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<mods:title id="CB6371E322A5C46E104C32E7E633B161">On cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from South India and Sri Lanka, with descriptions of new taxa</mods:title>
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<mods:namePart id="8F0FA36EBA54F7C85DF4EFA34AD4EC77">Anisyutkin, Leonid N.</mods:namePart>
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<treatment id="03CA2572FFD1FF82858017D1C0DFB1C1" ID-DOI="http://doi.org/10.5281/zenodo.6134130" ID-GBIF-Taxon="119515393" ID-Zenodo-Dep="6134130" LSID="urn:lsid:plazi:treatment:03CA2572FFD1FF82858017D1C0DFB1C1" httpUri="http://treatment.plazi.org/id/03CA2572FFD1FF82858017D1C0DFB1C1" lastPageNumber="316" pageId="15" pageNumber="316">
<subSubSection id="C379C7EFFFD1FF82858017D1C124B6CC" box="[151,555,440,466]" pageId="15" pageNumber="316" type="nomenclature">
<paragraph id="8BDC9464FFD1FF82858017D1C124B6CC" blockId="15.[151,555,440,466]" box="[151,555,440,466]" pageId="15" pageNumber="316">
<heading id="D0942308FFD1FF82858017D1C124B6CC" bold="true" box="[151,555,440,466]" fontSize="11" level="1" pageId="15" pageNumber="316" reason="1">
<emphasis id="B9174876FFD1FF82858017D1C124B6CC" bold="true" box="[151,555,440,466]" pageId="15" pageNumber="316">
Genus
<taxonomicName id="4C63EFE7FFD1FF8285F917D1C124B6CC" ID-CoL="8MTXV" authority="Kirby, 1903" authorityName="Kirby" authorityYear="1903" box="[238,555,440,466]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD1FF8285F917D1C29EB6CC" bold="true" box="[238,401,440,466]" italics="true" pageId="15" pageNumber="316">Rhabdoblatta</emphasis>
<bibRefCitation id="EFF2E995FFD1FF82848F17D1C124B6CC" author="Kirby, W. F." box="[408,555,440,466]" firstAuthor="Kirby" journalOrPublisher="Annals and Magazine of Natural History, Series" pageId="15" pageNumber="316" pagination="273 - 280" part="7" refId="ref19888" refString="Kirby, W. F. (1903) Notes on Blattidae &amp; c., with Description of new Genera and Species in the collection of the British Museum, South Kensington. No 2. Annals and Magazine of Natural History, Series 7, 12, 273 - 280. http: // dx. doi. org / 10.1080 / 00222930308678853" title="Notes on Blattidae &amp; c., with Description of new Genera and Species in the collection of the British Museum, South Kensington. No 2" type="journal article" year="1903">Kirby, 1903</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C379C7EFFFD1FF8285801797C07CB509" box="[151,883,510,535]" pageId="15" pageNumber="316" type="type_taxon">
<paragraph id="8BDC9464FFD1FF8285801797C07CB509" blockId="15.[151,1437,510,1759]" box="[151,883,510,535]" pageId="15" pageNumber="316">
<emphasis id="B9174876FFD1FF8285801797C224B509" bold="true" box="[151,299,510,535]" pageId="15" pageNumber="316">
<typeStatus id="54D82AC6FFD1FF8285801797C3DDB509" box="[151,210,510,535]" pageId="15" pageNumber="316">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="4C63EFE7FFD1FF82842D1797C1C6B509" ID-CoL="6G4T2" authority="Walker, 1868" authorityName="Walker" authorityYear="1868" box="[314,713,510,535]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="praecipua">
<emphasis id="B9174876FFD1FF82842D1797C125B508" box="[314,554,510,535]" italics="true" pageId="15" pageNumber="316">Epilampra praecipua</emphasis>
<bibRefCitation id="EFF2E995FFD1FF8287261797C1C6B509" author="Walker, F." bookContentInfo="239 pp." box="[561,713,510,535]" firstAuthor="Walker" journalOrPublisher="Printed for the Trustees of the British Museum, London" pageId="15" pageNumber="316" refId="ref20607" refString="Walker, F. (1868) Catalogue of the specimens of Blattariae in the collection of the British Museum. Printed for the Trustees of the British Museum, London, 239 pp." title="Catalogue of the specimens of Blattariae in the collection of the British Museum" type="book" year="1868">Walker, 1868</bibRefCitation>
</taxonomicName>
. By monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C379C7EFFFD1FF8285D0144BC7DAB32D" pageId="15" pageNumber="316" type="diagnosis">
<paragraph id="8BDC9464FFD1FF8285D0144BC67AB525" blockId="15.[151,1437,510,1759]" box="[199,1397,546,571]" pageId="15" pageNumber="316">
<emphasis id="B9174876FFD1FF8285D0144BC2C2B525" bold="true" box="[199,461,546,571]" pageId="15" pageNumber="316">Differential diagnosis.</emphasis>
The genus
<taxonomicName id="4C63EFE7FFD1FF828758144BC1E8B525" box="[591,743,546,571]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD1FF828758144BC1E8B525" box="[591,743,546,571]" italics="true" pageId="15" pageNumber="316">Rhabdoblatta</emphasis>
</taxonomicName>
can be diagnosed by the following complex of characters.
