treatments-xml/data/03/C2/87/03C287BBE858FFBF31A64E7EFDF6FA3A.xml
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<document id="29F3305061C6A65DA85E64032864E42C" ID-CLB-Dataset="29854" ID-DOI="10.11646/zootaxa.4438.1.1" ID-GBIF-Dataset="3338d9d4-c5fa-4f77-89fb-901b1e028237" ID-ISSN="1175-5326" ID-Zenodo-Dep="1294121" ID-ZooBank="55BEB1B1-DA14-4CC4-92EB-1EA98AD6277C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529535606996" checkinUser="plazi" docAuthor="Rakotonirina, Jean Claude &amp; Fisher, Brian L." docDate="2018" docId="03C287BBE858FFBF31A64E7EFDF6FA3A" docLanguage="en" docName="zootaxa.4438.1.1.pdf.imf" docOrigin="Zootaxa 4438 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Camponotus sada Rakotonirina &amp; Fisher 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="54" masterDocId="FFFBFFC3E86BFF8A31314822FFAAFFFA" masterDocTitle="Taxonomic revision of the Malagasy Camponotus subgenus Mayria (Hymenoptera, Formicidae) using qualitative and quantitative morphology" masterLastPageNumber="58" masterPageNumber="1" pageNumber="52" updateTime="1698550116491" updateUser="ExternalLinkService">
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<mods:title id="55B30A10A090616C5327E39388640D08">Taxonomic revision of the Malagasy Camponotus subgenus Mayria (Hymenoptera, Formicidae) using qualitative and quantitative morphology</mods:title>
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<mods:namePart id="B28B5BCD190556DE4FC4A36EF6680DF3">Rakotonirina, Jean Claude</mods:namePart>
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<mods:namePart id="7819681A169DEF277BF9DF866381D64C">Fisher, Brian L.</mods:namePart>
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<mods:title id="25ACD76D1A4FE8E2A589F8CD0987E24B">Zootaxa</mods:title>
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<mods:date id="E4C6BAEB650F0EA2FDF14780677F1FAC">2018</mods:date>
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<treatment id="03C287BBE858FFBF31A64E7EFDF6FA3A" ID-DOI="http://doi.org/10.5281/zenodo.5987986" ID-GBIF-Taxon="146015945" ID-Zenodo-Dep="5987986" LSID="urn:lsid:plazi:treatment:03C287BBE858FFBF31A64E7EFDF6FA3A" httpUri="http://treatment.plazi.org/id/03C287BBE858FFBF31A64E7EFDF6FA3A" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52">
<subSubSection id="C3716526E858FFB931A64E7EFED0F96D" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="8BD436ADE858FFB931A64E7EFD74F98C" blockId="51.[151,734,1628,1687]" box="[151,734,1628,1654]" pageId="51" pageNumber="52">
<heading id="D09C81C1E858FFB931A64E7EFD74F98C" bold="true" box="[151,734,1628,1654]" fontSize="11" level="1" pageId="51" pageNumber="52" reason="1">
<emphasis id="B91FEABFE858FFB931A64E7EFD74F98C" bold="true" box="[151,734,1628,1654]" pageId="51" pageNumber="52">
<taxonomicName id="4C6B4D2EE858FFB931A64E7EFD3FF98C" authority="Rakotonirina &amp; Fisher" authorityName="Rakotonirina &amp; Fisher" authorityYear="2018" box="[151,661,1628,1654]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="sada" status="sp. nov.">
<emphasis id="B91FEABFE858FFB931A64E7EFEC4F98C" bold="true" box="[151,366,1628,1654]" italics="true" pageId="51" pageNumber="52">Camponotus sada</emphasis>
Rakotonirina &amp; Fisher
</taxonomicName>
<taxonomicNameLabel id="A22C57C4E858FFB933AD4E7EFD74F98C" box="[668,734,1628,1654]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD436ADE858FFB931A64E5CFED0F96D" blockId="51.[151,734,1628,1687]" box="[151,378,1662,1687]" pageId="51" pageNumber="52">
(
<figureCitation id="13502A28E858FFB931AF4E5CFEB5F96D" box="[158,287,1662,1687]" captionStart="FIGURE 8" captionStartId="16.[151,250,774,796]" captionTargetBox="[152,1435,193,735]" captionTargetId="figure@16.[152,1435,193,752]" captionTargetPageId="16" captionText="FIGURE 8. Head in full-face view. A: C. sada (CASENT0498916). B: C. christi (CASENT0129791)." httpUri="https://zenodo.org/record/1294137/files/figure.png" pageId="51" pageNumber="52">Figures 8A</figureCitation>
,
<figureCitation id="13502A28E858FFB9301A4E5DFEE3F96D" box="[299,329,1663,1687]" captionStart="FIGURE 33" captionStartId="52.[151,250,1733,1755]" captionTargetBox="[188,1398,301,1697]" captionTargetId="figure@52.[188,1398,301,1712]" captionTargetPageId="52" captionText="FIGURE 33. Camponotus sada minor worker CASENT0498916. A: lateral view. B: head in full-face view. C: dorsal view." httpUri="https://zenodo.org/record/1294177/files/figure.png" pageId="51" pageNumber="52">33</figureCitation>
,
<figureCitation id="13502A28E858FFB930644E5DFED8F96D" box="[341,370,1663,1687]" captionStart="FIGURES 3542" captionStartId="55.[151,264,1963,1985]" captionTargetBox="[189,1370,193,1916]" captionTargetId="figure@55.[189,1370,193,1916]" captionTargetPageId="55" captionText="FIGURES 3542. Distribution maps of Camponotus subgenus Mayria in Madagascar. Fig. 35: C. maculiventris, Fig. 36: C. mainty, Fig. 37: C. manabo, Fig. 38: C. pulcher, Fig. 39: C. raina, Fig. 40: C. repens, Fig. 41: C. sada, Fig. 42: C. tanosy." httpUri="https://zenodo.org/record/1294181/files/figure.png" pageId="51" pageNumber="52">41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3716526E858FFB931A64EE4FB50F84D" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="8BD436ADE858FFB931A64EE4FCA6F8DD" blockId="51.[151,1437,1734,2011]" pageId="51" pageNumber="52">
<materialsCitation id="3B033CF0E858FFB931A64EE4FCA2F8DD" ID-GBIF-Occurrence="1851012646" collectingDate="2003-12-16" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="425" location="Foret d'Andavakoera" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<emphasis id="B91FEABFE858FFB931A64EE4FECDF925" bold="true" box="[151,359,1734,1759]" pageId="51" pageNumber="52">
<typeStatus id="54D0880FE858FFB931A64EE4FEAEF925" box="[151,260,1734,1759]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
worker.
