244 lines
27 KiB
XML
244 lines
27 KiB
XML
<document id="8559A64CD858EF71A509B81AE859B55D" ID-CLB-Dataset="46857" ID-DOI="10.11646/zootaxa.4072.3.3" ID-GBIF-Dataset="7d92035f-e4aa-40c7-a424-5e8047d30799" ID-ISSN="1175-5326" ID-Zenodo-Dep="261812" ID-ZooBank="C42DAE9A-4E94-4CA6-B9C8-A5FE15A47CD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460358144663" checkinUser="plazi" docAuthor="Génier, François & Josso, Jean-François" docDate="2016" docId="03B487F3FFCFFFBA9396FE5DFDA9F8A8" docLanguage="en" docName="zootaxa.4072.3.3.pdf" docOrigin="Zootaxa 4072 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Catharsius dux Harold 1878" docType="treatment" docVersion="8" lastPageNumber="338" masterDocId="FF8DFF8BFFCAFFBF9301FF81FF8CFFC7" masterDocTitle="Notes on Afrotropical Catharsius species described by Edgar von Harold (Coleoptera: Scarabaeidae, Scarabaeinae)" masterLastPageNumber="342" masterPageNumber="333" pageNumber="338" updateTime="1698294164494" updateUser="ExternalLinkService">
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<mods:title id="4429AEB737B5FA3D3E356C92CF154109">Notes on Afrotropical Catharsius species described by Edgar von Harold (Coleoptera: Scarabaeidae, Scarabaeinae)</mods:title>
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<mods:namePart id="F0F9F8743D3CC02E30B226B18C916261">Génier, François</mods:namePart>
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<mods:namePart id="F65CFEC2AF95E15F811420E95C88295E">Josso, Jean-François</mods:namePart>
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<mods:title id="888A27CA2DEE1C1F97982636E4CB8451">Zootaxa</mods:title>
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<mods:date id="A5AA23C8445126AC1A3635303FFD58C3">2016</mods:date>
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<treatment id="03B487F3FFCFFFBA9396FE5DFDA9F8A8" ID-DOI="http://doi.org/10.5281/zenodo.6080001" ID-GBIF-Taxon="119409137" ID-Zenodo-Dep="6080001" LSID="urn:lsid:plazi:treatment:03B487F3FFCFFFBA9396FE5DFDA9F8A8" httpUri="http://treatment.plazi.org/id/03B487F3FFCFFFBA9396FE5DFDA9F8A8" lastPageNumber="338" pageId="5" pageNumber="338">
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<subSubSection id="C307656EFFCFFFBA9396FE5DFEA2FDD0" pageId="5" pageNumber="338" type="nomenclature">
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<paragraph id="8BA236E5FFCFFFBA9396FE5DFE76FE31" blockId="5.[151,506,476,535]" box="[151,506,476,502]" pageId="5" pageNumber="338">
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<heading id="D0EA8189FFCFFFBA9396FE5DFE76FE31" bold="true" box="[151,506,476,502]" fontSize="11" level="1" pageId="5" pageNumber="338" reason="1">
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<taxonomicName id="4C1D4D66FFCFFFBA9396FE5DFE76FE31" ID-CoL="RTYK" authority="Harold, 1878" authorityName="Harold" authorityYear="1878" box="[151,506,476,502]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="dux">
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<emphasis id="B969EAF7FFCFFFBA9396FE5DFE76FE31" bold="true" box="[151,506,476,502]" pageId="5" pageNumber="338">
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<emphasis id="B969EAF7FFCFFFBA9396FE5DFEDCFE31" bold="true" box="[151,336,476,502]" italics="true" pageId="5" pageNumber="338">Catharsius dux</emphasis>
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<bibRefCitation id="EF8C4B14FFCFFFBA9256FE5DFE76FE31" author="Harold" box="[343,506,476,502]" pageId="5" pageNumber="338" refString="Harold, E. von (1878) Diagnosen neuer Coleopteren aus dem innern Africa. Mittheilungen der munchner entomologischen Verein Munchen, 2, 99 - 111." type="journal article" year="1878">Harold, 1878</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BA236E5FFCFFFBA9396FE7FFEA2FDD0" blockId="5.[151,506,476,535]" box="[151,302,510,535]" pageId="5" pageNumber="338">