</paragraph>
<paragraph id="8BDC9464FFD1FF8285D0142FC2A8B40D" blockId="15.[151,1437,510,1759]" pageId="15" pageNumber="316">
Sexual dimorphism poorly marked, both sexes with the completely developed tegmina and wings surpassing the abdominal apex. The anterior margin of fore femur is armed like the
<typeStatus id="54D82AC6FFD1FF8286E01402C728B59D" box="[1015,1063,619,643]" pageId="15" pageNumber="316">type</typeStatus>
B. Hind metatarsus equal to or slightly longer than other segments combined, with the small apical euplantula along its lower margin; the euplantula without spinules; the 2nd4th segments with euplantulae small; all segments of metatarsus with 2 more or less equal rows of spines along its lower margin. Claws symmetrical; arolium well-developed. Paraprocts of the blaberid
<typeStatus id="54D82AC6FFD1FF8284161493C23FB40D" box="[257,304,762,787]" pageId="15" pageNumber="316">type</typeStatus>
(
<figureCitation id="135888E1FFD1FF8284571493C28CB40D" box="[320,387,762,787]" captionStart="FIGURE 8" captionStartId="16.[151,250,1554,1576]" captionTargetBox="[174,1423,210,1519]" captionTargetId="figure@16.[151,1436,193,1533]" captionTargetPageId="16" captionText="FIGURE 8. Rhabdoblatta praecipua (Walker, 1868) (male). A—facial part of head; B—pronotum from above; C—abdominal apex from above; D—paraprocts and adjacent structures from below; E—hypandrium from below; F—right phallomere from above; G—outlines of sclerite L 2 D from above; H—apical part of sclerite L 2 D from above; I—sclerite L 3; J L—outlines of apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: X — 10 th abdominal tergite; ap. scl.; c. p. R 1 T; d. scl.; f. s.; R 2; R 3; R 4; R 5; par.; pv.; s. t. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I; j = J L." httpUri="https://zenodo.org/record/253202/files/figure.png" pageId="15" pageNumber="316">Fig. 8</figureCitation>
D).
</paragraph>
<paragraph id="8BDC9464FFD1FF8285D01577C7DAB32D" blockId="15.[151,1437,510,1759]" pageId="15" pageNumber="316">
Male genitalia (
<figureCitation id="135888E1FFD1FF82846A1577C2CDB429" box="[381,450,798,823]" captionStart="FIGURE 8" captionStartId="16.[151,250,1554,1576]" captionTargetBox="[174,1423,210,1519]" captionTargetId="figure@16.[151,1436,193,1533]" captionTargetPageId="16" captionText="FIGURE 8. Rhabdoblatta praecipua (Walker, 1868) (male). A—facial part of head; B—pronotum from above; C—abdominal apex from above; D—paraprocts and adjacent structures from below; E—hypandrium from below; F—right phallomere from above; G—outlines of sclerite L 2 D from above; H—apical part of sclerite L 2 D from above; I—sclerite L 3; J L—outlines of apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: X — 10 th abdominal tergite; ap. scl.; c. p. R 1 T; d. scl.; f. s.; R 2; R 3; R 4; R 5; par.; pv.; s. t. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I; j = J L." httpUri="https://zenodo.org/record/253202/files/figure.png" pageId="15" pageNumber="316">Fig. 8</figureCitation>
FL). Right phallomere (R+N) generally similar to those of
<taxonomicName id="4C63EFE7FFD1FF8281681577C7E8B429" box="[1151,1255,798,823]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD1FF8281681577C7E8B429" box="[1151,1255,798,823]" italics="true" pageId="15" pageNumber="316">Morphna</emphasis>
</taxonomicName>
and the related genera, i.e.
