="431" rowspan-1="1">
<th id="7698143EC029FCBB9885FB60BE52FB1C" box="[159,638,1208,1230]" gridcol="0" gridrow="22" pageId="6" pageNumber="431">Nuchalplate width</th>
<td id="7698143EC029FCBB9BB4FB60BFE4FB1C" box="[942,968,1208,1230]" gridcol="2" gridrow="22" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EFB60B881FB1C" box="[1028,1197,1208,1230]" gridcol="3" gridrow="22" pageId="6" pageNumber="431">6.2 (5.96.4)</td>
<td id="7698143EC029FCBB9D51FB60B95BFB1C" box="[1355,1399,1208,1230]" gridcol="4" gridrow="22" pageId="6" pageNumber="431">5.5</td>
</tr>
<tr id="35497D42C029FCBB9885FB39B95BFB25" box="[159,1399,1249,1271]" gridrow="23" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885FB39BE52FB25" box="[159,638,1249,1271]" gridcol="0" gridrow="23" pageId="6" pageNumber="431">Body depth</th>
<td id="7698143EC029FCBB9BB4FB39BFE4FB25" box="[942,968,1249,1271]" gridcol="2" gridrow="23" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFB39B881FB25" box="[1028,1197,1249,1271]" gridcol="3" gridrow="23" pageId="6" pageNumber="431">17.6 (16.620.0)</td>
</tr>
<tr id="35497D42C029FCBB9885FAD2B95BFAF2" box="[159,1399,1290,1312]" gridrow="24" pageId="6" pageNumber="431">
<th id="7698143EC029FCBB9885FAD2BE52FAF2" box="[159,638,1290,1312]" gridcol="0" gridrow="24" pageId="6" pageNumber="431">Body width</th>
<td id="7698143EC029FCBB9AFFFAD2BF66FAF2" box="[741,842,1290,1312]" gridcol="1" gridrow="24" pageId="6" pageNumber="431">19.019.2</td>
<td id="7698143EC029FCBB9BB4FAD2BFE4FAF2" box="[942,968,1290,1312]" gridcol="2" gridrow="24" pageId="6" pageNumber="431">10</td>
<td id="7698143EC029FCBB9C1EFAD2B881FAF2" box="[1028,1197,1290,1312]" gridcol="3" gridrow="24" pageId="6" pageNumber="431">18.3 (16.820.2)</td>
<td id="7698143EC029FCBB9D51FAD2B95BFAF2" box="[1355,1399,1290,1312]" gridcol="4" gridrow="24" pageId="6" pageNumber="431">19.0</td>
</tr>
<tr id="35497D42C029FCBB9885FAECB95BFA98" box="[159,1399,1332,1354]" gridrow="25" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885FAECBE52FA98" box="[159,638,1332,1354]" gridcol="0" gridrow="25" pageId="6" pageNumber="431">Distance from snout to pectoral fin</th>
<td id="7698143EC029FCBB9BB4FAECBFE4FA98" box="[942,968,1332,1354]" gridcol="2" gridrow="25" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EFAECB881FA98" box="[1028,1197,1332,1354]" gridcol="3" gridrow="25" pageId="6" pageNumber="431">23.3 (21.526.9)</td>
</tr>
<tr id="35497D42C029FCBB9885FA85B95BFAA1" box="[159,1399,1373,1395]" gridrow="26" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885FA85BE52FAA1" box="[159,638,1373,1395]" gridcol="0" gridrow="26" pageId="6" pageNumber="431">Distance from snout to dorsal fin</th>
<td id="7698143EC029FCBB9BB4FA85BFE4FAA1" box="[942,968,1373,1395]" gridcol="2" gridrow="26" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFA85B881FAA1" box="[1028,1197,1373,1395]" gridcol="3" gridrow="26" pageId="6" pageNumber="431">33.8 (24.737.8)</td>
</tr>
<tr id="35497D42C029FCBB9885FA5EB95BFA4E" box="[159,1399,1414,1436]" gridrow="27" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885FA5EBE52FA4E" box="[159,638,1414,1436]" gridcol="0" gridrow="27" pageId="6" pageNumber="431">Distance from snout to pelvic fin</th>
<td id="7698143EC029FCBB9BB4FA5EBFE4FA4E" box="[942,968,1414,1436]" gridcol="2" gridrow="27" pageId="6" pageNumber="431">9</td>
<td id="7698143EC029FCBB9C1EFA5EB881FA4E" box="[1028,1197,1414,1436]" gridcol="3" gridrow="27" pageId="6" pageNumber="431">50.0 (34.353.9)</td>
</tr>
<tr id="35497D42C029FCBB9885FA68B95BFA14" box="[159,1399,1456,1478]" gridrow="28" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885FA68BE52FA14" box="[159,638,1456,1478]" gridcol="0" gridrow="28" pageId="6" pageNumber="431">Distance from snout to anal fin</th>
<td id="7698143EC029FCBB9BB4FA68BFE4FA14" box="[942,968,1456,1478]" gridcol="2" gridrow="28" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EFA68B881FA14" box="[1028,1197,1456,1478]" gridcol="3" gridrow="28" pageId="6" pageNumber="431">69.5 (66.671.9)</td>
</tr>
<tr id="35497D42C029FCBB9885FA01B95BFA3D" box="[159,1399,1497,1519]" gridrow="29" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885FA01BE52FA3D" box="[159,638,1497,1519]" gridcol="0" gridrow="29" pageId="6" pageNumber="431">Distance from snout to adipose fin</th>
<td id="7698143EC029FCBB9BB4FA01BFE4FA3D" box="[942,968,1497,1519]" gridcol="2" gridrow="29" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EFA01B881FA3D" box="[1028,1197,1497,1519]" gridcol="3" gridrow="29" pageId="6" pageNumber="431">74.8 (72.877.0)</td>
</tr>
<tr id="35497D42C029FCBB9885F9DAB95BF9CA" box="[159,1399,1538,1560]" gridrow="30" pageId="6" pageNumber="431">
<th id="7698143EC029FCBB9885F9DABE52F9CA" box="[159,638,1538,1560]" gridcol="0" gridrow="30" pageId="6" pageNumber="431">Caudal-peduncle height</th>
<td id="7698143EC029FCBB9AFFF9DABF66F9CA" box="[741,842,1538,1560]" gridcol="1" gridrow="30" pageId="6" pageNumber="431">6.87.2</td>
<td id="7698143EC029FCBB9BB4F9DABFE4F9CA" box="[942,968,1538,1560]" gridcol="2" gridrow="30" pageId="6" pageNumber="431">10</td>
<td id="7698143EC029FCBB9C1EF9DAB881F9CA" box="[1028,1197,1538,1560]" gridcol="3" gridrow="30" pageId="6" pageNumber="431">6.5 (5.96.8)</td>
<td id="7698143EC029FCBB9D51F9DAB95BF9CA" box="[1355,1399,1538,1560]" gridcol="4" gridrow="30" pageId="6" pageNumber="431">6.7</td>
</tr>
<tr id="35497D42C029FCBB9885F9F4B95BF990" box="[159,1399,1580,1602]" gridrow="31" pageId="6" pageNumber="431" rowspan-1="1">
<th id="7698143EC029FCBB9885F9F4BE52F990" box="[159,638,1580,1602]" gridcol="0" gridrow="31" pageId="6" pageNumber="431">Pectoral-fin spine length</th>
<td id="7698143EC029FCBB9BB4F9F4BFE4F990" box="[942,968,1580,1602]" gridcol="2" gridrow="31" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EF9F4B881F990" box="[1028,1197,1580,1602]" gridcol="3" gridrow="31" pageId="6" pageNumber="431">19.5 (18.021.8)</td>
<td id="7698143EC029FCBB9D51F9F4B95BF990" box="[1355,1399,1580,1602]" gridcol="4" gridrow="31" pageId="6" pageNumber="431">18.0</td>
</tr>
<tr id="35497D42C029FCBB9885F98DB95BF9B9" box="[159,1399,1621,1643]" gridrow="32" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885F98DBE52F9B9" box="[159,638,1621,1643]" gridcol="0" gridrow="32" pageId="6" pageNumber="431">Dorsal-fin spine length</th>
