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<document id="136A1785BC21D317991C711D7B18B6DF" ID-DOI="10.11646/zootaxa.4759.1.1" ID-GBIF-Dataset="f75b13f6-a898-42ff-93a4-2d177da2207f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3735855" ID-ZooBank="8C3F94C9-3923-43F4-918A-6A1CA92F1F49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585730665729" checkinUser="plazi" docAuthor="Huynh, Cuong" docDate="2020" docId="03C287E9861BFFE4FF40FDC25703FF09" docLanguage="en" docName="zootaxa.4759.1.1.pdf" docOrigin="Zootaxa 4759 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monographis condorensis Huynh 2020, new species" docType="treatment" docVersion="5" lastPageNumber="18" masterDocId="FFFBFF918616FFF5FFD7FF8A547AFFDE" masterDocTitle="Three new species of penicillate millipedes from the Con Dao Islands of southeast Vietnam (Diplopoda, Lophoproctidae and Polyxenidae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="14" updateTime="1698734097456" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="7F3A3ED5E7E7EE184D20D3AB4297FCB8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E9ED3A880E8CE1491CA3953B01BAB243">
<mods:title id="B6AC35EAE52524FA37CF3B9D83504A5C">Three new species of penicillate millipedes from the Con Dao Islands of southeast Vietnam (Diplopoda, Lophoproctidae and Polyxenidae)</mods:title>
</mods:titleInfo>
<mods:name id="15D6F40C8342BA7BF421AC45F2D6CF73" type="personal">
<mods:role id="FBC4693BD658166B9A8C1749777C6A87">
<mods:roleTerm id="59B09E46575B2E5853575531A0303C1D">Author</mods:roleTerm>
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<mods:namePart id="5BCF172B1880A115F747A95CDC4E43CF">Huynh, Cuong</mods:namePart>
</mods:name>
<mods:typeOfResource id="22A6C917466FAD8834E6910255C8532C">text</mods:typeOfResource>
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<mods:title id="12269DB0F836EA2F335A411F8A552906">Zootaxa</mods:title>
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<mods:part id="2AC935B9FCF9D64BE1D2238E8C819C2A">
<mods:date id="CC49E4DAD663D5F77979AE3091ABC2A6">2020</mods:date>
<mods:detail id="C95A0E4468480034795DFCDDA9EE6296" type="pubDate">
<mods:number id="76D3BB67C5D9B2C42A6E7FCA7E9E9074">2020-04-01</mods:number>
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<mods:detail id="7347961DA82D4F7EEFDE8D800CF67891" type="volume">
<mods:number id="5B96FA63417598937EF79AC2B6264A91">4759</mods:number>
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<mods:number id="F1EC5612F660A25C664A78F7E3E98143">1</mods:number>
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<mods:start id="C92E24A098272DAB1EB253BFAD7350BE">1</mods:start>
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<mods:classification id="3FA16A30185778426DE005FAEE4CCF26">journal article</mods:classification>
<mods:identifier id="2781E6146F9116487153AB5443824B97" type="DOI">10.11646/zootaxa.4759.1.1</mods:identifier>
<mods:identifier id="0AD24B524EBB24BCCDEFD6C659D84DE3" type="GBIF-Dataset">f75b13f6-a898-42ff-93a4-2d177da2207f</mods:identifier>
<mods:identifier id="818B33CF36F673BFB599E1EEA67A120D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D66C46DB5F206F34241A9DDFA013C72B" type="Zenodo-Dep">3735855</mods:identifier>
<mods:identifier id="358FA19F7B91874FACE180878E6A4326" type="ZooBank">8C3F94C9-3923-43F4-918A-6A1CA92F1F49</mods:identifier>
</mods:mods>
<treatment id="03C287E9861BFFE4FF40FDC25703FF09" ID-DOI="http://doi.org/10.5281/zenodo.3812001" ID-GBIF-Taxon="163420470" ID-Zenodo-Dep="3812001" LSID="urn:lsid:plazi:treatment:03C287E9861BFFE4FF40FDC25703FF09" httpUri="http://treatment.plazi.org/id/03C287E9861BFFE4FF40FDC25703FF09" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14">
<subSubSection id="C3716574861BFFF8FF40FDC25610FDBC" box="[151,618,584,611]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BD436FF861BFFF8FF40FDC25610FDBC" blockId="13.[151,618,584,646]" box="[151,618,584,611]" pageId="13" pageNumber="14">
<heading id="D09C8193861BFFF8FF40FDC25610FDBC" bold="true" box="[151,618,584,611]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B91FEAED861BFFF8FF40FDC25610FDBC" bold="true" box="[151,618,584,611]" pageId="13" pageNumber="14">
<taxonomicName id="4C6B4D7C861BFFF8FF40FDC255A9FDBD" ID-CoL="681d0ea4-07bc-4c91-a8d4-840ca4874a0e" authority="Huynh, 2020" authorityName="Huynh" authorityYear="2020" box="[151,467,584,611]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B91FEAED861BFFF8FF40FDC255A9FDBD" bold="true" box="[151,467,584,611]" italics="true" pageId="13" pageNumber="14">Monographis condorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C5796861BFFF8FE0DFDC35610FDBC" box="[474,618,584,611]" pageId="13" pageNumber="14" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3716574861BFFF8FF40FDE6556AFD58" box="[151,272,620,646]" pageId="13" pageNumber="14" type="description">
<paragraph id="8BD436FF861BFFF8FF40FDE6556AFD58" blockId="13.[151,618,584,646]" box="[151,272,620,646]" pageId="13" pageNumber="14">
<figureCitation id="13502A7A861BFFF8FF40FDE6556AFD58" box="[151,272,620,646]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStart-4="FIGURE 15" captionStart-5="FIGURE 16" captionStartId-0="13.[151,250,1721,1746]" captionStartId-1="15.[151,250,1825,1850]" captionStartId-2="16.[151,250,1754,1779]" captionStartId-3="17.[151,250,1373,1398]" captionStartId-4="18.[151,250,1890,1915]" captionStartId-5="19.[151,250,1225,1250]" captionTargetBox-0="[205,1382,1028,1696]" captionTargetBox-1="[186,1412,391,1786]" captionTargetBox-2="[206,1383,181,1728]" captionTargetBox-3="[151,1435,241,1348]" captionTargetBox-4="[177,1408,351,1859]" captionTargetBox-5="[171,1410,186,1199]" captionTargetId-0="figure@13.[205,1382,1028,1696]" captionTargetId-1="figure@15.[175,1412,381,1800]" captionTargetId-2="figure@16.[205,1383,181,1728]" captionTargetId-3="figure@17.[151,1436,237,1348]" captionTargetId-4="figure@18.[168,1420,345,1865]" captionTargetId-5="figure@19.[171,1416,181,1199]" captionTargetPageId-0="13" captionTargetPageId-1="15" captionTargetPageId-2="16" captionTargetPageId-3="17" captionTargetPageId-4="18" captionTargetPageId-5="19" captionText-0="FIGURE 11. Monographis condorensis n. sp. showing the body cover with dense, long trichomes." captionText-1="FIGURE 12. Holotype Monographis condorensis n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. T2: Tergite 2; D. T10: Tergite 10—the last tergite; E. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; F. Gnathochilarium: the lateral palp (LP) with 11 sensilla and medial palp (MP) with 22 sensilla; G. Clypeo-labrum showing a smooth surface, asetose, with setae along both anterior edges. H. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d)." captionText-2="FIGURE 13. SEM images of Monographis condorensis n. sp., A. Dorsal view of the head capsule, showing 8 ommatidia and trichobothria a, b and c; B. Dorsal view of the body, co: Collum, Lp: lateral protuberances and pp: pleural projection, T2: tergite 2 and T3: tergite 3; C. Clypeo-labrum showing the smooth surface and asetose (l), s: setae along anterior edges, Se: posterior setae, bse: a minute backward-pointing setae; D. Right gnathochilarium showing the lateral palp (LP) with 5 sensilla and 6 sensilla sockets, 22 sensilla on the medial palp (MP); E. Bare caudal bundle in the female showed 2 distinctive structures: caudal trichome sockets (cts) which include barbate trichome sockets (bts) in the middle of the bundle, forming a dorsal single bundle, nest trichome sockets (nts) forming 2 ventral bundles; these bundles are separated by the trichome socket-free tissue (tsft); T: telson; F. Leg articles and arrows indicate spines which are typical characters of Monographis." captionText-3="FIGURE 14. Monographis condorensis n. sp., A. SEM images of antennomeres VI, VII and VIII with 4 sensory cones (sc), showing the arrangement of sensilla on antennomeres VI and VII; B. Antennomere VII with 2 thick sensilla (T), setiform sensillum (s) between them and a conical sensillum (c) located in posterior position; C. Antennomere VI with a group of 29 thin sensilla forming a triangular shape followed by a setiform sensillum." captionText-4="FIGURE 15. Holotype Monographis condorensis n. sp., A. Antennomeres VI, VII and VIII with sensory cones (sc), showing the arrangement of sensilla on antennomeres VI and VII; B. Antennomere VII with 2 thick sensilla (Ta (anterior) and Tp (posterior)), setiform sensillum (s) between them and a conical sensillum (c) located next to Tp; antennomere VI with a group of 26 sensilla (23 thin sensilla (t) and 3 thick sensilla (★)) forming a triangular shape followed by a setiform sensillum (s)." captionText-5="FIGURE 16. Holotype Monographis condorensis n. sp., A. 