<taxonomicName id="4C63EFE7FFD1FF828409152BC2E5B445" box="[286,490,834,859]" class="Insecta" family="Blaberidae" genus="Indoapterolampra" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9174876FFD1FF828409152BC2E5B445" box="[286,490,834,859]" italics="true" pageId="15" pageNumber="316">Indoapterolampra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224F50DFFD1FF8284E3152BC150B445" box="[500,607,834,859]" pageId="15" pageNumber="316" rank="genus">
<emphasis id="B9174876FFD1FF8284E3152BC150B445" bold="true" box="[500,607,834,859]" pageId="15" pageNumber="316">gen. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C63EFE7FFD1FF828767152BC016B445" box="[624,793,834,859]" class="Insecta" family="Blaberidae" genus="Comptolampra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD1FF828767152BC016B445" box="[624,793,834,859]" italics="true" pageId="15" pageNumber="316">Comptolampra</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63EFE7FFD1FF82863C152BC0A1B445" box="[811,942,834,859]" class="Insecta" family="Blaberidae" genus="Placoblatta" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD1FF82863C152BC0A1B445" box="[811,942,834,859]" italics="true" pageId="15" pageNumber="316">Placoblatta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63EFE7FFD1FF8286A9152BC741B445" box="[958,1102,834,859]" class="Insecta" family="Blaberidae" genus="Anisolampra" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD1FF8286A9152BC741B445" box="[958,1102,834,859]" italics="true" pageId="15" pageNumber="316">Anisolampra</emphasis>
</taxonomicName>
and
<taxonomicName id="4C63EFE7FFD1FF82819D152BC62BB445" box="[1162,1316,834,859]" class="Insecta" family="Blaberidae" genus="Opisthoplatia" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD1FF82819D152BC62BB445" box="[1162,1316,834,859]" italics="true" pageId="15" pageNumber="316">Opisthoplatia</emphasis>
</taxonomicName>
(see
<figureCitation id="135888E1FFD1FF828072152BC3ACB461" captionStart="FIGURE 3" captionStartId="7.[151,250,1635,1657]" captionTargetBox="[161,1422,202,1595]" captionTargetId="figure@7.[151,1436,193,1614]" captionTargetPageId="7" captionText="FIGURE 3. Indoapterolampra rugosiuscula sp. nov., hypandrium and male genitalia (A D, G J—holotype, E, F—paratype). A—hypandrium and genitalia from above; B, E—right phallomere from above; C—sclerite R 3 from below; D, F—cranial part of sclerite R 3 from outside; G, H—apical part of sclerite L 2 D from above (G) and outside (H); I, J—sclerite L 3. Dotted area shows membranous parts. Abbreviations: a. scl.; b. L 2 D; b. L 3; c. p. R 1 T; f. s.; f. scl.; L 4 U; m. l.; r. plm.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B F; c = G, H; d = I, J." httpUri="https://zenodo.org/record/253197/files/figure.png" pageId="15" pageNumber="316">Figs. 3</figureCitation>
BF, 4G, H, 5GI, 6L, 8F and
<bibRefCitation id="EFF2E995FFD1FF828719150FC1C0B461" author="Anisyutkin, L. N." box="[526,719,870,895]" firstAuthor="Anisyutkin" journalOrPublisher="Entomological Review" pageId="15" pageNumber="316" pagination="434 - 454" part="79" refId="ref19136" refString="Anisyutkin, L. N. (1999) Cockroaches of the subfamily Epilamprinae (Dictyoptera, Blaberidae) from the Indochina Peninsula. Entomological Review, 79 (4), 434 - 454." title="Cockroaches of the subfamily Epilamprinae (Dictyoptera, Blaberidae) from the Indochina Peninsula" type="journal article" year="1999">Anisyutkin 1999</bibRefCitation>
,
<bibRefCitation id="EFF2E995FFD1FF8287CA150EC017B461" author="Anisyutkin, L. N." box="[733,792,871,895]" firstAuthor="Anisyutkin" journalOrPublisher="Entomological Review" pageId="15" pageNumber="316" pagination="190 - 208" part="80" refId="ref19172" refString="Anisyutkin, L. N. (2000) New Cockroach Species of the Genus Rhabdoblatta Kirby (Dictyoptera, Blaberidae) from Southeast Asia: I. Entomological Review, 80 (2), 190 - 208." title="New Cockroach Species of the Genus Rhabdoblatta Kirby (Dictyoptera, Blaberidae) from Southeast Asia: I" type="journal article" year="2000">2000</bibRefCitation>
,
<bibRefCitation id="EFF2E995FFD1FF828631150EC06EB461" author="Anisyutkin, L. N." box="[806,865,871,895]" firstAuthor="Anisyutkin" journalOrPublisher="Entomological Review" pageId="15" pageNumber="316" pagination="540 - 556" part="82" refId="ref19212" refString="Anisyutkin, L. N. (2003) New and Little Known Cockroaches of the Genus Rhabdoblatta Kirby (Dictyoptera, Blaberidae) from Vietnam and Southern China: II. Entomological Review, 82 (5), 540 - 556." title="New and Little Known Cockroaches of the Genus Rhabdoblatta Kirby (Dictyoptera, Blaberidae) from Vietnam and Southern China: II" type="journal article" year="2003">2003</bibRefCitation>
;
<bibRefCitation id="EFF2E995FFD1FF828667150FC7C2B461" author="Anisyutkin, L. N. &amp; Gorochov, A. V." box="[880,1229,870,895]" firstAuthor="Anisyutkin" journalOrPublisher="Entomological Review" pageId="15" pageNumber="316" pagination="289 - 295" part="81" refId="ref19412" refString="Anisyutkin, L. N. &amp; Gorochov, A. V. (2001) New Data on the Genus Morphna Shelford (Dictyoptera, Blattida, Blaberidae) of Indochina. Entomological Review, 81 (3), 289 - 295." title="New Data on the Genus Morphna Shelford (Dictyoptera, Blattida, Blaberidae) of Indochina" type="journal article" year="2001">Anisyutkin &amp; Gorochov 2001</bibRefCitation>
