treatments-xml/data/03/B4/87/03B487C9A21EFF84FF54FCE6A41F7D74.xml
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<document id="FD9BE58F54F44042D9A5EB25E4E5AEF0" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629572761108" checkinUser="felipe" docAuthor="Grismer, L. Lee, Grismer, Jesse L., Wood, Perry L. &amp; Onn, Chan Kin" docDate="2008" docId="03B487C9A21EFF84FF54FCE6A41F7D74" docLanguage="en" docName="zt01931p024.pdf" docOrigin="Zootaxa 1931" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Cnemaspis bayuensis Grismer &amp; Grismer &amp; Wood &amp; Onn 2008, sp. nov." docType="treatment" docVersion="2" lastPageNumber="9" masterDocId="FF8DFFB1A218FF8CFFC3FFD9A0787816" masterDocTitle="The distribution, taxonomy, and redescription of the geckos Cnemaspis affinis (Stoliczka 1887) and C. flavolineata (Nicholls 1949) with descriptions of a new montane species and two new lowland, karst- dwelling species from Peninsular Malaysia" masterLastPageNumber="24" masterPageNumber="1" pageNumber="7" updateTime="1699077686533" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="460103C1FC4255315D32B0DBE364FD8F">The distribution, taxonomy, and redescription of the geckos Cnemaspis affinis (Stoliczka 1887) and C. flavolineata (Nicholls 1949) with descriptions of a new montane species and two new lowland, karst- dwelling species from Peninsular Malaysia</mods:title>
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<mods:namePart id="403B581C01263B6D7094C0F5A8C4B7AD">Grismer, L. Lee</mods:namePart>
<mods:affiliation id="FB4B37E20E66F79C85526E97D433F63A">Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California, 92515 - 8247 USA. &amp; Institute for Environment and Development (LESTARI), Universiti Kebangsaan Malaysia, 43600 Bangi, Selangor Darul Ehsan, Malaysia. E-mail: kin _ onn @ yahoo. com</mods:affiliation>
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<mods:namePart id="BEB2DAE38979009226B0A454E6CB7170">Grismer, Jesse L.</mods:namePart>
<mods:affiliation id="EBBFEA3078968EF678548EA10E38F543">Department of Biology, Villanova University, 800 Lancaster Avenue, Villanova, Pensylvania 92108.</mods:affiliation>
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<mods:namePart id="57E211FAE8FAF5219F874F0EE45D9601">Wood, Perry L.</mods:namePart>
<mods:affiliation id="80F7E50383FC3EC162AC52B7DFE8DA95">Department of Biology, Villanova University, 800 Lancaster Avenue, Villanova, Pensylvania 92108.</mods:affiliation>
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<mods:namePart id="76551B74BA40B56B3434C1A4BC4B9E59">Onn, Chan Kin</mods:namePart>
<mods:affiliation id="EFBEB8D2EAD89ADDD4F4268D091582D1">Institute for Environment and Development (LESTARI), Universiti Kebangsaan Malaysia, 43600 Bangi, Selangor Darul Ehsan, Malaysia. E-mail: kin _ onn @ yahoo. com</mods:affiliation>
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<mods:title id="D8FFEFD25F4393F0D3B8A3159DB5363C">Zootaxa</mods:title>
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<mods:date id="76EDF4D1A7FF7CAA182950D5FFC02CBA">2008</mods:date>
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<mods:number id="1A63506AD79FDCA53613E110D306E0B8">2008-11-12</mods:number>
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<mods:number id="586C61697756D116A0F900D2CF600BCD">1931</mods:number>
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<treatment id="03B487C9A21EFF84FF54FCE6A41F7D74" LSID="urn:lsid:plazi:treatment:03B487C9A21EFF84FF54FCE6A41F7D74" httpUri="http://treatment.plazi.org/id/03B487C9A21EFF84FF54FCE6A41F7D74" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="C3076554A21EFF8AFF54FCE6A1DD7B6A" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BA236DFA21EFF8AFF54FCE6A1857B4F" blockId="6.[151,509,831,932]" box="[151,509,831,857]" pageId="6" pageNumber="7">
<heading id="D0EA81B3A21EFF8AFF54FCE6A1857B4F" bold="true" box="[151,509,831,857]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B969EACDA21EFF8AFF54FCE6A1857B4F" bold="true" box="[151,509,831,857]" pageId="6" pageNumber="7">
<taxonomicName id="4C1D4D5CA21EFF8AFF54FCE6A1E17B4F" authority="Grismer &amp; Grismer &amp; Wood &amp; Onn, 2008" authorityName="Grismer &amp; Grismer &amp; Wood &amp; Onn" authorityYear="2008" box="[151,409,831,857]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="bayuensis" status="sp. nov.">
<emphasis id="B969EACDA21EFF8AFF54FCE6A1E17B4F" bold="true" box="[151,409,831,857]" italics="true" pageId="6" pageNumber="7">Cnemaspis bayuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57B6A21EFF8AFE63FCE6A1857B4F" box="[416,509,831,857]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="8BA236DFA21EFF8AFF54FCBBA1DD7B6A" blockId="6.[151,509,831,932]" box="[151,421,866,892]" pageId="6" pageNumber="7">