<td id="7698143EC029FCBB9BB4F98DBFE4F9B9" box="[942,968,1621,1643]" gridcol="2" gridrow="32" pageId="6" pageNumber="431">5</td>
<td id="7698143EC029FCBB9C1EF98DB881F9B9" box="[1028,1197,1621,1643]" gridcol="3" gridrow="32" pageId="6" pageNumber="431">19.2 (17.721.8)</td>
</tr>
<tr id="35497D42C029FCBB9885F9A6B95BF946" box="[159,1399,1662,1684]" gridrow="33" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885F9A6BE52F946" box="[159,638,1662,1684]" gridcol="0" gridrow="33" pageId="6" pageNumber="431">Pelvic-fin base length</th>
<td id="7698143EC029FCBB9BB4F9A6BFE4F946" box="[942,968,1662,1684]" gridcol="2" gridrow="33" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EF9A6B881F946" box="[1028,1197,1662,1684]" gridcol="3" gridrow="33" pageId="6" pageNumber="431">3.7 (3.53.9)</td>
</tr>
<tr id="35497D42C029FCBB9885F970B95BF96C" box="[159,1399,1704,1726]" gridrow="34" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885F970BE52F96C" box="[159,638,1704,1726]" gridcol="0" gridrow="34" pageId="6" pageNumber="431">Pelvic-fin height</th>
<td id="7698143EC029FCBB9BB4F970BFE4F96C" box="[942,968,1704,1726]" gridcol="2" gridrow="34" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EF970B881F96C" box="[1028,1197,1704,1726]" gridcol="3" gridrow="34" pageId="6" pageNumber="431">15.2 (12.817.5)</td>
</tr>
<tr id="35497D42C029FCBB9885F909B95BF935" box="[159,1399,1745,1767]" gridrow="35" pageId="6" pageNumber="431" rowspan-4="1">
<th id="7698143EC029FCBB9885F909BE52F935" box="[159,638,1745,1767]" gridcol="0" gridrow="35" pageId="6" pageNumber="431">Adipose-fin base length</th>
<td id="7698143EC029FCBB9AFFF909BF66F935" box="[741,842,1745,1767]" gridcol="1" gridrow="35" pageId="6" pageNumber="431">7.99.0</td>
<td id="7698143EC029FCBB9BB4F909BFE4F935" box="[942,968,1745,1767]" gridcol="2" gridrow="35" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EF909B881F935" box="[1028,1197,1745,1767]" gridcol="3" gridrow="35" pageId="6" pageNumber="431">6.9 (6.08.0)</td>
</tr>
<tr id="35497D42C029FCBB9885F922B95BF8C2" box="[159,1399,1786,1808]" gridrow="36" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885F922BE52F8C2" box="[159,638,1786,1808]" gridcol="0" gridrow="36" pageId="6" pageNumber="431">Adipose-fin height</th>
<td id="7698143EC029FCBB9BB4F922BFE4F8C2" box="[942,968,1786,1808]" gridcol="2" gridrow="36" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EF922B881F8C2" box="[1028,1197,1786,1808]" gridcol="3" gridrow="36" pageId="6" pageNumber="431">3.8 (3.54.3)</td>
</tr>
<tr id="35497D42C029FCBB9885F8FCB95BF8E8" box="[159,1399,1828,1850]" gridrow="37" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885F8FCBE52F8E8" box="[159,638,1828,1850]" gridcol="0" gridrow="37" pageId="6" pageNumber="431">Anal-fin base length</th>
<td id="7698143EC029FCBB9BB4F8FCBFE4F8E8" box="[942,968,1828,1850]" gridcol="2" gridrow="37" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EF8FCB881F8E8" box="[1028,1197,1828,1850]" gridcol="3" gridrow="37" pageId="6" pageNumber="431">14.8 (13.816.0)</td>
</tr>
<tr id="35497D42C029FCBB9885F895B95BF8B1" box="[159,1399,1869,1891]" gridrow="38" pageId="6" pageNumber="431" rowspan-1="1" rowspan-4="1">
<th id="7698143EC029FCBB9885F895BE52F8B1" box="[159,638,1869,1891]" gridcol="0" gridrow="38" pageId="6" pageNumber="431">Anal-fin height</th>
<td id="7698143EC029FCBB9BB4F895BFE4F8B1" box="[942,968,1869,1891]" gridcol="2" gridrow="38" pageId="6" pageNumber="431">8</td>
<td id="7698143EC029FCBB9C1EF895B881F8B1" box="[1028,1197,1869,1891]" gridcol="3" gridrow="38" pageId="6" pageNumber="431">15.8 (12.418.0)</td>
</tr>
<tr id="35497D42C029FCBB9885F8AEB95BF85E" box="[159,1399,1910,1932]" gridrow="39" pageId="6" pageNumber="431" rowspan-4="1">
<th id="7698143EC029FCBB9885F8AEBE52F85E" box="[159,638,1910,1932]" gridcol="0" gridrow="39" pageId="6" pageNumber="431">Caudal-fin dorsal lobe length</th>
<td id="7698143EC029FCBB9AFFF8AEBF66F85E" box="[741,842,1910,1932]" gridcol="1" gridrow="39" pageId="6" pageNumber="431">30.0</td>
<td id="7698143EC029FCBB9BB4F8AEBFE4F85E" box="[942,968,1910,1932]" gridcol="2" gridrow="39" pageId="6" pageNumber="431">3</td>
<td id="7698143EC029FCBB9C1EF8AEB881F85E" box="[1028,1197,1910,1932]" gridcol="3" gridrow="39" pageId="6" pageNumber="431">36.0 (32.738.1)</td>
</tr>
<tr id="35497D42C029FCBB9885F878B95BF864" box="[159,1399,1952,1974]" gridrow="40" pageId="6" pageNumber="431" rowspan-4="1">
<th id="7698143EC029FCBB9885F878BE52F864" box="[159,638,1952,1974]" gridcol="0" gridrow="40" pageId="6" pageNumber="431">Caudal-fin ventral lobe length</th>
<td id="7698143EC029FCBB9AFFF878BF66F864" box="[741,842,1952,1974]" gridcol="1" gridrow="40" pageId="6" pageNumber="431">24.925.2</td>
<td id="7698143EC029FCBB9BB4F878BFE4F864" box="[942,968,1952,1974]" gridcol="2" gridrow="40" pageId="6" pageNumber="431">6</td>
<td id="7698143EC029FCBB9C1EF878B881F864" box="[1028,1197,1952,1974]" gridcol="3" gridrow="40" pageId="6" pageNumber="431">30.6 (28.533.1)</td>
</tr>
</table>
</paragraph>
<caption id="DF062F88C0280343988DF91FBE4BF88A" httpUri="https://zenodo.org/record/2647206/files/figure.png" pageId="7" pageNumber="432" startId="7.[151,250,1735,1757]" targetBox="[151,1435,193,1714]" targetPageId="7">
<paragraph id="8BC67F00C0280343988DF91FBE4BF88A" blockId="7.[151,1436,1735,1880]" pageId="7" pageNumber="432">
<emphasis id="B90DA312C0280343988DF91FBD21F90E" bold="true" box="[151,269,1735,1757]" pageId="7" pageNumber="432">FIGURE 2</emphasis>
. Body and head in lateral and dorsal view of all
<taxonomicName id="4C790483C02803439AE3F91FBFD4F90F" box="[761,1016,1735,1757]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="432" phylum="Chordata" rank="species" species="gradoculis">
<emphasis id="B90DA312C02803439AE3F91FBFD4F90F" box="[761,1016,1735,1757]" italics="true" pageId="7" pageNumber="432">Paragenidens gradoculis</emphasis>
</taxonomicName>
specimens in zoological collections. (A). NMW 48272, rio Doce, Espírito Santo state. (B). MCZ 7681 (Museum of Comparative Zoology, Harvard University), rio Doce, Espírito Santo state, scale 50 mm. (C). MZUSP 2295, lagoa Feia, Rio de Janeiro, 251 mm SL. (D). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo. (E). MZUSP 1010414, 170 mm SL, lagoa Juparanã, Espírito Santo state. (F). LBP 24068, 240 mm SL, lagoa Nova, Espírito Santo state.
