165 lines
18 KiB
XML
165 lines
18 KiB
XML
<document id="A0FBE0D6602316888D6F21043DF54FCA" ID-CLB-Dataset="46857" ID-DOI="10.11646/zootaxa.4072.3.3" ID-GBIF-Dataset="7d92035f-e4aa-40c7-a424-5e8047d30799" ID-ISSN="1175-5326" ID-Zenodo-Dep="261812" ID-ZooBank="C42DAE9A-4E94-4CA6-B9C8-A5FE15A47CD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460358144663" checkinUser="plazi" docAuthor="Génier, François & Josso, Jean-François" docDate="2016" docId="03B487F3FFC3FFB69396FF16FA98FC46" docLanguage="en" docName="zootaxa.4072.3.3.pdf" docOrigin="Zootaxa 4072 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Catharsius brutus Harold 1881" docType="treatment" docVersion="8" lastPageNumber="342" masterDocId="FF8DFF8BFFCAFFBF9301FF81FF8CFFC7" masterDocTitle="Notes on Afrotropical Catharsius species described by Edgar von Harold (Coleoptera: Scarabaeidae, Scarabaeinae)" masterLastPageNumber="342" masterPageNumber="333" pageNumber="342" updateTime="1698294164494" updateUser="ExternalLinkService">
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<mods:titleInfo id="CFEA4C91EE4033745DE18EB2720EF7F4">
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<mods:title id="9BCC1259970B1DDC1CD4D593C8252EED">Notes on Afrotropical Catharsius species described by Edgar von Harold (Coleoptera: Scarabaeidae, Scarabaeinae)</mods:title>
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<mods:namePart id="BF91075B5096333AB8E6E7FD4329E20F">Génier, François</mods:namePart>
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<mods:namePart id="0B5BEF49E3801145DEF75D0635EAD138">Josso, Jean-François</mods:namePart>
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<mods:title id="DF6287E2715CA4F4FBA23E6F2B5FF84D">Zootaxa</mods:title>
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<mods:part id="9AAF2FA48D2D9B713E86D303D64B7E34">
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<mods:date id="AFB5D82654CAB19C9F1B206BD3770024">2016</mods:date>
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<mods:number id="7DC75708451DE4977CBE4CEE064F48D4">4072</mods:number>
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<treatment id="03B487F3FFC3FFB69396FF16FA98FC46" ID-DOI="http://doi.org/10.5281/zenodo.6080003" ID-GBIF-Taxon="119409141" ID-Zenodo-Dep="6080003" LSID="urn:lsid:plazi:treatment:03B487F3FFC3FFB69396FF16FA98FC46" httpUri="http://treatment.plazi.org/id/03B487F3FFC3FFB69396FF16FA98FC46" lastPageNumber="342" pageId="9" pageNumber="342">
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<subSubSection id="C307656EFFC3FFB69396FF16FEA2FF14" pageId="9" pageNumber="342" type="nomenclature">
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<paragraph id="8BA236E5FFC3FFB69396FF16FD96FF75" blockId="9.[151,538,151,211]" box="[151,538,151,178]" pageId="9" pageNumber="342">
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<heading id="D0EA8189FFC3FFB69396FF16FD96FF75" bold="true" box="[151,538,151,178]" fontSize="11" level="1" pageId="9" pageNumber="342" reason="1">
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<taxonomicName id="4C1D4D66FFC3FFB69396FF16FD96FF75" ID-CoL="RTXY" authority="Harold, 1881" authorityName="Harold" authorityYear="1881" box="[151,538,151,178]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="342" phylum="Arthropoda" rank="species" species="brutus">
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<emphasis id="B969EAF7FFC3FFB69396FF16FD96FF75" bold="true" box="[151,538,151,178]" pageId="9" pageNumber="342">
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<emphasis id="B969EAF7FFC3FFB69396FF16FEFCFF76" bold="true" box="[151,368,151,177]" italics="true" pageId="9" pageNumber="342">Catharsius brutus</emphasis>
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<bibRefCitation id="EF8C4B14FFC3FFB69276FF19FD96FF75" author="Harold" box="[375,538,152,178]" pageId="9" pageNumber="342" refString="Harold, E. von (1881) Beschreibung neuer, auf seiner, von der Akademie unterstutzten Reise in Ostafrika, vozuglich in den Districten von Taita und Ukamba auf einer Tour von Mombassa nach den Kenia, von Hrn. J. M. Hildebrandt gesammelten Coleopteren. Monatsbericht der Koniglich Preussischen Akademie des Wissenschaften zu Berlin, 1880, 260 - 270." type="journal article" year="1881">Harold, 1881</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BA236E5FFC3FFB69396FF3BFEA2FF14" blockId="9.[151,538,151,211]" box="[151,302,186,211]" pageId="9" pageNumber="342">