<collectingRegion id="49D9F83DA21EFF8AFF54FCBBA0B07B6A" box="[151,200,866,892]" country="India" name="Goa" pageId="6" pageNumber="7">Gua</collectingRegion>
Bayu Rock Gecko
</paragraph>
</subSubSection>
<subSubSection id="C3076554A21EFF8AFF54FC53A15A7BB2" box="[151,290,906,932]" pageId="6" pageNumber="7" type="description">
<paragraph id="8BA236DFA21EFF8AFF54FC53A15A7BB2" blockId="6.[151,509,831,932]" box="[151,290,906,932]" pageId="6" pageNumber="7">
<figureCitation id="13262A5AA21EFF8AFF54FC53A17E7BB2" box="[151,262,906,932]" captionStart="FIGURE 2" captionStartId="5.[151,255,1811,1835]" captionTargetBox="[213,1371,513,1787]" captionTargetId="figure-219@5.[213,1377,513,1787]" captionTargetPageId="5" captionText="FIGURE 2. Upper left; juvenile female Cnemaspis affinis from Pulau Pinang, Penang (LSUHC 8975). Middle left; adult male holotype of C. bayuensis from Kampung Bayu, Kelantan (ZRC 2.6759). Lower left; adult female non-striped form of C. flavolineata from Cameron Highlands, Pahang (LSUDPC 3456). Upper right; adult female paratype of C. karsticola from Gunung Reng, Kelantan (ZRC 2.6763). Middle right; adult female C. mcguirei in night color phase from Bukit Larut, Perak (LSUDPC 4441). Lower right; adult female striped form of C. flavolineata from Frasers Hill, Pahang (LSUHC 8079)." pageId="6" pageNumber="7">Figures 2</figureCitation>
,
<figureCitation id="13262A5AA21EFF8AFED0FC53A15A7BB2" box="[275,290,906,932]" captionStart="FIGURE 3" captionStartId="7.[151,255,1016,1040]" captionTargetBox="[151,1436,277,989]" captionTargetId="figure-400@7.[151,1436,277,989]" captionTargetPageId="7" captionText="FIGURE 3. Type series of Cnemaspis bayuensis. Left to right; holotype ZRC 2.6759, paratypes ZRC 2.676061, LSUHC 9073" pageId="6" pageNumber="7">3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3076554A21EFF8AFF54FC03A4B67C82" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BA236DFA21EFF8AFF54FC03A1317C52" blockId="6.[151,1437,986,2012]" pageId="6" pageNumber="7">
<materialsCitation id="3B753C82A21EFF8AFF54FC03A1317C52" collectingDate="2008-06-24" collectionCode="ZRC" collectorName="Adult &amp; Jesse L. Grismer &amp; Perry L. Wood, Jr." country="Malaysia" elevation="120" latitude="5.0941668" location="Gua Bayu" longLatPrecision="1" longitude="102.221085" municipality="Peninsular" pageId="6" pageNumber="7" specimenCode="ZRC 2.6759" specimenCount="1" stateProvince="Kelantan" typeStatus="holotype">
<emphasis id="B969EACDA21EFF8AFF54FC03A1687BE2" bold="true" box="[151,272,986,1012]" pageId="6" pageNumber="7">
<typeStatus id="54A6887DA21EFF8AFF54FC03A1737BE2" box="[151,267,986,1012]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectorName id="26E85309A21EFF8AFEDBFC03A1227BE2" box="[280,346,986,1012]" pageId="6" pageNumber="7">Adult</collectorName>
male (
<specimenCode id="DBBB9EA4A21EFF8AFE6FFC03A23A7BE2" box="[428,578,986,1012]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="7">ZRC 2.6759</specimenCode>
) collected on
<date id="FFA3101FA21EFF8AFD2AFC03A3FF7BE2" box="[745,903,986,1012]" pageId="6" pageNumber="7" value="2008-06-24">
<collectingDate id="EFE7E9F7A21EFF8AFD2AFC03A3FF7BE2" box="[745,903,986,1012]" pageId="6" pageNumber="7" value="2008-06-24">24 June 2008</collectingDate>
</date>
by
<collectorName id="26E85309A21EFF8AFC76FC03A4057BE2" box="[949,1149,986,1012]" pageId="6" pageNumber="7">Jesse L. Grismer</collectorName>
,
<collectorName id="26E85309A21EFF8AFB4AFC03A51C7BE2" box="[1161,1380,986,1012]" pageId="6" pageNumber="7">Perry L. Wood, Jr.</collectorName>
, and L. Lee Grismer at 1030 hrs at
<quantity id="4CE59B3AA21EFF8AFDEFFBDBA2067C0A" box="[556,638,1026,1052]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="7" unit="m" value="120.0">
<elevation id="0030D1ECA21EFF8AFDEFFBDBA2067C0A" box="[556,638,1026,1052]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="7" unit="m" value="120.0">120 m</elevation>
</quantity>
from
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,
<collectingRegion id="49D9F83DA21EFF8AFCAAFBDBA3AE7C0A" box="[873,982,1026,1052]" country="Malaysia" name="Kelantan" pageId="6" pageNumber="7">Kelantan</collectingRegion>
,
<collectingMunicipality id="6BC6ACA5A21EFF8AFC2BFBDBA41C7C0A" box="[1000,1124,1026,1052]" pageId="6" pageNumber="7">Peninsular</collectingMunicipality>
<collectingCountry id="F30A764FA21EFF8AFBB0FBDBA4A67C0A" box="[1139,1246,1026,1052]" name="Malaysia" pageId="6" pageNumber="7">Malaysia</collectingCountry>
(
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,
<geoCoordinate id="EE295018A21EFF8AFF54FBF3A1437C52" box="[151,315,1066,1092]" degrees="102" direction="east" minutes="13.265" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="102.221085">102°13.265 E</geoCoordinate>
).
