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<document id="19494822DDB79ACAD398EE0A7C201AF3" ID-CLB-Dataset="21895" ID-DOI="10.5852/ejt.2016.219" ID-GBIF-Dataset="f80b497f-b264-4b3f-86b7-0918ecbec68f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3840226" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590168918509" checkinUser="valdenar" docAuthor="Jiruskova, Alice, Motyka, Michal &amp; Bocak, Ladislav" docDate="2016" docId="03B487FEF035FFCCFDF3A032FE27F9CE" docLanguage="en" docName="ejt-2016-219.pdf.imf" docOrigin="European Journal of Taxonomy 219 (219)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cautires Waterhouse 1879" docType="treatment" docVersion="7" lastPageNumber="5" masterDocId="FF8DFF86F031FFC8FF8AA21CFFA7FFEB" masterDocTitle="High diversity and endemism in the genus Cautires Waterhouse, 1879 (Coleoptera: Lycidae) from the Malay mountain forests, with the descriptions of fourteen new species" masterLastPageNumber="29" masterPageNumber="1" pageNumber="5" updateTime="1698841051393" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="43CDC687C2DCD3DA0DC44390E0E30CFF">High diversity and endemism in the genus Cautires Waterhouse, 1879 (Coleoptera: Lycidae) from the Malay mountain forests, with the descriptions of fourteen new species</mods:title>
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<mods:namePart id="9E328EAE5B38B949D166440A0A30E9CE">Jiruskova, Alice</mods:namePart>
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<mods:affiliation id="6261FB5C83B6814DA29FC61174213A81">Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, 771 46 Olomouc, Czech Republic. &amp; Email: alice. jiruskova @ seznam. cz &amp; urn: lsid: zoobank. org: author: E 8 EA 3180 - 2399 - 4 EA 9 - 86 BD-DBFC 381 E 2 F 93</mods:affiliation>
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<mods:namePart id="C9C5C9246FEF0E5E80DCD46CB18E6EBA">Motyka, Michal</mods:namePart>
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<mods:affiliation id="E0679199F8187006011A5D5F25CD9578">Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, 771 46 Olomouc, Czech Republic. &amp; Email: michal. motyka @ upol. cz &amp; urn: lsid: zoobank. org: author: 242 DF 8 C 7 - 1 E 68 - 45 F 7 - AD 24 - C 4 BC 0 C 5 D 0 D 94</mods:affiliation>
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<mods:namePart id="FD66D16A2202AB397C47D848992D7282">Bocak, Ladislav</mods:namePart>
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<mods:affiliation id="081A77190660A5B5EE18146C9FAB2C9E">Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, 771 46 Olomouc, Czech Republic. &amp; urn: lsid: zoobank. org: author: 78383404 - FF 9 F- 46 EA-A 1 AE-ACF 3613 E 1 D 69 &amp; corresponding author: ladislav. bocak @ upol. cz</mods:affiliation>
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<subSubSection id="C3076563F035FFCCFDF3A032FC1CFDA2" box="[633,955,558,585]" pageId="4" pageNumber="5" type="nomenclature">
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<taxonomicName id="4C1D4D6BF035FFCCFDF3A032FC1CFDA2" authority="Waterhouse" authorityName="Waterhouse" authorityYear="1879" box="[633,955,558,585]" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFDF3A032FD47FDA3" bold="true" box="[633,736,558,584]" italics="true" pageId="4" pageNumber="5">Cautires</emphasis>
<bibRefCitation id="EF8C4B19F035FFCCFD6DA032FC1CFDA2" author="Waterhouse C. O." box="[743,955,558,585]" pageId="4" pageNumber="5" refId="ref15539" refString="Waterhouse C. O. 1879. Illustration of the Typical Specimens of Coleoptera in the Collection of the British Museum. Part I. Lycidae. British Museum, London." type="book" year="1879">Waterhouse, 1879</bibRefCitation>
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<figureCitation id="13262A6DF035FFCCFD39A04EFC8CFD87" box="[691,811,594,620]" captionStart="Figs 2–20" captionStartId="3.[189,241,1671,1697]" captionTargetBox="[189,1396,1131,1609]" captionTargetId="figure@3.[189,1398,1126,1615]" captionTargetPageId="3" captionText="Figs 2–20. Cautires sp. 2. Habitus in dorsal view. 3–20. Male antennae. 3. C. alexae sp. nov. 4. C. andujari sp. nov. 5. C. arribasae sp. nov. 6. C. berembanensis sp. nov. 7. C. campestris sp. nov. 8. C. communis sp. nov. 9. C. griseus Kleine, 1930. 10. C. jasarensis sp. nov. 11. C. katarinae sp. nov. 12. C. kirstenae sp. nov. 13. C. kotatinggensis sp. nov. 14. C. linardi sp. nov. 15. C. maseki sp. nov. 16. C. pahangensis sp. nov. 17. C. nervosus Kleine, 1926. 18. C. renatae sp. nov. 19. C. reverandi Pic, 1925. 20. C. simillimus Kleine, 1926. Scale bars: Fig. 2 = 2.0 mm; Figs 3–20 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3840230" httpUri="https://zenodo.org/record/3840230/files/figure.png" pageId="4" pageNumber="5">Figs 6–20</figureCitation>
,
<figureCitation id="13262A6DF035FFCCFCBDA04EFC27FD87" box="[823,896,594,620]" captionStart-0="Figs 41–58" captionStart-1="Figs 59–78" captionStartId-0="9.[190,242,1822,1848]" captionStartId-1="12.[189,241,1824,1850]" captionTargetBox-0="[193,1395,820,1772]" captionTargetBox-1="[192,1392,492,1770]" captionTargetId-0="figure@9.[189,1398,818,1776]" captionTargetId-1="figure@12.[189,1398,491,1781]" captionTargetPageId-0="9" captionTargetPageId-1="12" captionText-0="Figs 41–58. Male elytra. 41. Cautires alexae sp. nov. 42. C. andujari sp. nov. 43. C. arribasae sp. nov. 44. C. berembanensis sp. nov. 45. C. campestris sp. nov. 46. C. communis sp. nov. 47. C. griseus Kleine, 1930. 48. C. jasarensis sp. nov. 49. C. katarinae sp. nov. 50. C. kirstenae sp. nov. 51. C. kotatinggensis sp. nov. 52. C. linardi sp. nov. 53. C. maseki sp. nov. 54. C. pahangensis sp. nov. 55. C. nervosus Kleine, 1926. 56. C. renatae sp. nov. 57. C. reverandi Pic, 1925. 58. C. simillimus Kleine, 1926. Scale bars = 1.0 mm." captionText-1="Figs 59–78. Phallus. 59. Cautires alexae sp. nov. 60. C. andujari sp. nov. 61. C. arribasae sp. nov. 62. C. berembanensis sp. nov. 63. C. campestris sp. nov. 64. C. communis sp. nov. 65. C. griseus Kleine, 1930. 66. C. jasarensis sp. nov. 67. C. katarinae sp. nov. 68. C. kirstenae sp. nov. 69. C. kotatinggensis sp. nov. 70. C. linardi sp. nov. 71. C. maseki sp. nov. 72. C. pahangensis sp. nov. 73. C. nervosus Kleine, 1926. 74. C. renatae sp. nov. 75. C. reverandi Pic, 1925. 76. C. simillimus Kleine, 1926. 77. C. selangorensis Kleine, 1930. 78. C. tenebricus Kleine, 1926. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3840234" figureDoi-1="http://doi.org/10.5281/zenodo.3840236" httpUri-0="https://zenodo.org/record/3840234/files/figure.png" httpUri-1="https://zenodo.org/record/3840236/files/figure.png" pageId="4" pageNumber="5">41–78</figureCitation>
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<treatmentCitation id="0ABC10F9F035FFCCFF37A088FD81FD46" author="Waterhouse C. O." box="[189,550,659,686]" page="36" pageId="4" pageNumber="5" year="1879">
<taxonomicName id="4C1D4D6BF035FFCCFF37A088FD81FD46" authority="Waterhouse" authorityName="Waterhouse" authorityPageNumber="36" authorityYear="1879" box="[189,550,659,686]" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFF37A088FE86FD45" box="[189,289,660,686]" italics="true" pageId="4" pageNumber="5">Cautires</emphasis>
<bibRefCitation id="EF8C4B19F035FFCCFEACA08FFD81FD46" author="Waterhouse C. O." box="[294,550,659,685]" pageId="4" pageNumber="5" refId="ref15539" refString="Waterhouse C. O. 1879. Illustration of the Typical Specimens of Coleoptera in the Collection of the British Museum. Part I. Lycidae. British Museum, London." type="book" year="1879">Waterhouse, 1879: 36</bibRefCitation>
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species:
<taxonomicName id="4C1D4D6BF035FFCCFD58A088FCB1FD46" ID-CoL="5HML" box="[722,790,660,685]" class="Insecta" family="Lycidae" genus="Lycus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFD58A088FCB1FD46" box="[722,790,660,685]" italics="true" pageId="4" pageNumber="5">Lycus</emphasis>
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(
<date id="FFA31028F035FFCCFCAEA08FFCDCFD46" box="[804,891,659,685]" pageId="4" pageNumber="5" value="1922-01">gen. 22</date>
)
<emphasis id="B969EAFAF035FFCCFC03A08FFC52FD46" box="[905,1013,659,685]" italics="true" pageId="4" pageNumber="5">excellens</emphasis>
<bibRefCitation id="EF8C4B19F035FFCCFC70A08FFB6BFD46" author="Waterhouse C. O." box="[1018,1228,659,685]" pageId="4" pageNumber="5" pagination="95 - 118" refId="ref15481" refString="Waterhouse C. O. 1878. On the different forms occurring in the Colepterous family Lycidae, with descriptions of new genera and species. Transaction of the Entomological Society of London 1: 95 - 118. Available from http: // biodiversitylibrary. org / page / 12552290 [accessed 6 Jul. 2016]" type="journal article" year="1878">Waterhouse, 1878</bibRefCitation>
by subsequent designation:
<treatmentCitation id="0ABC10F9F035FFCCFEDEA0AAFDEAFD3A" author="Bourgeois J. M." box="[340,589,694,721]" page="345" pageId="4" pageNumber="5" year="1891">
<bibRefCitation id="EF8C4B19F035FFCCFEDEA0AAFDEAFD3A" author="Bourgeois J. M." box="[340,589,694,721]" pageId="4" pageNumber="5" pagination="337 - 364" refId="ref14383" refString="Bourgeois J. M. 1891. Etudes sur la distribution geographique des Malacodermes. I. Lycides. Annales de la Societe entomologique de France 60: 337 - 364. Available from http: // biodiversitylibrary. org / page / 32547697 [accessed 6 Jul. 2016]" type="journal article" year="1891">Bourgeois 1891: 345</bibRefCitation>
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<treatmentCitation id="0ABC10F9F035FFCCFF37A0C5FDF4FD1F" author="Waterhouse C. O." box="[189,595,729,756]" page="34" pageId="4" pageNumber="5" year="1879">
<taxonomicName id="4C1D4D6BF035FFCCFF37A0C5FDF4FD1F" authority="Waterhouse, 1879: 34" authorityName="Waterhouse" authorityPageNumber="34" authorityYear="1879" box="[189,595,729,756]" class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFF37A0C5FE96FD18" box="[189,305,729,755]" italics="true" pageId="4" pageNumber="5">Bulenides</emphasis>
<bibRefCitation id="EF8C4B19F035FFCCFECAA0C5FDF4FD1F" author="Waterhouse C. O." box="[320,595,729,756]" pageId="4" pageNumber="5" refId="ref15539" refString="Waterhouse C. O. 1879. Illustration of the Typical Specimens of Coleoptera in the Collection of the British Museum. Part I. Lycidae. British Museum, London." type="book" year="1879">Waterhouse, 1879: 34</bibRefCitation>
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<typeStatus id="54A6884AF035FFCCFDECA0C6FD07FD1F" box="[614,672,730,756]" pageId="4" pageNumber="5">Type</typeStatus>
species:
<taxonomicName id="4C1D4D6BF035FFCCFC96A0C6FCC7FD18" ID-CoL="5HML" box="[796,864,730,755]" class="Insecta" family="Lycidae" genus="Lycus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFC96A0C6FCC7FD18" box="[796,864,730,755]" italics="true" pageId="4" pageNumber="5">Lycus</emphasis>
</taxonomicName>
(gen. 21)
<taxonomicName id="4C1D4D6BF035FFCCFC7BA0C5FAEDFD1F" authority="Waterhouse, 1878" authorityName="Waterhouse" authorityYear="1878" box="[1009,1354,729,756]" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="obsoletus">
<emphasis id="B969EAFAF035FFCCFC7BA0C5FBC7FD18" box="[1009,1120,729,755]" italics="true" pageId="4" pageNumber="5">obsoletus</emphasis>
<bibRefCitation id="EF8C4B19F035FFCCFBE4A0C5FAEDFD1F" author="Waterhouse C. O." box="[1134,1354,729,756]" pageId="4" pageNumber="5" pagination="95 - 118" refId="ref15481" refString="Waterhouse C. O. 1878. On the different forms occurring in the Colepterous family Lycidae, with descriptions of new genera and species. Transaction of the Entomological Society of London 1: 95 - 118. Available from http: // biodiversitylibrary. org / page / 12552290 [accessed 6 Jul. 2016]" type="journal article" year="1878">Waterhouse, 1878</bibRefCitation>
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by subsequent designation:
<treatmentCitation id="0ABC10F9F035FFCCFE57A0E1FD70FCFC" author="Bourgeois J. M." box="[477,727,764,791]" page="345" pageId="4" pageNumber="5" year="1891">
<bibRefCitation id="EF8C4B19F035FFCCFE57A0E1FD70FCFC" author="Bourgeois J. M." box="[477,727,764,791]" pageId="4" pageNumber="5" pagination="337 - 364" refId="ref14383" refString="Bourgeois J. M. 1891. Etudes sur la distribution geographique des Malacodermes. I. Lycides. Annales de la Societe entomologique de France 60: 337 - 364. Available from http: // biodiversitylibrary. org / page / 32547697 [accessed 6 Jul. 2016]" type="journal article" year="1891">Bourgeois 1891: 345</bibRefCitation>
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.
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</paragraph>
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<treatmentCitationGroup id="AB0D11C6F035FFCCFF37A122FD07FCB3" box="[189,672,830,857]" pageId="4" pageNumber="5">
<taxonomicName id="4C1D4D6BF035FFCCFF37A122FE96FCB3" authorityName="Waterhouse" authorityYear="1879" box="[189,305,830,856]" class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFF37A122FE96FCB3" box="[189,305,830,856]" italics="true" pageId="4" pageNumber="5">Bulenides</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABC10F9F035FFCCFEC4A122FD3BFCB3" author="Dudkova P. &amp; Bocak L." box="[334,668,830,856]" page="34" pageId="4" pageNumber="5" year="2010">
<bibRefCitation id="EF8C4B19F035FFCCFEC4A122FD3BFCB3" author="Dudkova P. &amp; Bocak L." box="[334,668,830,856]" pageId="4" pageNumber="5" pagination="28 - 48" refId="ref14434" refString="Dudkova P. &amp; Bocak L. 2010. A review of the Cautires obsoletus species group from Indo-Burma (Coleoptera: Lycidae). Zootaxa 2527: 28 - 48." type="journal article" year="2010">Dudkova &amp; Bocak 2010: 34</bibRefCitation>
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.
