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<document id="90FC9EEFB2744B3C27121D9C92B55ED2" ID-CLB-Dataset="53324" ID-DOI="10.11646/phytotaxa.576.2.1" ID-GBIF-Dataset="1a9bd59f-7293-432f-b3c1-6df0bc659474" ID-ISSN="1179-3163" ID-Zenodo-Dep="7461280" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1671521402645" checkinUser="plazi" docAuthor="Hassan, Walaa A., Al-Shaye, Najla A., Alghamdi, Salma, Korany, Shereen M. &amp; Iamonico, Duilio" docDate="2022" docId="03C987964B4FFFA906B9037457C3618A" docLanguage="en" docName="phytotaxa.576.2.1.pdf" docOrigin="Phytotaxa 576 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.576.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Amaranthus graecizans subsp. sylvestris Brenan" docType="treatment" docVersion="7" lastPageNumber="143" masterDocId="FFF0FFEE4B47FFA106310226570A6226" masterDocTitle="Taxonomic revision of the genus Amaranthus (Amaranthaceae) in Saudi Arabia" masterLastPageNumber="157" masterPageNumber="135" pageNumber="143" updateTime="1699413697922" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AECC3711528CE1B2F92BC0F52309A9A4">Taxonomic revision of the genus Amaranthus (Amaranthaceae) in Saudi Arabia</mods:title>
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<mods:namePart id="92ED3213055B7F6C93F14F8B10B6E50F">Hassan, Walaa A.</mods:namePart>
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<mods:namePart id="FFCA74F18135CE4AB7C2D48D566904DB">Al-Shaye, Najla A.</mods:namePart>
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<mods:affiliation id="5B586485460D3152A845AC06A04ADB2E">Department of Biology, College of Science, Princess Nourah bint Abdulrahman University, P. O. Box 84428, Riyadh 11671, Saudi Arabia &amp; naaalshaye @ pnu. edu. sa; https: // orcid. org / 0000 - 0002 - 0447 - 8613</mods:affiliation>
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<mods:namePart id="5E6AA845218F6791C63084EB946D23AE">Alghamdi, Salma</mods:namePart>
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<mods:affiliation id="6A3ABC735E021B9C64875233F95DFCD9">Department of Biology, College of Science, Princess Nourah bint Abdulrahman University, P. O. Box 84428, Riyadh 11671, Saudi Arabia &amp; saatalghamdi @ pnu. edu. sa; https: // orcid. org / 0000 - 0001 - 7716 - 2233</mods:affiliation>
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<mods:namePart id="5E8ABBBFAAE520DED78E1A804737AF6E">Korany, Shereen M.</mods:namePart>
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<mods:affiliation id="CBAE17FEDF4B676E0F7234C83F820609">Department of Biology, College of Science, Princess Nourah bint Abdulrahman University, P. O. Box 84428, Riyadh 11671, Saudi Arabia &amp; smkorany @ pnu. edu. as; https: // orcid. org / 0000 - 0002 - 9355 - 3952</mods:affiliation>
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<mods:namePart id="0DA9E5683B72E052761BE0BC543E3E9D">Iamonico, Duilio</mods:namePart>
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<mods:affiliation id="6A3BA95104A0DDDE19012B3C081CF4EA">Ce. R. S. I. Te. S., University of Rome Sapienza, viale XXIV Maggio 7, 04100 - Latina, Italy &amp; duilio. iamonico @ uniroma 1. it; https: // orcid. org / 0000 - 0001 - 5491 - 7568 &amp; Authors that contributed equally to the work</mods:affiliation>
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<emphasis id="B914EA924B4FFFA906B9037457A7634A" bold="true" box="[136,173,338,364]" pageId="8" pageNumber="143">5b.</emphasis>
<treatmentCitation id="0AC110914B4FFFA906850374535F634A" author="Brenan" box="[180,1109,338,365]" journal="Watsonia" page="273" pageId="8" pageNumber="143" score="9.729594737742866" volume="4" year="1961">
<taxonomicName id="4C604D034B4FFFA9068503745446634B" ID-CoL="933W5" authority="(Vill.) Brenan" authorityName="Brenan" baseAuthorityName="Vill." box="[180,844,338,365]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="143" phylum="Tracheophyta" rank="subSpecies" species="graecizans" subSpecies="sylvestris">
<emphasis id="B914EA924B4FFFA90685037456C2634A" bold="true" box="[180,456,338,365]" italics="true" pageId="8" pageNumber="143">Amaranthus graecizans</emphasis>
