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<document id="D73F55D6681563C14D6CD4AEE52AE943" ID-DOI="10.1080/00222930902894096" ID-ISSN="1464-5262" ID-Zenodo-Dep="4651894" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617211343156" checkinUser="carolina" docAuthor="Hong, Yong &amp; James, Samuel W." docDate="2009" docId="03B487E8554EFF84FE54FA72FBBE5232" docLanguage="en" docName="JNATHIST.43.21-22.1229-1256.pdf" docOrigin="Journal of Natural History 43 (21 - 22)" docSource="http://dx.doi.org/10.1080/00222930902894096" docStyle="DocumentStyle:FCA295E821436EF9307F50C7AC261B53.5:JNatHist.2008-2012.journal_article.1cover" docStyleId="FCA295E821436EF9307F50C7AC261B53" docStyleName="JNatHist.2008-2012.journal_article.1cover" docStyleVersion="5" docTitle="Amynthas calculatus Hong &amp; James 2009, sp. nov." docType="treatment" docVersion="3" lastPageNumber="1248" masterDocId="FF8DFF90555CFF90FF99FFA5FF95573D" masterDocTitle="Some new Korean Megascolecoid earthworms (Oligochaeta)" masterLastPageNumber="1256" masterPageNumber="1229" pageNumber="1246" updateTime="1698897605693" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="5DBDF7F2A7D3ED577B4D35CD1F84EC68" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8EAA91800385444527CCC792FBD611F9">
<mods:title id="F34465BAEBBDBC1E374E58A67EEB1895">Some new Korean Megascolecoid earthworms (Oligochaeta)</mods:title>
</mods:titleInfo>
<mods:name id="D5902641AAE2DD7519B19C2FCF0E314F" type="personal">
<mods:role id="FABCDF25B0A9AFA598D2D20C40B64F14">
<mods:roleTerm id="D10DF23183EC08231A41622602F9699A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4D56ADC55F0EFA7D06B8394B57912E13">Hong, Yong</mods:namePart>
</mods:name>
<mods:name id="6B5291B5243D1441F54E60293F62EC21" type="personal">
<mods:role id="A36062F6E82B687D4B91F74BE9C029A1">
<mods:roleTerm id="AF6BC51CCB1D8E833EE57D4701E92C5E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A80B2EE83965A02B86DA066C2B7AF4CB">James, Samuel W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="DA31B562F5A5F3C69B62D79F9BC5993A">text</mods:typeOfResource>
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<mods:titleInfo id="D578290E8A3C4B3D60E9A971681B654F">
<mods:title id="89A5FED01D6D9D47DBF4526465D3727B">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="126CFA411865E05BD75BDD9A87EE2735">
<mods:date id="96BAADCD5F987766875052A994E2515D">2009</mods:date>
<mods:detail id="F8205EDEE9E4958B9BC0F472B09A5AC9" type="pubDate">
<mods:number id="AD234E316A1C983BC43DECE5B09BAD6B">2009-05-21</mods:number>
</mods:detail>
<mods:detail id="5B5DD3D9E23188A0EC1E1A09D9EC0C7D" type="volume">
<mods:number id="2B058A9ADD83DAE1F6F5F215C5F49077">43</mods:number>
</mods:detail>
<mods:detail id="B491BCB069460217E0CC8ED1D8C637AE" type="issue">
<mods:number id="F893C243230AB80E68C8129AC89789DA">21 - 22</mods:number>
</mods:detail>
<mods:extent id="090707FEDEBEE810D832C3A3335DA0F8" unit="page">
<mods:start id="EAB2029376D0045A0AF36C5D24C6062A">1229</mods:start>
<mods:end id="BC446A3852442E119A05CA2EBC093597">1256</mods:end>
</mods:extent>
</mods:part>
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<mods:location id="6059B7450DCEFFB9EB7615C5D6524878">
<mods:url id="BE9EFF927E5FFA90F6EDFAC8BFA838BF">http://dx.doi.org/10.1080/00222930902894096</mods:url>
</mods:location>
<mods:classification id="2F9E023EF4A98F4FC0D6A6A071AF6E13">journal article</mods:classification>
<mods:identifier id="CE0965FA42877B3AE9A6F0A3D496C808" type="DOI">10.1080/00222930902894096</mods:identifier>
<mods:identifier id="377BEC008456BA7531D8FFE6CC1025CF" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="BE10A04B199BA1067E973D6697387FCE" type="Zenodo-Dep">4651894</mods:identifier>
</mods:mods>
<treatment id="03B487E8554EFF84FE54FA72FBBE5232" LSID="urn:lsid:plazi:treatment:03B487E8554EFF84FE54FA72FBBE5232" httpUri="http://treatment.plazi.org/id/03B487E8554EFF84FE54FA72FBBE5232" lastPageId="20" lastPageNumber="1248" pageId="18" pageNumber="1246">
<subSubSection id="C3076575554EFF82FE54FA72FC8752D2" box="[461,786,1495,1519]" pageId="18" pageNumber="1246" type="nomenclature">
<paragraph id="8BA236FE554EFF82FE54FA72FC8752D2" blockId="18.[461,786,1495,1551]" box="[461,786,1495,1519]" pageId="18" pageNumber="1246">
<heading id="D0EA8192554EFF82FE54FA72FC8752D2" box="[461,786,1495,1519]" centered="true" fontSize="36" level="2" pageId="18" pageNumber="1246" reason="7">
<taxonomicName id="4C1D4D7D554EFF82FE54FA72FD2752D2" authority="Hong &amp; James, 2009" authorityName="Hong &amp; James" authorityYear="2009" box="[461,690,1495,1519]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="18" pageNumber="1246" phylum="Annelida" rank="species" species="calculatus" status="sp. nov.">
<emphasis id="B969EAEC554EFF82FE54FA72FD2752D2" bold="true" box="[461,690,1495,1519]" italics="true" pageId="18" pageNumber="1246">Amynthas calculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A5797554EFF82FD20FA7DFC8752D2" box="[697,786,1496,1519]" pageId="18" pageNumber="1246" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3076575554EFF84FDA8FA52FF66549A" lastPageId="20" lastPageNumber="1248" pageId="18" pageNumber="1246" type="description">
<paragraph id="8BA236FE554EFF82FDA8FA52FD3B5132" blockId="18.[461,786,1495,1551]" box="[561,686,1527,1551]" pageId="18" pageNumber="1246">
(
<figureCitation id="13262A7B554EFF82FDA1FA52FD335132" box="[568,678,1527,1551]" captionStart="Figure 10" captionStartId="19.[212,279,1109,1131]" captionTargetBox="[522,860,168,1077]" captionTargetId="figure-147@19.[521,861,168,1078]" captionTargetPageId="19" captionText="Figure 10. Amynthas calculatus sp. nov.: ventral view and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4651920" httpUri="https://zenodo.org/record/4651920/files/figure.png" pageId="18" pageNumber="1246">Figure 10</figureCitation>
)
</paragraph>
<paragraph id="8BA236FE554EFF82FF09F9E5FEBF5165" blockId="18.[144,298,1600,1624]" box="[144,298,1600,1624]" pageId="18" pageNumber="1246">
<emphasis id="B969EAEC554EFF82FF09F9E5FEBF5165" box="[144,298,1600,1624]" italics="true" pageId="18" pageNumber="1246">Type material</emphasis>
</paragraph>
<paragraph id="8BA236FE554EFF82FF09F9C2FC8F51A2" blockId="18.[144,1104,1639,1696]" pageId="18" pageNumber="1246">
<materialsCitation id="3B753CA3554EFF82FF09F9C2FC8F51A2" collectingDate="1971-08-18" collectingDateMax="1971-08-22" collectingDateMin="1971-08-18" collectorName="J. S. Son" country="South Korea" county="Clitellate" location="Mt Songni" municipality="Boeun-gun" pageId="18" pageNumber="1246" specimenCount="1" stateProvince="Chungbuk" typeStatus="holotype">
<typeStatus id="54A6885C554EFF82FF09F9C2FF6A5142" box="[144,255,1639,1663]" pageId="18" pageNumber="1246" type="holotype">Holotype</typeStatus>
:
<collectingCounty id="62C34E72554EFF82FE94F9C2FEE65142" box="[269,371,1639,1663]" pageId="18" pageNumber="1246">Clitellate</collectingCounty>
(NIBRIV0000120471) specimen:
<collectingCountry id="F30A766E554EFF82FD67F9CDFCDC5142" box="[766,841,1640,1663]" name="South Korea" pageId="18" pageNumber="1246">Korea</collectingCountry>
,
<collectingRegion id="49D9F81C554EFF82FCC1F9C2FBDE5142" box="[856,1099,1639,1663]" country="South Korea" name="Chungbuk" pageId="18" pageNumber="1246">Chungcheongbuk-do</collectingRegion>
,
<collectingMunicipality id="6BC6AC84554EFF82FF09F92DFE85519D" box="[144,272,1672,1696]" pageId="18" pageNumber="1246">Boeun-gun</collectingMunicipality>
,
<location id="8EC26025554EFF82FE82F92DFE0251A2" LSID="urn:lsid:plazi:treatment:03B487E8554EFF84FE54FA72FBBE5232:8EC26025554EFF82FE82F92DFE0251A2" box="[283,407,1671,1695]" country="South Korea" county="Clitellate" municipality="Boeun-gun" name="Mt Songni" pageId="18" pageNumber="1246" stateProvince="Chungbuk">Mt Songni</location>
,
<date id="FFA3103E554EFF82FE3BF922FDE951A2" box="[418,636,1671,1695]" pageId="18" pageNumber="1246" value="1971-08-18" valueMax="1971-08-22" valueMin="1971-08-18">
<collectingDate id="EFE7E9D6554EFF82FE3BF922FDE951A2" box="[418,636,1671,1695]" pageId="18" pageNumber="1246" value="1971-08-18" valueMax="1971-08-22" valueMin="1971-08-18">18–22 August 1971</collectingDate>
</date>
,
<collectorName id="26E85328554EFF82FD11F922FD7651A2" box="[648,739,1671,1695]" pageId="18" pageNumber="1246">J.S. Son</collectorName>
coll.
</materialsCitation>
</paragraph>
<caption id="DF626676554FFF83FF4DFBF0FC2D5356" ID-DOI="http://doi.org/10.5281/zenodo.4651920" ID-Zenodo-Dep="4651920" box="[212,952,1109,1131]" httpUri="https://zenodo.org/record/4651920/files/figure.png" pageId="19" pageNumber="1247" startId="19.[212,279,1109,1131]" targetBox="[522,860,168,1077]" targetPageId="19">
<paragraph id="8BA236FE554FFF83FF4DFBF0FC2D5356" blockId="19.[212,952,1109,1131]" box="[212,952,1109,1131]" pageId="19" pageNumber="1247">
Figure 10.
<taxonomicName id="4C1D4D7D554FFF83FED1FBF0FD805356" authority="Hong &amp; James, 2009" authorityName="Hong &amp; James" authorityYear="2009" box="[328,533,1109,1131]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="19" pageNumber="1247" phylum="Annelida" rank="species" species="calculatus" status="sp. nov.">
<emphasis id="B969EAEC554FFF83FED1FBF0FD805356" box="[328,533,1109,1131]" italics="true" pageId="19" pageNumber="1247">Amynthas calculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A5797554FFF83FD85FBF0FDFA5357" box="[540,623,1109,1130]" pageId="19" pageNumber="1247" rank="species">sp. nov.</taxonomicNameLabel>
: ventral view and spermathecae.
</paragraph>
</caption>
<paragraph id="8BA236FE554FFF83FF4DFB06FED85386" blockId="19.[212,333,1187,1211]" box="[212,333,1187,1211]" pageId="19" pageNumber="1247">
<emphasis id="B969EAEC554FFF83FF4DFB06FED85386" box="[212,333,1187,1211]" italics="true" pageId="19" pageNumber="1247">Etymology</emphasis>
</paragraph>
<paragraph id="8BA236FE554FFF83FF4DFB6EFE51523E" blockId="19.[212,1172,1227,1283]" pageId="19" pageNumber="1247">
The name
<taxonomicName id="4C1D4D7D554FFF83FED2FB6EFE2C53DE" authorityName="Hong &amp; James" authorityYear="2009" box="[331,441,1227,1251]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="19" pageNumber="1247" phylum="Annelida" rank="species" species="calculatus">
<emphasis id="B969EAEC554FFF83FED2FB6EFE2C53DE" box="[331,441,1227,1251]" italics="true" pageId="19" pageNumber="1247">calculatus</emphasis>
</taxonomicName>
is Latin for provided with pebbles, here referring to the shape of the male pore region.
</paragraph>
<paragraph id="8BA236FE554FFF83FF4DFA91FED45271" blockId="19.[212,321,1332,1356]" box="[212,321,1332,1356]" pageId="19" pageNumber="1247">
<emphasis id="B969EAEC554FFF83FF4DFA91FED45271" box="[212,321,1332,1356]" italics="true" pageId="19" pageNumber="1247">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BA236FE554FFF83FF4DFAFEFB8F528E" blockId="19.[212,1172,1371,1459]" pageId="19" pageNumber="1247">
Three pairs of spermathecal pores, presetal in VI–VIII just below mid-lateral,
<quantity id="4CE59B1B554FFF83FF4DFADEFEAA52AE" box="[212,319,1403,1427]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="1247" unit="mm" value="0.3">0.30 mm</quantity>
circumference apart; male field composed of two 2.7–3.8-mm diameter elevated discs with complex ornamentation,
<quantity id="4CE59B1B554FFF83FD50FA3EFCBE528E" box="[713,811,1435,1459]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="1247" unit="mm" value="0.25">0.25 mm</quantity>
circumference apart.
</paragraph>
<paragraph id="8BA236FE554FFF83FF4DFA41FEC152C1" blockId="19.[212,340,1508,1532]" box="[212,340,1508,1532]" pageId="19" pageNumber="1247">
<emphasis id="B969EAEC554FFF83FF4DFA41FEC152C1" box="[212,340,1508,1532]" italics="true" pageId="19" pageNumber="1247">Description</emphasis>
</paragraph>
<paragraph id="8BA236FE554FFF83FF4DF9A9FC26519E" blockId="19.[212,1172,1547,1699]" pageId="19" pageNumber="1247">
Brown dorsal pigment. Dimensions 115 by
<quantity id="4CE59B1B554FFF83FD50F9AEFCB5511E" box="[713,800,1547,1571]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="19" pageNumber="1247" unit="mm" value="4.6">4.6 mm</quantity>
at segment X,
<quantity id="4CE59B1B554FFF83FC48F9AEFBB2511E" box="[977,1063,1547,1571]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="19" pageNumber="1247" unit="mm" value="4.2">4.2 mm</quantity>
at XXX,
<quantity id="4CE59B1B554FFF83FF4DF98EFEA5517E" box="[212,304,1579,1603]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="19" pageNumber="1247" unit="mm" value="4.5">4.5 mm</quantity>
at clitellum; body cylindrical throughout, segments 99. Setae regularly distributed around segmental equators, numbering 36 at VIII, 47 at XX; four between male pores; setal formula AA: AB: YZ: ZZ = 5.5: 2: 4: 4 at XIII. Female pore single in XIV,
<quantity id="4CE59B1B554FFF83FE28F92EFD9D519E" box="[433,520,1675,1699]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="1247" unit="mm" value="0.7">0.7 mm</quantity>
, oval-shaped. First dorsal pore 12/13.
</paragraph>
<paragraph id="8BA236FE5548FF84FF23FF02FE745682" blockId="20.[144,1104,167,863]" pageId="20" pageNumber="1248">
Male field composed of two 2.7–3.8-mm diameter elevated discs with complex ornamentation,
<quantity id="4CE59B1B5548FF84FED1FF62FE3957E2" box="[328,428,199,223]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="1248" unit="mm" value="0.25">0.25 mm</quantity>
circumference apart. Paired epidermal thickenings centred on XVIII, extending from posterior quarter of XVII to anterior quarter of XIX, thickenings
<quantity id="4CE59B1B5548FF84FE83FEA2FE355622" box="[282,416,263,287]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.8" metricValueMin="2.7" pageId="20" pageNumber="1248" unit="mm" value="3.25" valueMax="3.8" valueMin="2.7">2.7–3.8 mm</quantity>
long; each with transverse pebble-textured ridge equatorial on XVIII and ring-shaped grooves anterior, posterior to ridge, two pairs of rounded quadrangular papillae within the paired epidermal thickenings. Three pairs of spermathecal pores
<quantity id="4CE59B1B5548FF84FEDDFEC2FE5C5642" box="[324,457,359,383]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="20" pageNumber="1248" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
conspicuous, presetal in VI–VIII just below mid-lateral, some furrowed, hardened areas on segmental equators VI–VIII, spermathecal pores
<quantity id="4CE59B1B5548FF84FF09FE02FF645682" box="[144,241,423,447]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="1248" unit="mm" value="0.3">0.30 mm</quantity>
circumference apart.
</paragraph>
<paragraph id="8BA236FE5548FF84FF23FE62FD0C5542" blockId="20.[144,1104,167,863]" pageId="20" pageNumber="1248">Septa 5/6, 6/7, 7/8 thick, 8/9, 9/10 absent, 10/11–13/14 thickly muscular. Gizzard globular in VIII–X. Intestine begins XV, lymph glands small from XXVII. Typhlosole medium size fold from XXVII. Hearts X–XIII oesophageal, IX lateral, X and XI inside testes sacs. Intestinal caeca simple, originating in XXVII, and extending anteriorly to about XXVI, each consisting of a finger-shaped sac, each finger with seven or eight serriform outgrowths on ventral margin.</paragraph>
<paragraph id="8BA236FE5548FF84FF23FD22FD7E5462" blockId="20.[144,1104,167,863]" pageId="20" pageNumber="1248">Ovaries in XIII. Three pairs of spermathecae VI–VIII; each ampulla medium size voluminous ovate pouch, ducts short, thin, diverticula club-shaped, longer than ducts, shorter than ampulla; no nephridia on spermathecal ducts. Male sexual system holandric, testes and funnels X, XI in ring-shaped sacs formed by dorsal and ventral fusion, enclosing other segmental contents of X, XI. Seminal vesicles in XI, XII with small dorsal appendage. Prostates XVIII, extending to XVII–XX, each divided into two or three main parts, ducts thick, short, muscular.</paragraph>
<paragraph id="8BA236FE5548FF84FF09FC2AFF66549A" blockId="20.[144,243,911,935]" box="[144,243,911,935]" pageId="20" pageNumber="1248">
<emphasis id="B969EAEC5548FF84FF09FC2AFF66549A" box="[144,243,911,935]" italics="true" pageId="20" pageNumber="1248">Remarks</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30765755548FF84FF09FC12FBBE5232" pageId="20" pageNumber="1248" type="reference_group">
<paragraph id="8BA236FE5548FF84FF09FC12FBBE5232" blockId="20.[144,1104,951,1295]" pageId="20" pageNumber="1248">
<taxonomicName id="4C1D4D7D5548FF84FF09FC12FEE254F2" authority="Hong &amp; James, 2009" authorityName="Hong &amp; James" authorityYear="2009" box="[144,375,951,975]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="calculatus" status="sp. nov.">
<emphasis id="B969EAEC5548FF84FF09FC12FEE254F2" box="[144,375,951,975]" italics="true" pageId="20" pageNumber="1248">Amynthas calculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57975548FF84FE19FC1DFE4954F2" box="[384,476,952,975]" pageId="20" pageNumber="1248" rank="species">sp. nov.</taxonomicNameLabel>
and the next three species, which have intrasegmental spermathecal pores on VI, VII and VIII, key out to the
<taxonomicName id="4C1D4D7D5548FF84FC87FC72FCE554D2" baseAuthorityName="Rosa" baseAuthorityYear="1890" box="[798,880,983,1007]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="bournei">
<emphasis id="B969EAEC5548FF84FC87FC72FCE554D2" box="[798,880,983,1007]" italics="true" pageId="20" pageNumber="1248">bournei</emphasis>
</taxonomicName>
-group in
<bibRefCitation id="EF8C4B0F5548FF84FC7FFC72FEBE5332" author="Sims RW &amp; Easton EG" pageId="20" pageNumber="1248" pagination="169 - 268" refId="ref11457" refString="Sims RW, Easton EG. 1972. A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biol J Linnean Soc, London. 4: 169 - 268." type="journal article" year="1972">Sims and Easton (1972)</bibRefCitation>
. The
<taxonomicName id="4C1D4D7D5548FF84FEFFFC52FE205332" baseAuthorityName="Rosa" baseAuthorityYear="1890" box="[358,437,1015,1039]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="bournei">
<emphasis id="B969EAEC5548FF84FEFFFC52FE205332" box="[358,437,1015,1039]" italics="true" pageId="20" pageNumber="1248">bournei</emphasis>
</taxonomicName>
-group is composed of seven species:
<taxonomicName id="4C1D4D7D5548FF84FCD1FC52FBD95332" authority="(Rosa, 1890)" baseAuthorityName="Rosa" baseAuthorityYear="1890" box="[840,1100,1015,1039]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="bournei">
<emphasis id="B969EAEC5548FF84FCD1FC52FC225332" box="[840,951,1015,1039]" italics="true" pageId="20" pageNumber="1248">A. bournei</emphasis>
(Rosa, 1890)
</taxonomicName>
,
<taxonomicName id="4C1D4D7D5548FF84FF09FBB2FE2C5312" authority="(Gates, 1932)" baseAuthorityName="Gates" baseAuthorityYear="1932" box="[144,441,1047,1071]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="sulcatus">
<emphasis id="B969EAEC5548FF84FF09FBB2FE9B5312" box="[144,270,1047,1071]" italics="true" pageId="20" pageNumber="1248">A. sulcatus</emphasis>
(Gates, 1932)
</taxonomicName>
,
<taxonomicName id="4C1D4D7D5548FF84FE50FBB2FD665312" authority="(Chen, 1946)" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[457,755,1047,1071]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="domosus">
<emphasis id="B969EAEC5548FF84FE50FBB2FDD85312" box="[457,589,1047,1071]" italics="true" pageId="20" pageNumber="1248">A. domosus</emphasis>
(
<bibRefCitation id="EF8C4B0F5548FF84FDF8FBB2FD7F5312" author="Chen Y" box="[609,746,1047,1071]" pageId="20" pageNumber="1248" pagination="83 - 141" refId="ref10776" refString="Chen Y. 1946. On the terrestrial Oligochaeta from Szechwan III. J West China Border Res Soc. 16: 83 - 141." type="journal article" year="1946">Chen, 1946</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C1D4D7D5548FF84FC9AFBB2FBDE5312" authority="(Chen, 1946)" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[771,1099,1047,1071]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="mucrorimus">
<emphasis id="B969EAEC5548FF84FC9AFBB2FC325312" box="[771,935,1047,1071]" italics="true" pageId="20" pageNumber="1248">A. mucrorimus</emphasis>
(
<bibRefCitation id="EF8C4B0F5548FF84FC23FBB2FBD65312" author="Chen Y" box="[954,1091,1047,1071]" pageId="20" pageNumber="1248" pagination="83 - 141" refId="ref10776" refString="Chen Y. 1946. On the terrestrial Oligochaeta from Szechwan III. J West China Border Res Soc. 16: 83 - 141." type="journal article" year="1946">Chen, 1946</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C1D4D7D5548FF84FF09FB92FE635372" authority="(Kobayashi, 1936)" baseAuthorityName="Kobayashi" baseAuthorityYear="1936" box="[144,502,1079,1103]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="koryoensis">
<emphasis id="B969EAEC5548FF84FF09FB92FE8A5372" box="[144,287,1079,1103]" italics="true" pageId="20" pageNumber="1248">A. koryoensis</emphasis>
(
<bibRefCitation id="EF8C4B0F5548FF84FEB6FB92FE7B5372" author="Kobayashi S" box="[303,494,1079,1103]" pageId="20" pageNumber="1248" pagination="139 - 184" refId="ref11271" refString="Kobayashi S. 1936. Earthworms from Koryo, Korea. Sci Rep Tohoku Imperial Univ, Tokyo. 11: 139 - 184." type="journal article" year="1936">Kobayashi, 1936</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C1D4D7D5548FF84FD9BFB92FCB85372" authority="(Kobayashi, 1936)" baseAuthorityName="Kobayashi" baseAuthorityYear="1936" box="[514,813,1079,1103]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="vallis">
<emphasis id="B969EAEC5548FF84FD9BFB92FDCF5372" box="[514,602,1079,1103]" italics="true" pageId="20" pageNumber="1248">A. vallis</emphasis>
(
<bibRefCitation id="EF8C4B0F5548FF84FDF2FB92FCB35372" author="Kobayashi S" box="[619,806,1079,1103]" pageId="20" pageNumber="1248" pagination="139 - 184" refId="ref11271" refString="Kobayashi S. 1936. Earthworms from Koryo, Korea. Sci Rep Tohoku Imperial Univ, Tokyo. 11: 139 - 184." type="journal article" year="1936">Kobayashi, 1936</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C1D4D7D5548FF84FCF1FB92FF5C5352" authority="Hong, 2001" authorityName="Hong" authorityYear="2001" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="pagyeiensis">
<emphasis id="B969EAEC5548FF84FCF1FB92FB975372" box="[872,1026,1079,1103]" italics="true" pageId="20" pageNumber="1248">A. pagyeiensis</emphasis>
Hong, 2001
</taxonomicName>
. Both
<taxonomicName id="4C1D4D7D5548FF84FE8FFBF2FE025352" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[278,407,1111,1135]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="domosus">
<emphasis id="B969EAEC5548FF84FE8FFBF2FE025352" box="[278,407,1111,1135]" italics="true" pageId="20" pageNumber="1248">A. domosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D7D5548FF84FE4AFBF2FDE25352" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[467,631,1111,1135]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="mucrorimus">
<emphasis id="B969EAEC5548FF84FE4AFBF2FDE25352" box="[467,631,1111,1135]" italics="true" pageId="20" pageNumber="1248">A. mucrorimus</emphasis>
</taxonomicName>
, from
<collectingCountry id="F30A766E5548FF84FD51FBF2FC855352" box="[712,784,1111,1135]" name="China" pageId="20" pageNumber="1248">China</collectingCountry>
, have genital markings, but
<taxonomicName id="4C1D4D7D5548FF84FF09FBD2FEB653B2" baseAuthorityName="Kobayashi" baseAuthorityYear="1936" box="[144,291,1143,1167]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="koryoensis">
<emphasis id="B969EAEC5548FF84FF09FBD2FEB653B2" box="[144,291,1143,1167]" italics="true" pageId="20" pageNumber="1248">A. koryoensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D7D5548FF84FEA8FBD2FE1C53B2" baseAuthorityName="Kobayashi" baseAuthorityYear="1936" box="[305,393,1143,1167]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="vallis">
<emphasis id="B969EAEC5548FF84FEA8FBD2FE1C53B2" box="[305,393,1143,1167]" italics="true" pageId="20" pageNumber="1248">A. vallis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D7D5548FF84FE01FBD2FDA653B2" authorityName="Hong" authorityYear="2001" box="[408,563,1143,1167]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="pagyeiensis">
<emphasis id="B969EAEC5548FF84FE01FBD2FDA653B2" box="[408,563,1143,1167]" italics="true" pageId="20" pageNumber="1248">A. pagyeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D7D5548FF84FDD8FBD2FD5B53B2" authority="Hong &amp; James, 2009" authorityName="Hong &amp; James" authorityYear="2009" box="[577,718,1143,1167]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="calculatus" status="sp. nov.">
<emphasis id="B969EAEC5548FF84FDD8FBD2FD5B53B2" box="[577,718,1143,1167]" italics="true" pageId="20" pageNumber="1248">A. calculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57975548FF84FD4CFBDDFCB853B2" box="[725,813,1144,1167]" pageId="20" pageNumber="1248" rank="species">sp. nov.</taxonomicNameLabel>
and the next three species of the
<taxonomicName id="4C1D4D7D5548FF84FF40FB32FEBD5392" baseAuthorityName="Rosa" baseAuthorityYear="1890" box="[217,296,1175,1199]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="bournei">
<emphasis id="B969EAEC5548FF84FF40FB32FEBD5392" box="[217,296,1175,1199]" italics="true" pageId="20" pageNumber="1248">bournei</emphasis>
</taxonomicName>
-group have no genital markings. All these Korean
<taxonomicName id="4C1D4D7D5548FF84FCFCFB32FC215392" baseAuthorityName="Rosa" baseAuthorityYear="1890" box="[869,948,1175,1199]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="bournei">
<emphasis id="B969EAEC5548FF84FCFCFB32FC215392" box="[869,948,1175,1199]" italics="true" pageId="20" pageNumber="1248">bournei</emphasis>
</taxonomicName>
-group species have male fields with disc-shaped porophores. The present new species differs from other
<taxonomicName id="4C1D4D7D5548FF84FF48FB72FEB453D2" baseAuthorityName="Rosa" baseAuthorityYear="1890" box="[209,289,1239,1263]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="20" pageNumber="1248" phylum="Annelida" rank="species" species="bournei">
<emphasis id="B969EAEC5548FF84FF48FB72FEB453D2" box="[209,289,1239,1263]" italics="true" pageId="20" pageNumber="1248">bournei</emphasis>
</taxonomicName>
-group species with respect to the male disc shape, and it has conspicuous spermathecal pores, presetal in hardened areas on VI–VIII just below mid-lateral.
</paragraph>
</subSubSection>
</treatment>
</document>
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