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</subSubSection>
<subSubSection id="C3076563F035FFCCFF37A163FE1DFB54" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BA236E8F035FFCCFF37A163FE91FC72" blockId="4.[189,310,895,921]" box="[189,310,895,921]" pageId="4" pageNumber="5">
<heading id="D0EA8184F035FFCCFF37A163FE91FC72" bold="true" box="[189,310,895,921]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B969EAFAF035FFCCFF37A163FE91FC72" bold="true" box="[189,310,895,921]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA236E8F035FFCCFF37A1B3FE1DFB54" blockId="4.[189,1399,942,1215]" pageId="4" pageNumber="5">
<taxonomicName id="4C1D4D6BF035FFCCFF37A1B3FE86FC22" authorityName="Waterhouse" authorityYear="1879" box="[189,289,943,969]" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFF37A1B3FE86FC22" box="[189,289,943,969]" italics="true" pageId="4" pageNumber="5">Cautires</emphasis>
</taxonomicName>
belongs to the subtribe
<taxonomicName id="4C1D4D6BF035FFCCFDA3A1B2FD31FC23" authorityName="Sklenarova, Kubecek &amp; Bocak" authorityYear="2014" box="[553,662,942,968]" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subTribe" subTribe="Cautirina">Cautirina</taxonomicName>
in close relationships to
<taxonomicName id="4C1D4D6BF035FFCCFC23A1B2FB82FC23" authorityName="Waterhouse" authorityYear="1879" box="[937,1061,942,968]" class="Insecta" family="Lycidae" genus="Xylobanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFC23A1B2FB82FC23" box="[937,1061,942,968]" italics="true" pageId="4" pageNumber="5">Xylobanus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C1D4D6BF035FFCCFBBBA1B2FB33FC23" authorityName="Laporte" authorityYear="1836" box="[1073,1172,942,968]" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Lycidae</taxonomicName>
:
<taxonomicName id="4C1D4D6BF035FFCCFB11A1B2FACDFC22" authorityName="Kleine" authorityYear="1926" box="[1179,1386,942,969]" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Metriorrhynchini">Metriorrhynchini</taxonomicName>
), from which it can be distinguished by continuous larval terga and a simple, usually slender phallus with a pair of sickle-shaped thorns in the internal sac (
<bibRefCitation id="EF8C4B19F035FFCCFC8FA1E8FBB3FBE4" author="Sklenarova K. &amp; Kubecek V. &amp; Bocak L." box="[773,1044,1012,1039]" pageId="4" pageNumber="5" pagination="37 - 54" refId="ref15307" refString="Sklenarova K., Kubecek V. &amp; Bocak L. 2014. Subtribal classification of Metriorrhynchini (Insecta: Coleoptera: Lycidae): an integrative approach using molecular phylogeny and morphology of adults and larvae. Arthropod Systematics and Phylogeny 72: 37 - 54." type="journal article" year="2014">
Sklenarova
<emphasis id="B969EAFAF035FFCCFC1BA1E9FC6BFBE4" box="[913,972,1013,1039]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
2014
</bibRefCitation>
). Almost all Oriental
<taxonomicName id="4C1D4D6BF035FFCCFA98A1E9FAD1FBE4" authorityName="Waterhouse" authorityYear="1879" box="[1298,1398,1013,1039]" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFA98A1E9FAD1FBE4" box="[1298,1398,1013,1039]" italics="true" pageId="4" pageNumber="5">Cautires</emphasis>
</taxonomicName>
have the male antennae flabellate (
