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<document id="774E87E014B15C333A84C8EEDE5C11B6" ID-CLB-Dataset="26946" ID-DOI="10.11646/zootaxa.4590.4.1" ID-GBIF-Dataset="64c1a738-381b-4ba2-bc35-317d756e6a29" ID-ISSN="1175-5326" ID-Zenodo-Dep="2656164" ID-ZooBank="CD2A364C-6CF9-4FF5-AD28-7AF388868B15" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1556698917896" checkinUser="plazi" docAuthor="Mohrig, Werner, Kauschke, Ellen &amp; Broadley, Adam" docDate="2019" docId="03C987F6FFDBFFA8BD885D19FB08711B" docLanguage="en" docName="zootaxa.4590.4.1.pdf" docOrigin="Zootaxa 4590 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cratyna (Cratyna) longicosta Mohrig &amp; Kauschke &amp; Broadley 2019, comb. n." docType="treatment" docVersion="8" lastPageNumber="416" masterDocId="FFF0FF8EFFD4FFA7BD1F590EFFB97777" masterDocTitle="Revision of black fungus gnat species (Diptera, Sciaridae) described from the Hawaiian Islands by D. E. Hardy and W. A. Steffan, and a contribution to the knowledge of the sciarid fauna of the Galápagos Islands" masterLastPageNumber="439" masterPageNumber="401" pageNumber="416" updateTime="1698724290689" updateUser="ExternalLinkService">
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<mods:title id="CCEC5C9F0290B6D0DEAA901A969CEEEE">Revision of black fungus gnat species (Diptera, Sciaridae) described from the Hawaiian Islands by D. E. Hardy and W. A. Steffan, and a contribution to the knowledge of the sciarid fauna of the Galápagos Islands</mods:title>
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<paragraph id="8BDF36E0FFDBFFA8BD885D19FCA77346" blockId="15.[151,798,1047,1107]" box="[151,798,1047,1073]" pageId="15" pageNumber="416">
<heading id="D097818CFFDBFFA8BD885D19FCA77346" bold="true" box="[151,798,1047,1073]" fontSize="11" level="1" pageId="15" pageNumber="416" reason="1">
<emphasis id="B914EAF2FFDBFFA8BD885D19FCA77346" bold="true" box="[151,798,1047,1073]" pageId="15" pageNumber="416">
<taxonomicName id="4C604D63FFDBFFA8BD885D19FD127346" ID-CoL="8BLNK" authority="(Hardy, 1956)" authorityName="Mohrig &amp; Kauschke &amp; Broadley" authorityYear="2019" baseAuthorityName="Hardy" baseAuthorityYear="1956" box="[151,683,1047,1073]" class="Insecta" family="Sciaridae" genus="Cratyna" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="longicosta" status="comb. nov." subGenus="Cratyna">
<emphasis id="B914EAF2FFDBFFA8BD885D19FF407346" bold="true" box="[151,249,1047,1073]" italics="true" pageId="15" pageNumber="416">Cratyna</emphasis>
(
<emphasis id="B914EAF2FFDBFFA8BC155D19FED57346" bold="true" box="[266,364,1047,1073]" italics="true" pageId="15" pageNumber="416">Cratyna</emphasis>
)
<emphasis id="B914EAF2FFDBFFA8BC635D19FE417346" bold="true" box="[380,504,1047,1073]" italics="true" pageId="15" pageNumber="416">longicosta</emphasis>
(
<bibRefCitation id="EFF14B11FFDBFFA8BF185D19FD1A7346" author="Hardy, D. E." box="[519,675,1047,1073]" pageId="15" pageNumber="416" pagination="72 - 90" refId="ref15081" refString="Hardy, D. E. (1956) New Hawaiian Sciaridae (Diptera). Proceedings of the Hawaiian Entomological Society, 16 (1), 72 - 90." type="journal article" year="1956">Hardy, 1956</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2275789FFDBFFA8BFAC5D19FCA77346" box="[691,798,1047,1073]" pageId="15" pageNumber="416" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C37A656BFFDBFFA8BD885D34FE8D7325" box="[151,308,1082,1107]" pageId="15" pageNumber="416" type="description">
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(
<figureCitation id="135B2A65FFDBFFA8BD805D34FE947325" box="[159,301,1082,1107]" captionStart="FIGURE 12" captionStartId="18.[151,250,1997,2019]" captionTargetBox="[221,1395,218,1916]" captionTargetId="figure@18.[179,1419,195,1982]" captionTargetPageId="18" captionText="FIGURE 12 AC. Cratyna longicosta (Hardy, 1956). Holotype. A. Hypopygium; B. Basal segments of antenna; C. Palpus." httpUri="https://zenodo.org/record/2656186/files/figure.png" pageId="15" pageNumber="416">