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta present in the middle of the femur; D. Seta on tibia; E. Smaller spine on tarsus 1; F. Larger spine on T2; G. Telotarsus—claw structures (SEM image of one specimen) and a drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi-0="http://doi.org/10.5281/zenodo.3735877" figureDoi-1="http://doi.org/10.5281/zenodo.3735879" figureDoi-2="http://doi.org/10.5281/zenodo.3735883" figureDoi-3="http://doi.org/10.5281/zenodo.3735885" figureDoi-4="http://doi.org/10.5281/zenodo.3735887" figureDoi-5="http://doi.org/10.5281/zenodo.3735889" httpUri-0="https://zenodo.org/record/3735877/files/figure.png" httpUri-1="https://zenodo.org/record/3735879/files/figure.png" httpUri-2="https://zenodo.org/record/3735883/files/figure.png" httpUri-3="https://zenodo.org/record/3735885/files/figure.png" httpUri-4="https://zenodo.org/record/3735887/files/figure.png" httpUri-5="https://zenodo.org/record/3735889/files/figure.png" pageId="13" pageNumber="14">Fig. 11–16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3716574861BFFF8FF40FD3E55F8FCE5" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BD436FF861BFFF8FF40FD3E55F8FCE5" blockId="13.[151,1436,692,1007]" pageId="13" pageNumber="14">
<emphasis id="B91FEAED861BFFF8FF40FD3E5503FD11" bold="true" box="[151,377,692,719]" pageId="13" pageNumber="14">Material examined</emphasis>
: Type specimens. Adult
<specimenCount id="9D6DFD76861BFFF8FD59FD3F56D9FD11" box="[654,675,693,719]" pageId="13" pageNumber="14" type="male">♂</specimenCount>
<typeStatus id="54D0885D861BFFF8FD7EFD3E5771FD10" box="[681,779,692,718]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED7AAE3A861BFFF8FCCDFD3F5722FD11" box="[794,856,693,719]" pageId="13" pageNumber="14">QMS</collectionCode>
109020),
<specimenCount id="9D6DFD76861BFFF8FC1DFD3F506DFD10" box="[970,1047,692,719]" pageId="13" pageNumber="14" type="adult">2 adult</specimenCount>
<specimenCount id="9D6DFD76861BFFF8FBC9FD3F5049FD11" box="[1054,1075,693,719]" pageId="13" pageNumber="14" type="male">♂</specimenCount>
<typeStatus id="54D0885D861BFFF8FBEDFD3F50DFFD11" box="[1082,1189,693,719]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED7AAE3A861BFFF8FB63FD3F5088FD11" box="[1204,1266,693,719]" pageId="13" pageNumber="14">QMS</collectionCode>
109021–22),
<specimenCount id="9D6DFD76861BFFF8FA59FD3E54B5FD2C" pageId="13" pageNumber="14" type="adult">5 adult</specimenCount>
<specimenCount id="9D6DFD76861BFFF8FF01FD535491FD2D" box="[214,235,729,755]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
<typeStatus id="54D0885D861BFFF8FF25FD535527FD2D" box="[242,349,729,755]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED7AAE3A861BFFF8FEBCFD5355D3FD2D" box="[363,425,729,755]" pageId="13" pageNumber="14">QMS</collectionCode>
109023–27) from
<collectingCountry id="F37C766F861BFFF8FDACFD5256A7FD2C" box="[635,733,728,754]" name="Vietnam" pageId="13" pageNumber="14">Vietnam</collectingCountry>
,
<collectingRegion id="49AFF81D861BFFF8FD30FD53506CFD2C" box="[743,1046,728,755]" country="Vietnam" name="Ba Ria-Vung Tau" pageId="13" pageNumber="14">Ba Ria-Vung Tau Province</collectingRegion>
, Con Dao Islands, Con Son Island, leaf litter from So Ray old fruit plantation,
<geoCoordinate id="EE5F5038861BFFF8FD54FD765775FCC8" box="[643,783,764,790]" degrees="8.691263" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="8.691263">8.691263°N</geoCoordinate>
,
<geoCoordinate id="EE5F5038861BFFF8FCCCFD7657BAFCC9" box="[795,960,764,791]" degrees="106.588949" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="106.58895">106.588949°E</geoCoordinate>
, elevation
<quantity id="4C939B1A861BFFF8FBEDFD7750FCFCC9" box="[1082,1158,765,791]" metricMagnitude="2" metricUnit="m" metricValue="2.23" pageId="13" pageNumber="14" unit="m" value="223.0">223 m</quantity>
; on
<date id="FFD5103F861BFFF8FB61FD765125FCC9" box="[1206,1375,764,791]" pageId="13" pageNumber="14" value="2015-07-25">25th July 2015</date>
(collected by C. Huynh).
</paragraph>
</subSubSection>
<subSubSection id="C3716574861BFFF8FF10FCCE55A6FC30" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8BD436FF861BFFF8FF10FCCE55A6FC30" blockId="13.[151,1436,692,1007]" pageId="13" pageNumber="14">
<emphasis id="B91FEAED861BFFF8FF10FCCE5540FC81" bold="true" box="[199,314,836,863]" pageId="13" pageNumber="14">Diagnosis</emphasis>
: Differs from congeners in having dense, long trichomes. Body length over
<quantity id="4C939B1A861BFFF8FB48FCCE5080FC81" box="[1183,1274,836,863]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="14" unit="mm" value="3.5">3.5 mm</quantity>
. Antennomere VI with 24–29 sensilla forming a triangular shape, setiform sensillum located in the anterior position, conical sensillum absent. Labrum asetose smooth, with long slender setae in the anterior margin. Telotarsus with posterior lateral process shorter than half length of the claw, anterior lateral process and lamella process present, anterior setiform process longer than the claw.
</paragraph>
</subSubSection>
<caption id="DF146677861BFFF8FF40F93350FEF90C" ID-DOI="http://doi.org/10.5281/zenodo.3735877" ID-Zenodo-Dep="3735877" box="[151,1156,1721,1746]" httpUri="https://zenodo.org/record/3735877/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1721,1746]" targetBox="[205,1382,1028,1696]" targetPageId="13">
<paragraph id="8BD436FF861BFFF8FF40F93350FEF90C" blockId="13.[151,1156,1721,1746]" box="[151,1156,1721,1746]" pageId="13" pageNumber="14">
<emphasis id="B91FEAED861BFFF8FF40F9335564F90C" bold="true" box="[151,286,1721,1746]" pageId="13" pageNumber="14">FIGURE 11.</emphasis>
<taxonomicName id="4C6B4D7C861BFFF8FEF2F9305656F90C" authority="Huynh, 2020" authorityName="Huynh" authorityYear="2020" box="[293,556,1722,1746]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B91FEAED861BFFF8FEF2F9305656F90C" box="[293,556,1722,1746]" italics="true" pageId="13" pageNumber="14">Monographis condorensis</emphasis>
</taxonomicName>
<emphasis id="B91FEAED861BFFF8FDE4F9305617F90C" bold="true" box="[563,621,1722,1746]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A22C5796861BFFF8FDE4F9305617F90C" box="[563,621,1722,1746]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
showing the body cover with dense, long trichomes.
</paragraph>
</caption>
<subSubSection id="C3716574861BFFFBFF10F88A51F1F961" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="8BD436FF861BFFFBFF10F88A51F1F961" blockId="13.[151,1437,1792,2035]" lastBlockId="14.[151,1437,152,2015]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B91FEAED861BFFF8FF10F88A552BF8C5" bold="true" box="[199,337,1792,1819]" pageId="13" pageNumber="14">Description</emphasis>
: Colour and trichome patterns dark grey on both sides with contrasting light green medial band on dorsal surface of the body. Body covered with dense, long trichomes. Caudal bundle black-silver (
<figureCitation id="13502A7A861BFFF8FADEF8AE5121F8E0" box="[1289,1371,1828,1854]" captionStart="FIGURE 11" captionStartId="13.[151,250,1721,1746]" captionTargetBox="[205,1382,1028,1696]" captionTargetId="figure@13.[205,1382,1028,1696]" captionTargetPageId="13" captionText="FIGURE 11. Monographis condorensis n. sp. showing the body cover with dense, long trichomes." figureDoi="http://doi.org/10.5281/zenodo.3735877" httpUri="https://zenodo.org/record/3735877/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
).
<typeStatus id="54D0885D861BFFF8FAA6F8AF54A7F8BC" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
male body
<quantity id="4C939B1A861BFFF8FEB2F8C255C5F8BD" box="[357,447,1864,1891]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="14" unit="mm" value="3.5">3.5 mm</quantity>
;
<typeStatus id="54D0885D861BFFF8FE1BF8C35646F8BD" box="[460,572,1865,1891]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
: males
<quantity id="4C939B1A861BFFF8FD46F8C25761F8BD" box="[657,795,1864,1891]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="13" pageNumber="14" unit="mm" value="3.55" valueMax="3.6" valueMin="3.5">3.5–3.6 mm</quantity>
, females
<quantity id="4C939B1A861BFFF8FC50F8C25069F8BD" box="[903,1043,1864,1891]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.5" metricValueMin="3.8" pageId="13" pageNumber="14" unit="mm" value="4.15" valueMax="4.5" valueMin="3.8">3.8–4.5 mm</quantity>
; caudal bundle in both sexes same length (
<quantity id="4C939B1A861BFFF8FF39F8E65536F859" box="[238,332,1900,1927]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="14" unit="mm" value="0.9">0.9 mm</quantity>
).