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</paragraph>
<paragraph id="8BA236DFA21EFF8AFF05FB8BA4B67C82" blockId="6.[151,1437,986,2012]" pageId="6" pageNumber="7">
<materialsCitation id="3B753C82A21EFF8AFF05FB8BA4B27C82" collectionCode="ZRC, LSUHC" pageId="6" pageNumber="7" specimenCode="ZRC 2.6760, LSUHC 9073, ZRC 2.6761" specimenCount="3" typeStatus="paratype">
<emphasis id="B969EACDA21EFF8AFF05FB8BA1337C7A" bold="true" box="[198,331,1106,1132]" pageId="6" pageNumber="7">
<typeStatus id="54A6887DA21EFF8AFF05FB8BA13E7C7A" box="[198,326,1106,1132]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Collection locality, date and time of collection, and collectors of the
<typeStatus id="54A6887DA21EFF8AFBB0FB8BA49B7C7A" box="[1139,1251,1106,1132]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
are the same as that for the
<typeStatus id="54A6887DA21EFF8AFEDDFBA3A1F07C82" box="[286,392,1146,1172]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
.
<specimenCode id="DBBB9EA4A21EFF8AFE57FBA3A25D7C82" box="[404,549,1146,1172]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="7">ZRC 2.6760</specimenCode>
and
<specimenCode id="DBBB9EA4A21EFF8AFD9DFBA3A2877C82" box="[606,767,1146,1172]" collectionCode="LSUHC" country="0" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="6" pageNumber="7">LSUHC 9073</specimenCode>
are males and
<specimenCode id="DBBB9EA4A21EFF8AFC6CFBA3A4397C82" box="[943,1089,1146,1172]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="7">ZRC 2.6761</specimenCode>
is a female
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.
</paragraph>
</subSubSection>
<subSubSection id="C3076554A21EFF8AFF05FB7BA2587E62" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BA236DFA21EFF8AFF05FB7BA2587E62" blockId="6.[151,1437,986,2012]" pageId="6" pageNumber="7">
<emphasis id="B969EACDA21EFF8AFF05FB7BA1467CAA" bold="true" box="[198,318,1186,1212]" pageId="6" pageNumber="7">Diagnosis</emphasis>
.
<taxonomicName id="4C1D4D5CA21EFF8AFE8CFB7AA2347CAA" authorityName="Grismer &amp; Grismer &amp; Wood &amp; Onn" authorityYear="2008" box="[335,588,1187,1212]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="bayuensis">
<emphasis id="B969EACDA21EFF8AFE8CFB7AA2347CAA" box="[335,588,1187,1212]" italics="true" pageId="6" pageNumber="7">Cnemaspis bayuensis</emphasis>
</taxonomicName>
differs from all other Southeast Asian species of
<taxonomicName id="4C1D4D5CA21EFF8AFB61FB7AA55A7CAA" authorityName="Grismer &amp; Grismer &amp; Wood &amp; Onn" authorityYear="2008" box="[1186,1314,1187,1212]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B969EACDA21EFF8AFB61FB7AA55A7CAA" box="[1186,1314,1187,1212]" italics="true" pageId="6" pageNumber="7">Cnemaspis</emphasis>
</taxonomicName>
in having the unique combination of adult males reaching
<quantity id="4CE59B3AA21EFF8AFD14FB13A3387CF2" box="[727,832,1226,1252]" metricMagnitude="-2" metricUnit="m" metricValue="4.61" pageId="6" pageNumber="7" unit="mm" value="46.1">46.1 mm</quantity>
SVL, adult females reaching
<quantity id="4CE59B3AA21EFF8AFB65FB13A5777CF2" box="[1190,1295,1226,1252]" metricMagnitude="-2" metricUnit="m" metricValue="4.52" pageId="6" pageNumber="7" unit="mm" value="45.2">45.2 mm</quantity>
SVL; 910 supralabials; large, lateral postmentals separated at midline by 13 smaller postmentals; eight or nine infralabials; forearm, subtibials, ventrals, subcaudals, and dorsal tubercles keeled; 2530 paravertebral tubercles; tubercles on flanks, relatively small and not linearly arranged; lateral caudal tubercles may or may not be within lateral caudal furrow; ventrolateral caudal tubercles present anteriorly; median subcaudal row not enlarged or keeled; subcaudals keeled; two postcloacal tubercles; femoral pores absent; discontinuous or continuous row of 59 precloacal scales bearing pores; subtibial scales not shield-like; no enlarged submetatarsal scales; 2730 subdigital lamellae on fourth toe; no distinct, large, dark spots on neck; no dark shoulder patch enclosing ocelli; no light, postscapular band; white markings on flanks; indistinct, dark, caudal bands; subcaudal region pigmented, not immaculate. These differences are summarized across all Southeast Asian species in