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(
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<figureCitation id="13262A60FFC3FFB6939EFF3BFEA9FF14" box="[159,293,186,211]" captionStart="FIGURES 27 – 33" captionStartId="8.[151,264,1741,1764]" captionTargetBox="[262,1321,203,1711]" captionTargetId="figure@8.[243,1344,193,1721]" captionTargetPageId="8" captionText="FIGURES 27 – 33. Catharsius brutus Harold (Figs. 27 – 31. holotype ♂, Figs. 32 – 33. non teneral specimen). 27. habitus, dorsal view; 28. head, dorsal view; 29. labels; 30. parameres, dorsal view; 31. aedeagus, lateral view; 32. parameres, dorsal view; 33. aedeagus, lateral view." httpUri="https://zenodo.org/record/261818/files/figure.png" pageId="9" pageNumber="342">Figs. 27–33</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C307656EFFC3FFB69396FE82FD80FEDF" box="[151,524,258,280]" pageId="9" pageNumber="342" type="reference_group">
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<paragraph id="8BA236E5FFC3FFB69396FE82FD80FEDF" blockId="9.[151,524,258,280]" box="[151,524,258,280]" pageId="9" pageNumber="342">
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<treatmentCitationGroup id="AB0D11CBFFC3FFB69396FE82FD80FEDF" box="[151,524,258,280]" pageId="9" pageNumber="342">
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<taxonomicName id="4C1D4D66FFC3FFB69396FE82FD80FEDF" ID-CoL="RTXY" authority="Harold, 1881: 263" authorityName="Harold" authorityPageNumber="263" authorityYear="1881" box="[151,524,258,280]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="342" phylum="Arthropoda" rank="species" species="brutus">
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<emphasis id="B969EAF7FFC3FFB69396FE82FEC1FEDF" box="[151,333,259,280]" italics="true" pageId="9" pageNumber="342">Catharsius brutus</emphasis>
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<treatmentCitation id="0ABC10F4FFC3FFB69252FE83FD80FEDF" author="Harold" box="[339,524,258,280]" page="263" pageId="9" pageNumber="342" year="1881">
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<bibRefCitation id="EF8C4B14FFC3FFB69252FE83FE51FEDF" author="Harold" box="[339,477,258,280]" pageId="9" pageNumber="342" refString="Harold, E. von (1881) Beschreibung neuer, auf seiner, von der Akademie unterstutzten Reise in Ostafrika, vozuglich in den Districten von Taita und Ukamba auf einer Tour von Mombassa nach den Kenia, von Hrn. J. M. Hildebrandt gesammelten Coleopteren. Monatsbericht der Koniglich Preussischen Akademie des Wissenschaften zu Berlin, 1880, 260 - 270." type="journal article" year="1881">Harold, 1881</bibRefCitation>
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: 263
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</treatmentCitation>
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</taxonomicName>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C307656EFFC3FFB69396FEC4FDF6FDD1" pageId="9" pageNumber="342" type="materials_examined">
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<paragraph id="8BA236E5FFC3FFB69396FEC4FD97FE99" blockId="9.[151,1437,325,897]" box="[151,539,325,350]" pageId="9" pageNumber="342">
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<emphasis id="B969EAF7FFC3FFB69396FEC4FEB9FE99" bold="true" box="[151,309,325,350]" pageId="9" pageNumber="342">
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<typeStatus id="54A68847FFC3FFB69396FEC4FF5EFE99" box="[151,210,325,350]" pageId="9" pageNumber="342">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F30A7675FFC3FFB6923DFEC4FE11FE99" box="[316,413,325,350]" name="Tanzania" pageId="9" pageNumber="342">Sansibar</collectingCountry>
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(
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<collectingCountry id="F30A7675FFC3FFB692AAFEC4FD9FFE99" box="[427,531,325,350]" name="Tanzania" pageId="9" pageNumber="342">Tanzania</collectingCountry>
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)
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</paragraph>
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<paragraph id="8BA236E5FFC3FFB693C6FEEDFDF6FDD1" blockId="9.[151,1437,325,897]" pageId="9" pageNumber="342">