<emphasis id="B91FEAED861BFFF8FEB6F8E755DBF859" bold="true" box="[353,417,1901,1927]" pageId="13" pageNumber="14">Head</emphasis>
: Each side 8 ommatidia: 4 dorsal, 4 lateral (1 anterior, 2 medial and 1 posterior). Vertex with 2 posterior trichome groups and a large medial gap. Each group with 2 rows: Anterior row curved slightly, on an oblique angle with large trichome sockets in central position, gradually reducing the size at both ends; posterior row with large sockets; a narrow medial space between these rows.
<typeStatus id="54D0885D861BFFF8FC72F8525077F82C" box="[933,1037,2008,2034]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
posterior trichome groups with 18 sockets (L) and 19 (R) sockets in anterior rows, posterior rows with 5 sockets both sides (
<figureCitation id="13502A7A8618FFFBFB7BFF13516EFF6D" box="[1196,1300,153,179]" captionStart="FIGURE 12" captionStartId="15.[151,250,1825,1850]" captionTargetBox="[186,1412,391,1786]" captionTargetId="figure@15.[175,1412,381,1800]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis condorensis n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. T2: Tergite 2; D. T10: Tergite 10—the last tergite; E. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; F. Gnathochilarium: the lateral palp (LP) with 11 sensilla and medial palp (MP) with 22 sensilla; G. Clypeo-labrum showing a smooth surface, asetose, with setae along both anterior edges. H. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d)." figureDoi="http://doi.org/10.5281/zenodo.3735879" httpUri="https://zenodo.org/record/3735879/files/figure.png" pageId="14" pageNumber="15">Fig. 12A</figureCitation>
) (
<typeStatus id="54D0885D8618FFFBFAFAFF1351E6FF6D" box="[1325,1436,153,179]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
showed that variation is common in this species, regardless of sex, ranging from 14–22 sockets in anterior rows,
<quantity id="4C939B1A8618FFFBFF40FF6B54A5FF25" box="[151,223,225,251]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.524" metricValueMin="0.508" pageId="14" pageNumber="15" unit="in" value="4.0" valueMax="6.0" valueMin="2.0">2–6 in</quantity>
posterior rows).
<emphasis id="B91FEAED8618FFFBFE75FF6A5633FF25" bold="true" box="[418,585,224,251]" pageId="14" pageNumber="15">Trichobothria</emphasis>
typically thin, sensory hairs with narrow cylindrical funicles; trichobothria equal in socket size forming an isosceles triangle with equal distance
<emphasis id="B91FEAED8618FFFBFC63FE8E57AAFEC1" bold="true" box="[948,976,260,287]" italics="true" pageId="14" pageNumber="15">ab</emphasis>
and
<emphasis id="B91FEAED8618FFFBFBDDFE8E505EFEC1" bold="true" box="[1034,1060,260,287]" italics="true" pageId="14" pageNumber="15">bc</emphasis>
(
<figureCitation id="13502A7A8618FFFBFBE2FE8F50E1FEC1" box="[1077,1179,261,287]" captionStart="FIGURE 12" captionStartId="15.[151,250,1825,1850]" captionTargetBox="[186,1412,391,1786]" captionTargetId="figure@15.[175,1412,381,1800]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis condorensis n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. T2: Tergite 2; D. T10: Tergite 10—the last tergite; E. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; F. Gnathochilarium: the lateral palp (LP) with 11 sensilla and medial palp (MP) with 22 sensilla; G. Clypeo-labrum showing a smooth surface, asetose, with setae along both anterior edges. H. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d)." figureDoi="http://doi.org/10.5281/zenodo.3735879" httpUri="https://zenodo.org/record/3735879/files/figure.png" pageId="14" pageNumber="15">Figs.12E</figureCitation>
and
<figureCitation id="13502A7A8618FFFBFB03FE8F517FFEC1" box="[1236,1285,261,287]" captionStart="FIGURE 13" captionStartId="16.[151,250,1754,1779]" captionTargetBox="[206,1383,181,1728]" captionTargetId="figure@16.[205,1383,181,1728]" captionTargetPageId="16" captionText="FIGURE 13. SEM images of Monographis condorensis n. sp., A. Dorsal view of the head capsule, showing 8 ommatidia and trichobothria a, b and c; B. Dorsal view of the body, co: Collum, Lp: lateral protuberances and pp: pleural projection, T2: tergite 2 and T3: tergite 3; C. Clypeo-labrum showing the smooth surface and asetose (l), s: setae along anterior edges, Se: posterior setae, bse: a minute backward-pointing setae; D. Right gnathochilarium showing the lateral palp (LP) with 5 sensilla and 6 sensilla sockets, 22 sensilla on the medial palp (MP); E. Bare caudal bundle in the female showed 2 distinctive structures: caudal trichome sockets (cts) which include barbate trichome sockets (bts) in the middle of the bundle, forming a dorsal single bundle, nest trichome sockets (nts) forming 2 ventral bundles; these bundles are separated by the trichome socket-free tissue (tsft); T: telson; F. Leg articles and arrows indicate spines which are typical characters of Monographis." figureDoi="http://doi.org/10.5281/zenodo.3735883" httpUri="https://zenodo.org/record/3735883/files/figure.png" pageId="14" pageNumber="15">13A</figureCitation>
) (trichobothrium
<emphasis id="B91FEAED8618FFFBFF02FEA35499FE9D" bold="true" box="[213,227,297,323]" italics="true" pageId="14" pageNumber="15">a</emphasis>
located in posterior position of the head capsule, trichobothrium
<emphasis id="B91FEAED8618FFFBFC05FEA2579AFE9D" bold="true" box="[978,992,296,323]" italics="true" pageId="14" pageNumber="15">b</emphasis>
in lateral position, trichobothrium
<emphasis id="B91FEAED8618FFFBFAA6FEA35107FE9D" bold="true" box="[1393,1405,297,323]" italics="true" pageId="14" pageNumber="15">c</emphasis>
in anterior position).
<emphasis id="B91FEAED8618FFFBFEBFFEC655A1FEB9" bold="true" box="[360,475,332,359]" pageId="14" pageNumber="15">Antennae</emphasis>
: Eight antennomeres, 4 sensory cones, typical characteristics of
<taxonomicName id="4C6B4D7C8618FFFBFB60FEC7513FFEB9" authorityName="Lucas" authorityYear="1840" box="[1207,1349,333,359]" class="Diplopoda" family="Polyxenidae" kingdom="Animalia" order="Polyxenida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="family">Polyxenidae</taxonomicName>
. Antennomere VI with 24–29 bacilliform sensilla (
<figureCitation id="13502A7A8618FFFBFD5EFEFB5710FE55" box="[649,874,369,395]" captionStart="FIGURE 14" captionStartId="17.[151,250,1373,1398]" captionTargetBox="[151,1435,241,1348]" captionTargetId="figure@17.[151,1436,237,1348]" captionTargetPageId="17" captionText="FIGURE 14. Monographis condorensis n. sp., A. SEM images of antennomeres VI, VII and VIII with 4 sensory cones (sc), showing the arrangement of sensilla on antennomeres VI and VII; B. Antennomere VII with 2 thick sensilla (T), setiform sensillum (s) between them and a conical sensillum (c) located in posterior position; C. Antennomere VI with a group of 29 thin sensilla forming a triangular shape followed by a setiform sensillum." figureDoi="http://doi.org/10.5281/zenodo.3735885" httpUri="https://zenodo.org/record/3735885/files/figure.png" pageId="14" pageNumber="15">Figs. 14A, B and C</figureCitation>
).
<typeStatus id="54D0885D8618FFFBFCA8FEFB579DFE55" box="[895,999,369,395]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
with 26 bacilliform sensilla forming a triangular shape (3 thick and 23 thin bacilliform sensilla), setiform sensillum located in anterior position, conical sensillum absent. Antennomere VII with 2 thick bacilliform sensilla (T, Ta and Tp), setiform sensillum (s) located in between these two sensilla (T), conical sensillum (c) located next to Tp (
<figureCitation id="13502A7A8618FFFBFBD2FE5750BDFE29" box="[1029,1223,476,503]" captionStart="FIGURE 15" captionStartId="18.[151,250,1890,1915]" captionTargetBox="[177,1408,351,1859]" captionTargetId="figure@18.[168,1420,345,1865]" captionTargetPageId="18" captionText="FIGURE 15. Holotype Monographis condorensis n. sp., A. Antennomeres VI, VII and VIII with sensory cones (sc), showing the arrangement of sensilla on antennomeres VI and VII; B. Antennomere VII with 2 thick sensilla (Ta (anterior) and Tp (posterior)), setiform sensillum (s) between them and a conical sensillum (c) located next to Tp; antennomere VI with a group of 26 sensilla (23 thin sensilla (t) and 3 thick sensilla (★)) forming a triangular shape followed by a setiform sensillum (s)." figureDoi="http://doi.org/10.5281/zenodo.3735887" httpUri="https://zenodo.org/record/3735887/files/figure.png" pageId="14" pageNumber="15">Figs. 15A and B</figureCitation>
).