</emphasis>
<collectingCountry id="F37C763DE858FFB9305C4EE4FE53F925" box="[365,505,1734,1759]" name="Madagascar" pageId="51" pageNumber="52">Madagascar</collectingCountry>
: Province
<collectingRegion id="49AFF84FE858FFB9335F4EE4FD52F925" box="[622,760,1734,1759]" country="Madagascar" name="Antsiranana" pageId="51" pageNumber="52">Antsiranana</collectingRegion>
,
<location id="8EB46076E858FFB932354EE5FC5FF925" LSID="urn:lsid:plazi:treatment:03C287BBE858FFBF31A64E7EFDF6FA3A:8EB46076E858FFB932354EE5FC5FF925" box="[772,1013,1734,1759]" country="Madagascar" name="Foret d'Andavakoera" pageId="51" pageNumber="52" stateProvince="Antsiranana">Forêt dAndavakoera</location>
,
<locationDeviation id="9A662F8FE858FFB935304EE5FAE9F925" box="[1025,1347,1734,1759]" pageId="51" pageNumber="52">
<quantity id="4C939B48E858FFB935304EE5FBF7F925" box="[1025,1117,1734,1759]" metricMagnitude="4" metricUnit="m" metricValue="2.1399999999999997" pageId="51" pageNumber="52" unit="km" value="21.4">21.4 km</quantity>
75° ENE Ambilobe
</locationDeviation>
;
<locationDeviation id="9A662F8FE858FFB9347F4EE4FEFAF8F9" pageId="51" pageNumber="52">
<quantity id="4C939B48E858FFB9347F4EE4FA36F925" box="[1358,1436,1734,1759]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="51" pageNumber="52" unit="km" value="4.6">4.6 km</quantity>
356° N Betsiaka
</locationDeviation>
, -13.11833, 49.23,
<quantity id="4C939B48E858FFB933174EC8FDC5F8F8" box="[550,623,1770,1795]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="51" pageNumber="52" unit="m" value="425.0">
<elevation id="0046D19EE858FFB933174EC8FDC5F8F8" box="[550,623,1770,1795]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="51" pageNumber="52" unit="m" value="425.0">425 m</elevation>
</quantity>
, rainforest, under stone,
<date id="FFD5106DE858FFB932B04EC8FBA5F8F9" box="[897,1039,1770,1795]" pageId="51" pageNumber="52" value="2003-12-16">
<collectingDate id="EF91E985E858FFB932B04EC8FBA5F8F9" box="[897,1039,1770,1795]" pageId="51" pageNumber="52" value="2003-12-16">16 Dec 2003</collectingDate>
</date>
(
<collectorName id="269E537BE858FFB9352E4EC9FB37F8F9" box="[1055,1181,1770,1795]" pageId="51" pageNumber="52">B.L. Fisher</collectorName>
<emphasis id="B91FEABFE858FFB935924ECEFB71F8F9" box="[1187,1243,1770,1795]" italics="true" pageId="51" pageNumber="52">et al.</emphasis>
) collection code: BLF10346, specimen code: CASENT0498916 (
<collectionCode id="ED7AAE68E858FFB933804F2DFCAAF8DD" box="[689,768,1807,1831]" pageId="51" pageNumber="52">CASC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD436ADE858FFB931F64F10FB50F84D" blockId="51.[151,1437,1734,2011]" pageId="51" pageNumber="52">
<materialsCitation id="3B033CF0E858FFB931F64F10FB50F84D" ID-GBIF-Occurrence="1851012744" collectingDate="2003-12-16" collectionCode="CASC, BMNH, MHNG, MSNG, PBZT" collectorName="B. L. Fisher" country="Madagascar" elevation="425" location="Foret d'Andavakoera" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<emphasis id="B91FEABFE858FFB931F64F10FEECF8B1" bold="true" box="[199,326,1842,1867]" pageId="51" pageNumber="52">
<typeStatus id="54D0880FE858FFB931F64F10FEEBF8B1" box="[199,321,1842,1867]" pageId="51" pageNumber="52" type="paratype">Paratypes</typeStatus>
.
</emphasis>
8 minor and 5 major workers with same data as holotype but with specimen codes: minors: CASENT0498917, CASENT0498918, CASENT0804651, CASENT0804652, CASENT0804654, CASENT0804655, CASENT0804657, CASENT0804658; majors: CASENT0498919, CASENT0498920, CASENT0498921, CASENT0804653, CASENT0804656 (
<collectionCode id="ED7AAE68E858FFB9321F4FBDFCD1F84D" box="[814,891,1951,1975]" pageId="51" pageNumber="52">CASC</collectionCode>
,
<collectionCode id="ED7AAE68E858FFB932B64FBDFC74F84C" box="[903,990,1951,1974]" pageId="51" pageNumber="52">BMNH</collectionCode>
,
<collectionCode id="ED7AAE68E858FFB932D84FBDFB94F84D" box="[1001,1086,1951,1975]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="51" pageNumber="52">MHNG</collectionCode>
,
<collectionCode id="ED7AAE68E858FFB935784FBDFB30F84D" box="[1097,1178,1951,1975]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="51" pageNumber="52">MSNG</collectionCode>
,
<collectionCode id="ED7AAE68E858FFB935944FBDFB44F84D" box="[1189,1262,1951,1975]" country="Madagascar" httpUri="http://grbio.org/cool/a6tz-5rcd" name="Parc Botanique et Zoologique de Tsimbazaza" pageId="51" pageNumber="52">PBZT</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3716526E858FFBE31F64FE0FE5AFF02" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="8BD436ADE858FFB931F64FE0FA36F821" blockId="51.[151,1437,1734,2011]" box="[199,1436,1986,2011]" pageId="51" pageNumber="52">
<emphasis id="B91FEABFE858FFB931F64FE0FEEBF821" bold="true" box="[199,321,1986,2011]" pageId="51" pageNumber="52">Diagnosis.</emphasis>
Median portion of clypeus without longitudinal carina; anteromedian margin of clypeus covered
</paragraph>
<paragraph id="8BD436ADE85FFFBE31A648B5FE5AFF02" blockId="52.[151,1437,151,248]" pageId="52" pageNumber="53">with lamella; mandible with six teeth; in full-face view, head distinctly longer than broad; antennal scape covered with appressed short hairs; in lateral view, junction of dorsal and declivity surfaces of propodeum broadly rounded; petiolar node longer than high.</paragraph>
</subSubSection>
<caption id="DF146625E85FFFBE31A64EE7FAD6F921" box="[151,1404,1733,1755]" httpUri="https://zenodo.org/record/1294177/files/figure.png" pageId="52" pageNumber="53" startId="52.[151,250,1733,1755]" targetBox="[188,1398,301,1697]" targetPageId="52">
<paragraph id="8BD436ADE85FFFBE31A64EE7FAD6F921" blockId="52.[151,1404,1733,1755]" box="[151,1404,1733,1755]" pageId="52" pageNumber="53">
<emphasis id="B91FEABFE85FFFBE31A64EE7FEB5F921" bold="true" box="[151,287,1733,1755]" pageId="52" pageNumber="53">FIGURE 33.</emphasis>
<taxonomicName id="4C6B4D2EE85FFFBE30144EE4FE76F921" authorityName="Rakotonirina &amp; Fisher" authorityYear="2018" box="[293,476,1733,1755]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sada">
<emphasis id="B91FEABFE85FFFBE30144EE4FE76F921" box="[293,476,1733,1755]" italics="true" pageId="52" pageNumber="53">Camponotus sada</emphasis>
</taxonomicName>
minor worker CASENT0498916. A: lateral view. B: head in full-face view. C: dorsal view.
</paragraph>
</caption>
<subSubSection id="C3716526E85FFFBF31F64F20FD03FD9A" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="description">
<paragraph id="8BD436ADE85FFFBF31F64F20FEC1FF4A" blockId="52.[151,1437,1794,2035]" lastBlockId="53.[151,1437,151,1472]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<emphasis id="B91FEABFE85FFFBE31F64F20FDB2F8E1" bold="true" box="[199,536,1794,1819]" pageId="52" pageNumber="53">Description. Minor worker.</emphasis>
In full-face view, head elongate, rectangular, with broadly convex posterior margin. Anteromedian clypeal margin broadly convex and with broadly rounded lamella. Eyes breaking lateral outlines of head, their posterior margins located well behind the mid-length of the head (PoOc/CL: 0.33±0.01; 0.310.35). Mandible triangular, apical margin with six sharp teeth. Antennal scape long, roughly its distal half extending beyond posterior cephalic border. In lateral view, promesonotum convex, propodeum straight, sometimes slightly concave, its dorsum joining the straight and anteriorly inclined declivity in a broad angle; propodeal dorsum approximately three times longer than height of declivity. Petiolar node longer than high and inclined anteriorly.
</paragraph>
<paragraph id="8BD436ADE85EFFBF31F6489FFF4CFE57" blockId="53.[151,1437,151,1472]" pageId="53" pageNumber="54">More than one pair of elongate, stout, erect hairs near posterior border of head; pronotum, junction of dorsum and declivity of propodeum and posterodorsal margin of petiole each with one pair of elongate, stout erect hairs; few short appressed hairs present on dorsum of mesosoma. Antennal scape with abundant appressed short hairs. Body color generally black, with whitish distal portion of coxae, trochanter, femur and posterior portion of first gastral sternite; leg black on proximal portion and becoming progressively lighter in color toward its apex; first and second gastral tergite with pair of white spots; antennal scape and funiculus basally light brown and apically dark brown.</paragraph>
<paragraph id="8BD436ADE85EFFBF31F64995FD03FD9A" blockId="53.[151,1437,151,1472]" pageId="53" pageNumber="54">
<emphasis id="B91FEABFE85EFFBF31F64995FED2FE2A" bold="true" box="[199,376,439,464]" pageId="53" pageNumber="54">Major worker.</emphasis>
Differing from minor worker in the following characters: larger head, absence of lamella on anterior clypeal margin, antennal scape barely surpassing posterior cephalic margin, much more robust mesosoma, with pronotum and mesonotum forming an even convexity, separated from propodeal dorsum by a shallow angle; higher than long petiolar node; much higher propodeal declivity; presence of at least three pairs of erect hairs on mesonotum. Other features as in minor worker.