L. subsp.
<emphasis id="B914EA924B4FFFA9040C037455AF634B" bold="true" box="[573,677,338,365]" italics="true" pageId="8" pageNumber="143">sylvestris</emphasis>
(Vill.) Brenan
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, Watsonia 4: 273. 1961
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<taxonomicName id="4C604D034B4FFFA902450374566563B6" ID-CoL="3F6TL" authority="Vill., Cat. Pl. Jard. Strasb." class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="143" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="B914EA924B4FFFA9024503745266634A" box="[1140,1388,338,364]" italics="true" pageId="8" pageNumber="143">Amaranthus sylvestris</emphasis>
Vill., Cat. Pl. Jard. Strasb.
</taxonomicName>
111. 1807 ≡
<treatmentCitation id="0AC110914B4FFFA90434035053E363B6" author="Poiret" box="[517,1257,374,403]" page="44" pageId="8" pageNumber="143" volumeTitle="Tabl. Ecole Bot." year="1804">
<taxonomicName id="4C604D034B4FFFA904340350537263B7" ID-CoL="3F6TL" authority="Desf. ex Poiret, Tabl. Ecole Bot." authorityName="Poiret, Tabl. Ecole Bot." box="[517,1144,374,403]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="143" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="B914EA924B4FFFA90434035055F763B6" box="[517,765,374,400]" italics="true" pageId="8" pageNumber="143">Amaranthus sylvestris</emphasis>
Desf. ex Poiret, Tabl. École Bot.
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: 44. 1804
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,
<emphasis id="B914EA924B4FFFA902C40351526C63B6" box="[1269,1382,374,400]" italics="true" pageId="8" pageNumber="143">nom. nud.</emphasis>
,
<emphasis id="B914EA924B4FFFA90345035157C26392" italics="true" pageId="8" pageNumber="143">nom. inval.</emphasis>
(Art. 38.2 Ex.1 of the ICN) ≡
<taxonomicName id="4C604D034B4FFFA9040403BC52676392" ID-CoL="933W5" authority="(Desf.) Asch., Beitr. Fl. Aethiop." baseAuthorityName="Desf." box="[565,1389,410,437]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="143" phylum="Tracheophyta" rank="variety" species="graecizans" variety="sylvestris">
<emphasis id="B914EA924B4FFFA9040403BC544B6392" box="[565,833,410,436]" italics="true" pageId="8" pageNumber="143">Amaranthus graecizans</emphasis>
var.
<emphasis id="B914EA924B4FFFA9054F03BC54EC6392" box="[894,998,410,436]" italics="true" pageId="8" pageNumber="143">sylvestris</emphasis>
(Desf.) Asch., Beitr. Fl. Aethiop.
</taxonomicName>
: 176. 1867,
<emphasis id="B914EA924B4FFFA906FE0398565963FE" box="[207,339,446,472]" italics="true" pageId="8" pageNumber="143">comb. illeg.</emphasis>
<treatmentCitation id="0AC110914B4FFFA90742039856A163DA" author="Vill." issue="1" journal="Inst. Catalana Hist. Nat., Secc. Bot." page="89" pageId="8" pageNumber="143" score="7.350453370179623" volume="38" year="1974">
<taxonomicName id="4C604D034B4FFFA90742039857F063DB" ID-CoL="933W5" authority="(Vill.) O. Bolos &amp; Vigo, Butl. Inst. Catalana Hist. Nat., Secc. Bot." baseAuthorityName="Vill." class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="143" phylum="Tracheophyta" rank="subSpecies" species="graecizans" subSpecies="sylvestris">
<emphasis id="B914EA924B4FFFA907420398557663FE" box="[371,636,446,472]" italics="true" pageId="8" pageNumber="143">Amaranthus graecizans</emphasis>
subsp.