<emphasis id="B91FEAED8618FFFBFB09FE5651EEFE29" bold="true" box="[1246,1428,476,503]" pageId="14" pageNumber="15">Clypeo-labrum</emphasis>
:
<typeStatus id="54D0885D8618FFFBFF40FD8B5578FDC5" box="[151,258,513,539]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
, labrum posterior margin with 10 setae, setae equal in length to labrum width (Setae on the
<typeStatus id="54D0885D8618FFFBFAE6FD8B51E6FDC5" box="[1329,1436,513,539]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
ranged from
<quantity id="4C939B1A8618FFFBFEF8FDAF55EFFDE1" box="[303,405,549,575]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="14" pageNumber="15" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">10–11 in</quantity>
both sexes). Labrum surface smooth and asetose, labrum posterior margin has a row of minute backward-pointing setae. Lateral lamella and single broad lamella each side median cleft, 20 slender setae, pair at the base, present on anterior margin of each side of labrum (
<figureCitation id="13502A7A8618FFFBFC19FDE7504CFD59" box="[974,1078,621,647]" captionStart="FIGURE 12" captionStartId="15.[151,250,1825,1850]" captionTargetBox="[186,1412,391,1786]" captionTargetId="figure@15.[175,1412,381,1800]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis condorensis n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. T2: Tergite 2; D. T10: Tergite 10—the last tergite; E. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; F. Gnathochilarium: the lateral palp (LP) with 11 sensilla and medial palp (MP) with 22 sensilla; G. Clypeo-labrum showing a smooth surface, asetose, with setae along both anterior edges. H. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d)." figureDoi="http://doi.org/10.5281/zenodo.3735879" httpUri="https://zenodo.org/record/3735879/files/figure.png" pageId="14" pageNumber="15">Fig. 12G</figureCitation>
) (The number of these anterior slender setae varies among
<typeStatus id="54D0885D8618FFFBFE00FD1B563FFD75" box="[471,581,657,683]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
, ranging from 18–24 on each side. (
<figureCitation id="13502A7A8618FFFBFC3AFD1B5031FD72" box="[1005,1099,657,684]" captionStart="FIGURE 13" captionStartId="16.[151,250,1754,1779]" captionTargetBox="[206,1383,181,1728]" captionTargetId="figure@16.[205,1383,181,1728]" captionTargetPageId="16" captionText="FIGURE 13. SEM images of Monographis condorensis n. sp., A. Dorsal view of the head capsule, showing 8 ommatidia and trichobothria a, b and c; B. Dorsal view of the body, co: Collum, Lp: lateral protuberances and pp: pleural projection, T2: tergite 2 and T3: tergite 3; C. Clypeo-labrum showing the smooth surface and asetose (l), s: setae along anterior edges, Se: posterior setae, bse: a minute backward-pointing setae; D. Right gnathochilarium showing the lateral palp (LP) with 5 sensilla and 6 sensilla sockets, 22 sensilla on the medial palp (MP); E. Bare caudal bundle in the female showed 2 distinctive structures: caudal trichome sockets (cts) which include barbate trichome sockets (bts) in the middle of the bundle, forming a dorsal single bundle, nest trichome sockets (nts) forming 2 ventral bundles; these bundles are separated by the trichome socket-free tissue (tsft); T: telson; F. Leg articles and arrows indicate spines which are typical characters of Monographis." figureDoi="http://doi.org/10.5281/zenodo.3735883" httpUri="https://zenodo.org/record/3735883/files/figure.png" pageId="14" pageNumber="15">Fig.13C</figureCitation>
)).
<emphasis id="B91FEAED8618FFFBFBBBFD1A5141FD75" bold="true" box="[1132,1339,656,683]" pageId="14" pageNumber="15">Gnathochilarium</emphasis>
: Lateral palp 1.25 times medial palp; 11 conical sensilla on the lateral palp and 22 on the medial palp. Same in
<typeStatus id="54D0885D8618FFFBFAEDFD3F51E6FD11" box="[1338,1436,693,719]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
and
<typeStatus id="54D0885D8618FFFBFF1EFD53554EFD2D" box="[201,308,729,755]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
(
<figureCitation id="13502A7A8618FFFBFE91FD5355CEFD2D" box="[326,436,729,755]" captionStart="FIGURE 12" captionStartId="15.[151,250,1825,1850]" captionTargetBox="[186,1412,391,1786]" captionTargetId="figure@15.[175,1412,381,1800]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis condorensis n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. T2: Tergite 2; D. T10: Tergite 10—the last tergite; E. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; F. Gnathochilarium: the lateral palp (LP) with 11 sensilla and medial palp (MP) with 22 sensilla; G. Clypeo-labrum showing a smooth surface, asetose, with setae along both anterior edges. H. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d)." figureDoi="http://doi.org/10.5281/zenodo.3735879" httpUri="https://zenodo.org/record/3735879/files/figure.png" pageId="14" pageNumber="15">Figs. 12F</figureCitation>
and
<figureCitation id="13502A7A8618FFFBFE38FD535658FD2D" box="[495,546,729,755]" captionStart="FIGURE 13" captionStartId="16.[151,250,1754,1779]" captionTargetBox="[206,1383,181,1728]" captionTargetId="figure@16.[205,1383,181,1728]" captionTargetPageId="16" captionText="FIGURE 13. SEM images of Monographis condorensis n. sp., A. Dorsal view of the head capsule, showing 8 ommatidia and trichobothria a, b and c; B. Dorsal view of the body, co: Collum, Lp: lateral protuberances and pp: pleural projection, T2: tergite 2 and T3: tergite 3; C. Clypeo-labrum showing the smooth surface and asetose (l), s: setae along anterior edges, Se: posterior setae, bse: a minute backward-pointing setae; D. Right gnathochilarium showing the lateral palp (LP) with 5 sensilla and 6 sensilla sockets, 22 sensilla on the medial palp (MP); E. Bare caudal bundle in the female showed 2 distinctive structures: caudal trichome sockets (cts) which include barbate trichome sockets (bts) in the middle of the bundle, forming a dorsal single bundle, nest trichome sockets (nts) forming 2 ventral bundles; these bundles are separated by the trichome socket-free tissue (tsft); T: telson; F. Leg articles and arrows indicate spines which are typical characters of Monographis." figureDoi="http://doi.org/10.5281/zenodo.3735883" httpUri="https://zenodo.org/record/3735883/files/figure.png" pageId="14" pageNumber="15">13D</figureCitation>
).
<emphasis id="B91FEAED8618FFFBFDEFFD5356FEFD2D" bold="true" box="[568,644,729,755]" pageId="14" pageNumber="15">Trunk</emphasis>
: Body with 10 tergites, 9 pleural projections, and a telson excluding caudal bundle. 13 pairs of legs. Collum (tergite 1) with lateral protuberances 4–6 trichome sockets each side. Other tergites with pair of pleural projections located in anterolateral positions. The tergal trichome socket arrangements typically have 2 broad oval shapes, slightly enlarged in lateral position, with the posterior row straight slightly to the centre, an intermedial gap between these rows (
<figureCitation id="13502A7A8618FFFBFC96FCE357D4FC5D" box="[833,942,873,899]" captionStart="FIGURE 12" captionStartId="15.[151,250,1825,1850]" captionTargetBox="[186,1412,391,1786]" captionTargetId="figure@15.[175,1412,381,1800]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis condorensis n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. T2: Tergite 2; D. T10: Tergite 10—the last tergite; E. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; F. Gnathochilarium: the lateral palp (LP) with 11 sensilla and medial palp (MP) with 22 sensilla; G. Clypeo-labrum showing a smooth surface, asetose, with setae along both anterior edges. H. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d)." figureDoi="http://doi.org/10.5281/zenodo.3735879" httpUri="https://zenodo.org/record/3735879/files/figure.png" pageId="14" pageNumber="15">Figs. 12C</figureCitation>
and
<figureCitation id="13502A7A8618FFFBFC34FCE3506FFC5D" box="[995,1045,873,899]" captionStart="FIGURE 13" captionStartId="16.[151,250,1754,1779]" captionTargetBox="[206,1383,181,1728]" captionTargetId="figure@16.[205,1383,181,1728]" captionTargetPageId="16" captionText="FIGURE 13. SEM images of Monographis condorensis n. sp., A. Dorsal view of the head capsule, showing 8 ommatidia and trichobothria a, b and c; B. Dorsal view of the body, co: Collum, Lp: lateral protuberances and pp: pleural projection, T2: tergite 2 and T3: tergite 3; C. Clypeo-labrum showing the smooth surface and asetose (l), s: setae along anterior edges, Se: posterior setae, bse: a minute backward-pointing setae; D. Right gnathochilarium showing the lateral palp (LP) with 5 sensilla and 6 sensilla sockets, 22 sensilla on the medial palp (MP); E. Bare caudal bundle in the female showed 2 distinctive structures: caudal trichome sockets (cts) which include barbate trichome sockets (bts) in the middle of the bundle, forming a dorsal single bundle, nest trichome sockets (nts) forming 2 ventral bundles; these bundles are separated by the trichome socket-free tissue (tsft); T: telson; F. Leg articles and arrows indicate spines which are typical characters of Monographis." figureDoi="http://doi.org/10.5281/zenodo.3735883" httpUri="https://zenodo.org/record/3735883/files/figure.png" pageId="14" pageNumber="15">13B</figureCitation>
).