<bibRefCitation id="EF8C4B2EA21EFF8AFF76F983A1C97E62" author="Grismer, L. L. &amp; Chan, K. O. &amp; Nurolhuda, N. &amp; Sumontha, M." box="[181,433,1626,1652]" pageId="6" pageNumber="7" pagination="51 - 68" refId="ref13238" refString="Grismer, L. L., Chan, K. O., Nurolhuda, N. &amp; Sumontha, M. (2008 b) A new species of karst dwelling gecko (genus Cnemaspis Strauch 1887) from the border region of Thailand and Peninsular Malaysia. Zootaxa, 1875, 51 - 68." type="journal article" year="2008">
Grismer
<emphasis id="B969EACDA21EFF8AFEDFF982A12F7E62" box="[284,343,1627,1652]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
(2008b
</bibRefCitation>
:
<tableCitation id="C69F0364A21EFF8AFE77F983A2777E62" box="[436,527,1626,1652]" captionStart="TABLE 1" captionStartId="4.[151,241,153,177]" captionText="TABLE 1. Descriptive measurements of the type series of Cnemaspis bayuensis. Holotype ZRC 2.6759." pageId="6" pageNumber="7">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
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<emphasis id="B969EACDA21EFF8AFF05F95BA1887E8A" bold="true" box="[198,496,1666,1692]" pageId="6" pageNumber="7">
Description of
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.
</emphasis>
Adult male; SVL
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; head oblong in dorsal profile, moderate in size (HL/ SVL 0.25), somewhat narrow (HW/SVL 0.17), flattened (HD/HL 0.42), distinct from neck; snout short (ES/ HL 0.49), slightly concave in lateral profile; postnasal region constricted medially, flat; scales of rostrum weakly keeled, slightly raised, larger than similarly shaped scales on occiput; low, supraorbital ridges; no frontorostral sulcus; canthus rostralis nearly absent, smoothly rounded; eye large (ED/HL 0.22); extra-brillar fringe scales largest anteriorly; pupil round; ear opening oval, taller than wide; rostral slightly concave, dorsal 80% divided by longitudinal groove; rostral bordered posteriorly by supranasals and one small, azygous scale and laterally by first supralabials; 10R, 9L raised supralabials of similar size; 9R, 8L infralabials, slightly decreasing in size posteriorly; nostrils elliptical, oriented posterodorsally; bordered posteriorly by small, gran- ular, postnasal scales; mental large, triangular, bordered posteriorly by three large postmentals, outer two largest, not contacting medially; gular scales slightly raised, weakly keeled; throat scales larger, keeled.
</paragraph>
<caption id="DF626657A21FFF8BFF54FC21A1527C24" pageId="7" pageNumber="8" startId="7.[151,255,1016,1040]" targetBox="[151,1436,277,989]" targetPageId="7">
<paragraph id="8BA236DFA21FFF8BFF54FC21A1527C24" blockId="7.[151,1436,1016,1074]" pageId="7" pageNumber="8">
<emphasis id="B969EACDA21FFF8BFF54FC21A1667C06" bold="true" box="[151,286,1016,1040]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
Type series of
<emphasis id="B969EACDA21FFF8BFE17FC21A2BE7C19" box="[468,710,1016,1039]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C1D4D5CA21FFF8BFE17FC21A2BA7C19" authorityName="Grismer &amp; Grismer &amp; Wood &amp; Onn" authorityYear="2008" box="[468,706,1016,1039]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bayuensis">Cnemaspis bayuensis</taxonomicName>
.