<figureCitation id="13262A6DF035FFCCFDDAA604FD6DFBD9" box="[592,714,1048,1074]" captionStart="Figs 2–20" captionStartId="3.[189,241,1671,1697]" captionTargetBox="[189,1396,1131,1609]" captionTargetId="figure@3.[189,1398,1126,1615]" captionTargetPageId="3" captionText="Figs 2–20. Cautires sp. 2. Habitus in dorsal view. 3–20. Male antennae. 3. C. alexae sp. nov. 4. C. andujari sp. nov. 5. C. arribasae sp. nov. 6. C. berembanensis sp. nov. 7. C. campestris sp. nov. 8. C. communis sp. nov. 9. C. griseus Kleine, 1930. 10. C. jasarensis sp. nov. 11. C. katarinae sp. nov. 12. C. kirstenae sp. nov. 13. C. kotatinggensis sp. nov. 14. C. linardi sp. nov. 15. C. maseki sp. nov. 16. C. pahangensis sp. nov. 17. C. nervosus Kleine, 1926. 18. C. renatae sp. nov. 19. C. reverandi Pic, 1925. 20. C. simillimus Kleine, 1926. Scale bars: Fig. 2 = 2.0 mm; Figs 3–20 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3840230" httpUri="https://zenodo.org/record/3840230/files/figure.png" pageId="4" pageNumber="5">Figs 7–20</figureCitation>
), some species have very short lamellae and the antenna is acutely serrate (
<figureCitation id="13262A6DF035FFCCFEFEA627FE1CFBBD" box="[372,443,1083,1110]" captionStart="Figs 2–20" captionStartId="3.[189,241,1671,1697]" captionTargetBox="[189,1396,1131,1609]" captionTargetId="figure@3.[189,1398,1126,1615]" captionTargetPageId="3" captionText="Figs 2–20. Cautires sp. 2. Habitus in dorsal view. 3–20. Male antennae. 3. C. alexae sp. nov. 4. C. andujari sp. nov. 5. C. arribasae sp. nov. 6. C. berembanensis sp. nov. 7. C. campestris sp. nov. 8. C. communis sp. nov. 9. C. griseus Kleine, 1930. 10. C. jasarensis sp. nov. 11. C. katarinae sp. nov. 12. C. kirstenae sp. nov. 13. C. kotatinggensis sp. nov. 14. C. linardi sp. nov. 15. C. maseki sp. nov. 16. C. pahangensis sp. nov. 17. C. nervosus Kleine, 1926. 18. C. renatae sp. nov. 19. C. reverandi Pic, 1925. 20. C. simillimus Kleine, 1926. Scale bars: Fig. 2 = 2.0 mm; Figs 3–20 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3840230" httpUri="https://zenodo.org/record/3840230/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
). The female antennae are always serrate. Each elytron bears four primary and five secondary longitudinal costae (
<figureCitation id="13262A6DF035FFCCFDA1A642FD15FB92" box="[555,690,1118,1145]" captionStart="Figs 41–58" captionStartId="9.[190,242,1822,1848]" captionTargetBox="[193,1395,820,1772]" captionTargetId="figure@9.[189,1398,818,1776]" captionTargetPageId="9" captionText="Figs 41–58. Male elytra. 41. Cautires alexae sp. nov. 42. C. andujari sp. nov. 43. C. arribasae sp. nov. 44. C. berembanensis sp. nov. 45. C. campestris sp. nov. 46. C. communis sp. nov. 47. C. griseus Kleine, 1930. 48. C. jasarensis sp. nov. 49. C. katarinae sp. nov. 50. C. kirstenae sp. nov. 51. C. kotatinggensis sp. nov. 52. C. linardi sp. nov. 53. C. maseki sp. nov. 54. C. pahangensis sp. nov. 55. C. nervosus Kleine, 1926. 56. C. renatae sp. nov. 57. C. reverandi Pic, 1925. 58. C. simillimus Kleine, 1926. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3840234" httpUri="https://zenodo.org/record/3840234/files/figure.png" pageId="4" pageNumber="5">Figs 41–58</figureCitation>
) and numerous transverse costae connecting them.
<taxonomicName id="4C1D4D6BF035FFCCFA98A643FAD1FB92" authorityName="Waterhouse" authorityYear="1879" box="[1298,1398,1119,1145]" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFA98A643FAD1FB92" box="[1298,1398,1119,1145]" italics="true" pageId="4" pageNumber="5">Cautires</emphasis>
</taxonomicName>
species have the lanceolate phallus with membranous internal sac bearing two sickle-shaped thorns at its base (
<figureCitation id="13262A6DF035FFCCFEAEA6B8FE09FB54" box="[292,430,1188,1215]" captionStart="Figs 59–78" captionStartId="12.[189,241,1824,1850]" captionTargetBox="[192,1392,492,1770]" captionTargetId="figure@12.[189,1398,491,1781]" captionTargetPageId="12" captionText="Figs 59–78. Phallus. 59. Cautires alexae sp. nov. 60. C. andujari sp. nov. 61. C. arribasae sp. nov. 62. C. berembanensis sp. nov. 63. C. campestris sp. nov. 64. C. communis sp. nov. 65. C. griseus Kleine, 1930. 66. C. jasarensis sp. nov. 67. C. katarinae sp. nov. 68. C. kirstenae sp. nov. 69. C. kotatinggensis sp. nov. 70. C. linardi sp. nov. 71. C. maseki sp. nov. 72. C. pahangensis sp. nov. 73. C. nervosus Kleine, 1926. 74. C. renatae sp. nov. 75. C. reverandi Pic, 1925. 76. C. simillimus Kleine, 1926. 77. C. selangorensis Kleine, 1930. 78. C. tenebricus Kleine, 1926. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3840236" httpUri="https://zenodo.org/record/3840236/files/figure.png" pageId="4" pageNumber="5">Figs 59–78</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3076563F035FFCCFF37A6FAFE27F9CE" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BA236E8F035FFCCFF37A6FAFE83FAEB" blockId="4.[189,292,1254,1280]" box="[189,292,1254,1280]" pageId="4" pageNumber="5">
<heading id="D0EA8184F035FFCCFF37A6FAFE83FAEB" bold="true" box="[189,292,1254,1280]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B969EAFAF035FFCCFF37A6FAFE83FAEB" bold="true" box="[189,292,1254,1280]" pageId="4" pageNumber="5">Remark</emphasis>
</heading>
</paragraph>
<paragraph id="8BA236E8F035FFCCFF37A708FE27F9CE" blockId="4.[189,1399,1300,1573]" pageId="4" pageNumber="5">
The revised concept of
<taxonomicName id="4C1D4D6BF035FFCCFE7CA709FDFDFAC4" authorityName="Waterhouse" authorityYear="1879" box="[502,602,1301,1327]" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFE7CA709FDFDFAC4" box="[502,602,1301,1327]" italics="true" pageId="4" pageNumber="5">Cautires</emphasis>
</taxonomicName>
merged the genera
<taxonomicName id="4C1D4D6BF035FFCCFCECA709FB60FAC4" authority="Waterhouse, 1879" authorityName="Waterhouse" authorityYear="1879" box="[870,1223,1300,1327]" class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFCECA709FC7DFAC4" box="[870,986,1301,1327]" italics="true" pageId="4" pageNumber="5">Bulenides</emphasis>
<bibRefCitation id="EF8C4B19F035FFCCFC60A708FB60FAC4" author="Waterhouse C. O." box="[1002,1223,1300,1327]" pageId="4" pageNumber="5" refId="ref15539" refString="Waterhouse C. O. 1879. Illustration of the Typical Specimens of Coleoptera in the Collection of the British Museum. Part I. Lycidae. British Museum, London." type="book" year="1879">Waterhouse, 1879</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1D4D6BF035FFCCFA98A709FE31FAB9" authority="Waterhouse, 1879" authorityName="Waterhouse" authorityYear="1879" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFA98A709FAD1FAC4" box="[1298,1398,1301,1327]" italics="true" pageId="4" pageNumber="5">Cautires</emphasis>
<bibRefCitation id="EF8C4B19F035FFCCFF37A724FE31FAB9" author="Waterhouse C. O." box="[189,406,1336,1362]" pageId="4" pageNumber="5" refId="ref15539" refString="Waterhouse C. O. 1879. Illustration of the Typical Specimens of Coleoptera in the Collection of the British Museum. Part I. Lycidae. British Museum, London." type="book" year="1879">Waterhouse, 1879</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF8C4B19F035FFCCFE20A724FD47FAB9" author="Dudkova P. &amp; Bocak L." box="[426,736,1336,1362]" pageId="4" pageNumber="5" pagination="28 - 48" refId="ref14434" refString="Dudkova P. &amp; Bocak L. 2010. A review of the Cautires obsoletus species group from Indo-Burma (Coleoptera: Lycidae). Zootaxa 2527: 28 - 48." type="journal article" year="2010">Dudkova &amp; Bocak 2010</bibRefCitation>
). Later, the terminal position of
<taxonomicName id="4C1D4D6BF035FFCCFBF0A724FB49FAB9" authorityName="Waterhouse" authorityYear="1879" box="[1146,1262,1336,1362]" class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFBF0A724FB49FAB9" box="[1146,1262,1336,1362]" italics="true" pageId="4" pageNumber="5">Bulenides</emphasis>
</taxonomicName>
within the
<taxonomicName id="4C1D4D6BF035FFCCFF37A747FE86FA9E" authorityName="Waterhouse" authorityYear="1879" box="[189,289,1371,1397]" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFF37A747FE86FA9E" box="[189,289,1371,1397]" italics="true" pageId="4" pageNumber="5">Cautires</emphasis>
</taxonomicName>
clade was confirmed by subsequent molecular phylogenetic study by
<bibRefCitation id="EF8C4B19F035FFCCFBDEA747FAD4FA9E" author="Sklenarova K. &amp; Kubecek V. &amp; Bocak L." box="[1108,1395,1371,1397]" pageId="4" pageNumber="5" pagination="37 - 54" refId="ref15307" refString="Sklenarova K., Kubecek V. &amp; Bocak L. 2014. Subtribal classification of Metriorrhynchini (Insecta: Coleoptera: Lycidae): an integrative approach using molecular phylogeny and morphology of adults and larvae. Arthropod Systematics and Phylogeny 72: 37 - 54." type="journal article" year="2014">
Sklenarova
<emphasis id="B969EAFAF035FFCCFB55A740FAB4FA9E" box="[1247,1299,1371,1397]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. (2014)
</bibRefCitation>
. The latter study also redefined the limits of
<taxonomicName id="4C1D4D6BF035FFCCFD34A762FBB1FA73" authority="Waterhouse, 1879" authorityName="Waterhouse" authorityYear="1879" box="[702,1046,1406,1432]" class="Insecta" family="Lycidae" genus="Xylobanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFD34A762FC9DFA73" box="[702,826,1406,1432]" italics="true" pageId="4" pageNumber="5">Xylobanus</emphasis>
<bibRefCitation id="EF8C4B19F035FFCCFCCAA762FBB1FA73" author="Waterhouse C. O." box="[832,1046,1406,1432]" pageId="4" pageNumber="5" refId="ref15539" refString="Waterhouse C. O. 1879. Illustration of the Typical Specimens of Coleoptera in the Collection of the British Museum. Part I. Lycidae. British Museum, London." type="book" year="1879">Waterhouse, 1879</bibRefCitation>
</taxonomicName>
, which was originally defined by the absence of secondary costae. Some
<taxonomicName id="4C1D4D6BF035FFCCFD4AA7BDFC9BFA50" authorityName="Waterhouse" authorityYear="1879" box="[704,828,1441,1467]" class="Insecta" family="Lycidae" genus="Xylobanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFD4AA7BDFC9BFA50" box="[704,828,1441,1467]" italics="true" pageId="4" pageNumber="5">Xylobanus</emphasis>
</taxonomicName>
have elytral costae similar to those of
<taxonomicName id="4C1D4D6BF035FFCCFA98A7BEFAD1FA57" authorityName="Waterhouse" authorityYear="1879" box="[1298,1398,1442,1468]" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFA98A7BEFAD1FA57" box="[1298,1398,1442,1468]" italics="true" pageId="4" pageNumber="5">Cautires</emphasis>
</taxonomicName>
and adults of these species can be identified using the shape of male genitalia. The species group of
<taxonomicName id="4C1D4D6BF035FFCCFAD1A7D9FE8BF9E9" authorityName="Waterhouse" authorityYear="1878" class="Insecta" family="Lycidae" genus="Cautires" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="obsoletus">
<emphasis id="B969EAFAF035FFCCFAD1A7D9FE8BF9E9" italics="true" pageId="4" pageNumber="5">C. obsoletus</emphasis>
</taxonomicName>
corresponds with the limits of
<taxonomicName id="4C1D4D6BF035FFCCFD15A7F4FCB4F9E9" authorityName="Waterhouse" authorityYear="1879" box="[671,787,1512,1538]" class="Insecta" family="Lycidae" genus="Bulenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAFAF035FFCCFD15A7F4FCB4F9E9" box="[671,787,1512,1538]" italics="true" pageId="4" pageNumber="5">Bulenides</emphasis>
</taxonomicName>
and these species differ in absence of lateral ridges in the pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>
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