<typeStatus id="54D0885D8618FFFBFBF0FCE350E8FC5D" box="[1063,1170,873,899]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
, collum with 73 sockets each side, lateral protuberance 5 (L) and 4 (R) sockets (
<figureCitation id="13502A7A8618FFFBFCCFFC07570FFC76" box="[792,885,909,936]" captionStart="FIGURE 12" captionStartId="15.[151,250,1825,1850]" captionTargetBox="[186,1412,391,1786]" captionTargetId="figure@15.[175,1412,381,1800]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis condorensis n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. T2: Tergite 2; D. T10: Tergite 10—the last tergite; E. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; F. Gnathochilarium: the lateral palp (LP) with 11 sensilla and medial palp (MP) with 22 sensilla; G. Clypeo-labrum showing a smooth surface, asetose, with setae along both anterior edges. H. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d)." figureDoi="http://doi.org/10.5281/zenodo.3735879" httpUri="https://zenodo.org/record/3735879/files/figure.png" pageId="14" pageNumber="15">Fig.12B</figureCitation>
) (
<typeStatus id="54D0885D8618FFFBFC59FC075787FC79" box="[910,1021,909,935]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
ranged 65–78 sockets in the collum and lateral protuberances 4–6 sockets). Tergite 2 had a similar structural pattern with the posterior row slightly longer and 73 (L) and 71 (R) trichome sockets (
<figureCitation id="13502A7A8618FFFBFD06FC5F574AFC2E" box="[721,816,981,1008]" captionStart="FIGURE 12" captionStartId="15.[151,250,1825,1850]" captionTargetBox="[186,1412,391,1786]" captionTargetId="figure@15.[175,1412,381,1800]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis condorensis n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. T2: Tergite 2; D. T10: Tergite 10—the last tergite; E. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; F. Gnathochilarium: the lateral palp (LP) with 11 sensilla and medial palp (MP) with 22 sensilla; G. Clypeo-labrum showing a smooth surface, asetose, with setae along both anterior edges. H. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d)." figureDoi="http://doi.org/10.5281/zenodo.3735879" httpUri="https://zenodo.org/record/3735879/files/figure.png" pageId="14" pageNumber="15">Fig.12C</figureCitation>
), the patterns are similar to characteristic large gaps. Tergite 10 is the exception with these trichome sockets being smaller and denser, a narrow space exists between the lateral rosette trichome sockets and posterior row. Trichome sockets of this tergite are over 100 and the tergite 10 of the
<typeStatus id="54D0885D8618FFFBFED5FBCB551EFB85" box="[258,356,1089,1115]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
was counted 115 (L) and 112 (R) sockets (
<figureCitation id="13502A7A8618FFFBFC91FBCB57D2FB82" box="[838,936,1089,1116]" captionStart="FIGURE 12" captionStartId="15.[151,250,1825,1850]" captionTargetBox="[186,1412,391,1786]" captionTargetId="figure@15.[175,1412,381,1800]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis condorensis n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. T2: Tergite 2; D. T10: Tergite 10—the last tergite; E. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; F. Gnathochilarium: the lateral palp (LP) with 11 sensilla and medial palp (MP) with 22 sensilla; G. Clypeo-labrum showing a smooth surface, asetose, with setae along both anterior edges. H. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d)." figureDoi="http://doi.org/10.5281/zenodo.3735879" httpUri="https://zenodo.org/record/3735879/files/figure.png" pageId="14" pageNumber="15">Fig.12D</figureCitation>
).
<emphasis id="B91FEAED8618FFFBFC6CFBCB5789FB85" bold="true" box="[955,1011,1089,1115]" pageId="14" pageNumber="15">Legs</emphasis>
: Leg segmentation following
<bibRefCitation id="EFFA4B0E8618FFFBFA92FBCB549FFBA1" author="Manton, S. M." pageId="14" pageNumber="15" pagination="153 - 187" refId="ref15174" refString="Manton, S. M. (1956) The Evolution of Arthropodan Locomotory Mechanisms-Part 5: The Structure, Habits and Evolution of the Pselaphognatha (Diplopoda). Journal of the Linnean Society London, 43, 153 - 187. https: // doi. org / 10.1111 / j. 1096 - 3642.1957. tb 02516. x" type="journal article" year="1956">Manton (1956)</bibRefCitation>
. Legs 1 and 2 without trochanter, leg 1 without tarsus 1. Chaetotaxy (setae on leg articles): coxa 1 and 2: 2 setae, coxae 3–13: 2–4 setae; pre-femur, post-femur, tibia with 1 seta, except femur with 1–4 setae, tarsus 1 and 2 with a spine (
<figureCitation id="13502A7A8618FFFBFEFDFB2755EBFB16" box="[298,401,1197,1224]" captionStart="FIGURE 16" captionStartId="19.[151,250,1225,1250]" captionTargetBox="[171,1410,186,1199]" captionTargetId="figure@19.[171,1416,181,1199]" captionTargetPageId="19" captionText="FIGURE 16. Holotype Monographis condorensis n. sp., A. 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta present in the middle of the femur; D. Seta on tibia; E. Smaller spine on tarsus 1; F. Larger spine on T2; G. Telotarsus—claw structures (SEM image of one specimen) and a drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi="http://doi.org/10.5281/zenodo.3735889" httpUri="https://zenodo.org/record/3735889/files/figure.png" pageId="14" pageNumber="15">Fig. 16A</figureCitation>
). Coxa, pre-femur and femur with biarticulated seta and ridged funicle (
<figureCitation id="13502A7A8618FFFBFB7EFB275172FB16" box="[1193,1288,1197,1224]" captionStart="FIGURE 16" captionStartId="19.[151,250,1225,1250]" captionTargetBox="[171,1410,186,1199]" captionTargetId="figure@19.[171,1416,181,1199]" captionTargetPageId="19" captionText="FIGURE 16. Holotype Monographis condorensis n. sp., A. 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta present in the middle of the femur; D. Seta on tibia; E. Smaller spine on tarsus 1; F. Larger spine on T2; G. Telotarsus—claw structures (SEM image of one specimen) and a drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi="http://doi.org/10.5281/zenodo.3735889" httpUri="https://zenodo.org/record/3735889/files/figure.png" pageId="14" pageNumber="15">Fig.16B</figureCitation>
). 1–3 smaller biarticulated setae in posterior position of femur, some also on penis (
<figureCitation id="13502A7A8618FFFBFC7FFB5B506AFB32" box="[936,1040,1233,1260]" captionStart="FIGURE 16" captionStartId="19.[151,250,1225,1250]" captionTargetBox="[171,1410,186,1199]" captionTargetId="figure@19.[171,1416,181,1199]" captionTargetPageId="19" captionText="FIGURE 16. Holotype Monographis condorensis n. sp., A. 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta present in the middle of the femur; D. Seta on tibia; E. Smaller spine on tarsus 1; F. Larger spine on T2; G. Telotarsus—claw structures (SEM image of one specimen) and a drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi="http://doi.org/10.5281/zenodo.3735889" httpUri="https://zenodo.org/record/3735889/files/figure.png" pageId="14" pageNumber="15">Fig. 16C</figureCitation>
). Post-femur and tibia distally with setiform seta (
<figureCitation id="13502A7A8618FFFBFEEDFB7F55DEFACE" box="[314,420,1269,1296]" captionStart="FIGURE 16" captionStartId="19.[151,250,1225,1250]" captionTargetBox="[171,1410,186,1199]" captionTargetId="figure@19.[171,1416,181,1199]" captionTargetPageId="19" captionText="FIGURE 16. Holotype Monographis condorensis n. sp., A. 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta present in the middle of the femur; D. Seta on tibia; E. Smaller spine on tarsus 1; F. Larger spine on T2; G. Telotarsus—claw structures (SEM image of one specimen) and a drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi="http://doi.org/10.5281/zenodo.3735889" httpUri="https://zenodo.org/record/3735889/files/figure.png" pageId="14" pageNumber="15">Fig. 16D</figureCitation>
). The spine on tarsus 2 is sharply pointed and twice as long as (
<figureCitation id="13502A7A8618FFFBFB55FB7F50A6FACE" box="[1154,1244,1269,1296]" captionStart="FIGURE 16" captionStartId="19.[151,250,1225,1250]" captionTargetBox="[171,1410,186,1199]" captionTargetId="figure@19.[171,1416,181,1199]" captionTargetPageId="19" captionText="FIGURE 16. Holotype Monographis condorensis n. sp., A. 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta present in the middle of the femur; D. Seta on tibia; E. Smaller spine on tarsus 1; F. Larger spine on T2; G. Telotarsus—claw structures (SEM image of one specimen) and a drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi="http://doi.org/10.5281/zenodo.3735889" httpUri="https://zenodo.org/record/3735889/files/figure.png" pageId="14" pageNumber="15">Fig.16F</figureCitation>
) the spine in the antero-sternal position of tarsus 1 (
<figureCitation id="13502A7A8618FFFBFDCDFA935605FAEA" box="[538,639,1305,1332]" captionStart="FIGURE 16" captionStartId="19.[151,250,1225,1250]" captionTargetBox="[171,1410,186,1199]" captionTargetId="figure@19.[171,1416,181,1199]" captionTargetPageId="19" captionText="FIGURE 16. Holotype Monographis condorensis n. sp., A. 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta present in the middle of the femur; D. Seta on tibia; E. Smaller spine on tarsus 1; F. Larger spine on T2; G. Telotarsus—claw structures (SEM image of one specimen) and a drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi="http://doi.org/10.5281/zenodo.3735889" httpUri="https://zenodo.org/record/3735889/files/figure.png" pageId="14" pageNumber="15">Fig. 16E</figureCitation>
). There 4 spines in tarsus 1 from the
<typeStatus id="54D0885D8618FFFBFBC0FA935007FAED" box="[1047,1149,1305,1331]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
. The posterior edge of the last sternite had 4 setae similar to those present on the coxa and the number of these setae are the same in the
<typeStatus id="54D0885D8618FFFBFAB6FAB754BDFAA5" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
and
<typeStatus id="54D0885D8618FFFBFED7FAEB5515FAA5" box="[256,367,1377,1403]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
.