</emphasis>
Left to right; holotype ZRC 2.6759, paratypes ZRC 2.676061, LSUHC 9073
</paragraph>
</caption>
<paragraph id="8BA236DFA21FFF8BFF05FBBCA1B27FF9" blockId="7.[151,1437,1125,2031]" pageId="7" pageNumber="8">Body slender, elongate; small, keeled, dorsal scales equal in size throughout body, intermixed with several large, multicarinate tubercles more or less randomly arranged; tubercles extend from occiput to base of tail; tubercles on flanks not enlarged, moderate in size; 27 paravertebral tubercles; pectoral and abdominal scales strongly keeled, not elongate, same size throughout; abdominal scales slightly larger than dorsals; eight continuous, precloacal scales bearing pores arranged in a chevron; precloacal depression absent; femoral pores absent; forelimbs moderately long, slender; dorsal scales of brachium not raised, weakly keeled; dorsal scales of forearm keeled, slightly raised; ventral scales of brachium and forearm smooth, raised, juxtaposed; palmar scales smooth, juxtaposed, raised; digits long with an inflected joint; claws recurved; subdigital lamellae unnotched; lamellae beneath first phalanges granular proximally, widened distally; lamellae beneath phalanx immediately following inflection granular, lamellae of distal phalanges wide; interdigital webbing present; fingers increase in length from first to fourth with fourth and fifth equal in length; hind limbs slightly longer and thicker than forelimbs; dorsal scales of thigh keeled, raised, juxtaposed; scales of anterior margin of thigh keeled; ventral scales of thigh weakly keeled; subtibial scales keeled, flat, imbricate, with no enlarged anterior row; plantar scales smooth, juxtaposed, raised; no enlarged submetatarsal scales beneath first metatarsal; digits elongate with an inflected jointed; claws recurved; subdigital lamellae unnotched; lamellae beneath first phalanges not granular proximally but wider distally; lamellae beneath phalanx immediately following inflection granular, lamellae of distal phalanges wide; interdigital webbing present; toes increase in length from first to fourth with fourth being slightly longer than fifth; 29 subdigital lamellae on fourth toe; dorsal caudal scales arranged in segmented whorls, flat anteriorly, weakly keeled, juxtaposed; shallow middorsal furrow; deeper, single lateral furrow; no enlarged, median subcaudal scales; subcaudal scales keeled; no median row of enlarged, keeled subcaudal scales; caudal tubercles do not encircle tail; caudal tubercles may or may not be present in lateral furrow; two enlarged, postcloacal tubercles on lateral surface of hemipenial swellings at base of tail; tail 0.82% of SVL,</paragraph>
<paragraph id="8BA236DFA210FF84FF05FF4EA19E79E4" blockId="8.[151,1437,151,1378]" pageId="8" pageNumber="9">
<emphasis id="B969EACDA210FF84FF05FF4EA27178A7" bold="true" box="[198,521,151,177]" pageId="8" pageNumber="9">
Coloration (in life,
<figureCitation id="13262A5AA210FF84FE7BFF4EA18778A7" box="[440,511,151,177]" captionStart="FIGURE 2" captionStartId="5.[151,255,1811,1835]" captionTargetBox="[213,1371,513,1787]" captionTargetId="figure-219@5.[213,1377,513,1787]" captionTargetPageId="5" captionText="FIGURE 2. Upper left; juvenile female Cnemaspis affinis from Pulau Pinang, Penang (LSUHC 8975). Middle left; adult male holotype of C. bayuensis from Kampung Bayu, Kelantan (ZRC 2.6759). Lower left; adult female non-striped form of C. flavolineata from Cameron Highlands, Pahang (LSUDPC 3456). Upper right; adult female paratype of C. karsticola from Gunung Reng, Kelantan (ZRC 2.6763). Middle right; adult female C. mcguirei in night color phase from Bukit Larut, Perak (LSUDPC 4441). Lower right; adult female striped form of C. flavolineata from Frasers Hill, Pahang (LSUHC 8079)." pageId="8" pageNumber="9">Fig. 2</figureCitation>
)
</emphasis>
. Dorsal ground color brown; head and body overlain with irregularly shaped dark spots; light markings on occiput; rostrum yellowish with dark streaks; single, thin, dark, postorbital stripes extending onto nape but not contacting medially; dark, anteriorly projecting, triangular marking between them; paravertebral, white markings on nape followed by distinct, white, alternating, paravertebral blotches extending to base of tail; distinct, transversely elongate, white markings on flanks; diffuse brown and mottled white bands encircle tail; irregularly shaped, dark and light markings on limbs; dark and light, diffuse bands encircling digits; gular region with faint, brown, reticulate pattern; ventral surfaces of body and limbs dull beige, immaculate, darkening laterally; subcaudal region suffused with pigment. There is no sexual dimorphism in color pattern.
</paragraph>
<paragraph id="8BA236DFA210FF84FF05FE26A0D47AF4" blockId="8.[151,1437,151,1378]" pageId="8" pageNumber="9">
<emphasis id="B969EACDA210FF84FF05FE26A13B7A0F" bold="true" box="[198,323,511,537]" pageId="8" pageNumber="9">Variation.</emphasis>
<typeStatus id="54A6887DA210FF84FE89FDD9A1C47A0C" box="[330,444,512,538]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
approach the
<typeStatus id="54A6887DA210FF84FDA3FDD9A2BF7A0C" box="[608,711,512,538]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
in coloration and pattern (
<figureCitation id="13262A5AA210FF84FC3FFDD9A43B7A0C" box="[1020,1091,512,538]" captionStart="FIGURE 3" captionStartId="7.[151,255,1016,1040]" captionTargetBox="[151,1436,277,989]" captionTargetId="figure-400@7.[151,1436,277,989]" captionTargetPageId="7" captionText="FIGURE 3. Type series of Cnemaspis bayuensis. Left to right; holotype ZRC 2.6759, paratypes ZRC 2.676061, LSUHC 9073" pageId="8" pageNumber="9">Fig. 3</figureCitation>
).
<materialsCitation id="3B753C82A210FF84FB99FDD9A3357A7C" collectionCode="ZRC" pageId="8" pageNumber="9" specimenCode="ZRC 2.6761" specimenCount="2" typeStatus="holotype">
<specimenCode id="DBBB9EA4A210FF84FB99FDD9A4937A0C" box="[1114,1259,512,538]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="9">ZRC 2.6761</specimenCode>
is darker overall with distinct, dark blotches on the body and thin, elongate, white markings on the flanks. Subcaudal keeling is faint in the
<typeStatus id="54A6887DA210FF84FEAAFD89A1AA7A7C" box="[361,466,592,618]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
, but prominent in the
<typeStatus id="54A6887DA210FF84FD1AFD89A3357A7C" box="[729,845,592,618]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
</materialsCitation>
.
<materialsCitation id="3B753C82A210FF84FC9AFD89A2B27A84" collectionCode="ZRC" pageId="8" pageNumber="9" specimenCode="ZRC 2.6760, 61" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBBB9EA4A210FF84FC9AFD89A3957A7C" box="[857,1005,592,618]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="9">ZRC 2.6760</specimenCode>
<specimenCode id="DBBB9EA4A210FF84FC3FFD89A46E7A7C" box="[1020,1046,592,618]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="9">61</specimenCode>
show evidence of diffuse caudal banding which is not present in the
<typeStatus id="54A6887DA210FF84FD81FDA1A2D07A84" box="[578,680,632,658]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
or
</materialsCitation>
<materialsCitation id="3B753C82A210FF84FD10FDA1A4D97A84" box="[723,1185,632,658]" collectionCode="LSUHC" pageId="8" pageNumber="9" specimenCode="LSUHC 9073" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBB9EA4A210FF84FD10FDA1A3027A84" box="[723,890,632,658]" collectionCode="LSUHC" country="0" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="8" pageNumber="9">LSUHC 9073</specimenCode>
. The tail is 1.20% of the
</materialsCitation>
SVL in
<materialsCitation id="3B753C82A210FF84FACAFDA1A1267AAC" collectionCode="ZRC" pageId="8" pageNumber="9" specimenCode="ZRC 2.6760" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBB9EA4A210FF84FACAFDA1A5E47A84" box="[1289,1436,632,658]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="9">ZRC 2.6760</specimenCode>
and 1.34% of the
</materialsCitation>
SVL in
<materialsCitation id="3B753C82A210FF84FE02FD79A0DF7AF4" collectionCode="ZRC" pageId="8" pageNumber="9" specimenCode="ZRC 2.6761" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBB9EA4A210FF84FE02FD79A22D7AAC" box="[449,597,672,698]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="9">ZRC 2.6761</specimenCode>
. Morphometric variation and variation in scalation is presented in
<tableCitation id="C69F0364A210FF84FA98FD79A0DF7AF4" captionStart="TABLE 1" captionStartId="4.[151,241,153,177]" captionText="TABLE 1. Descriptive measurements of the type series of Cnemaspis bayuensis. Holotype ZRC 2.6759." pageId="8" pageNumber="9">Table 1</tableCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3076554A210FF84FF05FD36A5E47CD4" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BA236DFA210FF84FF05FD36A21B7B24" blockId="8.[151,1437,151,1378]" pageId="8" pageNumber="9">
<emphasis id="B969EACDA210FF84FF05FD36A1277B1F" bold="true" box="[198,351,751,777]" pageId="8" pageNumber="9">Distribution</emphasis>
.