): the sclerite R1T robust and heavily sclerotized, with caudal part subrectangular or widely rounded in shape, densely covered with bristles. The sclerite L2D (L1) subdivided into basal and apical parts (
<figureCitation id="135888E1FFD1FF8286D715C7C70BB4D9" box="[960,1028,942,967]" captionStart="FIGURE 8" captionStartId="16.[151,250,1554,1576]" captionTargetBox="[174,1423,210,1519]" captionTargetId="figure@16.[151,1436,193,1533]" captionTargetPageId="16" captionText="FIGURE 8. Rhabdoblatta praecipua (Walker, 1868) (male). A—facial part of head; B—pronotum from above; C—abdominal apex from above; D—paraprocts and adjacent structures from below; E—hypandrium from below; F—right phallomere from above; G—outlines of sclerite L 2 D from above; H—apical part of sclerite L 2 D from above; I—sclerite L 3; J L—outlines of apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: X — 10 th abdominal tergite; ap. scl.; c. p. R 1 T; d. scl.; f. s.; R 2; R 3; R 4; R 5; par.; pv.; s. t. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I; j = J L." httpUri="https://zenodo.org/record/253202/files/figure.png" pageId="15" pageNumber="316">Fig. 8</figureCitation>
G), the apical part of L2D with flat &quot;dorsal sclerite&quot; (
<figureCitation id="135888E1FFD1FF82844915BBC2ADB4F5" box="[350,418,978,1003]" captionStart="FIGURE 8" captionStartId="16.[151,250,1554,1576]" captionTargetBox="[174,1423,210,1519]" captionTargetId="figure@16.[151,1436,193,1533]" captionTargetPageId="16" captionText="FIGURE 8. Rhabdoblatta praecipua (Walker, 1868) (male). A—facial part of head; B—pronotum from above; C—abdominal apex from above; D—paraprocts and adjacent structures from below; E—hypandrium from below; F—right phallomere from above; G—outlines of sclerite L 2 D from above; H—apical part of sclerite L 2 D from above; I—sclerite L 3; J L—outlines of apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: X — 10 th abdominal tergite; ap. scl.; c. p. R 1 T; d. scl.; f. s.; R 2; R 3; R 4; R 5; par.; pv.; s. t. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I; j = J L." httpUri="https://zenodo.org/record/253202/files/figure.png" pageId="15" pageNumber="316">Fig. 8</figureCitation>
H,
<emphasis id="B9174876FFD1FF8284DF15BBC10CB4F5" box="[456,515,978,1003]" italics="true" pageId="15" pageNumber="316">d.scl.</emphasis>
) free or more or less fused with the cap-like &quot;apical sclerite&quot; (
<figureCitation id="135888E1FFD1FF8281CE15BBC613B4F5" box="[1241,1308,978,1003]" captionStart="FIGURE 8" captionStartId="16.[151,250,1554,1576]" captionTargetBox="[174,1423,210,1519]" captionTargetId="figure@16.[151,1436,193,1533]" captionTargetPageId="16" captionText="FIGURE 8. Rhabdoblatta praecipua (Walker, 1868) (male). A—facial part of head; B—pronotum from above; C—abdominal apex from above; D—paraprocts and adjacent structures from below; E—hypandrium from below; F—right phallomere from above; G—outlines of sclerite L 2 D from above; H—apical part of sclerite L 2 D from above; I—sclerite L 3; J L—outlines of apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: X — 10 th abdominal tergite; ap. scl.; c. p. R 1 T; d. scl.; f. s.; R 2; R 3; R 4; R 5; par.; pv.; s. t. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I; j = J L." httpUri="https://zenodo.org/record/253202/files/figure.png" pageId="15" pageNumber="316">Fig. 8</figureCitation>
H,
<emphasis id="B9174876FFD1FF82805415BBC683B4F5" box="[1347,1420,978,1003]" italics="true" pageId="15" pageNumber="316">ap.scl.</emphasis>
), densely covered with recumbent bristles. The sclerite L3 (L2d) varies in shape, usually with the more or less expressed &quot;apical crest&quot; (
<figureCitation id="135888E1FFD1FF8284A61273C2FEB32D" box="[433,497,1050,1075]" captionStart="FIGURE 8" captionStartId="16.[151,250,1554,1576]" captionTargetBox="[174,1423,210,1519]" captionTargetId="figure@16.[151,1436,193,1533]" captionTargetPageId="16" captionText="FIGURE 8. Rhabdoblatta praecipua (Walker, 1868) (male). A—facial part of head; B—pronotum from above; C—abdominal apex from above; D—paraprocts and adjacent structures from below; E—hypandrium from below; F—right phallomere from above; G—outlines of sclerite L 2 D from above; H—apical part of sclerite L 2 D from above; I—sclerite L 3; J L—outlines of apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: X — 10 th abdominal tergite; ap. scl.; c. p. R 1 T; d. scl.; f. s.; R 2; R 3; R 4; R 5; par.; pv.; s. t. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I; j = J L." httpUri="https://zenodo.org/record/253202/files/figure.png" pageId="15" pageNumber="316">Fig. 8</figureCitation>
JL,
<emphasis id="B9174876FFD1FF82873C1275C166B32D" box="[555,617,1052,1075]" italics="true" pageId="15" pageNumber="316">ap.cr.</emphasis>
), the &quot;folded structure&quot; distinct, the groove hge absent.