<emphasis id="B91FEAED8618FFFBFEADFAEA5647FAA5" bold="true" box="[378,573,1376,1403]" pageId="14" pageNumber="15">Telotarsus-Claw</emphasis>
: Claw slender with anterior lateral process and posterior lateral process less than half length of the claw; the anterior setiform process is longer than the claw; lamella process is present (
<figureCitation id="13502A7A8618FFFBFAB8FA0F54B1FA1A" captionStart="FIGURE 16" captionStartId="19.[151,250,1225,1250]" captionTargetBox="[171,1410,186,1199]" captionTargetId="figure@19.[171,1416,181,1199]" captionTargetPageId="19" captionText="FIGURE 16. Holotype Monographis condorensis n. sp., A. 2nd leg showing seven leg articles (co: coxa, pf: pre-femur, fe: femur, pof: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, c: claw) and sex organ—penis (P); B. Biarticulated seta present on the coxa, pre-femur, the distal edge of femur and the posterior edge of the last sternite; C. Similar smaller version of this seta present in the middle of the femur; D. Seta on tibia; E. Smaller spine on tarsus 1; F. Larger spine on T2; G. Telotarsus—claw structures (SEM image of one specimen) and a drawing of the telotarsus from the holotype (lateral view), p: posterior lateral process, a: anterior lateral process, c: claw, l: lamella process and s: anterior setiform process." figureDoi="http://doi.org/10.5281/zenodo.3735889" httpUri="https://zenodo.org/record/3735889/files/figure.png" pageId="14" pageNumber="15">Fig. 16G</figureCitation>
).
<emphasis id="B91FEAED8618FFFBFF37FA225588FA1D" bold="true" box="[224,498,1448,1475]" pageId="14" pageNumber="15">Sex organs in the male</emphasis>
: A pair of penes on coxae 2 and 2 pairs of coxal glands are located on the 8th–9th coxal plates.
<emphasis id="B91FEAED8618FFFBFEF0FA47550EFA39" bold="true" box="[295,372,1485,1511]" pageId="14" pageNumber="15">Telson</emphasis>
: Dorsal ornamental trichome sockets symmetrically arranged on both sides of telson; 25 (L) and 29 (R) sockets of trichomes
<emphasis id="B91FEAED8618FFFBFE06FA7B55A5F9D5" bold="true" box="[465,479,1521,1547]" italics="true" pageId="14" pageNumber="15">a</emphasis>
on each side of the telson in the
<typeStatus id="54D0885D8618FFFBFC9CFA7B57D7F9D5" box="[843,941,1521,1547]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(
<typeStatus id="54D0885D8618FFFBFC6CFA7B5052F9D5" box="[955,1064,1521,1547]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
with 14–29). Trichome
<emphasis id="B91FEAED8618FFFBFAE2FA7B5139F9D5" bold="true" box="[1333,1347,1521,1547]" italics="true" pageId="14" pageNumber="15">a</emphasis>
sockets form 2 rows, top row with small sockets, bottom row with larger sockets. Socket of trichome
<emphasis id="B91FEAED8618FFFBFB66F99E50C5F9F1" bold="true" box="[1201,1215,1556,1583]" italics="true" pageId="14" pageNumber="15">b</emphasis>
and 3 large sockets trichomes
<emphasis id="B91FEAED8618FFFBFED8F9B35561F98D" bold="true" box="[271,283,1593,1619]" italics="true" pageId="14" pageNumber="15">c</emphasis>
with protruding base:
<emphasis id="B91FEAED8618FFFBFDF7F9B35640F98D" bold="true" box="[544,570,1593,1619]" italics="true" pageId="14" pageNumber="15">c1</emphasis>
,
<emphasis id="B91FEAED8618FFFBFD9CF9B3561FF98D" bold="true" box="[587,613,1593,1619]" italics="true" pageId="14" pageNumber="15">c2</emphasis>
and
<emphasis id="B91FEAED8618FFFBFD77F9B356C0F98D" bold="true" box="[672,698,1593,1619]" italics="true" pageId="14" pageNumber="15">c3</emphasis>
, forming triangular shape located in ventroposterior positions in each arrangement. Circular indentation
<emphasis id="B91FEAED8618FFFBFD99F9D65626F9A9" bold="true" box="[590,604,1628,1655]" italics="true" pageId="14" pageNumber="15">d</emphasis>
apparent each side near the exterior side of trichomes
<emphasis id="B91FEAED8618FFFBFB66F9D750C7F9A9" bold="true" box="[1201,1213,1629,1655]" italics="true" pageId="14" pageNumber="15">c</emphasis>
(
<figureCitation id="13502A7A8618FFFBFB1BF9D75149F9A9" box="[1228,1331,1629,1655]" captionStart="FIGURE 12" captionStartId="15.[151,250,1825,1850]" captionTargetBox="[186,1412,391,1786]" captionTargetId="figure@15.[175,1412,381,1800]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis condorensis n. sp., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria a, b and c); B. Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets; C. T2: Tergite 2; D. T10: Tergite 10—the last tergite; E. Trichobothria a, b and c (right hand side) showing the sensory hairs and sockets; F. Gnathochilarium: the lateral palp (LP) with 11 sensilla and medial palp (MP) with 22 sensilla; G. Clypeo-labrum showing a smooth surface, asetose, with setae along both anterior edges. H. Dorsal ornamental trichome sockets of trichome a (a), trichome b (b), trichome c (c) and the circular indentation d (d)." figureDoi="http://doi.org/10.5281/zenodo.3735879" httpUri="https://zenodo.org/record/3735879/files/figure.png" pageId="14" pageNumber="15">Fig. 12H</figureCitation>
).
<emphasis id="B91FEAED8618FFFBFA92F9D6548EF945" bold="true" pageId="14" pageNumber="15">Caudal bundles</emphasis>
: Similar to congener (
<bibRefCitation id="EFFA4B0E8618FFFBFE3DF90B5680F945" author="Huynh, C. &amp; Veenstra, A." box="[490,762,1664,1691]" pageId="14" pageNumber="15" pagination="460 - 474" refId="ref14596" refString="Huynh, C. &amp; Veenstra, A. (2015) Description of a new species of Penicillate Millipede from the genus Monographis (Diplopoda: Polyxenidae) found in Vietnam. Zootaxa, 3964 (4), 460 - 474. https: // doi. org / 10.11646 / zootaxa. 3964.4.5" type="journal article" year="2015">Huynh &amp; Veenstra 2015</bibRefCitation>
,
<bibRefCitation id="EFFA4B0E8618FFFBFCD3F90B5730F945" author="Huynh, C. &amp; Veenstra, A." box="[772,842,1665,1691]" pageId="14" pageNumber="15" pagination="283 - 302" refId="ref14755" refString="Huynh, C. &amp; Veenstra, A. (2018 c) Two new species pf penicillate millipedes (Diplopoda, Polyxenidae) from Phu Quoc Island in southern Vietnam. Zootaxa, 4402 (2), 283 - 302. https: // doi. org / 10.11646 / zootaxa. 4402.2.3" type="journal article" year="2018">2018c</bibRefCitation>
) (
<figureCitation id="13502A7A8618FFFBFCB7F90B57BBF945" box="[864,961,1665,1691]" captionStart="FIGURE 13" captionStartId="16.[151,250,1754,1779]" captionTargetBox="[206,1383,181,1728]" captionTargetId="figure@16.[205,1383,181,1728]" captionTargetPageId="16" captionText="FIGURE 13. SEM images of Monographis condorensis n. sp., A. Dorsal view of the head capsule, showing 8 ommatidia and trichobothria a, b and c; B. Dorsal view of the body, co: Collum, Lp: lateral protuberances and pp: pleural projection, T2: tergite 2 and T3: tergite 3; C. Clypeo-labrum showing the smooth surface and asetose (l), s: setae along anterior edges, Se: posterior setae, bse: a minute backward-pointing setae; D. Right gnathochilarium showing the lateral palp (LP) with 5 sensilla and 6 sensilla sockets, 22 sensilla on the medial palp (MP); E. Bare caudal bundle in the female showed 2 distinctive structures: caudal trichome sockets (cts) which include barbate trichome sockets (bts) in the middle of the bundle, forming a dorsal single bundle, nest trichome sockets (nts) forming 2 ventral bundles; these bundles are separated by the trichome socket-free tissue (tsft); T: telson; F. Leg articles and arrows indicate spines which are typical characters of Monographis." figureDoi="http://doi.org/10.5281/zenodo.3735883" httpUri="https://zenodo.org/record/3735883/files/figure.png" pageId="14" pageNumber="15">Fig. 13E</figureCitation>
) and is classified as the caudal bundle type I arrangement (
<bibRefCitation id="EFFA4B0E8618FFFBFE95F92F5777F961" author="Conde, B. &amp; Nguyen Duy-Jacquemin, M." box="[322,781,1701,1727]" pageId="14" pageNumber="15" pagination="291 - 302" refId="ref14429" refString="Conde, B. &amp; Nguyen Duy-Jacquemin, M. (2008) Classification actuelle des Diplopodes Penicillates (Myriapodes) avec nouvelles d e finition des taxa. Bulletin de la Societe Zoologique de France, 133 (4), 291 - 302." type="journal article" year="2008">Condé &amp; Nguyen Duy-Jacquemin, 2008</bibRefCitation>
). Caudal trichomes with a series of 2–6 backward hooks.