<emphasis id="B914EA924B4FFFA904E20398543163FE" box="[723,827,446,472]" italics="true" pageId="8" pageNumber="143">sylvestris</emphasis>
(Vill.) O.Bolòs &amp; Vigo, Butl. Inst. Catalana Hist. Nat., Secc. Bot.
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38(1): 89. 1974
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<emphasis id="B914EA924B4FFFA907F903C455E263DA" box="[456,744,482,508]" italics="true" pageId="8" pageNumber="143">Amaranthus angustifolius</emphasis>
proles
<taxonomicName id="4C604D034B4FFFA9050903C4520063DA" ID-CoL="3F6TL" authority="(Vill.) Thell., Syn. Mitteleur. Fl." authorityName="Thell., Syn. Mitteleur. Fl." baseAuthorityName="Vill." box="[824,1290,482,508]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="143" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="B914EA924B4FFFA9050903C454AA63DA" box="[824,928,482,508]" italics="true" pageId="8" pageNumber="143">sylvestris</emphasis>
(Vill.) Thell., Syn. Mitteleur. Fl.
</taxonomicName>
[Ascherson &amp; Graebner] 5(1(5)): 300. 1914 ≡
<treatmentCitation id="0AC110914B4FFFA907DD002056B36062" author="Heukels" page="170" pageId="8" pageNumber="143" score="0.8122984981307134" volumeTitle="Geillustreerde Schoolflora voor Nederland" year="1934">
<taxonomicName id="4C604D034B4FFFA907DD002052A66006" ID-CoL="933W5" authority="(Vill.) Heukels, Geillustreerde Schoolflora" authorityName="Heukels, Geillustreerde Schoolflora" baseAuthorityName="Vill." box="[492,1452,518,544]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="143" phylum="Tracheophyta" rank="subSpecies" species="angustifolius" subSpecies="sylvestris">
<emphasis id="B914EA924B4FFFA907DD002054066006" box="[492,780,518,544]" italics="true" pageId="8" pageNumber="143">Amaranthus angustifolius</emphasis>
subsp.
<emphasis id="B914EA924B4FFFA90551002054C26006" box="[864,968,518,544]" italics="true" pageId="8" pageNumber="143">sylvestris</emphasis>
(Vill.) Heukels, Geïllustreerde Schoolflora
</taxonomicName>
voor
<collectingCountry id="F37776104B4FFFA906F3000C56316062" box="[194,315,554,580]" name="Netherlands" pageId="8" pageNumber="143">Nederland</collectingCountry>
: 170. 1934
</treatmentCitation>
.
</paragraph>
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<paragraph id="8BDF36804B4FFFA906B9007454966041" blockId="8.[136,1452,338,616]" box="[136,924,594,616]" pageId="8" pageNumber="143">
Type (
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designated by
<bibRefCitation id="EFF14B714B4FFFA9079F007455616041" author="Townsend, C. C." box="[430,619,594,616]" pageId="8" pageNumber="143" pagination="1 - 136" refId="ref18438" refString="Townsend, C. C. (1985) Amaranthaceae. In: Polhill, R. M. (Ed.) Flora of Tropical East Africa. A. A. Balkema, Rotterdam, pp. 1 - 136." type="book chapter" year="1985">Townsend 1985: 31</bibRefCitation>
):—
<emphasis id="B914EA924B4FFFA904A0007454616041" box="[657,875,594,615]" italics="true" pageId="8" pageNumber="143">Herb. Tournefort 1849</emphasis>
(P!).