</paragraph>
</subSubSection>
<subSubSection id="C379C7EFFFD1FF8285D01257C1A1B2A1" pageId="15" pageNumber="316" type="discussion">
<paragraph id="8BDC9464FFD1FF8285D01257C3E3B365" blockId="15.[151,1437,510,1759]" pageId="15" pageNumber="316">
<emphasis id="B9174876FFD1FF8285D01257C250B349" bold="true" box="[199,351,1086,1111]" pageId="15" pageNumber="316">Composition</emphasis>
. Follows
<bibRefCitation id="EFF2E995FFD1FF8284C51257C197B349" accessDate="2013-05-24" author="Beccaloni, G. W." box="[466,664,1086,1111]" firstAuthor="Beccaloni" pageId="15" pageNumber="316" publicationUrl="http://Blattodea.SpeciesFile.org" refId="ref19486" refString="Beccaloni, G. W. (2007) Blattodea Species File Online. Version 1.2 / 4.0. Available from: http: // Blattodea. SpeciesFile. org (accessed 24 May 2013)" title="Blattodea Species File Online. Version 1.2 / 4.0." type="journal article" year="2007">Beccaloni (2007)</bibRefCitation>
, except for
<taxonomicName id="4C63EFE7FFD1FF8286351256C0BAB348" box="[802,949,1087,1110]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="praecipua">
<emphasis id="B9174876FFD1FF8286351256C0BAB348" box="[802,949,1087,1110]" italics="true" pageId="15" pageNumber="316">R. praecipua</emphasis>
</taxonomicName>
which is considered the valid species (see below).
</paragraph>
<paragraph id="8BDC9464FFD1FF8285D012EFC2C0B3F9" blockId="15.[151,1437,510,1759]" pageId="15" pageNumber="316">
<emphasis id="B9174876FFD1FF8285D012EFC205B381" bold="true" box="[199,266,1158,1183]" pageId="15" pageNumber="316">Notes</emphasis>
. In my opinion, none of the characters listed above in the 'Differential diagnosis' can be considered a synapomorphy, if taken separately. The genus
<taxonomicName id="4C63EFE7FFD1FF8287A012C3C040B3DD" box="[695,847,1194,1219]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD1FF8287A012C3C040B3DD" box="[695,847,1194,1219]" italics="true" pageId="15" pageNumber="316">Rhabdoblatta</emphasis>
</taxonomicName>
is likely to be paraphyletic, with respect to some genera of the
<taxonomicName id="4C63EFE7FFD1FF82842712A7C2C4B3F9" box="[304,459,1230,1255]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="subFamily" subFamily="Epilamprinae">Epilamprinae</taxonomicName>
.
</paragraph>
<paragraph id="8BDC9464FFD1FF8285D0129BC079B231" blockId="15.[151,1437,510,1759]" pageId="15" pageNumber="316">
The genus
<taxonomicName id="4C63EFE7FFD1FF82845F129BC2EEB215" box="[328,481,1266,1291]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD1FF82845F129BC2EEB215" box="[328,481,1266,1291]" italics="true" pageId="15" pageNumber="316">Rhabdoblatta</emphasis>
</taxonomicName>
is one of the largest in the
<taxonomicName id="4C63EFE7FFD1FF82863C129BC0CAB215" box="[811,965,1266,1291]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="subFamily" subFamily="Epilamprinae">Epilamprinae</taxonomicName>
, numbering over 100 species (
<bibRefCitation id="EFF2E995FFD1FF82803D129BC3DEB231" accessDate="2013-05-24" author="Beccaloni, G. W." firstAuthor="Beccaloni" pageId="15" pageNumber="316" publicationUrl="http://Blattodea.SpeciesFile.org" refId="ref19486" refString="Beccaloni, G. W. (2007) Blattodea Species File Online. Version 1.2 / 4.0. Available from: http: // Blattodea. SpeciesFile. org (accessed 24 May 2013)" title="Blattodea Species File Online. Version 1.2 / 4.0." type="journal article" year="2007">Beccaloni 2007</bibRefCitation>
) and it is evident that more species remain to be discovered.