</paragraph>
</subSubSection>
<subSubSection id="C37165748618FFFAFF10F94357A3FEB9" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BD436FF8618FFFAFF10F94357A3FEB9" blockId="14.[151,1437,152,2015]" lastBlockId="15.[151,1436,153,359]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B91FEAED8618FFFBFF10F943554EF93D" bold="true" box="[199,308,1737,1763]" pageId="14" pageNumber="15">Remarks</emphasis>
:
<taxonomicName id="4C6B4D7C8618FFFBFE93F9435610F93D" authority="Huynh, 2020" authorityName="Huynh" authorityYear="2020" box="[324,618,1737,1763]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B91FEAED8618FFFBFE93F9435610F93D" box="[324,618,1737,1763]" italics="true" pageId="14" pageNumber="15">Monographis condorensis</emphasis>
</taxonomicName>
<emphasis id="B91FEAED8618FFFBFDA5F94356C8F93D" bold="true" box="[626,690,1737,1763]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A22C57968618FFFBFDA5F94356C8F93D" box="[626,690,1737,1763]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has a longer body, with dense, long trichomes that are dark grey, which differs from the yellowish-brown colour seen in live specimens of other
<taxonomicName id="4C6B4D7C8618FFFBFC29F96750EFF8D9" authorityName="Attems" authorityYear="1907" box="[1022,1173,1773,1799]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAED8618FFFBFC29F96750EFF8D9" box="[1022,1173,1773,1799]" italics="true" pageId="14" pageNumber="15">Monographis</emphasis>
</taxonomicName>
species. The number of sensilla (24–29) on the antennomere VI also differs from congeners that have fewer sensilla (12–20 (Nguyen Duy- Jacquemin and Condé, 1967)) forming a triangular shape. A row of the slender setae present on lamellae along the anterior margin of the labrum. Morphological and genetic differences indicate that
<taxonomicName id="4C6B4D7C8618FFFBFBE5F8D050A7F8AD" authorityName="Huynh" authorityYear="2020" box="[1074,1245,1881,1907]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="condorensis">
<emphasis id="B91FEAED8618FFFBFBE5F8D050A7F8AD" box="[1074,1245,1881,1907]" italics="true" pageId="14" pageNumber="15">M. condorensis</emphasis>
</taxonomicName>
is a new species. An updated diagnosis of genus
<taxonomicName id="4C6B4D7C8618FFFBFE23F8F756F1F849" authorityName="Attems" authorityYear="1907" box="[500,651,1917,1943]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAED8618FFFBFE23F8F756F1F849" box="[500,651,1917,1943]" italics="true" pageId="14" pageNumber="15">Monographis</emphasis>
</taxonomicName>
is also required based on the morphology of all described species. The arrangement of the sensilla on antennomere VI divides
<taxonomicName id="4C6B4D7C8618FFFBFD2AF82B57EEF865" authorityName="Attems" authorityYear="1907" box="[765,916,1953,1979]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAED8618FFFBFD2AF82B57EEF865" box="[765,916,1953,1979]" italics="true" pageId="14" pageNumber="15">Monographis</emphasis>
</taxonomicName>
into 2 groups: 1) forming a crescent shape and 2) triangular shape (
<bibRefCitation id="EFFA4B0E8618FFFBFEACF84F56DDF801" author="Huynh, C. &amp; Veenstra, A." box="[379,679,1989,2015]" pageId="14" pageNumber="15" pagination="283 - 302" refId="ref14755" refString="Huynh, C. &amp; Veenstra, A. (2018 c) Two new species pf penicillate millipedes (Diplopoda, Polyxenidae) from Phu Quoc Island in southern Vietnam. Zootaxa, 4402 (2), 283 - 302. https: // doi. org / 10.11646 / zootaxa. 4402.2.3" type="journal article" year="2018">Huynh &amp; Veenstra, 2018c</bibRefCitation>
).
<taxonomicName id="4C6B4D7C8618FFFBFD6CF84F579AF801" authority="Huynh, 2020" authorityName="Huynh" authorityYear="2020" box="[699,992,1989,2015]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B91FEAED8618FFFBFD6CF84F579AF801" box="[699,992,1989,2015]" italics="true" pageId="14" pageNumber="15">Monographis condorensis</emphasis>
</taxonomicName>
<emphasis id="B91FEAED8618FFFBFC3EF84F5053F801" bold="true" box="[1001,1065,1989,2015]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A22C57968618FFFBFC3EF84F5053F801" box="[1001,1065,1989,2015]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
belongs to group 2 with sensilla arranged in a triangular shape, which is found in
<emphasis id="B91FEAED8619FFFAFD17FF105177FF6D" box="[704,1293,153,179]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C6B4D7C8619FFFAFD17FF105706FF6D" authorityName="Huynh and Veenstra" authorityYear="2015" box="[704,892,153,179]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dongnaiensis">M. dongnaiensis</taxonomicName>
,
<taxonomicName id="4C6B4D7C8619FFFAFC5FFF105028FF6D" authorityName="Huynh and Veenstra" authorityYear="2013" box="[904,1106,153,179]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="queenslandica">M. queenslandica</taxonomicName>
,
<taxonomicName id="4C6B4D7C8619FFFAFB8AFF105177FF6D" authorityName="Conde and Nguyen Duy-Jacquemin" authorityYear="1963" box="[1117,1293,154,179]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="tamoyoensis">M. tamoyoensis</taxonomicName>
</emphasis>
and
<taxonomicName id="4C6B4D7C8619FFFAFA92FF105657FF09" authority="(Huynh &amp; Veenstra, 2015)" baseAuthorityName="Huynh &amp; Veenstra" baseAuthorityYear="2015" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="B91FEAED8619FFFAFA92FF105483FF09" italics="true" pageId="15" pageNumber="16">M. yunnanensis</emphasis>
(
<bibRefCitation id="EFFA4B0E8619FFFAFEDFFF37565FFF09" author="Huynh, C. &amp; Veenstra, A." box="[264,549,188,215]" pageId="15" pageNumber="16" pagination="460 - 474" refId="ref14596" refString="Huynh, C. &amp; Veenstra, A. (2015) Description of a new species of Penicillate Millipede from the genus Monographis (Diplopoda: Polyxenidae) found in Vietnam. Zootaxa, 3964 (4), 460 - 474. https: // doi. org / 10.11646 / zootaxa. 3964.4.5" type="journal article" year="2015">Huynh &amp; Veenstra, 2015</bibRefCitation>
)
</taxonomicName>
. The differences between
<taxonomicName id="4C6B4D7C8619FFFAFC85FF345784FF09" authority="Huynh, 2020" authorityName="Huynh" authorityYear="2020" box="[850,1022,189,215]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B91FEAED8619FFFAFC85FF345784FF09" box="[850,1022,189,215]" italics="true" pageId="15" pageNumber="16">M. condorensis</emphasis>
</taxonomicName>
<emphasis id="B91FEAED8619FFFAFBD2FF37503EFF09" bold="true" box="[1029,1092,189,215]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="A22C57968619FFFAFBD2FF37503EFF09" box="[1029,1092,189,215]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and other species in the group are the presence of setae on lamellae along the anterior margin of labrum, which is a character absent in the group; a large number of sensilla on antennomere VI (&gt;24 sensilla) and the claw structure (the claw is slender in structure with a shorter posterior lateral process, and the anterior setiform process is longer than the claw). All these characteristics make
<taxonomicName id="4C6B4D7C8619FFFAFEEEFEC4559EFEB9" authorityName="Huynh" authorityYear="2020" box="[313,484,333,359]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="condorensis">
<emphasis id="B91FEAED8619FFFAFEEEFEC4559EFEB9" box="[313,484,333,359]" italics="true" pageId="15" pageNumber="16">M. condorensis</emphasis>
</taxonomicName>
a distinctively new species in
<taxonomicName id="4C6B4D7C8619FFFAFCECFEC757A8FEB9" authorityName="Attems" authorityYear="1907" box="[827,978,333,359]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAED8619FFFAFCECFEC757A8FEB9" box="[827,978,333,359]" italics="true" pageId="15" pageNumber="16">Monographis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF1466778619FFFAFF40F8AB5165F830" ID-DOI="http://doi.org/10.5281/zenodo.3735879" ID-Zenodo-Dep="3735879" httpUri="https://zenodo.org/record/3735879/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1825,1850]" targetBox="[186,1412,391,1786]" targetPageId="15">
<paragraph id="8BD436FF8619FFFAFF40F8AB5165F830" blockId="15.[151,1437,1825,2030]" pageId="15" pageNumber="16">
<emphasis id="B91FEAED8619FFFAFF40F8AB555AF8E4" bold="true" box="[151,288,1825,1850]" pageId="15" pageNumber="16">FIGURE 12.</emphasis>
Holotype