Fig. 12 A
<emphasis id="B914EAF2FFDBFFA8BC145D34FEA07324" bold="true" box="[267,281,1082,1107]" pageId="15" pageNumber="416"></emphasis>
C
</figureCitation>
)
</paragraph>
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<subSubSection id="C37A656BFFDBFFA8BD885D8CFBBE73EF" box="[151,1031,1154,1176]" pageId="15" pageNumber="416" type="reference_group">
<paragraph id="8BDF36E0FFDBFFA8BD885D8CFBBE73EF" blockId="15.[151,1031,1154,1176]" box="[151,1031,1154,1176]" pageId="15" pageNumber="416">
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<taxonomicName id="4C604D63FFDBFFA8BD885D8CFD1E73EF" ID-CoL="88YDY" authority="Hardy, 1956" authorityName="Hardy" authorityYear="1956" box="[151,679,1154,1176]" class="Insecta" family="Sciaridae" genus="Plastosciara" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="longicosta" subGenus="Cosmosciara">
<emphasis id="B914EAF2FFDBFFA8BD885D8CFEAE73EF" box="[151,279,1154,1176]" italics="true" pageId="15" pageNumber="416">Plastosciara</emphasis>
(
<emphasis id="B914EAF2FFDBFFA8BC3A5D8CFE1273E0" box="[293,427,1154,1175]" italics="true" pageId="15" pageNumber="416">Cosmosciara</emphasis>
)
<emphasis id="B914EAF2FFDBFFA8BCA55D8CFD9D73EF" box="[442,548,1154,1176]" italics="true" pageId="15" pageNumber="416">longicosta</emphasis>
<bibRefCitation id="EFF14B11FFDBFFA8BF355D8CFD1E73EF" author="Hardy, D. E." box="[554,679,1154,1176]" pageId="15" pageNumber="416" pagination="72 - 90" refId="ref15081" refString="Hardy, D. E. (1956) New Hawaiian Sciaridae (Diptera). Proceedings of the Hawaiian Entomological Society, 16 (1), 72 - 90." type="journal article" year="1956">Hardy, 1956</bibRefCitation>
</taxonomicName>
[
<treatmentCitation id="0AC110F1FFDBFFA8BFAD5D8CFCDC73EF" author="Hardy, D. E." box="[690,869,1154,1176]" page="75" pageId="15" pageNumber="416" year="1956">
<bibRefCitation id="EFF14B11FFDBFFA8BFAD5D8CFCFB73EF" author="Hardy, D. E." box="[690,834,1154,1176]" pageId="15" pageNumber="416" pagination="72 - 90" refId="ref15081" refString="Hardy, D. E. (1956) New Hawaiian Sciaridae (Diptera). Proceedings of the Hawaiian Entomological Society, 16 (1), 72 - 90." type="journal article" year="1956">Hardy (1956)</bibRefCitation>
: 75
</treatmentCitation>
<emphasis id="B914EAF2FFDBFFA8BE7A5D8CFCC873EF" bold="true" box="[869,881,1154,1176]" pageId="15" pageNumber="416"></emphasis>
77, fig. 3 a
<emphasis id="B914EAF2FFDBFFA8BEC05D8CFC5273EF" bold="true" box="[991,1003,1154,1176]" pageId="15" pageNumber="416"></emphasis>
d].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37A656BFFDBFFA8BD885DCAFC62723E" pageId="15" pageNumber="416" type="materials_examined">
<paragraph id="8BDF36E0FFDBFFA8BD885DCAFEA97276" blockId="15.[151,1436,1220,1644]" pageId="15" pageNumber="416">
<emphasis id="B914EAF2FFDBFFA8BD885DCAFE6473AA" bold="true" box="[151,477,1220,1245]" pageId="15" pageNumber="416">
Material studied:
<materialsCitation id="3B083CBDFFDBFFA8BC6F5DCAFE6473AA" ID-GBIF-Occurrence="2237390862" box="[368,477,1220,1245]" collectingDate="1952-11-24" collectionCode="BPBM" collectorName="C. P. Hoyt" location="Hawai'i" pageId="15" pageNumber="416" specimenCount="1" stateProvince="Hawaii" typeStatus="holotype">
<typeStatus id="54DB8842FFDBFFA8BC6F5DCAFE6473AA" box="[368,477,1220,1245]" pageId="15" pageNumber="416" type="holotype">Holotype</typeStatus>
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</emphasis>
: Male,
<date id="FFDE1020FFDBFFA8BF295DCAFD0973AA" box="[566,688,1220,1245]" pageId="15" pageNumber="416" value="1952-11-24">
<collectingDate id="EF9AE9C8FFDBFFA8BF295DCAFD0973AA" box="[566,688,1220,1245]" pageId="15" pageNumber="416" value="1952-11-24">24.xi.1952</collectingDate>
</date>
,
<collectingRegion id="49A4F802FFDBFFA8BFA25DCAFCA373AA" box="[701,794,1220,1245]" country="United States of America" name="Hawaii" pageId="15" pageNumber="416">Hawaii</collectingRegion>
, Oahu, Hālawa Ridge, ex moss, leg.