</paragraph>
</caption>
<paragraph id="8BC67F00C028034C98DDF85EB962FF2A" blockId="7.[151,1437,1924,2022]" lastBlockId="8.[151,1437,151,789]" lastPageId="8" lastPageNumber="433" pageId="7" pageNumber="432">Bony bridge formed by lateral ethmoid and frontal relatively long and slender, readily discernible under skin. Cephalic shield exposed, moderately rough and granulated, long and relatively straight on lateral ethmoid, frontal, and supra cleithrum areas. Anterior portion of dorsomedial groove of neurocranium fleshy, inconspicuous to level of posterior nares; posterior portion of dorsomedial groove of neurocranium osseous, deep and conspicuous, with straight margins tapering posteriorly. Parieto-supraoccipital process slightly tapered antero-posteriorly, long and narrow on posterior portion, with profile straight. Nuchal plate crescent-shaped, moderately short and wide.</paragraph>
<paragraph id="8BC67F00C027034C98DDFEDCBD85FD7A" blockId="8.[151,1437,151,789]" pageId="8" pageNumber="433">Body width slightly greater than body depth at pectoral-girdle area, progressively compressed from pectoral fin to caudal peduncle. Lateral line sloping ventrally on anterior one-third, extending posteriorly to caudal peduncle, bending abruptly onto dorsal lobe of caudal fin. Dorsal-fin spine long and thick; anterior margin with granules on basal two-thirds, distal one-third with short serrations; posterior margin with serrations along almost its entire length. Seven dorsal-fin soft rays. Pectoral-fin spine long, as long as dorsal-fin spine; anterior margin with granules on basal two-thirds and distal one-third with short serrations; posterior margin straight on basal onefourth, distal three-fourths with conspicuous serrations. Nine to 10 pectoral-fin soft rays. Posterior cleithral process exposed, smooth and triangular shaped, long and pointed posteriorly. Pelvic fin deep and short at base, with six rays. Adipose-fin base short, less than one-half the length of anal-fin base, anterior origin at level of anterior half of anal fin. Anal fin very deep and relatively short at base, with 15-17 rays and ventral profile markedly concave. Caudal peduncle low. Caudal fin forked, dorsal and ventral lobes relatively long, posteriorly pointed; dorsal lobe longer than ventral lobe.</paragraph>
<paragraph id="8BC67F00C027034C98DDFD6CBDABFCC7" blockId="8.[151,1437,151,789]" pageId="8" pageNumber="433">
<emphasis id="B90DA312C027034C98DDFD6CBDC1FD1F" bold="true" box="[199,493,692,717]" pageId="8" pageNumber="433">Morphological variation</emphasis>
. The well-developed gill rakers on mesial surfaces of first and second gill arches, present in the specimens from
<collectingRegion id="49BDB1E2C027034C99E8FD0FBEBEFD22" box="[498,658,727,752]" country="Brazil" name="Espirito Santo" pageId="8" pageNumber="433">Espírito Santo</collectingRegion>
[
<figureCitation id="13426385C027034C9ABAFD0FBF2FFD22" box="[672,771,727,752]" captionStart="FIGURE 3" captionStartId="8.[151,250,1718,1740]" captionTargetBox="[151,1435,841,1697]" captionTargetId="figure@8.[151,1435,841,1697]" captionTargetPageId="8" captionText="FIGURE 3. Accessory tooth plates and mesial face of the first and second branchial arches. (A). holotype NMW 48272, rio Doce, Espírito Santo. (B). MZUSP 1010414, 156 mm SL, lagoa Juparanã, Espírito Santo state. (C). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo state. (D). MNRJ 2295, 251 mm SL, lagoa Feia, São João da Barra, Rio de Janeiro state." httpUri="https://zenodo.org/record/2647208/files/figure.png" pageId="8" pageNumber="433">Figure 3</figureCitation>
(c)], were not observed in the single specimen from
<collectingRegion id="49BDB1E2C027034C9D4BFD0FBCC4FCC7" country="Brazil" name="Rio de Janeiro" pageId="8" pageNumber="433">Rio de Janeiro</collectingRegion>
[
<figureCitation id="13426385C027034C98EFFD24BD7AFCC7" box="[245,342,764,789]" captionStart="FIGURE 3" captionStartId="8.[151,250,1718,1740]" captionTargetBox="[151,1435,841,1697]" captionTargetId="figure@8.[151,1435,841,1697]" captionTargetPageId="8" captionText="FIGURE 3. Accessory tooth plates and mesial face of the first and second branchial arches. (A). holotype NMW 48272, rio Doce, Espírito Santo. (B). MZUSP 1010414, 156 mm SL, lagoa Juparanã, Espírito Santo state. (C). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo state. (D). MNRJ 2295, 251 mm SL, lagoa Feia, São João da Barra, Rio de Janeiro state." httpUri="https://zenodo.org/record/2647208/files/figure.png" pageId="8" pageNumber="433">Figure 3</figureCitation>
(d)].