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(
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<figureCitation id="13262A60FFCFFFBA939EFE7FFEA9FDD0" box="[159,293,510,535]" captionStart-0="FIGURES 18 – 21" captionStart-1="FIGURES 22 – 26" captionStartId-0="6.[151,264,1776,1799]" captionStartId-1="7.[151,264,1768,1791]" captionTargetBox-0="[261,1320,204,1741]" captionTargetBox-1="[262,1321,206,1734]" captionTargetId-0="figure@6.[243,1344,193,1756]" captionTargetId-1="figure@7.[243,1344,193,1747]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 18 – 21. Catharsius dux Harold (lectotype ♀). 18. habitus, dorsal view; 19. head, dorsal view; 20. labels; 21. head and pronotum, frontal view." captionText-1="FIGURES 22 – 26. Catharsius dux Harold (alloreferent ♂). 22. habitus, dorsal view; 23. head, dorsal view; 24. labels; 25. parameres, dorsal view; 26. aedeagus, lateral view." httpUri-0="https://zenodo.org/record/261816/files/figure.png" httpUri-1="https://zenodo.org/record/261817/files/figure.png" pageId="5" pageNumber="338">Figs. 18–26</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C307656EFFCFFFBA9396FDC6FE7CFD9B" box="[151,496,582,604]" pageId="5" pageNumber="338" type="reference_group">
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<paragraph id="8BA236E5FFCFFFBA9396FDC6FE7CFD9B" blockId="5.[151,496,582,604]" box="[151,496,582,604]" pageId="5" pageNumber="338">
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<treatmentCitationGroup id="AB0D11CBFFCFFFBA9396FDC6FE7CFD9B" box="[151,496,582,604]" pageId="5" pageNumber="338">
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<taxonomicName id="4C1D4D66FFCFFFBA9396FDC6FE7CFD9B" ID-CoL="RTYK" authority="Harold, 1878: 101" authorityName="Harold" authorityPageNumber="101" authorityYear="1878" box="[151,496,582,604]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="dux">
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<emphasis id="B969EAF7FFCFFFBA9396FDC6FEBCFD9B" box="[151,304,583,604]" italics="true" pageId="5" pageNumber="338">Catharsius dux</emphasis>
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<treatmentCitation id="0ABC10F4FFCFFFBA9236FDC7FE7CFD9B" author="Harold" box="[311,496,582,604]" page="101" pageId="5" pageNumber="338" year="1878">
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<bibRefCitation id="EF8C4B14FFCFFFBA9236FDC7FE4DFD9B" author="Harold" box="[311,449,582,604]" pageId="5" pageNumber="338" refString="Harold, E. von (1878) Diagnosen neuer Coleopteren aus dem innern Africa. Mittheilungen der munchner entomologischen Verein Munchen, 2, 99 - 111." type="journal article" year="1878">Harold, 1878</bibRefCitation>
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: 101
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</treatmentCitation>
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</taxonomicName>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C307656EFFCFFFBA9396FD08FBECFC59" pageId="5" pageNumber="338" type="materials_examined">
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<paragraph id="8BA236E5FFCFFFBA9396FD08FDB5FD65" blockId="5.[151,1437,649,1903]" box="[151,569,649,674]" pageId="5" pageNumber="338">
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<emphasis id="B969EAF7FFCFFFBA9396FD08FEB9FD65" bold="true" box="[151,309,649,674]" pageId="5" pageNumber="338">
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<typeStatus id="54A68847FFCFFFBA9396FD08FF5EFD65" box="[151,210,649,674]" pageId="5" pageNumber="338">Type</typeStatus>
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locality.