<taxonomicName id="4C1D4D5CA210FF84FEB3FD29A2167B1F" authorityName="Grismer &amp; Grismer &amp; Wood &amp; Onn" authorityYear="2008" box="[368,622,752,777]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="bayuensis">
<emphasis id="B969EACDA210FF84FEB3FD29A2167B1F" box="[368,622,752,777]" italics="true" pageId="8" pageNumber="9">Cnemaspis bayuensis</emphasis>
</taxonomicName>
is known only from the Gua Bayu Cave region at Kampung Bayu,
<collectingRegion id="49D9F83DA210FF84FF54FCC1A17C7B24" box="[151,260,792,818]" country="Malaysia" name="Kelantan" pageId="8" pageNumber="9">Kelantan</collectingRegion>
, Peninsular
<collectingCountry id="F30A764FA210FF84FE51FCC1A1857B24" box="[402,509,792,818]" name="Malaysia" pageId="8" pageNumber="9">Malaysia</collectingCountry>
(
<figureCitation id="13262A5AA210FF84FDCEFCC1A22A7B24" box="[525,594,792,818]" captionStart="FIGURE 1" captionStartId="2.[151,255,1292,1316]" captionTargetBox="[157,1428,203,1260]" captionTargetId="figure-376@2.[151,1436,197,1265]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of Cnemaspis affinis, C. bayuensis, C. flavolineata, C. karsticola, and C. mcguirei in Peninsular Malaysia and Thailand and the general contours and location of the Bintang, Titi Wangsa, and Timur ranges." pageId="8" pageNumber="9">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BA236DFA210FF84FF05FCE6A5E47CD4" blockId="8.[151,1437,151,1378]" pageId="8" pageNumber="9">
<emphasis id="B969EACDA210FF84FF05FCE6A1F77B4F" bold="true" box="[198,399,831,857]" pageId="8" pageNumber="9">Natural History</emphasis>
.
<taxonomicName id="4C1D4D5CA210FF84FE63FC99A2E57B4F" authorityName="Grismer &amp; Grismer &amp; Wood &amp; Onn" authorityYear="2008" box="[416,669,832,857]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="bayuensis">
<emphasis id="B969EACDA210FF84FE63FC99A2E57B4F" box="[416,669,832,857]" italics="true" pageId="8" pageNumber="9">Cnemaspis bayuensis</emphasis>
</taxonomicName>
<materialsCitation id="3B753C82A210FF84FD6BFC99A5E07CD4" collectionCode="ZRC" county="Bayu region" elevation="5" location="One" municipality="Specimens" pageId="8" pageNumber="9" specimenCode="ZRC 2.6760, ZRC 2.6759, ZRC 2.6761" specimenCount="1" specimenCount-egg="1" stateProvince="Goa" typeStatus="holotype">
is a saxicolous species that appears to be restricted to the cone karst outcroppings of the
<collectingRegion id="49D9F83DA210FF84FE01FCB1A18B7B94" box="[450,499,872,898]" country="India" name="Goa" pageId="8" pageNumber="9">Gua</collectingRegion>
<collectingCounty id="62C34E53A210FF84FE38FCB1A2F37B94" box="[507,651,872,898]" pageId="8" pageNumber="9">Bayu region</collectingCounty>
surrounding the village of Bayu. Here, specimens were found during the day along the periphery of the karst formations as opposed to being deep within the cave systems.
<collectingMunicipality id="6BC6ACA5A210FF84FF54FC61A16E7BC4" box="[151,278,952,978]" pageId="8" pageNumber="9">Specimens</collectingMunicipality>
were seen in cracks (
<figureCitation id="13262A5AA210FF84FDDDFC61A21F7BC4" box="[542,615,952,978]" captionStart="FIGURE 4" captionStartId="10.[151,255,1997,2021]" captionTargetBox="[151,1436,1117,1971]" captionTargetId="figure-375@10.[151,1436,1117,1971]" captionTargetPageId="10" captionText="FIGURE 4. Karst habitat of Cnemapsis bayuensis at Kampung Bayu, Kelantan." pageId="8" pageNumber="9">Fig. 4</figureCitation>
) as well as on the cave walls and ceilings as high as
<quantity id="4CE59B3AA210FF84FB33FC61A5677BC4" box="[1264,1311,952,978]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="m" value="5.0">
<elevation id="0030D1ECA210FF84FB33FC61A5677BC4" box="[1264,1311,952,978]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="m" value="5.0">5 m</elevation>
</quantity>
above the ground.
<location id="8EC26004A210FF84FF39FC39A1537BEC" LSID="urn:lsid:plazi:treatment:03B487C9A21EFF84FF54FCE6A41F7D74:8EC26004A210FF84FF39FC39A1537BEC" box="[250,299,992,1018]" county="Bayu region" municipality="Specimens" name="One" pageId="8" pageNumber="9" stateProvince="Goa">One</location>
specimen (
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) was found on the side of an isolated piece of karst on the forest floor.
<location id="8EC26004A210FF84FF54FBD1A0A87C34" LSID="urn:lsid:plazi:treatment:03B487C9A21EFF84FF54FCE6A41F7D74:8EC26004A210FF84FF54FBD1A0A87C34" box="[151,208,1032,1058]" county="Bayu region" municipality="Specimens" name="Nine" pageId="8" pageNumber="9" stateProvince="Goa">Nine</location>
individuals were observed and all matched the coloration of the substrate upon which they were sitting.
<specimenCode id="DBBB9EA4A210FF84FF54FBE9A1507C5C" box="[151,296,1072,1098]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="9">ZRC 2.6760</specimenCode>
was nearly black, matching the color of the lichen on the isolated karst fragment on which it was collected.
<location id="8EC26004A210FF84FED1FB81A1BB7C64" LSID="urn:lsid:plazi:treatment:03B487C9A21EFF84FF54FCE6A41F7D74:8EC26004A210FF84FED1FB81A1BB7C64" box="[274,451,1112,1138]" county="Bayu region" municipality="Specimens" name="Approximately" pageId="8" pageNumber="9" stateProvince="Goa">Approximately</location>
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</quantity>
away on the nearly white cave wall, the very light-colored
<typeStatus id="54A6887DA210FF84FB2BFB81A5367C64" box="[1256,1358,1112,1138]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
(
<specimenCode id="DBBB9EA4A210FF84FAA2FB81A0917C8C" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="9">ZRC 2.6759</specimenCode>
) was collected. The color pattern of all
<taxonomicName id="4C1D4D5CA210FF84FD13FB59A3287C8F" authorityName="Grismer &amp; Grismer &amp; Wood &amp; Onn" authorityYear="2008" box="[720,848,1152,1177]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B969EACDA210FF84FD13FB59A3287C8F" box="[720,848,1152,1177]" italics="true" pageId="8" pageNumber="9">Cnemaspis</emphasis>
</taxonomicName>
lightens significantly at night, however, it is not known how the color pattern of this species may change.
<specimenCode id="DBBB9EA4A210FF84FCFBFB71A3B17CD4" box="[824,969,1192,1218]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="9">ZRC 2.6761</specimenCode>
was a gravid female carrying
<specimenCount id="9D1BFD56A210FF84FAE8FB71A5E07CD4" box="[1323,1432,1192,1218]" pageId="8" pageNumber="9" type="egg">two eggs</specimenCount>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3076554A210FF84FF06FB16A3D87D2F" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BA236DFA210FF84FF06FB16A3D87D2F" blockId="8.[151,1437,151,1378]" pageId="8" pageNumber="9">
<emphasis id="B969EACDA210FF84FF06FB16A12A7CFF" bold="true" box="[197,338,1231,1257]" pageId="8" pageNumber="9">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C1D4D5CA210FF84FD95FB09A2B17CFF" authorityName="Grismer &amp; Grismer &amp; Wood &amp; Onn" authorityYear="2008" box="[598,713,1232,1257]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="bayuensis">
<emphasis id="B969EACDA210FF84FD95FB09A2B17CFF" box="[598,713,1232,1257]" italics="true" pageId="8" pageNumber="9">bayuensis</emphasis>
</taxonomicName>
is in reference to the
<typeStatus id="54A6887DA210FF84FC1EFB09A4777CFC" box="[989,1039,1232,1258]" pageId="8" pageNumber="9">type</typeStatus>
locality of Kampung Bayu. The Latin suffix -
<emphasis id="B969EACDA210FF84FEF3FB21A1137D07" box="[304,363,1272,1297]" italics="true" pageId="8" pageNumber="9">ensis</emphasis>
is a derivation meaning “from” or “inhabiting” and renders the specific epithet an adjective that must be in grammatical accord with the gender of
<emphasis id="B969EACDA210FF84FCDBFAF9A3D87D2F" box="[792,928,1312,1337]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C1D4D5CA210FF84FCDBFAF9A3E47D2F" authorityName="Grismer &amp; Grismer &amp; Wood &amp; Onn" authorityYear="2008" box="[792,924,1312,1337]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">Cnemaspis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3076554A210FF84FF06FA9EA41F7D74" box="[197,1127,1351,1378]" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BA236DFA210FF84FF06FA9EA41F7D74" blockId="8.[151,1437,151,1378]" box="[197,1127,1351,1378]" pageId="8" pageNumber="9">
<emphasis id="B969EACDA210FF84FF06FA9EA1C17D77" bold="true" box="[197,441,1351,1377]" pageId="8" pageNumber="9">Material examined.</emphasis>
<materialsCitation id="3B753C82A210FF84FE02FA91A41B7D74" box="[449,1123,1352,1378]" collectionCode="LSUHC" location="Kampung Bayu" pageId="8" pageNumber="9" specimenCode="LSUHC 9070-73" specimenCount="1" stateProvince="Kelantan">
<collectingRegion id="49D9F83DA210FF84FE02FA91A2577D74" box="[449,559,1352,1378]" country="Malaysia" name="Kelantan" pageId="8" pageNumber="9">Kelantan</collectingRegion>
:
<location id="8EC26004A210FF84FDF9FA91A28B7D74" LSID="urn:lsid:plazi:treatment:03B487C9A21EFF84FF54FCE6A41F7D74:8EC26004A210FF84FDF9FA91A28B7D74" box="[570,755,1352,1378]" name="Kampung Bayu" pageId="8" pageNumber="9" stateProvince="Kelantan">Kampung Bayu</location>
<specimenCode id="DBBB9EA4A210FF84FD39FA91A3B17D74" box="[762,969,1352,1378]" collectionCode="LSUHC" country="0" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="8" pageNumber="9">LSUHC 907073</specimenCode>
(
<typeStatus id="54A6887DA210FF84FC1BFA91A4747D74" box="[984,1036,1352,1378]" pageId="8" pageNumber="9">type</typeStatus>
series)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>