</paragraph>
<paragraph id="8BDC9464FFD1FF8285D01353C1A1B2A1" blockId="15.[151,1437,510,1759]" pageId="15" pageNumber="316">
This genus was described as monotypic to include the single species
<taxonomicName id="4C63EFE7FFD1FF8286C71353C651B24D" authority="Walker, 1868" authorityName="Walker" authorityYear="1868" box="[976,1374,1338,1363]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="praecipua">
<emphasis id="B9174876FFD1FF8286C71353C7CEB24D" box="[976,1217,1338,1363]" italics="true" pageId="15" pageNumber="316">Epilampra praecipua</emphasis>
<bibRefCitation id="EFF2E995FFD1FF8281DF1353C651B24D" author="Walker, F." bookContentInfo="239 pp." box="[1224,1374,1338,1363]" firstAuthor="Walker" journalOrPublisher="Printed for the Trustees of the British Museum, London" pageId="15" pageNumber="316" refId="ref20607" refString="Walker, F. (1868) Catalogue of the specimens of Blattariae in the collection of the British Museum. Printed for the Trustees of the British Museum, London, 239 pp." title="Catalogue of the specimens of Blattariae in the collection of the British Museum" type="book" year="1868">Walker, 1868</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F374D4F4FFD1FF8285801337C208B269" box="[151,263,1374,1399]" name="Sri Lanka" pageId="15" pageNumber="316">Sri Lanka</collectingCountry>
(
<bibRefCitation id="EFF2E995FFD1FF8284001337C290B269" author="Kirby, W. F." box="[279,415,1374,1399]" firstAuthor="Kirby" journalOrPublisher="Annals and Magazine of Natural History, Series" pageId="15" pageNumber="316" pagination="273 - 280" part="7" refId="ref19888" refString="Kirby, W. F. (1903) Notes on Blattidae &amp; c., with Description of new Genera and Species in the collection of the British Museum, South Kensington. No 2. Annals and Magazine of Natural History, Series 7, 12, 273 - 280. http: // dx. doi. org / 10.1080 / 00222930308678853" title="Notes on Blattidae &amp; c., with Description of new Genera and Species in the collection of the British Museum, South Kensington. No 2" type="journal article" year="1903">Kirby 1903</bibRefCitation>
). Later, additional species from Old and New World, mostly from South-East Asia, were added to the genus
<taxonomicName id="4C63EFE7FFD1FF82846F13EBC11EB285" box="[376,529,1410,1435]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD1FF82846F13EBC11EB285" box="[376,529,1410,1435]" italics="true" pageId="15" pageNumber="316">Rhabdoblatta</emphasis>
</taxonomicName>
by different authors (
<bibRefCitation id="EFF2E995FFD1FF82861C13EBC0BEB285" author="Shelford, R." box="[779,945,1410,1435]" editor="Wytsman, P." firstAuthor="Shelford" journalOrPublisher="Bruxelles" pageId="15" pageNumber="316" pagination="1 - 21" part="101" refId="ref20566" refString="Shelford, R. (1910) Orthoptera. Fam. Blattidae. Subfam. Epilamprinae. In: Wytsman, P. (Ed.), Genera Insectorum, 101. Bruxelles, pp. 1 - 21." title="Orthoptera. Fam. Blattidae. Subfam. Epilamprinae" type="journal article" volumeTitle="Genera Insectorum" year="1910">Shelford 1910</bibRefCitation>
;
<bibRefCitation id="EFF2E995FFD1FF8286A813EBC768B285" author="Hanitsch, R." box="[959,1127,1410,1435]" firstAuthor="Hanitsch" journalOrPublisher="Journal Straits Branch Royal Asiatic Society" pageId="15" pageNumber="316" pagination="17 - 178" part="69" refId="ref19804" refString="Hanitsch, R. (1915) Malayan Blattidae. Journal Straits Branch Royal Asiatic Society, 69, 17 - 178." title="Malayan Blattidae" type="journal article" year="1915">Hanitsch 1915</bibRefCitation>
;
<bibRefCitation id="EFF2E995FFD1FF82816213EBC64FB285" author="Bey-Bienko, G. Ya" bookContentInfo="343 pp." box="[1141,1344,1410,1435]" firstAuthor="Bey-Bienko" journalOrPublisher="Leningrad, Moscow" pageId="15" pageNumber="316" refId="ref19522" refString="Bey-Bienko, G. Ya. (1950) Nasekomye Tarakanovye [Cockroach insects]. Fauna USSR, New Series 40. Leningrad, Moscow, 343 pp." title="Nasekomye Tarakanovye [Cockroach insects]. Fauna USSR, New Series 40" type="book" year="1950">Bey-Bienko 1950</bibRefCitation>
;
<bibRefCitation id="EFF2E995FFD1FF82805913EBC3DCB2A1" author="Princis, K." editor="Beier, M." firstAuthor="Princis" pageId="15" pageNumber="316" pagination="615 - 710" part="11" refId="ref20181" refString="Princis, K. (1967) Blattariae: Subordo Epilamproidea. Fam.: Nyctiboridae, Epilampridae. In: Beier, M. (Ed.), Orthopterorum Catalogus, 11, 615 - 710." title="Blattariae: Subordo Epilamproidea. Fam.: Nyctiboridae, Epilampridae" type="book chapter" volumeTitle="Orthopterorum Catalogus" year="1967">Princis 1967</bibRefCitation>
;
<bibRefCitation id="EFF2E995FFD1FF8285CA13CFC293B2A1" author="Anisyutkin, L. N." box="[221,412,1446,1471]" firstAuthor="Anisyutkin" journalOrPublisher="Entomological Review" pageId="15" pageNumber="316" pagination="190 - 208" part="80" refId="ref19172" refString="Anisyutkin, L. N. (2000) New Cockroach Species of the Genus Rhabdoblatta Kirby (Dictyoptera, Blaberidae) from Southeast Asia: I. Entomological Review, 80 (2), 190 - 208." title="New Cockroach Species of the Genus Rhabdoblatta Kirby (Dictyoptera, Blaberidae) from Southeast Asia: I" type="journal article" year="2000">Anisyutkin 2000</bibRefCitation>
,
<bibRefCitation id="EFF2E995FFD1FF8284B013CEC2EDB2A1" author="Anisyutkin, L. N." box="[423,482,1447,1471]" firstAuthor="Anisyutkin" journalOrPublisher="Entomological Review" pageId="15" pageNumber="316" pagination="540 - 556" part="82" refId="ref19212" refString="Anisyutkin, L. N. (2003) New and Little Known Cockroaches of the Genus Rhabdoblatta Kirby (Dictyoptera, Blaberidae) from Vietnam and Southern China: II. Entomological Review, 82 (5), 540 - 556." title="New and Little Known Cockroaches of the Genus Rhabdoblatta Kirby (Dictyoptera, Blaberidae) from Vietnam and Southern China: II" type="journal article" year="2003">2003</bibRefCitation>
;
<bibRefCitation id="EFF2E995FFD1FF8284F913CFC1ADB2A1" accessDate="2013-05-24" author="Beccaloni, G. W." box="[494,674,1446,1471]" firstAuthor="Beccaloni" pageId="15" pageNumber="316" publicationUrl="http://Blattodea.SpeciesFile.org" refId="ref19486" refString="Beccaloni, G. W. (2007) Blattodea Species File Online. Version 1.2 / 4.0. Available from: http: // Blattodea. SpeciesFile. org (accessed 24 May 2013)" title="Blattodea Species File Online. Version 1.2 / 4.0." type="journal article" year="2007">Beccaloni 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C379C7EFFFD1FF8285D013A5C0DFB1C1" pageId="15" pageNumber="316" type="discussion">
<paragraph id="8BDC9464FFD1FF8285D013A5C0DFB1C1" blockId="15.[151,1437,510,1759]" pageId="15" pageNumber="316">
<taxonomicName id="4C63EFE7FFD1FF8285D013A5C252B2FD" box="[199,349,1484,1507]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="praecipua">
<emphasis id="B9174876FFD1FF8285D013A5C252B2FD" box="[199,349,1484,1507]" italics="true" pageId="15" pageNumber="316">R. praecipua</emphasis>
</taxonomicName>
, the type species of the genus
<taxonomicName id="4C63EFE7FFD1FF8287CA13A3C07AB2FD" box="[733,885,1482,1507]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD1FF8287CA13A3C07AB2FD" box="[733,885,1482,1507]" italics="true" pageId="15" pageNumber="316">Rhabdoblatta</emphasis>
</taxonomicName>
, was synonymized with
<taxonomicName id="4C63EFE7FFD1FF8281B413A3C23EB119" authority="Walker, 1868" authorityName="Walker" authorityYear="1868" class="Insecta" family="Blattidae" genus="Polyzosteria" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="terranea">
<emphasis id="B9174876FFD1FF8281B413A3C693B2FD" box="[1187,1436,1482,1507]" italics="true" pageId="15" pageNumber="316">Polyzosteria terranea</emphasis>
<bibRefCitation id="EFF2E995FFD1FF8285801387C23EB119" author="Walker, F." bookContentInfo="239 pp." box="[151,305,1518,1543]" firstAuthor="Walker" journalOrPublisher="Printed for the Trustees of the British Museum, London" pageId="15" pageNumber="316" refId="ref20607" refString="Walker, F. (1868) Catalogue of the specimens of Blattariae in the collection of the British Museum. Printed for the Trustees of the British Museum, London, 239 pp." title="Catalogue of the specimens of Blattariae in the collection of the British Museum" type="book" year="1868">Walker, 1868</bibRefCitation>
</taxonomicName>
by K. Princis, without any argumentation: &quot;
<taxonomicName id="4C63EFE7FFD1FF8286521386C0A4B118" box="[837,939,1519,1542]" class="Insecta" family="Blattidae" genus="Polyzosteria" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="terranea">
<emphasis id="B9174876FFD1FF8286521386C0A4B118" box="[837,939,1519,1542]" italics="true" pageId="15" pageNumber="316">Terranea</emphasis>