<collectorName id="26955336FFDBFFA8B9D75DCBFA8F73AA" box="[1224,1334,1221,1245]" pageId="15" pageNumber="416">C.P. Hoyt</collectorName>
(BPBM, No. 2482).
</paragraph>
<paragraph id="8BDF36E0FFDBFFA8BDD85C02FC62723E" blockId="15.[151,1436,1220,1644]" pageId="15" pageNumber="416">
<emphasis id="B914EAF2FFDBFFA8BDD85C02FE027252" bold="true" box="[199,443,1292,1317]" pageId="15" pageNumber="416">Conservation status.</emphasis>
Embedded in
<collectingCountry id="F3777670FFDBFFA8BF425C02FD087252" box="[605,689,1292,1317]" name="Canada" pageId="15" pageNumber="416">Canada</collectingCountry>
balsam. Head, palpus, antennae and wings isolated, wings under a separate cover slip. All morphological structures are rather well preserved.
</paragraph>
</subSubSection>
<subSubSection id="C37A656BFFDBFFA8BDD85C5AFB08711B" pageId="15" pageNumber="416" type="discussion">
<paragraph id="8BDF36E0FFDBFFA8BDD85C5AFB08711B" blockId="15.[151,1436,1220,1644]" pageId="15" pageNumber="416">
<emphasis id="B914EAF2FFDBFFA8BDD85C5AFEF1721A" bold="true" box="[199,328,1364,1389]" pageId="15" pageNumber="416">Comments</emphasis>
. The species is characterized by a 1-segmented palpus with a small rudimentary second segment, the basal segment not swollen, short oval and without a deepened sensory pit; antenna rather short, 4
<superScript id="7C159BA8FFDBFFA8B9E35C76FAB172F1" attach="left" box="[1276,1288,1400,1414]" fontSize="6" pageId="15" pageNumber="416">th</superScript>
flagellomere with a l/w index of 1.2, hairs fine and somewhat shorter than the diameter of the basal node; legs slender, fore tibia without a distinct bristle patch; hypopygium with short and rather robust gonocoxites; gonocoxites with short and sparse hairs on the ventral inner side, gonostylus elongate, somewhat bulbous, apically with 2 fine and slightly curved spines and 2 fine subapical spines at the inner side. Hardy put the species into the genus
<taxonomicName id="4C604D63FFDBFFA8B8185F02FA2D7153" authorityName="Berg" authorityYear="1899" box="[1287,1428,1548,1572]" class="Insecta" family="Sciaridae" genus="Plastosciara" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAF2FFDBFFA8B8185F02FA2D7153" box="[1287,1428,1548,1572]" italics="true" pageId="15" pageNumber="416">Plastosciara</emphasis>
</taxonomicName>
, subgenus
<taxonomicName id="4C604D63FFDBFFA8BC135F3EFE197130" box="[268,416,1584,1607]" class="Insecta" family="Sciaridae" genus="Plastosciara" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="subGenus" subGenus="Cosmosciara">
<emphasis id="B914EAF2FFDBFFA8BC135F3EFE197130" box="[268,416,1584,1607]" italics="true" pageId="15" pageNumber="416">Cosmosciara</emphasis>
</taxonomicName>
. We agree with Hardy concerning the position in the genus
<taxonomicName id="4C604D63FFDBFFA8B99C5F3EFB667130" authorityName="Winnertz" authorityYear="1867" box="[1155,1247,1584,1607]" class="Insecta" family="Sciaridae" genus="Cratyna" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAF2FFDBFFA8B99C5F3EFB667130" box="[1155,1247,1584,1607]" italics="true" pageId="15" pageNumber="416">Cratyna</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C604D63FFDBFFA8B81A5F21FA2B7130" authorityName="Berg" authorityYear="1899" box="[1285,1426,1583,1607]" class="Insecta" family="Sciaridae" genus="Plastosciara" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAF2FFDBFFA8B81A5F21FA2B7130" box="[1285,1426,1583,1607]" italics="true" pageId="15" pageNumber="416">Plastosciara</emphasis>
</taxonomicName>
) because of a shortened palpus and two pairs of fine spines on the inner side of the gonostylus.
</paragraph>
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</treatment>
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