</paragraph>
</subSubSection>
<subSubSection id="C3716526E85EFFBF31F64A4EFC84FCA6" pageId="53" pageNumber="54" type="discussion">
<paragraph id="8BD436ADE85EFFBF31F64A4EFC66FD0A" blockId="53.[151,1437,151,1472]" pageId="53" pageNumber="54">
<emphasis id="B91FEABFE85EFFBF31F64A4EFEE6FD7F" bold="true" box="[199,332,620,645]" pageId="53" pageNumber="54">Discussion.</emphasis>
<taxonomicName id="4C6B4D2EE85EFFBF30644A4FFD8AFD7F" authorityName="Rakotonirina &amp; Fisher" authorityYear="2018" box="[341,544,620,645]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sada">
<emphasis id="B91FEABFE85EFFBF30644A4FFD8AFD7F" box="[341,544,620,645]" italics="true" pageId="53" pageNumber="54">Camponotus sada</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B4D2EE85EFFBF336B4A4FFD10FD7E" authorityName="Rakotonirina &amp; Fisher" authorityYear="2018" box="[602,698,621,644]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="raina">
<emphasis id="B91FEABFE85EFFBF336B4A4FFD10FD7E" box="[602,698,621,644]" italics="true" pageId="53" pageNumber="54">C. raina</emphasis>
</taxonomicName>
look similar but the latter has a short and high mesosoma and a generally reddish-orange head and mesosoma.
<taxonomicName id="4C6B4D2EE85EFFBF33954AB2FCC7FD52" authorityName="Rakotonirina &amp; Fisher" authorityYear="2018" box="[676,877,655,680]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sada">
<emphasis id="B91FEABFE85EFFBF33954AB2FCC7FD52" box="[676,877,655,680]" italics="true" pageId="53" pageNumber="54">Camponotus sada</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="4C6B4D2EE85EFFBF35414AB2FB72FD52" authorityName=", Emery" authorityYear="1896" box="[1136,1240,655,680]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="christi">
<emphasis id="B91FEABFE85EFFBF35414AB2FB72FD52" box="[1136,1240,655,680]" italics="true" pageId="53" pageNumber="54">C. christi</emphasis>
</taxonomicName>
,
<emphasis id="B91FEABFE85EFFBF35D74AB2FAC6FD52" box="[1254,1388,656,680]" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="4C6B4D2EE85EFFBF35D74AB2FAC2FD52" authorityName="Forel" authorityYear="1886" box="[1254,1384,656,680]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="foersteri">C. foersteri</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C6B4D2EE85EFFBF31A64A97FEFBFD37" box="[151,337,692,717]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="maculiventris">
<emphasis id="B91FEABFE85EFFBF31A64A97FEFBFD37" box="[151,337,692,717]" italics="true" pageId="53" pageNumber="54">C. maculiventris</emphasis>
</taxonomicName>
by the presence of a median longitudinal carina on the clypeus.
<taxonomicName id="4C6B4D2EE85EFFBF351F4A97FAA7FD36" authorityName="Rakotonirina &amp; Fisher" authorityYear="2018" box="[1070,1293,693,716]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tanosy">
<emphasis id="B91FEABFE85EFFBF351F4A97FAA7FD36" box="[1070,1293,693,716]" italics="true" pageId="53" pageNumber="54">Camponotus tanosy</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C6B4D2EE85EFFBF31A64AFAFF46FD0A" authorityName="Rakotonirina &amp; Fisher" authorityYear="2018" box="[151,236,727,752]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sada">
<emphasis id="B91FEABFE85EFFBF31A64AFAFF46FD0A" box="[151,236,727,752]" italics="true" pageId="53" pageNumber="54">C. sada</emphasis>
</taxonomicName>
by the absence of a lamella on the anteromediam clypeal margin.
</paragraph>
<paragraph id="8BD436ADE85EFFBF31F64ADEFC84FCA6" blockId="53.[151,1437,151,1472]" pageId="53" pageNumber="54">
The NC-clustering showed quantitative morphological differences between
<emphasis id="B91FEABFE85EFFBF352B4ADFFB57FCEE" box="[1050,1277,764,789]" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="4C6B4D2EE85EFFBF352B4ADFFBD9FCEF" authorityName="Rakotonirina &amp; Fisher" authorityYear="2018" box="[1050,1139,764,789]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sada">C. sada</taxonomicName>
,
<taxonomicName id="4C6B4D2EE85EFFBF354C4ADFFB57FCEE" authorityName="Forel" authorityYear="1886" box="[1149,1277,764,788]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="foersteri">C. foersteri</taxonomicName>
</emphasis>
,
<emphasis id="B91FEABFE85EFFBF343D4ADFFF4EFCC2" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="4C6B4D2EE85EFFBF343D4ADFFADCFCEF" authorityName=", Emery" authorityYear="1896" box="[1292,1398,764,789]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="christi">C. christi</taxonomicName>
,
<taxonomicName id="4C6B4D2EE85EFFBF34B34ADFFF4EFCC2" authorityName="Rakotonirina &amp; Fisher" authorityYear="2018" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="mainty">C. mainty</taxonomicName>
</emphasis>
and
<taxonomicName id="4C6B4D2EE85EFFBF302F4B02FE70FCC2" box="[286,474,799,824]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="maculiventris">
<emphasis id="B91FEABFE85EFFBF302F4B02FE70FCC2" box="[286,474,799,824]" italics="true" pageId="53" pageNumber="54">C. maculiventris</emphasis>
</taxonomicName>
that are confirmed by the partitioning method using hclust and kmeans algorithms.
<taxonomicName id="4C6B4D2EE85EFFBF31A64B67FECAFCA7" authorityName="Rakotonirina &amp; Fisher" authorityYear="2018" box="[151,352,836,861]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sada">
<emphasis id="B91FEABFE85EFFBF31A64B67FECAFCA7" box="[151,352,836,861]" italics="true" pageId="53" pageNumber="54">Camponotus sada</emphasis>
</taxonomicName>
is 100% successfully classified by LDA.
</paragraph>
</subSubSection>
<subSubSection id="C3716526E85EFFBF31F64B45FC38FBEA" pageId="53" pageNumber="54" type="distribution">
<paragraph id="8BD436ADE85EFFBF31F64B45FC38FBEA" blockId="53.[151,1437,151,1472]" pageId="53" pageNumber="54">
<emphasis id="B91FEABFE85EFFBF31F64B45FE53FC7A" bold="true" box="[199,505,871,896]" pageId="53" pageNumber="54">Distribution and biology.</emphasis>
One of the
<taxonomicName id="4C6B4D2EE85EFFBF33BF4B4BFD4BFC7A" authorityName="Forel" authorityYear="1878" box="[654,737,873,896]" class="Insecta" family="Formicidae" genus="Mayria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEABFE85EFFBF33BF4B4BFD4BFC7A" box="[654,737,873,896]" italics="true" pageId="53" pageNumber="54">Mayria</emphasis>
</taxonomicName>
group that occur only in the north of
<collectingCountry id="F37C763DE85EFFBF359B4B45FA99FC7A" box="[1194,1331,871,896]" name="Madagascar" pageId="53" pageNumber="54">Madagascar</collectingCountry>
,
<taxonomicName id="4C6B4D2EE85EFFBF34734B4AFA36FC7A" authorityName="Rakotonirina &amp; Fisher" authorityYear="2018" box="[1346,1436,871,896]" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sada">
<emphasis id="B91FEABFE85EFFBF34734B4AFA36FC7A" box="[1346,1436,871,896]" italics="true" pageId="53" pageNumber="54">C. sada</emphasis>
</taxonomicName>
inhabits the montane rainforests of Parc National Ambre, the rainforests of Andavakoera, and the dry forests of Bekaraoka, Montagne des Français, Réserves Spéciales dAnalamerana and de lAnkarana. Across these habitats, nests are built in the ground, under stones, or in root mats in the ground. Foraging behavior has been observed in leaf litter, on the forest floor, or on the lower parts of the vegetation.
</paragraph>
</subSubSection>
<subSubSection id="C3716526E85EFFBF31F64C3EFDF6FA3A" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="8BD436ADE85EFFBF31F64C3EFDF6FA3A" blockId="53.[151,1437,151,1472]" pageId="53" pageNumber="54">
<materialsCitation id="3B033CF0E85EFFBF31F64C3EFBEBFBA2" ID-GBIF-Occurrence="1851012595" collectionCode="CASC" collectorName="B. L. Fisher" elevation="425" location="Foret d'Andavakoera" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Antsiranana">
<emphasis id="B91FEABFE85EFFBF31F64C3EFD9DFBCF" bold="true" box="[199,567,1052,1077]" pageId="53" pageNumber="54">Additional material examined.</emphasis>
Province
<emphasis id="B91FEABFE85EFFBF339A4C3EFCE3FBCF" bold="true" box="[683,841,1052,1077]" pageId="53" pageNumber="54">
<collectingRegion id="49AFF84FE85EFFBF339A4C3EFCEFFBCF" box="[683,837,1052,1077]" country="Madagascar" name="Antsiranana" pageId="53" pageNumber="54">Antsiranana</collectingRegion>
:
</emphasis>
<location id="8EB46076E85EFFBF32634C3EFBEEFBCF" LSID="urn:lsid:plazi:treatment:03C287BBE858FFBF31A64E7EFDF6FA3A:8EB46076E85EFFBF32634C3EFBEEFBCF" box="[850,1092,1052,1077]" name="Foret d'Andavakoera" pageId="53" pageNumber="54" stateProvince="Antsiranana">Forêt dAndavakoera</location>
,
<quantity id="4C939B48E85EFFBF35604C3EFB04FBCF" box="[1105,1198,1052,1077]" metricMagnitude="4" metricUnit="m" metricValue="2.1399999999999997" pageId="53" pageNumber="54" unit="km" value="21.4">21.4 km</quantity>
75° ENE Ambilobe,
<quantity id="4C939B48E85EFFBF31A64C1DFF4FFBA2" box="[151,229,1087,1112]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="53" pageNumber="54" unit="km" value="4.6">4.6 km</quantity>
356° N Betsiaka,
<quantity id="4C939B48E85EFFBF30814C1DFE53FBAD" box="[432,505,1087,1112]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="53" pageNumber="54" unit="m" value="425.0">
<elevation id="0046D19EE85EFFBF30814C1DFE53FBAD" box="[432,505,1087,1112]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="53" pageNumber="54" unit="m" value="425.0">425 m</elevation>
</quantity>
, -13.11833, 49.23, rainforest, (
<collectorName id="269E537BE85EFFBF32624C62FC79FBA2" box="[851,979,1087,1112]" pageId="53" pageNumber="54">B.L. Fisher</collectorName>
) (
<collectionCode id="ED7AAE68E85EFFBF32D84C62FB92FBA2" box="[1001,1080,1088,1112]" pageId="53" pageNumber="54">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CF0E85EFFBF357D4C62FB83FB86" ID-GBIF-Occurrence="1851012808" collectionCode="CASC" collectorName="B. L. Fisher" elevation="175" location="Foret d'Ampondrabe" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Antsiranana">
<location id="8EB46076E85EFFBF357D4C62FA9CFBA2" LSID="urn:lsid:plazi:treatment:03C287BBE858FFBF31A64E7EFDF6FA3A:8EB46076E85EFFBF357D4C62FA9CFBA2" box="[1100,1334,1087,1112]" name="Foret d'Ampondrabe" pageId="53" pageNumber="54" stateProvince="Antsiranana">Forêt d'Ampondrabe</location>
,
<locationDeviation id="9A662F8FE85EFFBF34704C1DFECBFB87" pageId="53" pageNumber="54">
<quantity id="4C939B48E85EFFBF34704C1DFA36FBA2" box="[1345,1436,1087,1112]" metricMagnitude="4" metricUnit="m" metricValue="2.63" pageId="53" pageNumber="54" unit="km" value="26.3">26.3 km</quantity>
10° NNE Daraina
</locationDeviation>
,
<quantity id="4C939B48E85EFFBF305C4C46FE1DFB86" box="[365,439,1124,1149]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="53" pageNumber="54" unit="m" value="175.0">
<elevation id="0046D19EE85EFFBF305C4C46FE1DFB86" box="[365,439,1124,1149]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="53" pageNumber="54" unit="m" value="175.0">175 m</elevation>
</quantity>
, -12.97, 49.7, tropical dry forest, (
<collectorName id="269E537BE85EFFBF320B4C46FC11FB87" box="[826,955,1124,1149]" pageId="53" pageNumber="54">B.L. Fisher</collectorName>
) (
<collectionCode id="ED7AAE68E85EFFBF32E34C46FB8BFB86" box="[978,1057,1124,1148]" pageId="53" pageNumber="54">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CF0E85EFFBF35044C46FB85FB5A" ID-GBIF-Occurrence="1851012648" collectionCode="CASC" collectorName="B. L. Fisher" elevation="150" location="Foret de Bekaraoka" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Antsiranana">
<location id="8EB46076E85EFFBF35044C46FABEFB87" LSID="urn:lsid:plazi:treatment:03C287BBE858FFBF31A64E7EFDF6FA3A:8EB46076E85EFFBF35044C46FABEFB87" box="[1077,1300,1124,1149]" name="Foret de Bekaraoka" pageId="53" pageNumber="54" stateProvince="Antsiranana">Forêt de Bekaraoka</location>
,
<locationDeviation id="9A662F8FE85EFFBF342E4C46FE9AFB5A" pageId="53" pageNumber="54">
<quantity id="4C939B48E85EFFBF342E4C46FAC4FB87" box="[1311,1390,1124,1149]" metricMagnitude="3" metricUnit="m" metricValue="6.8" pageId="53" pageNumber="54" unit="km" value="6.8">6.8 km</quantity>
60° ENE Daraina
</locationDeviation>
,
<quantity id="4C939B48E85EFFBF300D4CA5FE2FFB5A" box="[316,389,1159,1184]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="53" pageNumber="54" unit="m" value="150.0">
<elevation id="0046D19EE85EFFBF300D4CA5FE2FFB5A" box="[316,389,1159,1184]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="53" pageNumber="54" unit="m" value="150.0">150 m</elevation>
</quantity>
, -13.16667, 49.71, tropical dry forest, (
<collectorName id="269E537BE85EFFBF32714CAAFC6BFB5A" box="[832,961,1159,1184]" pageId="53" pageNumber="54">B.L. Fisher</collectorName>
) (
<collectionCode id="ED7AAE68E85EFFBF32E94CAAFB8DFB5A" box="[984,1063,1160,1184]" pageId="53" pageNumber="54">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CF0E85EFFBF350A4CABFF5CFB12" ID-GBIF-Occurrence="1851012730" collectionCode="CASC" elevation="180" location="Diego Suarez" municipality="Montagne des Francais" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Antsiranana">
<collectingMunicipality id="6BB0ACD7E85EFFBF350A4CABFAE9FB5A" box="[1083,1347,1159,1185]" pageId="53" pageNumber="54">Montagne des Français</collectingMunicipality>
,
<locationDeviation id="9A662F8FE85EFFBF347F4CAAFE2FFB3F" pageId="53" pageNumber="54">
<quantity id="4C939B48E85EFFBF347F4CAAFA36FB5A" box="[1358,1436,1159,1184]" metricMagnitude="3" metricUnit="m" metricValue="7.2" pageId="53" pageNumber="54" unit="km" value="7.2">7.2 km</quantity>
142° SE
<collectingRegion id="49AFF84FE85EFFBF31CF4C8EFE2FFB3F" box="[254,389,1196,1221]" country="Madagascar" name="Antsiranana" pageId="53" pageNumber="54">Antsiranana</collectingRegion>
</locationDeviation>
(=
<location id="8EB46076E85EFFBF30944C8EFDEFFB3E" LSID="urn:lsid:plazi:treatment:03C287BBE858FFBF31A64E7EFDF6FA3A:8EB46076E85EFFBF30944C8EFDEFFB3E" box="[421,581,1196,1221]" municipality="Montagne des Francais" name="Diego Suarez" pageId="53" pageNumber="54" stateProvince="Antsiranana">Diego Suarez</location>
),
<quantity id="4C939B48E85EFFBF336B4C8EFD0FFB3E" box="[602,677,1196,1220]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="53" pageNumber="54" unit="m" value="180.0">
<elevation id="0046D19EE85EFFBF336B4C8EFD0FFB3E" box="[602,677,1196,1220]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="53" pageNumber="54" unit="m" value="180.0">180 m</elevation>
</quantity>
, -12.32278, 49.33817, tropical dry forest, (Fisher, Griswold
<emphasis id="B91FEABFE85EFFBF34684C8FFA39FB3F" box="[1369,1427,1196,1221]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
) (
<collectionCode id="ED7AAE68E85EFFBF31AE4CF2FF44FB12" box="[159,238,1232,1256]" pageId="53" pageNumber="54">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CF0E85EFFBF30334CF3FD31FAF6" ID-GBIF-Occurrence="1851012742" collectionCode="CASC" elevation="1300" location="Parc National Montagne d'Ambre" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Antsiranana">
<location id="8EB46076E85EFFBF30334CF3FD2AFB12" LSID="urn:lsid:plazi:treatment:03C287BBE858FFBF31A64E7EFDF6FA3A:8EB46076E85EFFBF30334CF3FD2AFB12" box="[258,640,1231,1257]" name="Parc National Montagne d'Ambre" pageId="53" pageNumber="54" stateProvince="Antsiranana">Parc National Montagne d'Ambre</location>
,
<locationDeviation id="9A662F8FE85EFFBF33BD4CF2FC48FB12" box="[652,994,1231,1256]" pageId="53" pageNumber="54">
<quantity id="4C939B48E85EFFBF33BD4CF2FD42FB12" box="[652,744,1231,1256]" metricMagnitude="4" metricUnit="m" metricValue="1.22" pageId="53" pageNumber="54" unit="km" value="12.2">12.2 km</quantity>
211° SSW Joffreville
</locationDeviation>
,
<quantity id="4C939B48E85EFFBF32DC4CF2FBEFFB12" box="[1005,1093,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="53" pageNumber="54" unit="m" value="1300.0">
<elevation id="0046D19EE85EFFBF32DC4CF2FBEFFB12" box="[1005,1093,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="53" pageNumber="54" unit="m" value="1300.0">1300 m</elevation>
</quantity>
, -12.59639, 49.1595, montane rainforest, (Fisher, Griswold
<emphasis id="B91FEABFE85EFFBF30DB4CD7FD8DFAF7" box="[490,551,1268,1293]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
) (
<collectionCode id="ED7AAE68E85EFFBF33754CD6FD39FAF6" box="[580,659,1268,1292]" pageId="53" pageNumber="54">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CF0E85EFFBF339A4CD6FCC2FACA" ID-GBIF-Occurrence="1851012738" collectionCode="CASC" collectorName="B. L. Fisher" elevation="225" location="Anivorano-Nord" municipality="Reserve Analamerana" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Antsiranana">
<collectingMunicipality id="6BB0ACD7E85EFFBF339A4CD6FC03FAF7" box="[683,937,1268,1293]" pageId="53" pageNumber="54">Réserve Analamerana</collectingMunicipality>
,
<quantity id="4C939B48E85EFFBF32884CD6FBB0FAF7" box="[953,1050,1268,1293]" metricMagnitude="4" metricUnit="m" metricValue="1.67" pageId="53" pageNumber="54" unit="km" value="16.7">16.7 km</quantity>
123°
<location id="8EB46076E85EFFBF35564CD6FA8FFAF7" LSID="urn:lsid:plazi:treatment:03C287BBE858FFBF31A64E7EFDF6FA3A:8EB46076E85EFFBF35564CD6FA8FFAF7" box="[1127,1317,1268,1293]" municipality="Reserve Analamerana" name="Anivorano-Nord" pageId="53" pageNumber="54" stateProvince="Antsiranana">Anivorano-Nord</location>
,
<quantity id="4C939B48E85EFFBF34054CD6FA28FAF6" box="[1332,1410,1268,1293]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="53" pageNumber="54" unit="m" value="225.0">
<elevation id="0046D19EE85EFFBF34054CD6FA28FAF6" box="[1332,1410,1268,1293]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="53" pageNumber="54" unit="m" value="225.0">225 m</elevation>
</quantity>
, - 12.80467, 49.37383, tropical dry forest, (
<collectorName id="269E537BE85EFFBF33454D3AFD5DFACA" box="[628,759,1303,1328]" pageId="53" pageNumber="54">B.L. Fisher</collectorName>
) (
<collectionCode id="ED7AAE68E85EFFBF32204D3AFCCAFACA" box="[785,864,1304,1328]" pageId="53" pageNumber="54">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CF0E85EFFBF32474D3AFC44FAAE" ID-GBIF-Occurrence="1851012750" collectionCode="CASC" collectorName="B. L. Fisher" elevation="60" location="Nord" municipality="Reserve Analamerana" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Antsiranana">
<collectingMunicipality id="6BB0ACD7E85EFFBF32474D3AFBD9FACA" box="[886,1139,1303,1328]" pageId="53" pageNumber="54">Réserve Analamerana</collectingMunicipality>
,
<quantity id="4C939B48E85EFFBF35B04D3AFB4AFACA" box="[1153,1248,1303,1328]" metricMagnitude="4" metricUnit="m" metricValue="2.84" pageId="53" pageNumber="54" unit="km" value="28.4">28.4 km</quantity>
99° Anivorano-
<location id="8EB46076E85EFFBF31A64D1EFF7EFAAF" LSID="urn:lsid:plazi:treatment:03C287BBE858FFBF31A64E7EFDF6FA3A:8EB46076E85EFFBF31A64D1EFF7EFAAF" box="[151,212,1340,1365]" municipality="Reserve Analamerana" name="Nord" pageId="53" pageNumber="54" stateProvince="Antsiranana">Nord</location>
,
<quantity id="4C939B48E85EFFBF31EE4D1EFEB0FAAE" box="[223,282,1340,1365]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="53" pageNumber="54" unit="m" value="60.0">
<elevation id="0046D19EE85EFFBF31EE4D1EFEB0FAAE" box="[223,282,1340,1365]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="53" pageNumber="54" unit="m" value="60.0">60 m</elevation>
</quantity>
, -12.74667, 49.49483, tropical dry forest, (
<collectorName id="269E537BE85EFFBF33CE4D1EFC2AFAAF" box="[767,896,1340,1365]" pageId="53" pageNumber="54">B.L. Fisher</collectorName>
) (
<collectionCode id="ED7AAE68E85EFFBF32A64D1EFC4CFAAE" box="[919,998,1340,1364]" pageId="53" pageNumber="54">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CF0E85EFFBF32CB4D1EFA3DFA82" ID-GBIF-Occurrence="1851012740" collectionCode="CASC" elevation="210" location="Reserve Speciale de l'Ankarana" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Antsiranana">
<location id="8EB46076E85EFFBF32CB4D1EFACAFAAF" LSID="urn:lsid:plazi:treatment:03C287BBE858FFBF31A64E7EFDF6FA3A:8EB46076E85EFFBF32CB4D1EFACAFAAF" box="[1018,1376,1340,1365]" name="Reserve Speciale de l'Ankarana" pageId="53" pageNumber="54" stateProvince="Antsiranana">Réserve Spéciale de l'Ankarana</location>
,
<locationDeviation id="9A662F8FE85EFFBF345A4D1EFE53FA82" pageId="53" pageNumber="54">
<quantity id="4C939B48E85EFFBF345A4D1EFF11FA82" metricMagnitude="4" metricUnit="m" metricValue="1.3599999999999999" pageId="53" pageNumber="54" unit="km" value="13.6">13.6 km</quantity>
192° SSW Anivorano Nord
</locationDeviation>
,
<quantity id="4C939B48E85EFFBF33354D42FDE7FA82" box="[516,589,1376,1400]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="53" pageNumber="54" unit="m" value="210.0">
<elevation id="0046D19EE85EFFBF33354D42FDE7FA82" box="[516,589,1376,1400]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="53" pageNumber="54" unit="m" value="210.0">210 m</elevation>
</quantity>
, -12.86361, 49.22583, tropical dry forest, (Fisher, Griswold
<emphasis id="B91FEABFE85EFFBF35C14D43FA82FA82" box="[1264,1320,1375,1400]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
) (
<collectionCode id="ED7AAE68E85EFFBF34714D42FA25FA82" box="[1344,1423,1376,1400]" pageId="53" pageNumber="54">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CF0E85EFFBF31A64DA6FDF2FA3A" ID-GBIF-Occurrence="1851012599" collectionCode="CASC" elevation="80" location="Reserve Speciale de l'Ankarana" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Antsiranana">
<location id="8EB46076E85EFFBF31A64DA6FDA3FA67" LSID="urn:lsid:plazi:treatment:03C287BBE858FFBF31A64E7EFDF6FA3A:8EB46076E85EFFBF31A64DA6FDA3FA67" box="[151,521,1412,1437]" name="Reserve Speciale de l'Ankarana" pageId="53" pageNumber="54" stateProvince="Antsiranana">Réserve Spéciale de l'Ankarana</location>
,
<locationDeviation id="9A662F8FE85EFFBF33294DA6FC12FA67" box="[536,952,1412,1437]" pageId="53" pageNumber="54">
<quantity id="4C939B48E85EFFBF33294DA6FDD2FA67" box="[536,632,1412,1437]" metricMagnitude="4" metricUnit="m" metricValue="2.29" pageId="53" pageNumber="54" unit="km" value="22.9">22.9 km</quantity>
224° SW Anivorano Nord
</locationDeviation>
,
<quantity id="4C939B48E85EFFBF32F64DA6FBACFA66" box="[967,1030,1412,1436]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="53" pageNumber="54" unit="m" value="80.0">
<elevation id="0046D19EE85EFFBF32F64DA6FBACFA66" box="[967,1030,1412,1436]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="53" pageNumber="54" unit="m" value="80.0">80 m</elevation>
</quantity>
, -12.90889, 49.10983, tropical dry forest, (Fisher, Griswold
<emphasis id="B91FEABFE85EFFBF30814D8BFE42FA3A" box="[432,488,1447,1472]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
) (
<collectionCode id="ED7AAE68E85EFFBF33314D8AFDE5FA3A" box="[512,591,1448,1472]" pageId="53" pageNumber="54">CASC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>