</taxonomicName>
ist auf das Larvenstadium von
<taxonomicName id="4C63EFE7FFD1FF82803D1386C693B118" box="[1322,1436,1519,1542]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="praecipua">
<emphasis id="B9174876FFD1FF82803D1386C693B118" box="[1322,1436,1519,1542]" italics="true" pageId="15" pageNumber="316">praecipua</emphasis>
</taxonomicName>
gegründet, welch letzterer Name also als Synonym eingezogen werden muss.&quot; (
<bibRefCitation id="EFF2E995FFD1FF828125107BC61BB135" author="Princis, K." box="[1074,1300,1554,1579]" journalOrPublisher="Opuscula entomologica" pageId="15" pageNumber="316" pagination="59 - 75" part="23" refId="ref20123" refString="Princis, K. (1958) Revision der Walkerschen und Kirbyschen Blattarientypen im British Museum of Natural History, London. II. Opuscula entomologica, 23 (1 - 2), 59 - 75." title="Revision der Walkerschen und Kirbyschen Blattarientypen im British Museum of Natural History, London. II" type="journal article" year="1958" yearSuffix="p">Princis 1958, p. 66</bibRefCitation>
). I have reexamined the
<typeStatus id="54D82AC6FFD1FF828429105FC2AFB151" box="[318,416,1590,1615]" pageId="15" pageNumber="316" type="holotype">holotype</typeStatus>
of '
<taxonomicName id="4C63EFE7FFD1FF8284C4105EC15BB150" box="[467,596,1591,1614]" class="Insecta" family="Blattidae" genus="Polyzosteria" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="terranea">
<emphasis id="B9174876FFD1FF8284C4105EC15BB150" box="[467,596,1591,1614]" italics="true" pageId="15" pageNumber="316">P. terranea</emphasis>
</taxonomicName>
', which turned out to be a cockroach larva belonging to the subfamily
<taxonomicName id="4C63EFE7FFD1FF8285801033C23DB16D" box="[151,306,1626,1651]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="subFamily" subFamily="Epilamprinae">Epilamprinae</taxonomicName>
. At the current state of knowledge, it can only be identified as
<taxonomicName id="4C63EFE7FFD1FF8286FE1033C78FB16D" box="[1001,1152,1626,1651]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="subFamily" subFamily="Epilamprinae">Epilamprinae</taxonomicName>
gen. sp. It is obvious that more than one
<taxonomicName id="4C63EFE7FFD1FF8284521017C2D3B189" box="[325,476,1662,1687]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="subFamily" subFamily="Epilamprinae">Epilamprinae</taxonomicName>
species occur in
<collectingCountry id="F374D4F4FFD1FF8287B11017C016B189" box="[678,793,1662,1687]" name="Sri Lanka" pageId="15" pageNumber="316">Sri Lanka</collectingCountry>
. Furthermore,
<emphasis id="B9174876FFD1FF8286D51016C74AB188" box="[962,1093,1663,1686]" italics="true" pageId="15" pageNumber="316">
'
<taxonomicName id="4C63EFE7FFD1FF8286D21016C74AB188" box="[965,1093,1663,1686]" class="Insecta" family="Blattidae" genus="Polyzosteria" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="terranea">P. terranea</taxonomicName>
</emphasis>
' is characterized by the black facial part of the head (
<bibRefCitation id="EFF2E995FFD1FF82848910CBC15EB1A5" accessDate="2013-05-24" author="Beccaloni, G. W." box="[414,593,1698,1723]" firstAuthor="Beccaloni" pageId="15" pageNumber="316" publicationUrl="http://Blattodea.SpeciesFile.org" refId="ref19486" refString="Beccaloni, G. W. (2007) Blattodea Species File Online. Version 1.2 / 4.0. Available from: http: // Blattodea. SpeciesFile. org (accessed 24 May 2013)" title="Blattodea Species File Online. Version 1.2 / 4.0." type="journal article" year="2007">Beccaloni 2007</bibRefCitation>
) as compared to the pale yellowish head in
<taxonomicName id="4C63EFE7FFD1FF82815410CDC7DBB1A5" box="[1091,1236,1700,1723]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="praecipua">
<emphasis id="B9174876FFD1FF82815410CDC7DBB1A5" box="[1091,1236,1700,1723]" italics="true" pageId="15" pageNumber="316">R. praecipua</emphasis>
</taxonomicName>
. Taking the latter difference into account, it is safe to consider
<taxonomicName id="4C63EFE7FFD1FF82879A10AEC011B1C0" box="[653,798,1735,1758]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="praecipua">
<emphasis id="B9174876FFD1FF82879A10AEC011B1C0" box="[653,798,1735,1758]" italics="true" pageId="15" pageNumber="316">R. praecipua</emphasis>
</taxonomicName>
a valid species.
</paragraph>
</subSubSection>
</treatment>
</document>