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The unique specimen on which Harold based his description is a well-developed male, it is very teneral and missing both posterior legs. The
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<typeStatus id="54A68847FFC3FFB69113FE10FDF8FE6D" box="[530,628,401,426]" pageId="9" pageNumber="342" type="holotype">holotype</typeStatus>
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bear the following labels (
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<figureCitation id="13262A60FFC3FFB690ADFE10FB8FFE6E" box="[940,1027,401,426]" captionStart="FIGURES 27 – 33" captionStartId="8.[151,264,1741,1764]" captionTargetBox="[262,1321,203,1711]" captionTargetId="figure@8.[243,1344,193,1721]" captionTargetPageId="8" captionText="FIGURES 27 – 33. Catharsius brutus Harold (Figs. 27 – 31. holotype ♂, Figs. 32 – 33. non teneral specimen). 27. habitus, dorsal view; 28. head, dorsal view; 29. labels; 30. parameres, dorsal view; 31. aedeagus, lateral view; 32. parameres, dorsal view; 33. aedeagus, lateral view." httpUri="https://zenodo.org/record/261818/files/figure.png" pageId="9" pageNumber="342">Fig. 29</figureCitation>
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): [Hildabrdt.] handwriting on green card; [60753] white card; [
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<typeStatus id="54A68847FFC3FFB692D2FE37FD9CFE09" box="[467,528,438,462]" pageId="9" pageNumber="342">Type</typeStatus>
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] red card; [
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<taxonomicName id="4C1D4D66FFC3FFB6919DFE34FD65FE0A" box="[668,745,437,461]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="342" phylum="Arthropoda" rank="species" species="brutus">Brutus</taxonomicName>
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/ Harold/
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<collectingCountry id="F30A7675FFC3FFB6905FFE34FC4EFE09" box="[862,962,437,462]" name="Tanzania" pageId="9" pageNumber="342">Sansibar</collectingCountry>
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] handwriting on green card; [
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<taxonomicName id="4C1D4D66FFC3FFB69622FE34FF50FE35" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="342" phylum="Arthropoda" rank="species" species="brutus">Catharsius brutus</taxonomicName>
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/ HAR./ M.C. Ferreira det.,1960] part handwriting on white card; [
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<typeStatus id="54A68847FFC3FFB690BFFE58FBEBFE36" box="[958,1127,473,497]" pageId="9" pageNumber="342" type="holotype">HOLOTYPUS</typeStatus>
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/
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<taxonomicName id="4C1D4D66FFC3FFB69777FE58FB60FE35" box="[1142,1260,473,498]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="342" phylum="Arthropoda" rank="genus">Catharsius</taxonomicName>
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/
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<taxonomicName id="4C1D4D66FFC3FFB697FAFE58FF42FDD2" authority="Harold, 1881" authorityName="Harold" authorityYear="1881" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="342" phylum="Arthropoda" rank="species" species="brutus">
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brutus
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<bibRefCitation id="EF8C4B14FFC3FFB69646FE58FF42FDD2" author="Harold" pageId="9" pageNumber="342" refString="Harold, E. von (1881) Beschreibung neuer, auf seiner, von der Akademie unterstutzten Reise in Ostafrika, vozuglich in den Districten von Taita und Ukamba auf einer Tour von Mombassa nach den Kenia, von Hrn. J. M. Hildebrandt gesammelten Coleopteren. Monatsbericht der Koniglich Preussischen Akademie des Wissenschaften zu Berlin, 1880, 260 - 270." type="journal article" year="1881">Harold, 1881</bibRefCitation>
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</taxonomicName>
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/ labelled by
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<collectionCode id="ED0CAE20FFC3FFB69261FE7FFE44FDD2" box="[352,456,510,533]" pageId="9" pageNumber="342">MNHUB</collectionCode>
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2014] red card.
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</paragraph>
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</subSubSection>
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<subSubSection id="C307656EFFC3FFB693C6FDA5FA98FC46" pageId="9" pageNumber="342" type="discussion">
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<paragraph id="8BA236E5FFC3FFB693C6FDA5FA98FC46" blockId="9.[151,1437,325,897]" pageId="9" pageNumber="342">
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<emphasis id="B969EAF7FFC3FFB693C6FDA5FEBCFDFA" bold="true" box="[199,304,548,573]" pageId="9" pageNumber="342">Remark.</emphasis>
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This is the last species of
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<taxonomicName id="4C1D4D66FFC3FFB6916BFDA4FD69FDFA" box="[618,741,549,573]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="342" phylum="Arthropoda" rank="genus">
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<emphasis id="B969EAF7FFC3FFB6916BFDA4FD69FDFA" box="[618,741,549,573]" italics="true" pageId="9" pageNumber="342">Catharsius</emphasis>
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</taxonomicName>
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described by Harold. The original description is based on a specimen collected during the expedition of J.M. Hildebrandt in the “Districten von Taita und Ukamba auf einer Tour von
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<collectingCountry id="F30A7675FFC3FFB69203FDEDFEF7FD42" box="[258,379,620,645]" name="Kenya" pageId="9" pageNumber="342">Mombassa</collectingCountry>
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nach dem
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<collectingCountry id="F30A7675FFC3FFB692F2FDEDFDB7FD42" box="[499,571,620,645]" name="Kenya" pageId="9" pageNumber="342">Kenia</collectingCountry>
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”. The
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<typeStatus id="54A68847FFC3FFB69183FDEDFD68FD42" box="[642,740,620,645]" pageId="9" pageNumber="342" type="holotype">holotype</typeStatus>
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label states “
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<collectingCountry id="F30A7675FFC3FFB69077FDEDFC57FD42" box="[886,987,620,645]" name="Tanzania" pageId="9" pageNumber="342">Sansibar</collectingCountry>
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” as the specimen locality which is south of the area described in the title of the publication. However, the title also state “vorzüglich” (especially) which seems to indicate that the expedition went to some other nearby area as well. The island of
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<collectingCountry id="F30A7675FFC3FFB69616FD35FAF7FD0A" box="[1303,1403,692,717]" name="Tanzania" pageId="9" pageNumber="342">Zanzibar</collectingCountry>
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is approximately
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<quantity id="4CE59B00FFC3FFB69244FD58FE13FD36" box="[325,415,728,753]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="342" unit="km" value="200.0">200 km</quantity>
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South of
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<collectingCountry id="F30A7675FFC3FFB6911AFD59FD14FD36" box="[539,664,728,753]" name="Kenya" pageId="9" pageNumber="342">Mombassa</collectingCountry>
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.
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<taxonomicName id="4C1D4D66FFC3FFB691A6FD58FCFAFD36" box="[679,886,729,753]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="342" phylum="Arthropoda" rank="species" species="brutus">
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<emphasis id="B969EAF7FFC3FFB691A6FD58FCFAFD36" box="[679,886,729,753]" italics="true" pageId="9" pageNumber="342">Catharsius brutus</emphasis>
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</taxonomicName>
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is cited from
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<collectingCountry id="F30A7675FFC3FFB69724FD58FBFDFD36" box="[1061,1137,729,753]" name="Kenya" pageId="9" pageNumber="342">Kenya</collectingCountry>
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,
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<collectingCountry id="F30A7675FFC3FFB69780FD59FB6AFD36" box="[1153,1254,728,753]" name="Tanzania" pageId="9" pageNumber="342">Tanzania</collectingCountry>
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(
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<collectingCountry id="F30A7675FFC3FFB697F8FD59FAECFD36" box="[1273,1376,728,753]" name="Tanzania" pageId="9" pageNumber="342">Zanzibar</collectingCountry>
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) and
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<collectingCountry id="F30A7675FFC3FFB69396FD7DFF7EFCD2" box="[151,242,764,789]" name="Uganda" pageId="9" pageNumber="342">Uganda</collectingCountry>
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. Recent collecting in Cabo Delgado Province (
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<collectingCountry id="F30A7675FFC3FFB6900CFD7DFC28FCD2" box="[781,932,764,789]" name="Mozambique" pageId="9" pageNumber="342">Mozambique</collectingCountry>
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) indicates that the species is present further south as well. Unconfirmed records indicate that the species occurs also in central
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<collectingCountry id="F30A7675FFC3FFB6973EFCA1FB5AFCFE" box="[1087,1238,800,825]" name="Mozambique" pageId="9" pageNumber="342">Mozambique</collectingCountry>
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, Mt. Mulange (S.
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<collectingCountry id="F30A7675FFC3FFB69396FCC5FF62FC9A" box="[151,238,836,861]" name="Malawi" pageId="9" pageNumber="342">Malawi</collectingCountry>
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) and Chirinda Forest (E.
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<collectingCountry id="F30A7675FFC3FFB69127FCC5FD2CFC9A" box="[550,672,836,861]" name="Zimbabwe" pageId="9" pageNumber="342">Zimbabwe</collectingCountry>
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) (pers. comm. A. Davis). We are adding photographs of a well sclerotized aedeagus (
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<figureCitation id="13262A60FFC3FFB6928CFCE9FD9FFC46" box="[397,531,872,897]" captionStart="FIGURES 27 – 33" captionStartId="8.[151,264,1741,1764]" captionTargetBox="[262,1321,203,1711]" captionTargetId="figure@8.[243,1344,193,1721]" captionTargetPageId="8" captionText="FIGURES 27 – 33. Catharsius brutus Harold (Figs. 27 – 31. holotype ♂, Figs. 32 – 33. non teneral specimen). 27. habitus, dorsal view; 28. head, dorsal view; 29. labels; 30. parameres, dorsal view; 31. aedeagus, lateral view; 32. parameres, dorsal view; 33. aedeagus, lateral view." httpUri="https://zenodo.org/record/261818/files/figure.png" pageId="9" pageNumber="342">Figs. 32–33</figureCitation>
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) to accompany the distorted aedeagus of the
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<typeStatus id="54A68847FFC3FFB6970CFCE9FBE3FC46" box="[1037,1135,872,897]" pageId="9" pageNumber="342" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13262A60FFC3FFB6977EFCE9FA85FC46" box="[1151,1289,872,897]" captionStart="FIGURES 27 – 33" captionStartId="8.[151,264,1741,1764]" captionTargetBox="[262,1321,203,1711]" captionTargetId="figure@8.[243,1344,193,1721]" captionTargetPageId="8" captionText="FIGURES 27 – 33. Catharsius brutus Harold (Figs. 27 – 31. holotype ♂, Figs. 32 – 33. non teneral specimen). 27. habitus, dorsal view; 28. head, dorsal view; 29. labels; 30. parameres, dorsal view; 31. aedeagus, lateral view; 32. parameres, dorsal view; 33. aedeagus, lateral view." httpUri="https://zenodo.org/record/261818/files/figure.png" pageId="9" pageNumber="342">Figs. 30–31</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |