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<document ID-DOI="http://doi.org/10.5281/zenodo.5193638" ID-GBIF-Dataset="a02f182b-32e4-4649-afe0-983cb21ceb9e" ID-GBIF-Taxon="160690030" ID-Zenodo-Dep="5193638" approvalRequired="5" approvalRequired_for_illustrations="3" approvalRequired_for_treatments="2" checkinTime="1576077498408" checkinUser="jeremy" docAuthor="M. A. Lieftinck" docDate="1940" docId="03B487FFFFB9FFDDFEF3B67DFD09AC3F" docLanguage="en" docName="Lieftinck - 1940 - Descriptions and records of South-east Asiatic Odonata (II) ABBYYed.pdf.imf" docOrigin="Treubia 17 (4)" docStyle="DocumentStyle{}" docTitle="Amphicnemis mariae M.A. Lieftinck 1940, sp. n." docType="treatment" docVersion="14" lastPageId="41" lastPageNumber="376" masterDocId="FF8DFF87FF9EFFF4FFF0BE36FF92AF52" masterDocTitle="Descriptions and records of South-East Asiatic Odonata (II)" masterLastPageNumber="392" masterPageNumber="337" pageId="39" pageNumber="374" updateTime="1632324461921" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Descriptions and records of South-East Asiatic Odonata (II)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>M. A. Lieftinck</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Treubia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>1940</mods:date>
<mods:detail type="pubDate">
<mods:number>1940-01-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>17</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>337</mods:start>
<mods:end>392</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3571116</mods:identifier>
<mods:identifier type="GBIF-Dataset">a02f182b-32e4-4649-afe0-983cb21ceb9e</mods:identifier>
<mods:identifier type="Zenodo-Dep">3571116</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5193638" ID-GBIF-Taxon="160690030" ID-Zenodo-Dep="5193638" LSID="urn:lsid:plazi:treatment:03B487FFFFB9FFDDFEF3B67DFD09AC3F" httpUri="http://treatment.plazi.org/id/03B487FFFFB9FFDDFEF3B67DFD09AC3F" lastPageId="41" lastPageNumber="376" pageId="39" pageNumber="374">
<subSubSection box="[259,1107,2123,2163]" pageId="39" pageNumber="374" type="nomenclature">
<paragraph blockId="39.[253,1753,198,2645]" box="[259,1107,2123,2163]" pageId="39" pageNumber="374">
<taxonomicName authority="M.A. Lieftinck, 1940" authorityName="M.A. Lieftinck" authorityYear="1940" box="[259,681,2123,2161]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="374" phylum="Arthropoda" rank="species" species="mariae" status="sp . n .">
<emphasis bold="true" box="[259,681,2123,2161]" pageId="39" pageNumber="374">Amphicnemis mariae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[721,814,2129,2162]" pageId="39" pageNumber="374">sp. n.</taxonomicNameLabel>
(
<figureCitation box="[854,1076,2124,2163]" captionStart="Pl" captionStartId="28.[749,785,1761,1789]" captionTargetBox="[510,2495,140,1742]" captionTargetPageId="28" captionText="Pl . 15 . — Structural characters of Malaysian species of Amphicnemis . 1 - 3 . Male anal appendages , dorsal view and right side . 4 . Right lateral view of prothorax ." figureDoi="http://doi.org/10.5281/zenodo.3571151" httpUri="https://zenodo.org/record/3571151/files/figure.png" pageId="39" pageNumber="374">pl. 15 fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="374" type="materials_examined">
<paragraph blockId="39.[253,1753,198,2645]" pageId="39" pageNumber="374">
<materialsCitation ID-GBIF-Occurrence="2467684771" collectingCode="Buitenzorg Museum" collectingDate="1937-05" collectingDateMax="1937-06" collectingDateMin="1937-05" collectorName="M. E. WALSH" country="Malaysia" location="N. Koetai, Sangkoelirang distr., Pelawan Besar &amp; Kariorang" pageId="39" pageNumber="374" specimenCount="3" specimenCount-male="3" stateProvince="E. Borneo">
Material studied: —
<specimenCount box="[740,802,2181,2217]" pageId="39" pageNumber="374" type="male">
<emphasis bold="true" box="[740,759,2188,2217]" pageId="39" pageNumber="374">3</emphasis>
</specimenCount>
(one ad.),
<collectingRegion box="[1014,1249,2187,2217]" pageId="39" pageNumber="374">E. Borneo</collectingRegion>
,
<collectingMunicipality box="[1280,1462,2186,2217]" pageId="39" pageNumber="374">N. Koetai</collectingMunicipality>
,
<collectingMunicipality pageId="39" pageNumber="374">Sangkoelirang distr</collectingMunicipality>
.,
<location LSID="urn:lsid:plazi:treatment:03B487FFFFB9FFDDFEF3B67DFD09AC3F:8EC26032FFB9FFD3FE8AB68BFC1DA7B7" box="[378,911,2237,2277]" country="Borneo" municipality="N. Koetai" name="Pelawan Besar &amp; Kariorang" pageId="39" pageNumber="374" stateProvince="E. Borneo">Pelawan Besar &amp; Kariorang</location>
,
<collectingDate box="[942,1242,2240,2278]" pageId="39" pageNumber="374">May -June, 1937</collectingDate>
,
<collectorName box="[1275,1528,2240,2271]" pageId="39" pageNumber="374">
M. E.
<emphasis box="[1405,1528,2240,2270]" italics="true" pageId="39" pageNumber="374">WALSH</emphasis>
</collectorName>
leg., in the Buitenzorg Museum.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2467684777" box="[665,1506,2292,2332]" collectingDate="1937-05" collectorName="M. E. WALSH" country="Malaysia" location="Pelawan Besar" pageId="39" pageNumber="374" specimenCount="1" specimenCount-male="1" stateProvince="E. Borneo" typeStatus="holotype">
<typeStatus box="[665,909,2292,2331]" pageId="39" pageNumber="374">Holotype</typeStatus>
<specimenCount box="[932,956,2292,2322]" pageId="39" pageNumber="374" type="male"></specimenCount>
:
<location LSID="urn:lsid:plazi:treatment:03B487FFFFB9FFDDFEF3B67DFD09AC3F:8EC26032FFB9FFD3FC17B6C3FA92A647" box="[999,1280,2293,2325]" country="Borneo" municipality="N. Koetai" name="Pelawan Besar" pageId="39" pageNumber="374" stateProvince="E. Borneo">Pelawan Besar</location>
,
<date box="[1316,1506,2294,2332]" pageId="39" pageNumber="374" value="1937-05">
<collectingDate box="[1316,1506,2294,2332]" pageId="39" pageNumber="374" value="1937-05">May 1937</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="40" lastPageNumber="375" pageId="39" pageNumber="374" type="description">
<paragraph blockId="39.[253,1753,198,2645]" pageId="39" pageNumber="374">
<emphasis box="[338,430,2389,2421]" italics="true" pageId="39" pageNumber="374">Male</emphasis>
(ad.), - Head coloured as described for
<taxonomicName authorityName="M.A. Lieftinck" authorityYear="1940" box="[1284,1389,2401,2422]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="374" phylum="Arthropoda" rank="species" species="annae">
<emphasis box="[1284,1389,2401,2422]" italics="true" pageId="39" pageNumber="374">annae</emphasis>
</taxonomicName>
, but labrum with anterior, ochreous, margin broader and with the yellow fascia in front of frons barely interrupted in the middle line.
</paragraph>
<paragraph blockId="39.[253,1753,198,2645]" lastBlockId="40.[194,1699,174,2611]" lastPageId="40" lastPageNumber="375" pageId="39" pageNumber="374">Colouring of pro- and synthorax as described in the key. In all three specimens there is a fine comma-shaped yellow spot lying close against the upper margin of the mesepimeron, on each side of the dorsal end of the humeral suture.</paragraph>
<paragraph blockId="40.[194,1699,174,2611]" pageId="40" pageNumber="375">
M.
<emphasis bold="true" box="[387,415,177,206]" pageId="40" pageNumber="375">A</emphasis>
. Lieftinck: Descriptions
<emphasis box="[901,1110,179,211]" italics="true" pageId="40" pageNumber="375">and records</emphasis>
0
<emphasis box="[1148,1202,181,218]" italics="true" pageId="40" pageNumber="375">ƒ S</emphasis>
.
<emphasis box="[1213,1241,182,212]" italics="true" pageId="40" pageNumber="375">E</emphasis>
.
<emphasis box="[1268,1565,182,217]" italics="true" pageId="40" pageNumber="375">
Asiatic
<taxonomicName authorityName="Fabricius" authorityYear="1793" box="[1414,1565,185,217]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="375" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
</emphasis>
,
<emphasis bold="true" box="[1635,1696,189,219]" pageId="40" pageNumber="375">
<pageNumber ambiguity="12" box="[1635,1696,189,219]" fuzzyness="0" pageId="40" pageNumber="375" score="146.0" value="375">375</pageNumber>
</emphasis>
ÎÎ
</paragraph>
<paragraph blockId="40.[194,1699,174,2611]" pageId="40" pageNumber="375">The squarish lustrous spot covering the upper part of the mesepisternum is separated from the dark colour on the mesepimeron by a fine yellow ling Sides and under surfaces pale yellow.</paragraph>
<paragraph blockId="40.[194,1699,174,2611]" pageId="40" pageNumber="375">Legs pale yellow, apices of all femora and tibiae, spines and tips of tarsal joints, black.</paragraph>
<paragraph blockId="40.[194,1699,174,2611]" box="[286,1699,648,721]" pageId="40" pageNumber="375">Neuration identical to the other species of the same group. Postnodals13-14/11-12,</paragraph>
<paragraph blockId="40.[194,1699,174,2611]" pageId="40" pageNumber="375">
<emphasis box="[202,418,702,739]" italics="true" pageId="40" pageNumber="375">Pterostigma</emphasis>
greyish-brown, with pale yellow margin. Abdomen coloured as in
<taxonomicName authorityName="Selys" authorityYear="1863" box="[739,892,761,803]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="375" phylum="Arthropoda" rank="species" species="wallacei">
<emphasis box="[739,892,761,803]" italics="true" pageId="40" pageNumber="375">wallacei</emphasis>
</taxonomicName>
; distal margin of segment 10 whitish. Anal appendages yellowish-white, shaped as shown on
<figureCitation box="[1169,1403,819,860]" captionStart="Pl" captionStartId="28.[749,785,1761,1789]" captionTargetBox="[510,2495,140,1742]" captionTargetPageId="28" captionText="Pl . 15 . — Structural characters of Malaysian species of Amphicnemis . 1 - 3 . Male anal appendages , dorsal view and right side . 4 . Right lateral view of prothorax ." figureDoi="http://doi.org/10.5281/zenodo.3571151" httpUri="https://zenodo.org/record/3571151/files/figure.png" pageId="40" pageNumber="375">pl. 15 fig. 1</figureCitation>
.
</paragraph>
<paragraph blockId="40.[194,1699,174,2611]" box="[283,609,862,898]" pageId="40" pageNumber="375">
<emphasis box="[283,414,862,892]" italics="true" pageId="40" pageNumber="375">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="375" type="discussion">
<paragraph blockId="40.[194,1699,174,2611]" pageId="40" pageNumber="375">
this
<emphasis box="[389,455,990,1041]" italics="true" pageId="40" pageNumber="375">new</emphasis>
Species is easily distinguished from its allies by the reduction of the metallic-green colour on the dorsum of the pro- and synthorax, and by the anal appendages, which appear to resemble those of
<taxonomicName authorityName="Laidlaw" authorityYear="1912" box="[1281,1417,1108,1147]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="375" phylum="Arthropoda" rank="species" species="remiger">
<emphasis box="[1281,1417,1108,1147]" italics="true" pageId="40" pageNumber="375">remiger</emphasis>
</taxonomicName>
fairly closely. Named in honour of Mrs. M. E. Walsh, in appreciation of her constant help in collecting
<taxonomicName authorityName="Fabricius" authorityYear="1793" box="[551,704,1212,1249]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="375" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
.
<taxonomicName authorityName="Ris" authorityYear="1911" box="[200,635,1332,1376]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="375" phylum="Arthropoda" rank="species" species="martini">
<emphasis bold="true" box="[200,635,1332,1376]" pageId="40" pageNumber="375">Amphicnemis martini</emphasis>
</taxonomicName>
Ris.
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="375" type="reference_group">
<paragraph blockId="40.[194,1699,174,2611]" pageId="40" pageNumber="375">
1911. Ris, Ann. Soc. ent. Belg. 55, p. 236 (key ♂), 237 238,
<figureCitation box="[1262,1353,1414,1445]" captionStart="Fig" captionStartId="15.[264,324,1892,1924]" captionTargetBox="[243,1351,1155,1881]" captionTargetPageId="15" captionText="Fig. 6. Drepanosticta barbatula, sp. n. ♂. Anal appendages, dorsal view and right side." figureDoi="http://doi.org/10.5281/zenodo.3571130" httpUri="https://zenodo.org/record/3571130/files/figure.png" pageId="40" pageNumber="375">fig. 6</figureCitation>
(apps. ♂). — ♂ W. Borneo
</paragraph>
<paragraph blockId="40.[194,1699,174,2611]" box="[202,1379,1499,1536]" pageId="40" pageNumber="375">
1913.
<emphasis box="[314,455,1499,1527]" italics="true" pageId="40" pageNumber="375">LAIDLAW</emphasis>
, P.
<emphasis box="[519,544,1503,1528]" italics="true" pageId="40" pageNumber="375">Z</emphasis>
. S. London, p. 72 73; 74 (key). - ♂♀ Sarawak.
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="375" type="materials_examined">
<paragraph blockId="40.[194,1699,174,2611]" pageId="40" pageNumber="375">
<materialsCitation ID-GBIF-Occurrence="2467684743" collectingCode="Buitenzorg Museum" collectingDate="1931-12-07" collectingDateMax="1932-07-19" collectingDateMin="1931-12-07" collectorName="L. Coomans DE RUITER" country="Malaysia" location="Singkawang, forest-marsh near Bakoean" pageId="40" pageNumber="375" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="W. Borneo">
Material studied: -
<specimenCount box="[694,757,1605,1636]" pageId="40" pageNumber="375" type="male">5 ♂</specimenCount>
(semiad.-ad.),
<specimenCount box="[1052,1112,1605,1637]" pageId="40" pageNumber="375" type="female">6 ♀</specimenCount>
(2 juv., red, 4 ad., blue),
<collectingRegion pageId="40" pageNumber="375">W. Borneo</collectingRegion>
,
<collectingMunicipality box="[407,623,1659,1698]" pageId="40" pageNumber="375">Singkawang</collectingMunicipality>
, forest-marsh near
<location LSID="urn:lsid:plazi:treatment:03B487FFFFB9FFDDFEF3B67DFD09AC3F:8EC26032FFB6FFDCFC17B84BFB16A9C9" box="[999,1156,1661,1691]" country="Borneo" municipality="Singkawang" name="Bakoean" pageId="40" pageNumber="375" stateProvince="W. Borneo">Bakoean</location>
,
<date box="[1187,1415,1662,1700]" pageId="40" pageNumber="375" value="1931-null-07">
<collectingDate box="[1187,1415,1662,1700]" pageId="40" pageNumber="375" value="1931-null-07">Dec. 7, 1931</collectingDate>
</date>
,
<collectingDate pageId="40" pageNumber="375">
Febr.
<emphasis bold="true" box="[1565,1681,1664,1697]" pageId="40" pageNumber="375">17 - 19</emphasis>
, July 19
</collectingDate>
,
<collectingDate box="[378,454,1718,1747]" pageId="40" pageNumber="375">1932</collectingDate>
,
<collectorName box="[489,924,1716,1747]" pageId="40" pageNumber="375">
L. Coomans
<emphasis box="[731,772,1725,1746]" italics="true" pageId="40" pageNumber="375">DE</emphasis>
RUITER
</collectorName>
leg., in the Buitenzorg Museum
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="41" lastPageNumber="376" pageId="40" pageNumber="375" type="description">
<paragraph blockId="40.[194,1699,174,2611]" pageId="40" pageNumber="375">
<emphasis box="[280,372,1770,1801]" italics="true" pageId="40" pageNumber="375">Male</emphasis>
. — These specimens agree exactly with Riss account of
<taxonomicName authorityName="Ris" authorityYear="1911" box="[1485,1681,1772,1803]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="375" phylum="Arthropoda" rank="species" species="martini">
<emphasis box="[1485,1511,1773,1802]" italics="true" pageId="40" pageNumber="375">A</emphasis>
.
<emphasis box="[1546,1681,1772,1803]" italics="true" pageId="40" pageNumber="375">martini</emphasis>
</taxonomicName>
, save that in the fully adult examples the colour of the pterostigmata of the fore wings is deep greenish-black centred with dark brown and with the border yellow, whilst those of the hinder wings are throughout bright orange. There are 14-15/13 postnodal cross-nerves. In the semi-adult males the thoracic sides and legs are yellow.
</paragraph>
<paragraph blockId="40.[194,1699,174,2611]" pageId="40" pageNumber="375">
The colour of the tenth abdominal segment varies from yellow to pale blue. Anal appendages wax-yellow to pale brown. Superior pair decidedly shorter than in
<taxonomicName authorityName="Selys" authorityYear="1863" box="[346,494,2241,2273]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="375" phylum="Arthropoda" rank="species" species="wallacei">
<emphasis box="[346,494,2241,2273]" italics="true" pageId="40" pageNumber="375">wallacei</emphasis>
</taxonomicName>
and allied species, each with a distinct triangular protuberance at about the middle of their length, directed obliquely inwards and upwards, followed by a similar (slightly more robust) triangular expansion, which projects horizontally inwards; the median incision between these two projections (Ris, loc.
<emphasis box="[272,317,2463,2494]" italics="true" pageId="40" pageNumber="375">cit</emphasis>
.
<figureCitation box="[344,436,2463,2501]" captionStart="Fig" captionStartId="15.[264,324,1892,1924]" captionTargetBox="[243,1351,1155,1881]" captionTargetPageId="15" captionText="Fig. 6. Drepanosticta barbatula, sp. n. ♂. Anal appendages, dorsal view and right side." figureDoi="http://doi.org/10.5281/zenodo.3571130" httpUri="https://zenodo.org/record/3571130/files/figure.png" pageId="40" pageNumber="375">fig. 6</figureCitation>
) is visible only in oblique dorsal view. Inferior pair of appendages only half a long as the superiors, cylindrical, tapering, slightly divergent, tips evenly and but slightly upcurved.
</paragraph>
<paragraph blockId="41.[251,1770,196,2645]" box="[251,1301,196,234]" pageId="41" pageNumber="376">
<pageNumber ambiguity="12" box="[251,313,198,227]" fuzzyness="0" pageId="41" pageNumber="376" score="147.0" value="376">376</pageNumber>
TREUBIA Deel 17, 1940,
<emphasis box="[1170,1241,196,226]" italics="true" pageId="41" pageNumber="376">AFL</emphasis>
.
<emphasis bold="true" box="[1273,1291,197,226]" pageId="41" pageNumber="376">4</emphasis>
.
</paragraph>
<paragraph blockId="41.[251,1770,196,2645]" pageId="41" pageNumber="376">Female (ad.). - Legs pale yellow, femora striped with black exteriorly and with the extremities also finely black. Anterior pair of tibiae with the basal one-third of the outer surfaces black.</paragraph>
<paragraph blockId="41.[251,1770,196,2645]" box="[340,1347,462,502]" pageId="41" pageNumber="376">Valves not projecting beyond the apex of abdomen.</paragraph>
<paragraph blockId="41.[251,1770,196,2645]" pageId="41" pageNumber="376">Female (juv.). - The red colour-phase of this species is peculiar for the brilliant metallic-green episterna of the mesothorax, which, in the red-bodied females of other species, are of the same colour as the rest of the thorax. There is also a median bronze-green spot on the middle of the prothorax that covers the whole of the posterior lobe as well.</paragraph>
<paragraph blockId="41.[251,1770,196,2645]" pageId="41" pageNumber="376">The coxae, tibiae and tarsi are yellowish whilst the femora are red, lacking black exterior stripes.</paragraph>
</subSubSection>
</treatment>
</document>