</paragraph>
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<subSubSection id="C37A650B4B4FFFA906B900B053DD6096" box="[136,1239,662,689]" pageId="8" pageNumber="143" type="description">
<paragraph id="8BDF36804B4FFFA906B900B053DD6096" blockId="8.[136,1451,662,940]" box="[136,1239,662,689]" pageId="8" pageNumber="143">
<emphasis id="B914EA924B4FFFA906B900B0564C6096" bold="true" box="[136,326,662,688]" pageId="8" pageNumber="143">Iconography:—</emphasis>
<bibRefCitation id="EFF14B714B4FFFA9077700B055046096" author="Willdenow, C. L." box="[326,526,662,688]" pageId="8" pageNumber="143" refId="ref18857" refString="Willdenow, C. L. (1790) Historia Amaranthorum. Impensis Ziegleri et fil., Turici, 38 pp." type="book" year="1790">Willdenow (1790</bibRefCitation>
: Tab. VIII, fig. 16 sub
<taxonomicName id="4C604D034B4FFFA9053A00B154786096" box="[779,882,662,689]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="143" phylum="Tracheophyta" rank="species" species="viridis">
<emphasis id="B914EA924B4FFFA9053A00B154786096" box="[779,882,662,689]" italics="true" pageId="8" pageNumber="143">A. viridis</emphasis>
</taxonomicName>
),
<bibRefCitation id="EFF14B714B4FFFA905B800B0535C6096" author="Bayon, N. D." box="[905,1110,662,689]" pageId="8" pageNumber="143" pagination="261 - 383" refId="ref15004" refString="Bayon, N. D. (2015) Revision taxonomica de las especies monoicas de Amaranthus (Amaranthaceae): Amaranthus subg. Amaranthus and Amaranthus subg. Albersia. Ann. Missouri Bot. Garden 101 (2): 261 - 383. https: // doi. org / 10.3417 / 2010080" type="journal article" year="2015">Bayón (2015: 329</bibRefCitation>
, Figura 34)
</paragraph>
</subSubSection>
<subSubSection id="C37A650B4B4FFFA9068C009C554F613A" pageId="8" pageNumber="143" type="biology_ecology">
<paragraph id="8BDF36804B4FFFA9068C009C557D60F2" blockId="8.[136,1451,662,940]" box="[189,631,698,724]" pageId="8" pageNumber="143">
<emphasis id="B914EA924B4FFFA9068C009C565560F2" bold="true" box="[189,351,698,724]" pageId="8" pageNumber="143">Phenology:—</emphasis>
Flowering time february.
</paragraph>
<paragraph id="8BDF36804B4FFFA9068C00F8531F60DE" blockId="8.[136,1451,662,940]" box="[189,1045,734,761]" pageId="8" pageNumber="143">
<emphasis id="B914EA924B4FFFA9068C00F856EF60DE" bold="true" box="[189,485,734,760]" pageId="8" pageNumber="143">Habitat and elevation:—</emphasis>
Human-made habitat (coastal plain),
<quantity id="4C989B654B4FFFA905B400F854D060DF" box="[901,986,734,761]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="8" pageNumber="143" unit="m" value="10.0" valueMax="15.0" valueMin="5.0">515 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8BDF36804B4FFFA9068C0124554F613A" blockId="8.[136,1451,662,940]" box="[189,581,770,797]" pageId="8" pageNumber="143">
<emphasis id="B914EA924B4FFFA9068C012456C6613A" bold="true" box="[189,460,770,796]" pageId="8" pageNumber="143">Chromosome number:</emphasis>
—2n = 32.