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</emphasis>
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Regn. Lunda (
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<collectingCountry id="F30A7675FFCFFFBA92DAFD08FDBDFD65" box="[475,561,649,674]" name="Angola" pageId="5" pageNumber="338">Angola</collectingCountry>
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)
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</paragraph>
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<paragraph id="8BA236E5FFCFFFBA93C6FD2FFBECFC59" blockId="5.[151,1437,649,1903]" pageId="5" pageNumber="338">
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In the very brief description given by Harold there is a length range which indicates that more than one specimen was studied. The range is “38–40 mill.” and most likely fits the size of the two female
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<typeStatus id="54A68847FFCFFFBA9632FD53FA15FD2D" box="[1331,1433,722,746]" pageId="5" pageNumber="338" type="syntype">syntypes</typeStatus>
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. Furthermore, Harold states that the sex described is female. Both specimens are labelled as “Lunda, Pogge”. The largest female specimen is selected as
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<typeStatus id="54A68847FFCFFFBA9158FC98FD4CFCF5" box="[601,704,793,818]" pageId="5" pageNumber="338" type="lectotype">lectotype</typeStatus>
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(
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<emphasis id="B969EAF7FFCFFFBA91D2FC98FC33FCF5" bold="true" box="[723,959,793,818]" pageId="5" pageNumber="338">present designation</emphasis>
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) and bear the following labels (
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<figureCitation id="13262A60FFCFFFBA9636FC98FA03FCF5" box="[1335,1423,793,818]" captionStart="FIGURES 18 – 21" captionStartId="6.[151,264,1776,1799]" captionTargetBox="[261,1320,204,1741]" captionTargetId="figure@6.[243,1344,193,1756]" captionTargetPageId="6" captionText="FIGURES 18 – 21. Catharsius dux Harold (lectotype ♀). 18. habitus, dorsal view; 19. head, dorsal view; 20. labels; 21. head and pronotum, frontal view." httpUri="https://zenodo.org/record/261816/files/figure.png" pageId="5" pageNumber="338">Fig. 20</figureCitation>
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): [Pogge!] handwriting on green card; [
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<taxonomicName id="4C1D4D66FFCFFFBA9142FCBCFDE3FC91" box="[579,623,829,854]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="dux">dux</taxonomicName>
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/ Harold/ Regn. Lunda] handwriting on green card; [60062] white card; [
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<typeStatus id="54A68847FFCFFFBA939CFCE3FF56FCBD" box="[157,218,866,890]" pageId="5" pageNumber="338">Type</typeStatus>
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] red card; [1285] green card; [
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<typeStatus id="54A68847FFCFFFBA9137FCE3FD45FCBD" box="[566,713,866,890]" pageId="5" pageNumber="338" type="syntype">SYNTYPUS</typeStatus>
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/
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<taxonomicName id="4C1D4D66FFCFFFBA91DBFCE0FCC3FCBD" box="[730,847,865,890]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="genus">Catharsius</taxonomicName>
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/
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<taxonomicName id="4C1D4D66FFCFFFBA9060FCE0FBABFCBD" authority="Harold, 1878" authorityName="Harold" authorityYear="1878" box="[865,1063,865,890]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="dux">
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dux
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<bibRefCitation id="EF8C4B14FFCFFFBA9092FCE0FBABFCBD" author="Harold" box="[915,1063,865,890]" pageId="5" pageNumber="338" refString="Harold, E. von (1878) Diagnosen neuer Coleopteren aus dem innern Africa. Mittheilungen der munchner entomologischen Verein Munchen, 2, 99 - 111." type="journal article" year="1878">Harold, 1878</bibRefCitation>
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</taxonomicName>
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/ labelled by
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<collectionCode id="ED0CAE20FFCFFFBA97BFFCE3FAAAFCBE" box="[1214,1318,866,889]" pageId="5" pageNumber="338">MNHUB</collectionCode>
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2014] red card; [
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<typeStatus id="54A68847FFCFFFBA93DDFC07FEF3FC59" box="[220,383,902,926]" pageId="5" pageNumber="338" type="lectotype">LECTOTYPE</typeStatus>
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♀/
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<taxonomicName id="4C1D4D66FFCFFFBA92ABFC04FDA9FC5A" box="[426,549,901,925]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="genus">
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<emphasis id="B969EAF7FFCFFFBA92ABFC04FDA9FC5A" box="[426,549,901,925]" italics="true" pageId="5" pageNumber="338">Catharsius</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C1D4D66FFCFFFBA9132FC04FDD7FC5A" box="[563,603,901,925]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="dux">
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<emphasis id="B969EAF7FFCFFFBA9132FC04FDD7FC5A" box="[563,603,901,925]" italics="true" pageId="5" pageNumber="338">dux</emphasis>
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</taxonomicName>
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/
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<bibRefCitation id="EF8C4B14FFCFFFBA916AFC04FD71FC59" author="Harold" box="[619,765,901,926]" pageId="5" pageNumber="338" refString="Harold, E. von (1878) Diagnosen neuer Coleopteren aus dem innern Africa. Mittheilungen der munchner entomologischen Verein Munchen, 2, 99 - 111." type="journal article" year="1878">Harold, 1878</bibRefCitation>
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/ dés. F. Génier, 2014] red card.
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</paragraph>
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</subSubSection>
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<subSubSection id="C307656EFFCFFFBA93C6FC28FDA9F8A8" pageId="5" pageNumber="338" type="discussion">
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<paragraph id="8BA236E5FFCFFFBA93C6FC28FD66FB20" blockId="5.[151,1437,649,1903]" pageId="5" pageNumber="338">
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<emphasis id="B969EAF7FFCFFFBA93C6FC28FEBCFC05" bold="true" box="[199,304,937,962]" pageId="5" pageNumber="338">Remark.</emphasis>
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The
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<typeStatus id="54A68847FFCFFFBA926EFC2BFE13FC05" box="[367,415,938,962]" pageId="5" pageNumber="338">type</typeStatus>
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locality was previously cited in error as “
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<collectingCountry id="F30A7675FFCFFFBA908BFC28FC5EFC05" box="[906,978,937,962]" name="Guinea-Bissau" pageId="5" pageNumber="338">Guiné</collectingCountry>
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” by
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<bibRefCitation id="EF8C4B14FFCFFFBA970CFC28FB48FC05" box="[1037,1220,937,962]" pageId="5" pageNumber="338" refString="Ferreira, M. C. (1960) Revisao das especies Africanas de " Catharsius " s. str. do " grupo Adamastor " e descricao de especies novas. Revista de Entomologia de Mocambique, 3 (1), 1 - 73" type="journal article">Ferreira, (1960)</bibRefCitation>
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. This
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<typeStatus id="54A68847FFCFFFBA960FFC2BFAB2FC05" box="[1294,1342,938,962]" pageId="5" pageNumber="338">type</typeStatus>
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locality most likely came from a misinterpretation of “im inneren
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<collectingCountry id="F30A7675FFCFFFBA9024FC4CFCF4FC21" box="[805,888,973,998]" name="Guinea-Bissau" pageId="5" pageNumber="338">Guinea</collectingCountry>
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” as cited in Harold’s original description. At the end of the 19th century “
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<collectingCountry id="F30A7675FFCFFFBA92A4FC75FE76FBCA" box="[421,506,1012,1037]" name="Guinea-Bissau" pageId="5" pageNumber="338">Guinea</collectingCountry>
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” was a very vague term for the “Gulf of
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<collectingCountry id="F30A7675FFCFFFBA90CDFC75FBAEFBCA" box="[972,1058,1012,1037]" name="Guinea-Bissau" pageId="5" pageNumber="338">Guinea</collectingCountry>
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”. Harold’s (1879) gave a detailed account of A.V. Homeyer and P. Pogge’s journey into “
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<collectingCountry id="F30A7675FFCFFFBA901FFB99FCFFFBF6" box="[798,883,1048,1073]" name="Angola" pageId="5" pageNumber="338">Angola</collectingCountry>
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und im Luanda-Reiche” or former pre-colonial confederation of the Lunda Kingdom (
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<bibRefCitation id="EF8C4B14FFCFFFBA915BFBBDFC88FB93" author="Moretto" box="[602,772,1084,1108]" pageId="5" pageNumber="338" refString="Moretto, Ph. (2014) Description de deux nouveaux Heliocopris Hope, 1837, d'Afrique tropicale (Coleoptera, Scarabaeidae). Catharsius, La Revue, 9, 1 - 9" type="journal article" year="2014">Moretto, 2014</bibRefCitation>
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). The original
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<typeStatus id="54A68847FFCFFFBA90B2FBBCFB86FB92" box="[947,1034,1085,1109]" pageId="5" pageNumber="338" type="syntype">syntype</typeStatus>
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labels state “Regn. Lunda” as the locality where Pogge collected the specimens. Harold gives the collecting sites for Pogge on a line approximately along the 10th parallel from 17 to 22° of longitude. Most of this itinerary is located within
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<collectingCountry id="F30A7675FFCFFFBA979AFB07FB61FB58" box="[1179,1261,1158,1183]" name="Angola" pageId="5" pageNumber="338">Angola</collectingCountry>
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and for a short distance in
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<collectingCountry id="F30A7675FFCFFFBA9223FB2AFD7EFB04" box="[290,754,1195,1220]" name="Democratic Republic of the Congo" pageId="5" pageNumber="338">the Democratic Republic of the Congo</collectingCountry>
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. Therefore, the exact
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<typeStatus id="54A68847FFCFFFBA90FAFB2DFBA7FB03" box="[1019,1067,1196,1220]" pageId="5" pageNumber="338">type</typeStatus>
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locality lies somewhere in the Malanje, North or South Lunda provinces of
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<collectingCountry id="F30A7675FFCFFFBA9191FB4FFD6AFB20" box="[656,742,1230,1255]" name="Angola" pageId="5" pageNumber="338">Angola</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BA236E5FFCFFFBA93C6FB72FCF5FA78" blockId="5.[151,1437,649,1903]" pageId="5" pageNumber="338">
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The current cited distribution range of this species is from
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<collectingCountry id="F30A7675FFCFFFBA9054FB72FC2AFACB" box="[853,934,1267,1292]" name="Guinea-Bissau" pageId="5" pageNumber="338">Guinea</collectingCountry>
|
||
in West Africa to
|
||
<collectingCountry id="F30A7675FFCFFFBA9773FB72FA8AFACB" box="[1138,1286,1267,1292]" name="Mozambique" pageId="5" pageNumber="338">Mozambique</collectingCountry>
|
||
and
|
||
<collectingCountry id="F30A7675FFCFFFBA963DFB72FA10FACB" box="[1340,1436,1267,1292]" name="Namibia" pageId="5" pageNumber="338">Namibia</collectingCountry>
|
||
in Southern Africa. Such wide distribution is rare among Afrotropical dung beetle species. For now, we can confirm the presence of this species from
|
||
<collectingCountry id="F30A7675FFCFFFBA9189FABAFD71FA94" box="[648,765,1339,1363]" name="Cameroon" pageId="5" pageNumber="338">Cameroon</collectingCountry>
|
||
(Northwest Region),
|
||
<collectingCountry id="F30A7675FFCFFFBA90FCFABAFA10FA93" box="[1021,1436,1339,1364]" name="Democratic Republic of the Congo" pageId="5" pageNumber="338">Congo, Democratic Republic of the</collectingCountry>
|
||
<collectingCountry id="F30A7675FFCFFFBA9396FADEFF68FAB0" box="[151,228,1375,1399]" name="Republic of the Congo" pageId="5" pageNumber="338">Congo</collectingCountry>
|
||
,
|
||
<collectingCountry id="F30A7675FFCFFFBA93F0FADFFEC8FAB0" box="[241,324,1374,1399]" name="Angola" pageId="5" pageNumber="338">Angola</collectingCountry>
|
||
and
|
||
<collectingCountry id="F30A7675FFCFFFBA927CFADFFE5FFAB0" box="[381,467,1374,1399]" name="Zambia" pageId="5" pageNumber="338">Zambia</collectingCountry>
|
||
(examined specimens and pers. comm. P. Moretto). We consider records outside this range dubious and probably based on mislabeled or misidentified specimens. All other countries cited by Ferreira are most likely misidentifications of
|
||
<taxonomicName id="4C1D4D66FFCFFFBA9133FA26FCF5FA78" authority="Thomson." authorityName="Thomson." box="[562,889,1446,1471]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="gorilla">
|
||
<emphasis id="B969EAF7FFCFFFBA9133FA26FC8CFA78" box="[562,768,1447,1471]" italics="true" pageId="5" pageNumber="338">Catharsius gorilla</emphasis>
|
||
Thomson.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BA236E5FFCFFFBA93C6FA4AFDA9F8A8" blockId="5.[151,1437,649,1903]" pageId="5" pageNumber="338">
|
||
<bibRefCitation id="EF8C4B14FFCFFFBA93C6FA4AFE2FFA24" author="Quedenfeldt" box="[199,419,1483,1508]" pageId="5" pageNumber="338" refString="Quedenfeldt, G. (1884) Verzeichniss der von Herrn Major a. D. von Mechow in Angola und am Quango-strom 1878 - 1881 gesammelten Pectinicornen und Lamellicornen. Berliner Entomologische Zeitschrift, 28 (2), 265 - 340. http: // dx. doi. org / 10.1002 / mmnd. 18840280207" type="journal article" year="1884">Quedenfeldt (1884)</bibRefCitation>
|
||
described the male of
|
||
<taxonomicName id="4C1D4D66FFCFFFBA91A3FA4AFD60FA24" box="[674,748,1483,1507]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="dux">
|
||
<emphasis id="B969EAF7FFCFFFBA91A3FA4AFD60FA24" box="[674,748,1483,1507]" italics="true" pageId="5" pageNumber="338">C. dux</emphasis>
|
||
</taxonomicName>
|
||
from a series of specimens collected in Malange (=Malanje),
|
||
<collectingCountry id="F30A7675FFCFFFBA9396FA6FFF61F9C0" box="[151,237,1518,1543]" name="Angola" pageId="5" pageNumber="338">Angola</collectingCountry>
|
||
. In order to remain as conservative as possible and conform to the current accepted species concept we retain this interpretation which does not challenge the valid status of
|
||
<taxonomicName id="4C1D4D66FFCFFFBA90BFF992FBD7F9EC" box="[958,1115,1555,1579]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="duciformis">
|
||
<emphasis id="B969EAF7FFCFFFBA90BFF992FBD7F9EC" box="[958,1115,1555,1579]" italics="true" pageId="5" pageNumber="338">C. duciformis</emphasis>
|
||
</taxonomicName>
|
||
. Although not regulated by ICZN, we “link” a male specimen with specific diagnostic characters to the female
|
||
<typeStatus id="54A68847FFCFFFBA9754F9B7FB30F988" box="[1109,1212,1590,1615]" pageId="5" pageNumber="338" type="lectotype">lectotype</typeStatus>
|
||
of this species (see
|
||
<bibRefCitation id="EF8C4B14FFCFFFBA9396F9DAFEEFF9B4" author="Dechambre" box="[151,355,1627,1652]" pageId="5" pageNumber="338" refString="Dechambre, R. - P. (2002) Neallotype ou Alloreferent: de l'utilite de ce concept et de son bon usage. Le Coleopteriste, 43, 163 - 164" type="journal article" year="2002">Dechambre, 2002</bibRefCitation>
|
||
). This is especially important for nomenclatural stability as the female specimens of this species are undistinguishable from females of
|
||
<taxonomicName id="4C1D4D66FFCFFFBA9159F9FEFB89F950" authority="Ferreira, 1959" authorityName="Ferreira" authorityYear="1959" box="[600,1029,1662,1687]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="duciformis">
|
||
<emphasis id="B969EAF7FFCFFFBA9159F9FEFCDAF950" box="[600,854,1663,1687]" italics="true" pageId="5" pageNumber="338">Catharsius duciformis</emphasis>
|
||
Ferreira, 1959
|
||
</taxonomicName>
|
||
. To our knowledge,
|
||
<taxonomicName id="4C1D4D66FFCFFFBA97F9F9FEFAC9F950" box="[1272,1349,1663,1687]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="dux">
|
||
<emphasis id="B969EAF7FFCFFFBA97F9F9FEFAC9F950" box="[1272,1349,1663,1687]" italics="true" pageId="5" pageNumber="338">C. dux</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D66FFCFFFBA9683F9FEFE83F97C" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="duciformis">
|
||