<taxonomicName id="4C6B4D7C8619FFFAFE5BF8A856EFF8E4" authority="Huynh, 2020" authorityName="Huynh" authorityYear="2020" box="[396,661,1826,1850]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B91FEAED8619FFFAFE5BF8A856EFF8E4" box="[396,661,1826,1850]" italics="true" pageId="15" pageNumber="16">Monographis condorensis</emphasis>
</taxonomicName>
<emphasis id="B91FEAED8619FFFAFD4BF8A856ACF8E4" bold="true" box="[668,726,1826,1850]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="A22C57968619FFFAFD4BF8A856ACF8E4" box="[668,726,1826,1850]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B91FEAED8619FFFAFD36F8AB5680F8E4" bold="true" box="[737,762,1825,1850]" pageId="15" pageNumber="16">A.</emphasis>
Head capsule with 8 ommatidia on each side, the posterior vertex trichome group (pv) and 3 sockets of trichobothria (trichobothria
<emphasis id="B91FEAED8619FFFAFC97F8CC5737F883" bold="true" box="[832,845,1862,1885]" italics="true" pageId="15" pageNumber="16">a</emphasis>
,
<emphasis id="B91FEAED8619FFFAFC8CF8CF5712F880" bold="true" box="[859,872,1861,1886]" italics="true" pageId="15" pageNumber="16">b</emphasis>
and
<emphasis id="B91FEAED8619FFFAFC4AF8CC57D2F883" bold="true" box="[925,936,1862,1885]" italics="true" pageId="15" pageNumber="16">c</emphasis>
);
<emphasis id="B91FEAED8619FFFAFC17F8CC57ADF880" bold="true" box="[960,983,1862,1886]" pageId="15" pageNumber="16">B.</emphasis>
Collum (co) and lateral protuberances (Lp) showing the arrangement of trichomes sockets;
<emphasis id="B91FEAED8619FFFAFDABF8E356EFF85C" bold="true" box="[636,661,1897,1922]" pageId="15" pageNumber="16">C.</emphasis>
T2: Tergite 2;
<emphasis id="B91FEAED8619FFFAFCFCF8E0573EF85C" bold="true" box="[811,836,1898,1922]" pageId="15" pageNumber="16">D.</emphasis>
T10: Tergite 10—the last tergite;
<emphasis id="B91FEAED8619FFFAFB4AF8E050CEF85C" bold="true" box="[1181,1204,1898,1922]" pageId="15" pageNumber="16">E.</emphasis>
Trichobothria
<emphasis id="B91FEAED8619FFFAFA9CF8E05122F85F" box="[1355,1368,1898,1921]" italics="true" pageId="15" pageNumber="16">a</emphasis>
,
<emphasis id="B91FEAED8619FFFAFAB3F8E0510BF85C" box="[1380,1393,1898,1922]" italics="true" pageId="15" pageNumber="16">b</emphasis>
and
<emphasis id="B91FEAED8619FFFAFF40F80454D8F87B" bold="true" box="[151,162,1934,1957]" italics="true" pageId="15" pageNumber="16">c</emphasis>
(right hand side) showing the sensory hairs and sockets;
<emphasis id="B91FEAED8619FFFAFD32F8045682F878" bold="true" box="[741,760,1934,1958]" pageId="15" pageNumber="16">F.</emphasis>
Gnathochilarium: the lateral palp (LP) with 11 sensilla and medial palp (MP) with 22 sensilla;
<emphasis id="B91FEAED8619FFFAFE78F83B55B3F814" bold="true" box="[431,457,1969,1994]" pageId="15" pageNumber="16">G.</emphasis>
Clypeo-labrum showing a smooth surface, asetose, with setae along both anterior edges.
<emphasis id="B91FEAED8619FFFAFA9AF838511DF814" bold="true" box="[1357,1383,1970,1994]" pageId="15" pageNumber="16">H.</emphasis>
Dorsal ornamental trichome sockets of trichome a (
<emphasis id="B91FEAED8619FFFAFDAEF85C56FCF833" bold="true" box="[633,646,2006,2029]" italics="true" pageId="15" pageNumber="16">a</emphasis>
), trichome b (
<emphasis id="B91FEAED8619FFFAFCC0F85F575EF830" bold="true" box="[791,804,2005,2030]" italics="true" pageId="15" pageNumber="16">b</emphasis>
), trichome c (
<emphasis id="B91FEAED8619FFFAFC64F85C57C4F833" bold="true" box="[947,958,2006,2029]" italics="true" pageId="15" pageNumber="16">c</emphasis>
) and the circular indentation d (
<emphasis id="B91FEAED8619FFFAFAD3F85F516BF830" bold="true" box="[1284,1297,2005,2030]" italics="true" pageId="15" pageNumber="16">d</emphasis>
).
</paragraph>
</caption>
<caption id="DF1466778606FFE5FF40F950502CF831" ID-DOI="http://doi.org/10.5281/zenodo.3735883" ID-Zenodo-Dep="3735883" httpUri="https://zenodo.org/record/3735883/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1754,1779]" targetBox="[206,1383,181,1728]" targetPageId="16">
<paragraph id="8BD436FF8606FFE5FF40F950502CF831" blockId="16.[151,1437,1754,2032]" pageId="16" pageNumber="17">
<emphasis id="B91FEAED8606FFE5FF40F950555AF92D" bold="true" box="[151,288,1754,1779]" pageId="16" pageNumber="17">FIGURE 13.</emphasis>
SEM images of
<taxonomicName id="4C6B4D7C8606FFE5FE1BF95156AEF92D" authority="Huynh, 2020" authorityName="Huynh" authorityYear="2020" box="[460,724,1755,1779]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B91FEAED8606FFE5FE1BF95156AEF92D" box="[460,724,1755,1779]" italics="true" pageId="16" pageNumber="17">Monographis condorensis</emphasis>
</taxonomicName>
<emphasis id="B91FEAED8606FFE5FD0CF951576FF92D" bold="true" box="[731,789,1755,1779]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A22C57968606FFE5FD0CF951576FF92D" box="[731,789,1755,1779]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B91FEAED8606FFE5FCF5F9505741F92D" bold="true" box="[802,827,1754,1779]" pageId="16" pageNumber="17">A.</emphasis>
Dorsal view of the head capsule, showing 8 ommatidia and trichobothria
<emphasis id="B91FEAED8606FFE5FEF7F9755557F8C8" bold="true" box="[288,301,1791,1814]" italics="true" pageId="16" pageNumber="17">a</emphasis>
,
<emphasis id="B91FEAED8606FFE5FEEFF974553FF8C9" bold="true" box="[312,325,1790,1815]" italics="true" pageId="16" pageNumber="17">b</emphasis>
and
<emphasis id="B91FEAED8606FFE5FEA2F97555FAF8C8" bold="true" box="[373,384,1791,1814]" italics="true" pageId="16" pageNumber="17">c</emphasis>
;
<emphasis id="B91FEAED8606FFE5FE5AF97555DEF8C9" bold="true" box="[397,420,1791,1815]" pageId="16" pageNumber="17">B.</emphasis>
Dorsal view of the body, co: Collum, Lp: lateral protuberances and pp: pleural projection, T2: tergite 2 and T3: tergite 3;
<emphasis id="B91FEAED8606FFE5FEB4F8A85506F8E5" bold="true" box="[355,380,1826,1851]" pageId="16" pageNumber="17">C.</emphasis>
Clypeo-labrum showing the smooth surface and asetose (l), s: setae along anterior edges, Se: posterior setae, bse: a minute backward-pointing setae;
<emphasis id="B91FEAED8606FFE5FDBFF8CD56FAF881" bold="true" box="[616,640,1863,1887]" pageId="16" pageNumber="17">D.</emphasis>
Right gnathochilarium showing the lateral palp (LP) with 5 sensilla and 6 sensilla sockets, 22 sensilla on the medial palp (MP);
<emphasis id="B91FEAED8606FFE5FD4CF8E156C8F85D" bold="true" box="[667,690,1899,1923]" pageId="16" pageNumber="17">E.</emphasis>
Bare caudal bundle in the female showed 2 distinctive structures: caudal trichome sockets (cts) which include barbate trichome sockets (bts) in the middle of the bundle, forming a dorsal single bundle, nest trichome sockets (nts) forming 2 ventral bundles; these bundles are separated by the trichome socket-free tissue (tsft); T: telson;
<emphasis id="B91FEAED8606FFE5FF36F85D548EF831" bold="true" box="[225,244,2007,2031]" pageId="16" pageNumber="17">F.</emphasis>
Leg articles and arrows indicate spines which are typical characters of
<taxonomicName id="4C6B4D7C8606FFE5FC1FF85D502BF831" authorityName="Attems" authorityYear="1907" box="[968,1105,2007,2031]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAED8606FFE5FC1FF85D502BF831" box="[968,1105,2007,2031]" italics="true" pageId="16" pageNumber="17">Monographis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C37165748607FFE4FF10FF135703FF09" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BD436FF8607FFE4FF10FF1351E6FF6D" blockId="17.[151,1436,153,215]" box="[199,1436,153,179]" pageId="17" pageNumber="18">
<emphasis id="B91FEAED8607FFE4FF10FF135532FF6D" bold="true" box="[199,328,153,179]" pageId="17" pageNumber="18">Etymology</emphasis>
: The name of
<taxonomicName id="4C6B4D7C8607FFE4FE24FF135763FF6D" authorityName="Huynh" authorityYear="2020" box="[499,793,153,179]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="condorensis">
<emphasis id="B91FEAED8607FFE4FE24FF135763FF6D" box="[499,793,153,179]" italics="true" pageId="17" pageNumber="18">Monographis condorensis</emphasis>
</taxonomicName>
refers to the old name of Con Dao Archipelago as Pulo
</paragraph>
<paragraph id="8BD436FF8607FFE4FF40FF375703FF09" blockId="17.[151,1436,153,215]" box="[151,889,189,215]" pageId="17" pageNumber="18">
Condore in southeast
<collectingCountry id="F37C766F8607FFE4FE5DFF375596FF09" box="[394,492,189,215]" name="Vietnam" pageId="17" pageNumber="18">Vietnam</collectingCountry>
, where this species was first found.
</paragraph>
</subSubSection>
</treatment>
</document>
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