</paragraph>
</subSubSection>
<caption id="DF062F88C027034C988DF96EB917F8D8" httpUri="https://zenodo.org/record/2647208/files/figure.png" pageId="8" pageNumber="433" startId="8.[151,250,1718,1740]" targetBox="[151,1435,841,1697]" targetPageId="8">
<paragraph id="8BC67F00C027034C988DF96EB917F8D8" blockId="8.[151,1436,1718,1802]" pageId="8" pageNumber="433">
<emphasis id="B90DA312C027034C988DF96EBD22F919" bold="true" box="[151,270,1718,1740]" pageId="8" pageNumber="433">FIGURE 3</emphasis>
. Accessory tooth plates and mesial face of the first and second branchial arches. (A). holotype NMW 48272, rio Doce, Espírito Santo. (B). MZUSP 1010414, 156 mm SL, lagoa Juparanã, Espírito Santo state. (C). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo state. (D). MNRJ 2295, 251 mm SL, lagoa Feia, São João da Barra, Rio de Janeiro state.
</paragraph>
</caption>
<subSubSection id="C3632C8BC027034D98DDF8E9BEEAFECF" lastPageId="9" lastPageNumber="434" pageId="8" pageNumber="433" type="distribution">
<paragraph id="8BC67F00C027034D98DDF8E9BEEAFECF" blockId="8.[151,1436,1841,2010]" lastBlockId="9.[151,1437,151,1400]" lastPageId="9" lastPageNumber="434" pageId="8" pageNumber="433">
<emphasis id="B90DA312C027034C98DDF8E9BDDFF898" bold="true" box="[199,499,1841,1866]" pageId="8" pageNumber="433">Distribution and habitat</emphasis>
.
<taxonomicName id="4C790483C027034C9A1CF8E9BF1FF89B" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[518,819,1841,1865]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="433" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C027034C9A1CF8E9BF1FF89B" box="[518,819,1841,1865]" italics="true" pageId="8" pageNumber="433">Paragenidens grandoculis</emphasis>
</taxonomicName>
is known from lagoa Juparanã [MZUSP 79459 and MZUSP 1010410114,
<figureCitation id="13426385C027034C99B0F88DBE34F8BC" box="[426,536,1877,1902]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1757]" captionTargetBox="[151,1435,193,1714]" captionTargetId="figure@7.[151,1435,193,1714]" captionTargetPageId="7" captionText="FIGURE 2. Body and head in lateral and dorsal view of all Paragenidens gradoculis specimens in zoological collections. (A). NMW 48272, rio Doce, Espírito Santo state. (B). MCZ 7681 (Museum of Comparative Zoology, Harvard University), rio Doce, Espírito Santo state, scale 50 mm. (C). MZUSP 2295, lagoa Feia, Rio de Janeiro, 251 mm SL. (D). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo. (E). MZUSP 1010414, 170 mm SL, lagoa Juparanã, Espírito Santo state. (F). LBP 24068, 240 mm SL, lagoa Nova, Espírito Santo state." httpUri="https://zenodo.org/record/2647206/files/figure.png" pageId="8" pageNumber="433">Figures 2</figureCitation>
(d), (e)] and lagoa Nova [LBP 24068,
<figureCitation id="13426385C027034C9BC5F88DB86EF8BC" box="[991,1090,1877,1902]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1757]" captionTargetBox="[151,1435,193,1714]" captionTargetId="figure@7.[151,1435,193,1714]" captionTargetPageId="7" captionText="FIGURE 2. Body and head in lateral and dorsal view of all Paragenidens gradoculis specimens in zoological collections. (A). NMW 48272, rio Doce, Espírito Santo state. (B). MCZ 7681 (Museum of Comparative Zoology, Harvard University), rio Doce, Espírito Santo state, scale 50 mm. (C). MZUSP 2295, lagoa Feia, Rio de Janeiro, 251 mm SL. (D). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo. (E). MZUSP 1010414, 170 mm SL, lagoa Juparanã, Espírito Santo state. (F). LBP 24068, 240 mm SL, lagoa Nova, Espírito Santo state." httpUri="https://zenodo.org/record/2647206/files/figure.png" pageId="8" pageNumber="433">Figure 2</figureCitation>
(f)], municipality of Linhares,
<collectingRegion id="49BDB1E2C027034C988DF8A1BD14F843" box="[151,312,1913,1938]" country="Brazil" name="Espirito Santo" pageId="8" pageNumber="433">Espírito Santo</collectingRegion>
state, both natural lakes that connect with lower river Doce basin only in floods periods, and from lagoa Feia [MZUSP 2295,
<figureCitation id="13426385C027034C99DDF845BE0BF864" box="[455,551,1949,1974]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1757]" captionTargetBox="[151,1435,193,1714]" captionTargetId="figure@7.[151,1435,193,1714]" captionTargetPageId="7" captionText="FIGURE 2. Body and head in lateral and dorsal view of all Paragenidens gradoculis specimens in zoological collections. (A). NMW 48272, rio Doce, Espírito Santo state. (B). MCZ 7681 (Museum of Comparative Zoology, Harvard University), rio Doce, Espírito Santo state, scale 50 mm. (C). MZUSP 2295, lagoa Feia, Rio de Janeiro, 251 mm SL. (D). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo. (E). MZUSP 1010414, 170 mm SL, lagoa Juparanã, Espírito Santo state. (F). LBP 24068, 240 mm SL, lagoa Nova, Espírito Santo state." httpUri="https://zenodo.org/record/2647206/files/figure.png" pageId="8" pageNumber="433">Figure 2</figureCitation>
(c)] at Campos dos Goytacazes municipality,
<collectingRegion id="49BDB1E2C027034C9C3FF845B8EBF864" box="[1061,1223,1949,1974]" country="Brazil" name="Rio de Janeiro" pageId="8" pageNumber="433">Rio de Janeiro</collectingRegion>
state, southeastern
<collectingCountry id="F36E3F90C027034C988DF819BCF7F808" box="[151,219,1985,2010]" name="Brazil" pageId="8" pageNumber="433">Brazil</collectingCountry>
(
<figureCitation id="13426385C027034C98F0F819BD60F808" box="[234,332,1985,2010]" captionStart="FIGURE 5" captionStartId="13.[151,250,1175,1197]" captionTargetBox="[161,1425,205,1138]" captionTargetId="figure@13.[151,1435,193,1154]" captionTargetPageId="13" captionText="FIGURE 5. Sampling localities of the material examined in the present study. Lagoa Juparanã, Espírito Santo State (green), lagoa Nova, Espírito Santo State (red), lagoa Feia, Rio de Janeiro State (yellow), (A) rio Doce between Aimorés (Minas Gerais State) and Linhares (Espírito Santo State) (light blue), and lagoons not sampled in the Espírito Santo Sate (pink)." httpUri="https://zenodo.org/record/2647212/files/figure.png" pageId="8" pageNumber="433">Figure 5</figureCitation>
). Recent collect efforts at the lower rio Doce at the municipality of Linhares (including the district of Regência) have shown that the species is not present in the main channel of rio Doce, where only specimens of
<taxonomicName id="4C790483C026034D988DFF64BD41FF06" authority="genidens" authorityName="genidens (Cuvier" authorityYear="1829" box="[151,365,188,212]" class="Actinopterygii" family="Ariidae" genus="Genidens" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="species" species="genidens">
<emphasis id="B90DA312C026034D988DFF64BD41FF06" box="[151,365,188,212]" italics="true" pageId="9" pageNumber="434">Genidens genidens</emphasis>
</taxonomicName>
were collected (CZNC 2467; specimens not deposited). Based on the know distribution of the species, supported by museum specimens, we infer that the distribution of
<taxonomicName id="4C790483C026034D9BFFFF38B922FF2A" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[997,1294,224,248]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C026034D9BFFFF38B922FF2A" box="[997,1294,224,248]" italics="true" pageId="9" pageNumber="434">Paragenidens grandoculis</emphasis>
</taxonomicName>
is restrict to freshwater coastal lakes from southeastern
<collectingCountry id="F36E3F90C026034D9A60FEDCBEEEFECF" box="[634,706,260,285]" name="Brazil" pageId="9" pageNumber="434">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3632C8BC026034D98DDFEFFBF75FB72" pageId="9" pageNumber="434" type="discussion">
<paragraph id="8BC67F00C026034D98DDFEFFBEB2FDB2" blockId="9.[151,1437,151,1400]" pageId="9" pageNumber="434">
<emphasis id="B90DA312C026034D98DDFEFFBD18FE92" bold="true" box="[199,308,295,320]" pageId="9" pageNumber="434">Remarks</emphasis>
.
<taxonomicName id="4C790483C026034D9951FEF0BE31FE92" authorityName="Steindachner" authorityYear="1877" box="[331,541,296,320]" class="Actinopterygii" family="Ariidae" genus="Arius" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C026034D9951FEF0BE31FE92" box="[331,541,296,320]" italics="true" pageId="9" pageNumber="434">Arius grandoculis</emphasis>
</taxonomicName>
was described by Steindachner (1877) based on specimens deposited at Naturhistorisches Museum, Wien [
<typeStatus id="54C2C1A2C026034D9A03FE94BEADFEB7" box="[537,641,332,357]" pageId="9" pageNumber="434" type="holotype">holotype</typeStatus>
, NMW 48272,
<figureCitation id="13426385C026034D9B37FE94BFA0FEB6" box="[813,908,332,357]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1757]" captionTargetBox="[151,1435,193,1714]" captionTargetId="figure@7.[151,1435,193,1714]" captionTargetPageId="7" captionText="FIGURE 2. Body and head in lateral and dorsal view of all Paragenidens gradoculis specimens in zoological collections. (A). NMW 48272, rio Doce, Espírito Santo state. (B). MCZ 7681 (Museum of Comparative Zoology, Harvard University), rio Doce, Espírito Santo state, scale 50 mm. (C). MZUSP 2295, lagoa Feia, Rio de Janeiro, 251 mm SL. (D). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo. (E). MZUSP 1010414, 170 mm SL, lagoa Juparanã, Espírito Santo state. (F). LBP 24068, 240 mm SL, lagoa Nova, Espírito Santo state." httpUri="https://zenodo.org/record/2647206/files/figure.png" pageId="9" pageNumber="434">Figure 2</figureCitation>
(a)] and two other specimens from the Museum of Comparative Zoology [MCZ 768081,
<figureCitation id="13426385C026034D9A9BFEB7BEC8FE5A" box="[641,740,367,392]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1757]" captionTargetBox="[151,1435,193,1714]" captionTargetId="figure@7.[151,1435,193,1714]" captionTargetPageId="7" captionText="FIGURE 2. Body and head in lateral and dorsal view of all Paragenidens gradoculis specimens in zoological collections. (A). NMW 48272, rio Doce, Espírito Santo state. (B). MCZ 7681 (Museum of Comparative Zoology, Harvard University), rio Doce, Espírito Santo state, scale 50 mm. (C). MZUSP 2295, lagoa Feia, Rio de Janeiro, 251 mm SL. (D). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo. (E). MZUSP 1010414, 170 mm SL, lagoa Juparanã, Espírito Santo state. (F). LBP 24068, 240 mm SL, lagoa Nova, Espírito Santo state." httpUri="https://zenodo.org/record/2647206/files/figure.png" pageId="9" pageNumber="434">Figure 2</figureCitation>
(b)], all collected by Charles F. Hartt and Edward Copeland during their travel as members of the Thayer Expedition across the lower rio Doce, between Aimorés (
<collectingRegion id="49BDB1E2C026034D9D4EFE4CBCCCFE02" country="Brazil" name="Minas Gerais" pageId="9" pageNumber="434">Minas Gerais</collectingRegion>
State) and Linhares (
<collectingRegion id="49BDB1E2C026034D99CEFE6FBE5AFE02" box="[468,630,439,464]" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="434">Espírito Santo</collectingRegion>
State). Additional specimens were collected by Ernst Garbe (naturalist traveler from the Museu Paulista, currently MZUSP) probably in 1906, in rio Doce at Linhares,
<collectingRegion id="49BDB1E2C026034D9CE3FE04B9B0FE26" box="[1273,1436,476,501]" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="434">Espírito Santo</collectingRegion>
State (MZUSP 2294) and at lagoa Feia at Campos dos Goytacazes,
<collectingRegion id="49BDB1E2C026034D9BB2FE27B87CFDCA" box="[936,1104,511,536]" country="Brazil" name="Rio de Janeiro" pageId="9" pageNumber="434">Rio de Janeiro</collectingRegion>
State [MZUSP 2295,
<figureCitation id="13426385C026034D9D49FE27BC8BFDEE" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1757]" captionTargetBox="[151,1435,193,1714]" captionTargetId="figure@7.[151,1435,193,1714]" captionTargetPageId="7" captionText="FIGURE 2. Body and head in lateral and dorsal view of all Paragenidens gradoculis specimens in zoological collections. (A). NMW 48272, rio Doce, Espírito Santo state. (B). MCZ 7681 (Museum of Comparative Zoology, Harvard University), rio Doce, Espírito Santo state, scale 50 mm. (C). MZUSP 2295, lagoa Feia, Rio de Janeiro, 251 mm SL. (D). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo. (E). MZUSP 1010414, 170 mm SL, lagoa Juparanã, Espírito Santo state. (F). LBP 24068, 240 mm SL, lagoa Nova, Espírito Santo state." httpUri="https://zenodo.org/record/2647206/files/figure.png" pageId="9" pageNumber="434">Figure 2</figureCitation>
(c)]. Garbes specimen from rio Doce was not found in the MZUSP collection and is possibly lost, while the specimen from lagoa Feia was examined here.
</paragraph>
<paragraph id="8BC67F00C026034D98DDFDB4BDCBFC8F" blockId="9.[151,1437,151,1400]" pageId="9" pageNumber="434">
During the 1960s decade three other putative records of
<taxonomicName id="4C790483C026034D9B41FDB4B8AAFD56" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[859,1158,620,644]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C026034D9B41FDB4B8AAFD56" box="[859,1158,620,644]" italics="true" pageId="9" pageNumber="434">Paragenidens grandoculis</emphasis>
</taxonomicName>
were reported. In 1965, Heraldo A. Britski and Izaurio A. Dias collected
<specimenCount id="9D7FB489C026034D9ACDFD48BF5EFD7A" box="[727,882,655,680]" pageId="9" pageNumber="434" type="generic">20 specimens</specimenCount>
of
<taxonomicName id="4C790483C026034D9B84FD49B86FFD7A" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[926,1091,656,680]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C026034D9B84FD49B86FFD7A" box="[926,1091,656,680]" italics="true" pageId="9" pageNumber="434">P. grandoculis</emphasis>
</taxonomicName>
at lagoa Juparanã (Linhares), which is the largest and most complete record of the species in scientific collections [MZUSP 1010410114,
<figureCitation id="13426385C026034D9D49FD6CBC8BFD22" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1757]" captionTargetBox="[151,1435,193,1714]" captionTargetId="figure@7.[151,1435,193,1714]" captionTargetPageId="7" captionText="FIGURE 2. Body and head in lateral and dorsal view of all Paragenidens gradoculis specimens in zoological collections. (A). NMW 48272, rio Doce, Espírito Santo state. (B). MCZ 7681 (Museum of Comparative Zoology, Harvard University), rio Doce, Espírito Santo state, scale 50 mm. (C). MZUSP 2295, lagoa Feia, Rio de Janeiro, 251 mm SL. (D). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo. (E). MZUSP 1010414, 170 mm SL, lagoa Juparanã, Espírito Santo state. (F). LBP 24068, 240 mm SL, lagoa Nova, Espírito Santo state." httpUri="https://zenodo.org/record/2647206/files/figure.png" pageId="9" pageNumber="434">Figure 2</figureCitation>
(e)]. Most of the specimens from that expedition were examined here and had their identity confirmed, but some of them (MZUSP 22995) were not found at the museum collection and are possibly lost. Another lot, possibly collected with the specimens from lagoa Juparanã, was discovered at MZUSP and identified as
<taxonomicName id="4C790483C026034D9CEDFCF9B9B0FCEA" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[1271,1436,800,824]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C026034D9CEDFCF9B9B0FCEA" box="[1271,1436,800,824]" italics="true" pageId="9" pageNumber="434">P. grandoculis</emphasis>
</taxonomicName>
[MZUSP 79459,
<figureCitation id="13426385C026034D9942FC9CBD95FC8F" box="[344,441,836,861]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1757]" captionTargetBox="[151,1435,193,1714]" captionTargetId="figure@7.[151,1435,193,1714]" captionTargetPageId="7" captionText="FIGURE 2. Body and head in lateral and dorsal view of all Paragenidens gradoculis specimens in zoological collections. (A). NMW 48272, rio Doce, Espírito Santo state. (B). MCZ 7681 (Museum of Comparative Zoology, Harvard University), rio Doce, Espírito Santo state, scale 50 mm. (C). MZUSP 2295, lagoa Feia, Rio de Janeiro, 251 mm SL. (D). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo. (E). MZUSP 1010414, 170 mm SL, lagoa Juparanã, Espírito Santo state. (F). LBP 24068, 240 mm SL, lagoa Nova, Espírito Santo state." httpUri="https://zenodo.org/record/2647206/files/figure.png" pageId="9" pageNumber="434">Figure 2</figureCitation>
(d)].
</paragraph>
<paragraph id="8BC67F00C026034D98DDFCB0BF75FB72" blockId="9.[151,1437,151,1400]" pageId="9" pageNumber="434">
Attempts of finding
<taxonomicName id="4C790483C026034D99B0FCB0BEFEFC52" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[426,722,872,896]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C026034D99B0FCB0BEFEFC52" box="[426,722,872,896]" italics="true" pageId="9" pageNumber="434">Paragenidens grandoculis</emphasis>
</taxonomicName>
in nature may have been hindered by the difficulties inherent in the recognition of the members of the family
<taxonomicName id="4C790483C026034D9A85FC54BFE4FC77" authority="(Marceniuk 2005)" authorityName="Marceniuk" authorityYear="2005" baseAuthorityName="Marceniuk" baseAuthorityYear="2005" box="[671,968,908,933]" class="Actinopterygii" family="Ariidae" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="family">
Ariidae (
<bibRefCitation id="EFE802F1C026034D9B18FC54BFECFC77" author="Marceniuk, A. P." box="[770,960,908,933]" pageId="9" pageNumber="434" pagination="89 - 101" refId="ref11913" refString="Marceniuk, A. P. (2005) Chave para identificacao das especies de bagres marinhos (Siluriformes, Ariidae) da costa Brasileira. Boletim do Instituto do Pesca, 31, 89 - 101." type="journal article" year="2005">Marceniuk 2005</bibRefCitation>
)
</taxonomicName>
and, particularly, the lack of information about the species itself. Additionally, the historic low number of records of
<taxonomicName id="4C790483C026034D9C22FC69B8CCFC1A" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[1080,1248,944,968]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C026034D9C22FC69B8CCFC1A" box="[1080,1248,944,968]" italics="true" pageId="9" pageNumber="434">P. grandoculis</emphasis>
</taxonomicName>
may also be a consequence of the poor knowledge of the fish fauna of the areas where the species occurs (
<bibRefCitation id="EFE802F1C026034D9C8EFC0CB930FC3E" author="Vieira, F." box="[1172,1308,980,1005]" pageId="9" pageNumber="434" pagination="5 - 22" refId="ref13890" refString="Vieira, F. (2010) Distribuicao, impactos ambientais e conservacao da fauna de peixes da bacia do rio Doce. MG Biota, 2, 5 - 22." type="journal article" year="2010">Vieira 2010</bibRefCitation>
; Sarmento- Soares
<emphasis id="B90DA312C026034D98F3FC21BD0EFBC2" box="[233,290,1016,1040]" italics="true" pageId="9" pageNumber="434">et al.</emphasis>
2012, 2013, 2014). The external morphological resemblance between
<taxonomicName id="4C790483C026034D9C25FC21B8CEFBC2" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[1087,1250,1016,1040]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C026034D9C25FC21B8CEFBC2" box="[1087,1250,1016,1040]" italics="true" pageId="9" pageNumber="434">P. grandoculis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C790483C026034D9D03FC20B9B7FBC2" authorityName="genidens (Cuvier" authorityYear="1829" box="[1305,1435,1016,1040]" class="Actinopterygii" family="Ariidae" genus="Genidens" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="species" species="genidens">
<emphasis id="B90DA312C026034D9D03FC20B9B7FBC2" box="[1305,1435,1016,1040]" italics="true" pageId="9" pageNumber="434">G. genidens</emphasis>
</taxonomicName>
certainly contributed to the mistaken identifications detected with the museum specimens (
<figureCitation id="13426385C026034D9CB6FBC4B93DFBE7" box="[1196,1297,1052,1077]" captionStart="FIGURE 6" captionStartId="14.[151,250,1215,1237]" captionTargetBox="[151,1435,301,1194]" captionTargetId="figure@14.[151,1435,301,1194]" captionTargetPageId="14" captionText="FIGURE 6. Body and head in lateral and dorsal view of Genidens genidens specimens wrongly identified as Paragenidens grandoculis. (A) MNRJ 3581, Campos dos Goytacazes, Rio de Janeiro. (B). MNRJ 9732, ilha Santana, Macaé, Rio de Janeiro. (C) NUPEM 1793, 230 mm SL, Macaé, Rio de Janeiro state], and the specimen collected in the main channel of the rio Doce in Linhares, Espírito Santo State. (D) CZNC 2467, specimen collected in the main channel of the rio Doce in Linhares, Espírito Santo State." httpUri="https://zenodo.org/record/2647214/files/figure.png" pageId="9" pageNumber="434">Figure 6</figureCitation>
; e.g. MNRJ 3581, MNRJ 9732, NPM 318). Nevertheless, local artisanal fishermen recognize both species as rabo seco (
<emphasis id="B90DA312C026034D9D6CFB98BC90FBAE" italics="true" pageId="9" pageNumber="434">dry tail</emphasis>
), but distinguish them as two different varieties, one from the main channel of the river [
<taxonomicName id="4C790483C026034D9CA5FBBCB96FFBAE" authorityName="genidens (Cuvier" authorityYear="1829" box="[1215,1347,1124,1148]" class="Actinopterygii" family="Ariidae" genus="Genidens" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="species" species="genidens">
<emphasis id="B90DA312C026034D9CA5FBBCB96FFBAE" box="[1215,1347,1124,1148]" italics="true" pageId="9" pageNumber="434">G. genidens</emphasis>
</taxonomicName>
,
<figureCitation id="13426385C026034D9D49FBBCBC8AFB72" captionStart="FIGURE 6" captionStartId="14.[151,250,1215,1237]" captionTargetBox="[151,1435,301,1194]" captionTargetId="figure@14.[151,1435,301,1194]" captionTargetPageId="14" captionText="FIGURE 6. Body and head in lateral and dorsal view of Genidens genidens specimens wrongly identified as Paragenidens grandoculis. (A) MNRJ 3581, Campos dos Goytacazes, Rio de Janeiro. (B). MNRJ 9732, ilha Santana, Macaé, Rio de Janeiro. (C) NUPEM 1793, 230 mm SL, Macaé, Rio de Janeiro state], and the specimen collected in the main channel of the rio Doce in Linhares, Espírito Santo State. (D) CZNC 2467, specimen collected in the main channel of the rio Doce in Linhares, Espírito Santo State." httpUri="https://zenodo.org/record/2647214/files/figure.png" pageId="9" pageNumber="434">Figure 6</figureCitation>
(d)] and the other from the lakes [
<taxonomicName id="4C790483C026034D9A05FB51BEEDFB72" authorityName="Marceniuk &amp; Ingenito &amp; Lima &amp; Gasparini &amp; Oliveira" authorityYear="2019" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[543,705,1160,1184]" class="Actinopterygii" family="Ariidae" genus="Paragenidens" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="434" phylum="Chordata" rank="species" species="grandoculis">
<emphasis id="B90DA312C026034D9A05FB51BEEDFB72" box="[543,705,1160,1184]" italics="true" pageId="9" pageNumber="434">P. grandoculis</emphasis>
</taxonomicName>
,
<figureCitation id="13426385C026034D9AD5FB5FBF03FB72" box="[719,815,1159,1184]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1757]" captionTargetBox="[151,1435,193,1714]" captionTargetId="figure@7.[151,1435,193,1714]" captionTargetPageId="7" captionText="FIGURE 2. Body and head in lateral and dorsal view of all Paragenidens gradoculis specimens in zoological collections. (A). NMW 48272, rio Doce, Espírito Santo state. (B). MCZ 7681 (Museum of Comparative Zoology, Harvard University), rio Doce, Espírito Santo state, scale 50 mm. (C). MZUSP 2295, lagoa Feia, Rio de Janeiro, 251 mm SL. (D). MZUSP 79459, 247 mm SL, rio Doce, Espírito Santo. (E). MZUSP 1010414, 170 mm SL, lagoa Juparanã, Espírito Santo state. (F). LBP 24068, 240 mm SL, lagoa Nova, Espírito Santo state." httpUri="https://zenodo.org/record/2647206/files/figure.png" pageId="9" pageNumber="434">Figure 2</figureCitation>
(f)].
</paragraph>
</subSubSection>
<subSubSection id="C3632C8BC026034D98DDFB74BD9CFAAA" pageId="9" pageNumber="434" type="materials_examined">
<paragraph id="8BC67F00C026034D98DDFB74BD9CFAAA" blockId="9.[151,1437,151,1400]" pageId="9" pageNumber="434">
<emphasis id="B90DA312C026034D98DDFB74BD86FB17" bold="true" box="[199,426,1196,1221]" pageId="9" pageNumber="434">Material examined</emphasis>
. Type-specimens:
<materialsCitation id="3B11755DC026034D9A9AFB74BDD3FB3A" ID-GBIF-Occurrence="2236620088" collectionCode="MCZ" location="rio Doce" pageId="9" pageNumber="434" specimenCode="MCZ 7680" specimenCount="1" stateProvince="Minas Gerais">
<specimenCode id="DBDFD77BC026034D9A9AFB74BED1FB17" box="[640,765,1196,1221]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="9" pageNumber="434" type="Museum">MCZ 7680</specimenCode>
(
<quantity id="4C81D2E5C026034D9B0BFB74BFA7FB16" box="[785,907,1196,1221]" metricMagnitude="0" metricUnit="m" metricValue="1.196" pageId="9" pageNumber="434" unit="mm" value="1196.0">1, 196 mm</quantity>
SL),
<collectingRegion id="49BDB1E2C026034D9BD7FB74B848FB17" box="[973,1124,1196,1221]" country="Brazil" name="Minas Gerais" pageId="9" pageNumber="434">Minas Gerais</collectingRegion>
e
<collectingRegion id="49BDB1E2C026034D9C98FB74B909FB16" box="[1154,1317,1196,1221]" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="434">Espirito Santo</collectingRegion>
,
<location id="8EA629DBC026034D9D28FB74B9B4FB16" LSID="urn:lsid:plazi:treatment:03D0CE16C02A034D988DFC57BD9CFAAA:8EA629DBC026034D9D28FB74B9B4FB16" box="[1330,1432,1196,1221]" name="rio Doce" pageId="9" pageNumber="434" stateProvince="Minas Gerais">rio Doce</location>
,
<locationDeviation id="9A746622C026034D988DFB17BDD3FB3A" box="[151,511,1231,1256]" firstLocation="Linhares" pageId="9" pageNumber="434" secondLocation="Aimores">between Linhares and Aimorés</locationDeviation>
</materialsCitation>
;
<materialsCitation id="3B11755DC026034D9A17FB08BDFAFADE" ID-GBIF-Occurrence="2236620083" collectionCode="MCZ" location="rio Doce" pageId="9" pageNumber="434" specimenCode="MCZ 7681" specimenCount="1" stateProvince="Minas Gerais">
<specimenCode id="DBDFD77BC026034D9A17FB08BEA7FB3A" box="[525,651,1231,1256]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="9" pageNumber="434" type="Museum">MCZ 7681</specimenCode>
(
<quantity id="4C81D2E5C026034D9ABAFB08BF30FB3A" box="[672,796,1232,1256]" metricMagnitude="0" metricUnit="m" metricValue="1.193" pageId="9" pageNumber="434" unit="mm" value="1193.0">1, 193 mm</quantity>
SL),
<collectingRegion id="49BDB1E2C026034D9B7AFB17BFD4FB3A" box="[864,1016,1231,1256]" country="Brazil" name="Minas Gerais" pageId="9" pageNumber="434">Minas Gerais</collectingRegion>
e
<collectingRegion id="49BDB1E2C026034D9C02FB17B890FB3A" box="[1048,1212,1231,1256]" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="434">Espirito Santo</collectingRegion>
,
<location id="8EA629DBC026034D9CD0FB17B91DFB3A" LSID="urn:lsid:plazi:treatment:03D0CE16C02A034D988DFC57BD9CFAAA:8EA629DBC026034D9CD0FB17B91DFB3A" box="[1226,1329,1231,1256]" name="rio Doce" pageId="9" pageNumber="434" stateProvince="Minas Gerais">rio Doce</location>
,
<locationDeviation id="9A746622C026034D9D25FB17BDFAFADE" firstLocation="Linhares" pageId="9" pageNumber="434" secondLocation="Aimores. Non">between Linhares and Aimorés. Non</locationDeviation>
</materialsCitation>
type-specimens:
<materialsCitation id="3B11755DC026034D9A85FB2CBDAFFAE2" ID-GBIF-Occurrence="2236620084" collectionCode="MZUSP" location="Linhares" municipality="Juparana" pageId="9" pageNumber="434" specimenCode="MZUSP 10104-10114" specimenCount="1" stateProvince="Espirito Santo">
<specimenCode id="DBDFD77BC026034D9A85FB2CBFB3FADE" box="[671,927,1268,1292]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="434">MZUSP 1010410114</specimenCode>
(9, 156
<quantity id="4C81D2E5C026034D9C12FB2CB845FADE" box="[1032,1129,1268,1293]" metricMagnitude="-1" metricUnit="m" metricValue="2.04" pageId="9" pageNumber="434" unit="mm" value="204.0">204 mm</quantity>
SL),
<collectingRegion id="49BDB1E2C026034D9CB7FB2CB97DFADE" box="[1197,1361,1268,1293]" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="434">Espírito Santo</collectingRegion>
, lagoa
<collectingMunicipality id="6BA2E57AC026034D988DFAC1BCD7FAE2" box="[151,251,1305,1328]" pageId="9" pageNumber="434">Juparanã</collectingMunicipality>
at
<location id="8EA629DBC026034D9904FACFBDAFFAE2" LSID="urn:lsid:plazi:treatment:03D0CE16C02A034D988DFC57BD9CFAAA:8EA629DBC026034D9904FACFBDAFFAE2" box="[286,387,1303,1328]" municipality="Juparana" name="Linhares" pageId="9" pageNumber="434" stateProvince="Espirito Santo">Linhares</location>
</materialsCitation>
;
<materialsCitation id="3B11755DC026034D998AFAC0B8C4FAE2" ID-GBIF-Occurrence="2236620085" box="[400,1256,1303,1328]" collectionCode="MZUSP" location="Linhares" municipality="Juparana" pageId="9" pageNumber="434" specimenCode="MZUSP 79459" specimenCount="1" stateProvince="Espirito Santo">
<specimenCode id="DBDFD77BC026034D998AFAC0BE17FAE2" box="[400,571,1303,1328]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="434">MZUSP 79459</specimenCode>
(
<quantity id="4C81D2E5C026034D9A55FAC0BEEBFAE2" box="[591,711,1304,1328]" metricMagnitude="0" metricUnit="m" metricValue="1.247" pageId="9" pageNumber="434" unit="mm" value="1247.0">1, 247 mm</quantity>
SL),
<collectingRegion id="49BDB1E2C026034D9B1DFACFBF86FAE2" box="[775,938,1303,1328]" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="434">Espírito Santo</collectingRegion>
, lagoa
<collectingMunicipality id="6BA2E57AC026034D9BE1FAC1B873FAE2" box="[1019,1119,1305,1328]" pageId="9" pageNumber="434">Juparanã</collectingMunicipality>
at
<location id="8EA629DBC026034D9C99FACFB8C4FAE2" LSID="urn:lsid:plazi:treatment:03D0CE16C02A034D988DFC57BD9CFAAA:8EA629DBC026034D9C99FACFB8C4FAE2" box="[1155,1256,1303,1328]" municipality="Juparana" name="Linhares" pageId="9" pageNumber="434" stateProvince="Espirito Santo">Linhares</location>
</materialsCitation>
;
<materialsCitation id="3B11755DC026034D9CEFFAC1BEC7FA87" ID-GBIF-Occurrence="2236620086" collectionCode="LBP" location="Linhares" municipality="Nova" pageId="9" pageNumber="434" specimenCount="1" stateProvince="Espirito Santo">
<collectionCode id="ED68E7C5C026034D9CEFFAC1B904FAE2" box="[1269,1320,1305,1328]" pageId="9" pageNumber="434">LBP</collectionCode>
24068 (
<quantity id="4C81D2E5C026034D9D93FAC0BCD8FA86" metricMagnitude="0" metricUnit="m" metricValue="1.24" pageId="9" pageNumber="434" unit="mm" value="1240.0">1, 240 mm</quantity>
SL),
<collectingRegion id="49BDB1E2C026034D9929FAE4BDF9FA86" box="[307,469,1340,1365]" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="434">Espírito Santo</collectingRegion>
, lagoa
<collectingMunicipality id="6BA2E57AC026034D9A3FFAE5BE4EFA86" box="[549,610,1341,1364]" pageId="9" pageNumber="434">Nova</collectingMunicipality>
at
<location id="8EA629DBC026034D9A9FFAE4BEC7FA87" LSID="urn:lsid:plazi:treatment:03D0CE16C02A034D988DFC57BD9CFAAA:8EA629DBC026034D9A9FFAE4BEC7FA87" box="[645,747,1340,1365]" municipality="Nova" name="Linhares" pageId="9" pageNumber="434" stateProvince="Espirito Santo">Linhares</location>
</materialsCitation>
;
<materialsCitation id="3B11755DC026034D9AECFAE4BD80FAAA" ID-GBIF-Occurrence="2236620087" collectionCode="MZUSP" location="Campos dos Goytacazes" municipality="Feia" pageId="9" pageNumber="434" specimenCode="MZUSP 2295" specimenCount="1" stateProvince="Rio de Janeiro">
<specimenCode id="DBDFD77BC026034D9AECFAE4BFBEFA87" box="[758,914,1340,1365]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="434">MZUSP 2295</specimenCode>
(
<quantity id="4C81D2E5C026034D9BBFFAE4B830FA86" box="[933,1052,1340,1365]" metricMagnitude="0" metricUnit="m" metricValue="1.251" pageId="9" pageNumber="434" unit="mm" value="1251.0">1, 251 mm</quantity>
SL),
<collectingRegion id="49BDB1E2C026034D9C46FAE4B92DFA87" box="[1116,1281,1340,1365]" country="Brazil" name="Rio de Janeiro" pageId="9" pageNumber="434">Rio de Janeiro</collectingRegion>
, lagoa
<collectingMunicipality id="6BA2E57AC026034D9D4AFAE4B9ACFA87" box="[1360,1408,1340,1365]" pageId="9" pageNumber="434">Feia</collectingMunicipality>
at
<location id="8EA629DBC026034D988DFAB8BD80FAAA" LSID="urn:lsid:plazi:treatment:03D0CE16C02A034D988DFC57BD9CFAAA:8EA629DBC026034D988DFAB8BD80FAAA" box="[151,428,1375,1400]" municipality="Feia" name="Campos dos Goytacazes" pageId="9" pageNumber="434" stateProvince="Rio de Janeiro">Campos dos Goytacazes</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>