</paragraph>
</subSubSection>
<subSubSection id="C37A650B4B4FFFA9068C010054336166" box="[189,825,806,832]" pageId="8" pageNumber="143" type="distribution">
<paragraph id="8BDF36804B4FFFA9068C010054336166" blockId="8.[136,1451,662,940]" box="[189,825,806,832]" pageId="8" pageNumber="143">
<emphasis id="B914EA924B4FFFA9068C010055786166" bold="true" box="[189,626,806,832]" pageId="8" pageNumber="143">
Occurrence in Saudi Arabia (
<figureCitation id="135B2A054B4FFFA9042A010055686166" box="[539,610,806,832]" captionStart="FIGURE 5" captionStartId="8.[136,229,1598,1620]" captionTargetId="figure-269@8.[340,1247,965,1573]" captionTargetPageId="8" captionText="FIGURE 5. Map of Amaranthus graecizans subsp. sylvestris." figureDoi="http://doi.org/10.5281/zenodo.7465628" httpUri="https://zenodo.org/record/7465628/files/figure.png" pageId="8" pageNumber="143">Fig. 5</figureCitation>
):
</emphasis>
<collectingRegion id="49A4F8624B4FFFA904A1010055C16166" box="[656,715,806,832]" country="Saudi Arabia" name="Jizan" pageId="8" pageNumber="143">Jizan</collectingRegion>
,
<collectingRegion id="49A4F8624B4FFFA904E70100543C6166" box="[726,822,806,832]" country="Saudi Arabia" name="Makkah" pageId="8" pageNumber="143">Makkah</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A650B4B4FFFA9068C016C57C3618A" pageId="8" pageNumber="143" type="materials_examined">
<paragraph id="8BDF36804B4FFFA9068C016C57C3618A" blockId="8.[136,1451,662,940]" pageId="8" pageNumber="143">
<emphasis id="B914EA924B4FFFA9068C016C56E66142" bold="true" box="[189,492,842,868]" pageId="8" pageNumber="143">Specimina visa selecta:—</emphasis>
<collectingCountry id="F37776104B4FFFA907DD016C55BC6142" box="[492,694,842,868]" name="Saudi Arabia" pageId="8" pageNumber="143">SAUDI ARABIA</collectingCountry>
,
<collectingRegion id="49A4F8624B4FFFA904F3016C542B6142" box="[706,801,842,868]" country="Saudi Arabia" name="Makkah" pageId="8" pageNumber="143">Makkah</collectingRegion>
, Jeddah,
<emphasis id="B914EA924B4FFFA905B8016C54A56142" box="[905,943,842,868]" italics="true" pageId="8" pageNumber="143">s.d.</emphasis>
,
<emphasis id="B914EA924B4FFFA9058F016D532E6142" box="[958,1060,842,868]" italics="true" pageId="8" pageNumber="143">Kruijt 48</emphasis>
(L 1684182!);
<collectingRegion id="49A4F8624B4FFFA902E0016C52076142" box="[1233,1293,842,868]" country="Saudi Arabia" name="Jizan" pageId="8" pageNumber="143">Jizan</collectingRegion>
, human-made habitat (coastal plain),
<quantity id="4C989B654B4FFFA907B3014856DF61AF" box="[386,469,878,905]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="8" pageNumber="143" unit="m" value="10.0" valueMax="15.0" valueMin="5.0">515 m</quantity>
a.s.l.,
<date id="FFDE10404B4FFFA9042B014855EA61AE" box="[538,736,878,904]" pageId="8" pageNumber="143" value="2021-02-17">17 February 2021</date>
, leg.
<emphasis id="B914EA924B4FFFA90526014854FF61AE" box="[791,1013,878,904]" italics="true" pageId="8" pageNumber="143">Masrhai et Al-shaye</emphasis>
, det.
<emphasis id="B914EA924B4FFFA9021E0148538761AE" box="[1071,1165,878,904]" italics="true" pageId="8" pageNumber="143">Masrhai</emphasis>
, conf.
<emphasis id="B914EA924B4FFFA902E60149524B61AE" box="[1239,1345,879,904]" italics="true" pageId="8" pageNumber="143">Iamonico</emphasis>
(PNUH!,
<collectionCode id="ED71AE454B4FFFA906B901B457B2618A" box="[136,184,914,940]" country="Italy" lsid="urn:lsid:biocol.org:col:15061" name="Università degli Studi di Roma La Sapienza" pageId="8" pageNumber="143" type="Herbarium">RO</collectionCode>
!).
</paragraph>
</subSubSection>
</treatment>
</document>