<emphasis id="B969EAF7FFCFFFBA9683F9FEFE83F97C" italics="true" pageId="5" pageNumber="338">C. duciformis</emphasis>
|
||
</taxonomicName>
|
||
have never been collected from the same locality. Furthermore, the parapatric distribution pattern of
|
||
<taxonomicName id="4C1D4D66FFCFFFBA9683F922FF4CF918" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="dux">
|
||
<emphasis id="B969EAF7FFCFFFBA9683F922FF4CF918" italics="true" pageId="5" pageNumber="338">C. dux</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D66FFCFFFBA9202F946FE2EF918" box="[259,418,1735,1759]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="duciformis">
|
||
<emphasis id="B969EAF7FFCFFFBA9202F946FE2EF918" box="[259,418,1735,1759]" italics="true" pageId="5" pageNumber="338">C. duciformis</emphasis>
|
||
</taxonomicName>
|
||
is also problematic for species recognition. This alloreferent, a well-developed male specimen (fig. 22–26), is from northwest
|
||
<collectingCountry id="F30A7675FFCFFFBA917FF96AFD58F8C3" box="[638,724,1771,1796]" name="Zambia" pageId="5" pageNumber="338">Zambia</collectingCountry>
|
||
and bears the following label data (
|
||
<figureCitation id="13262A60FFCFFFBA977FF96AFB5AF8C4" box="[1150,1238,1771,1796]" captionStart="FIGURES 22 – 26" captionStartId="7.[151,264,1768,1791]" captionTargetBox="[262,1321,206,1734]" captionTargetId="figure@7.[243,1344,193,1747]" captionTargetPageId="7" captionText="FIGURES 22 – 26. Catharsius dux Harold (alloreferent ♂). 22. habitus, dorsal view; 23. head, dorsal view; 24. labels; 25. parameres, dorsal view; 26. aedeagus, lateral view." httpUri="https://zenodo.org/record/261817/files/figure.png" pageId="5" pageNumber="338">Fig. 24</figureCitation>
|
||
): [
|
||
<collectingCountry id="F30A7675FFCFFFBA97F5F96AFAEEF8C4" box="[1268,1378,1771,1795]" name="Zambia" pageId="5" pageNumber="338">ZAMBIE</collectingCountry>
|
||
NW Prov./ 10Km N Mwinilunga/
|
||
<date id="FFA31025FFCFFFBA92EFF88FFDF7F8E0" box="[494,635,1806,1831]" pageId="5" pageNumber="338" value="2012-12-06">06-XII-2012</date>
|
||
/
|
||
<geoCoordinate id="EE295022FFCFFFBA918FF88EFD65F8E0" box="[654,745,1807,1831]" direction="south" orientation="latitude" pageId="5" pageNumber="338" precision="925" value="-11.566667">S11°34’</geoCoordinate>
|
||
<geoCoordinate id="EE295022FFCFFFBA91F5F88EFCDCF8E0" box="[756,848,1807,1831]" direction="east" orientation="longitude" pageId="5" pageNumber="338" precision="925" value="24.4">E24°24’</geoCoordinate>
|
||
/ leg. Josso Juhel Minetti]; [ALLOREFERENT ♂/
|
||
<taxonomicName id="4C1D4D66FFCFFFBA9396F8B2FE9EF88C" box="[151,274,1843,1867]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAF7FFCFFFBA9396F8B2FE9EF88C" box="[151,274,1843,1867]" italics="true" pageId="5" pageNumber="338">Catharsius</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C1D4D66FFCFFFBA9222F8B2FEC7F88C" box="[291,331,1843,1867]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="dux">
|
||
<emphasis id="B969EAF7FFCFFFBA9222F8B2FEC7F88C" box="[291,331,1843,1867]" italics="true" pageId="5" pageNumber="338">dux</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<bibRefCitation id="EF8C4B14FFCFFFBA925CF8B2FE7FF88C" author="Harold" box="[349,499,1843,1868]" pageId="5" pageNumber="338" refString="Harold, E. von (1878) Diagnosen neuer Coleopteren aus dem innern Africa. Mittheilungen der munchner entomologischen Verein Munchen, 2, 99 - 111." type="journal article" year="1878">Harold, 1878</bibRefCitation>
|
||
/ dés.: F. Génier & J.-F. Josso, 2015]. The alloreferent male is deposited together with the
|
||
<typeStatus id="54A68847FFCFFFBA93F8F8D7FEECF8A8" box="[249,352,1878,1903]" pageId="5" pageNumber="338" type="lectotype">lectotype</typeStatus>
|
||
female (ZMHB).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |