848 lines
116 KiB
XML
848 lines
116 KiB
XML
<document id="98D86AC72C153C2F6DA71E0E954B5AE4" ID-CLB-Dataset="7439" ID-DOI="10.11646/zootaxa.4934.1.1" ID-GBIF-Dataset="300bac18-55f5-4e34-90e1-7fccd02d68c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4558796" ID-ZooBank="79895443-4597-42A5-AF8A-023EACB20E10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614157470936" checkinUser="plazi" docAuthor="Archibald, Bruce, Cannings, Robert A., Erickson, Robert J., Bybee, Seth M. & Mathewes, Rolf W." docDate="2021" docId="03B487C2003AFFBDFF5BFD33FDE014A8" docLanguage="en" docName="zootaxa.4934.1.1.pdf" docOrigin="Zootaxa 4934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Okanagrion Archibald & Cannings & Erickson & Bybee & Mathewes 2021, new genus" docType="treatment" docVersion="7" lastPageNumber="43" masterDocId="FF8DFFBA0012FF97FFCCFFE5FFAE1215" masterDocTitle="The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America" masterLastPageNumber="133" masterPageNumber="1" pageNumber="41" updateTime="1698931040708" updateUser="ExternalLinkService">
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<mods:title id="43E595907B71FAE3714AB3ED9E8A8325">The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America</mods:title>
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<mods:namePart id="78D8AA2F434A3ED92B1B089B7A7BD341">Archibald, Bruce</mods:namePart>
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<mods:namePart id="D6503624E9014A44CCCBD006E609465F">Cannings, Robert A.</mods:namePart>
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<mods:nameIdentifier id="43CDFEC22D889C4212DD3D980D5D9F15" type="ORCID">0000-0001-9491-8186</mods:nameIdentifier>
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<mods:nameIdentifier id="0477F27F384CE10142A6906204F9FBB3" type="email">annings@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:namePart id="35BC31242EFF3BCADA9826C5A0375939">Erickson, Robert J.</mods:namePart>
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<mods:nameIdentifier id="E0B8CF72CDF556E1EA4426241F141DB1" type="ORCID">0000-0003-1162-0355</mods:nameIdentifier>
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<mods:nameIdentifier id="75140A14FF663AE6280645B1466023EC" type="email">1990platypus@gmail.com</mods:nameIdentifier>
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<mods:namePart id="22562A581984A890A579B78738D7EA82">Bybee, Seth M.</mods:namePart>
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<mods:nameIdentifier id="A88AEAB0C1362A375DFECF54C91BE1F0" type="email">seth.bybee@gmail.com</mods:nameIdentifier>
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<mods:namePart id="51DCD1E401D4961D76ED0717E6112A15">Mathewes, Rolf W.</mods:namePart>
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<mods:nameIdentifier id="FE8D636CA29A4BBC8834CEAA2D40C15E" type="email">r_mathewes@sfu.ca</mods:nameIdentifier>
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<mods:title id="96E922D5156DB743FA271D69AF8C0B65">Zootaxa</mods:title>
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<mods:date id="85A23FE9E5CCFA65A9B8FABF13502895">2021</mods:date>
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<mods:number id="11F34A4E36F7487B7D856087B09FA01D">2021-02-24</mods:number>
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<mods:number id="18AB6AA80C94DF3E78CC2E89CE1FC841">4934</mods:number>
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<mods:identifier id="9E0FDA9A3D739D67BCF3B7F3E62469C4" type="DOI">10.11646/zootaxa.4934.1.1</mods:identifier>
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<treatment id="03B487C2003AFFBDFF5BFD33FDE014A8" ID-DOI="http://doi.org/10.5281/zenodo.4672736" ID-GBIF-Taxon="180881805" ID-Zenodo-Dep="4672736" LSID="urn:lsid:plazi:treatment:03B487C2003AFFBDFF5BFD33FDE014A8" httpUri="http://treatment.plazi.org/id/03B487C2003AFFBDFF5BFD33FDE014A8" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
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<subSubSection id="C307655F003AFFBFFF5BFD33FD7310E4" box="[151,733,726,753]" pageId="40" pageNumber="41" type="nomenclature">
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<paragraph id="8BA236D4003AFFBFFF5BFD33FD7310E4" blockId="40.[151,733,726,789]" box="[151,733,726,753]" pageId="40" pageNumber="41">
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<heading id="D0EA81B8003AFFBFFF5BFD33FD7310E4" bold="true" box="[151,733,726,753]" fontSize="11" level="2" pageId="40" pageNumber="41" reason="4">
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<emphasis id="B969EAC6003AFFBFFF5BFD33FD7310E4" bold="true" box="[151,733,726,753]" pageId="40" pageNumber="41">
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<taxonomicName id="4C1D4D57003AFFBFFF5BFD33FDE010E5" authority="Archibald & Cannings" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[151,590,726,753]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B969EAC6003AFFBFFF5BFD33FE8410E4" bold="true" box="[151,298,726,753]" italics="true" pageId="40" pageNumber="41">Okanagrion</emphasis>
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Archibald & Cannings
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25A57BD003AFFBFFD97FD32FD7310E4" box="[603,733,727,753]" pageId="40" pageNumber="41" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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Okanagrion threadgillae holotype, SR 98-12-10. Photograph. A and B refer to drawings in Fig. 20. Scale bar is 5 mm." captionText-1="FIGURE 20. Okanagrion threadgillae holotype, SR 98-12-10. A, drawing of wing A of Fig. 19; B, drawing of wing B of Fig. 19. Both to scale, 5 mm." captionText-10="FIGURE 29. Okanagrion hobani paratype 5, F-1140. A, photograph; B, drawing. Both to scale, 5 mm." captionText-11="FIGURE 30. Okanagrion hobani. Paratype 6, SR 00-04-05:A, photograph; B, drawing; SR 06-62-06:A, photograph; B, drawing. Both to scale, 5 mm" captionText-12="FIGURE 31. Okanagrion hobani paratype 7, F-1044. A, photograph of complete specimen; B, close-up of anal appendages; C, drawing of abdominal showing segments 8 to 10 (10 apparently displaced and rotated somewhat clockwise in the view), paraprocts and one cercus; D, close-up showing proximal portions of wings; E, colour-coded wings referring to 22F through H; F, G, H: drawings of wings indicated in 22E: F, green wing, G, red wing; H, blue wing (purple wing very damaged, not informative). A to scale, B–H to scale; scale bars are 5 mm" captionText-13="FIGURE 32. Okanagrion hobani. Paratype 9, RBCM P1548: A, photograph; B, drawing. Paratype 8, RBCM P1547: C, photograph; D, drawing. SR 17-006-001: E, photograph; F, drawing. All to scale, 5 mm." captionText-14="FIGURE 33. Okanagrion hobani. SR 10-72-03: A, photograph; B, drawing. SR 95-03-30: C, photograph; D, drawing. All to scale, 5 mm." captionText-15="FIGURE 34. Okanagrion hobani. SRUI 99-71-07: A, photograph; B, drawing. RBCM P1549: C, photograph; D, drawing. All to scale, 5 mm." captionText-16="FIGURE 35. Okanagrion hobani relatively complete wings, comparative forewings (FW) and hind wings (HW). Red, blue arrows, see text. All to scale, 5 mm." captionText-17="FIGURE 36. Okanagrion angustum holotype, RBCM.EH2017.050.11.2491. A, photograph; B, drawing. Both to scale, 5 mm." captionText-18="FIGURE 37. Okanagrion dorrellae holotype, SR 93-11-02. A, photograph; B, drawing. Both to scale, 5 mm." captionText-19="FIGURE 38. Okanagrion liquetoalatum holotype, SR 06-69-17. A, photograph; B, drawing. Both to scale, 5 mm." captionText-2="FIGURE 21. Okanagrion beardi holotype, F-791. A, photograph; B, drawing. Small green arrow indicates IR2 originating on RP1-2. Both to scale, 5 mm." captionText-20="FIGURE 39. Okanagrion worleyae holotype, SR 99-14-02: A, photograph; B, drawing. Both to scale, 5 mm." captionText-21="FIGURE 40. Okanagrion worleyae paratype 1, SR 16-006-001. A, photograph; B, drawing. Both to scale, 5 mm." captionText-22="FIGURE 41. Okanagrion worleyae paratype 2, SR 11-21-09. A, photograph; B, drawing. Both to scale, 5 mm." captionText-23="FIGURE 42. Okanagrion incertae sedis. SR 11-60-17. A, photograph; B, drawing; SR 06-08-18: C, photograph; D, drawing; UWBM-74307: E, photograph; F, drawing. All to scale, 5 mm." captionText-24="FIGURE 43. Okanagrion incertae sedis. SR 94-05-30: A, photograph; B, drawing; SR 11-51-07: C, photograph; D, drawing; RBCM P1550: E, photograph of top wing in G; F, drawing of wing in E; G, all three wings. A–D to scale; E, F to scale; G to scale; all scales 5 mm." captionText-3="FIGURE 22. Okanagrion beardi paratype 1. RBCM P1546. A, drawing; B, photograph. Both to scale, 5 mm." captionText-4="FIGURE 23. Okanagrion lochmum holotype, RBCM 11799.001. A, photograph; B, drawing. Both to scale, 5 mm." captionText-5="FIGURE 24. Okanagrion hobani holotype, F-1052. A, photograph; B, drawing. Both to scale, 5 mm." captionText-6="FIGURE 25. Okanagrion hobani paratype 1, SR 11-21-33. A, photograph; B, drawing. Both to scale, 5 mm." captionText-7="FIGURE 26. Okanagrion hobani paratype 2, SR 02-24-14. A, photograph; B, drawing. Both to scale, 5 mm." captionText-8="FIGURE 27. Okanagrion hobani paratype 3, SR 94-05-22. A, photograph; B, drawing. Both to scale, 5 mm." captionText-9="FIGURE 28. Okanagrion hobani. Paratype 4, SRUI 99-96-55: A, photograph; B, drawing; SR 13-005-017: C, photograph; D, drawing.All to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4558846" figureDoi-1="http://doi.org/10.5281/zenodo.4558848" figureDoi-10="http://doi.org/10.5281/zenodo.4558874" figureDoi-11="http://doi.org/10.5281/zenodo.4558876" figureDoi-12="http://doi.org/10.5281/zenodo.4558878" figureDoi-13="http://doi.org/10.5281/zenodo.4558880" figureDoi-14="http://doi.org/10.5281/zenodo.4558882" figureDoi-15="http://doi.org/10.5281/zenodo.4558884" figureDoi-16="http://doi.org/10.5281/zenodo.4558886" figureDoi-17="http://doi.org/10.5281/zenodo.4558888" figureDoi-18="http://doi.org/10.5281/zenodo.4558890" figureDoi-19="http://doi.org/10.5281/zenodo.4558892" figureDoi-2="http://doi.org/10.5281/zenodo.4558852" figureDoi-20="http://doi.org/10.5281/zenodo.4558894" figureDoi-21="http://doi.org/10.5281/zenodo.4558896" figureDoi-22="http://doi.org/10.5281/zenodo.4558900" figureDoi-23="http://doi.org/10.5281/zenodo.4558902" figureDoi-24="http://doi.org/10.5281/zenodo.4558904" figureDoi-3="http://doi.org/10.5281/zenodo.4558854" figureDoi-4="http://doi.org/10.5281/zenodo.4558858" figureDoi-5="http://doi.org/10.5281/zenodo.4558862" figureDoi-6="http://doi.org/10.5281/zenodo.4558864" figureDoi-7="http://doi.org/10.5281/zenodo.4558868" figureDoi-8="http://doi.org/10.5281/zenodo.4558870" figureDoi-9="http://doi.org/10.5281/zenodo.4558872" httpUri-0="https://zenodo.org/record/4558846/files/figure.png" httpUri-1="https://zenodo.org/record/4558848/files/figure.png" httpUri-10="https://zenodo.org/record/4558874/files/figure.png" httpUri-11="https://zenodo.org/record/4558876/files/figure.png" httpUri-12="https://zenodo.org/record/4558878/files/figure.png" httpUri-13="https://zenodo.org/record/4558880/files/figure.png" httpUri-14="https://zenodo.org/record/4558882/files/figure.png" httpUri-15="https://zenodo.org/record/4558884/files/figure.png" httpUri-16="https://zenodo.org/record/4558886/files/figure.png" httpUri-17="https://zenodo.org/record/4558888/files/figure.png" httpUri-18="https://zenodo.org/record/4558890/files/figure.png" httpUri-19="https://zenodo.org/record/4558892/files/figure.png" httpUri-2="https://zenodo.org/record/4558852/files/figure.png" httpUri-20="https://zenodo.org/record/4558894/files/figure.png" httpUri-21="https://zenodo.org/record/4558896/files/figure.png" httpUri-22="https://zenodo.org/record/4558900/files/figure.png" httpUri-23="https://zenodo.org/record/4558902/files/figure.png" httpUri-24="https://zenodo.org/record/4558904/files/figure.png" httpUri-3="https://zenodo.org/record/4558854/files/figure.png" httpUri-4="https://zenodo.org/record/4558858/files/figure.png" httpUri-5="https://zenodo.org/record/4558862/files/figure.png" httpUri-6="https://zenodo.org/record/4558864/files/figure.png" httpUri-7="https://zenodo.org/record/4558868/files/figure.png" httpUri-8="https://zenodo.org/record/4558870/files/figure.png" httpUri-9="https://zenodo.org/record/4558872/files/figure.png" pageId="40" pageNumber="41">Figs. 19–43</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F003AFFBEFF5BFCA7FABA13DA" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="diagnosis">
|
||
<paragraph id="8BA236D4003AFFBEFF5BFCA7FABA13DA" blockId="40.[151,1437,834,2014]" lastBlockId="41.[151,1437,151,2036]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
|
||
<emphasis id="B969EAC6003AFFBFFF5BFCA7FEBC1149" bold="true" box="[151,274,834,860]" pageId="40" pageNumber="41">Diagnosis.</emphasis>
|
||
Wings distinct from all other genera of
|
||
<taxonomicName id="4C1D4D57003AFFBFFD1CFCA6FCDD1148" box="[720,883,835,861]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Dysagrioninae">Dysagrioninae</taxonomicName>
|
||
by: pterostigma 3–3.5, rarely 4 times longer than wide [separated from
|
||
<taxonomicName id="4C1D4D57003AFFBFFE5CFC82FD8F1194" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[400,545,871,897]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFE5CFC82FD8F1194" box="[400,545,871,897]" italics="true" pageId="40" pageNumber="41">Okanopteryx</emphasis>
|
||
</taxonomicName>
|
||
: 2.5 times;
|
||
<taxonomicName id="4C1D4D57003AFFBFFD64FC82FCE91194" authorityName="Archibald & Cannings" authorityYear="2021" box="[680,839,871,897]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFD64FC82FCE91194" box="[680,839,871,897]" italics="true" pageId="40" pageNumber="41">Stenodiafanus</emphasis>
|
||
</taxonomicName>
|
||
: 4;
|
||
<taxonomicName id="4C1D4D57003AFFBFFCB9FC82FBBB1194" authorityName="Cockerell" authorityYear="1908" box="[885,1045,871,897]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFCB9FC82FBBB1194" box="[885,1045,871,897]" italics="true" pageId="40" pageNumber="41">Phenacolestes</emphasis>
|
||
</taxonomicName>
|
||
: 5; not separated from
|
||
<taxonomicName id="4C1D4D57003AFFBFFAECFC82FA3B1195" box="[1312,1429,871,896]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFAECFC82FA3B1195" box="[1312,1429,871,896]" italics="true" pageId="40" pageNumber="41">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
: 3;
|
||
<taxonomicName id="4C1D4D57003AFFBFFF7FFC6EFEE411B0" box="[179,330,907,933]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFF7FFC6EFEE411B0" box="[179,330,907,933]" italics="true" pageId="40" pageNumber="41">Primorilestes</emphasis>
|
||
</taxonomicName>
|
||
: 3.5–4;
|
||
<taxonomicName id="4C1D4D57003AFFBFFE69FC6EFDB511B1" box="[421,539,907,932]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFE69FC6EFDB511B1" box="[421,539,907,932]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
</taxonomicName>
|
||
: 4–7.5;
|
||
<taxonomicName id="4C1D4D57003AFFBFFDBAFC6EFCBD11B1" box="[630,787,907,932]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFDBAFC6EFCBD11B1" box="[630,787,907,932]" italics="true" pageId="40" pageNumber="41">Dysagrionites</emphasis>
|
||
</taxonomicName>
|
||
species: unknown]; no oblique brace vein [separated from
|
||
<taxonomicName id="4C1D4D57003AFFBFFF5BFC4AFEA311DD" box="[151,269,943,968]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFF5BFC4AFEA311DD" box="[151,269,943,968]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFED7FC4AFD9411DC" authority=", Okanoptyeryx" authorityName="Okanoptyeryx" box="[283,570,943,969]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFED7FC4AFE3E11DD" box="[283,400,943,968]" italics="true" pageId="40" pageNumber="41">Furagrion</emphasis>
|
||
,
|
||
<emphasis id="B969EAC6003AFFBFFE51FC4AFD9411DC" box="[413,570,943,969]" italics="true" pageId="40" pageNumber="41">Okanoptyeryx</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFD84FC4AFD4911DC" authorityName="Archibald & Cannings" authorityYear="2021" box="[584,743,943,969]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFD84FC4AFD4911DC" box="[584,743,943,969]" italics="true" pageId="40" pageNumber="41">Stenodiafanus</emphasis>
|
||
</taxonomicName>
|
||
; wing region not preserved in
|
||
<taxonomicName id="4C1D4D57003AFFBFFBF1FC4AFA8511DC" authorityName="Nel & Arillo" authorityYear="2006" box="[1085,1323,943,969]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFBF1FC4AFA8511DC" box="[1085,1323,943,969]" italics="true" pageId="40" pageNumber="41">Electrophenacolestes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFAF5FC4AFF7411F9" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFAF5FC4AFF7411F9" italics="true" pageId="40" pageNumber="41">Dysagrionites</emphasis>
|
||
</taxonomicName>
|
||
species; not separated from
|
||
<taxonomicName id="4C1D4D57003AFFBFFDD1FC36FD1311F8" authorityName="Cockerell" authorityYear="1908" box="[541,701,979,1005]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFDD1FC36FD1311F8" box="[541,701,979,1005]" italics="true" pageId="40" pageNumber="41">Phenacolestes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFD00FC36FCCD11F8" box="[716,867,979,1005]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFD00FC36FCCD11F8" box="[716,867,979,1005]" italics="true" pageId="40" pageNumber="41">Primorilestes</emphasis>
|
||
</taxonomicName>
|
||
]; costal space 3–5 cells wide distal to pterostigma [distinct from
|
||
<taxonomicName id="4C1D4D57003AFFBFFEF4FC12FE611604" box="[312,463,1015,1041]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFEF4FC12FE611604" box="[312,463,1015,1041]" italics="true" pageId="40" pageNumber="41">Primorilestes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFE11FC12FDC01604" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[477,622,1015,1041]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFE11FC12FDC01604" box="[477,622,1015,1041]" italics="true" pageId="40" pageNumber="41">Okanopteryx</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFDB0FC12FCB51604" authorityName="Archibald & Cannings" authorityYear="2021" box="[636,795,1015,1041]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFDB0FC12FCB51604" box="[636,795,1015,1041]" italics="true" pageId="40" pageNumber="41">Stenodiafanus</emphasis>
|
||
</taxonomicName>
|
||
: all 2;
|
||
<taxonomicName id="4C1D4D57003AFFBFFCA6FC12FC4E1605" box="[874,992,1015,1040]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFCA6FC12FC4E1605" box="[874,992,1015,1040]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFC23FC12FBCA1605" box="[1007,1124,1015,1040]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFC23FC12FBCA1605" box="[1007,1124,1015,1040]" italics="true" pageId="40" pageNumber="41">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFBBEFC12FABC1604" authorityName="Cockerell" authorityYear="1908" box="[1138,1298,1015,1041]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFBBEFC12FABC1604" box="[1138,1298,1015,1041]" italics="true" pageId="40" pageNumber="41">Phenacolestes</emphasis>
|
||
</taxonomicName>
|
||
: all 1;
|
||
<taxonomicName id="4C1D4D57003AFFBFFAADFC12FEFA1620" authorityName="Nel & Arillo" authorityYear="2006" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFAADFC12FEFA1620" italics="true" pageId="40" pageNumber="41">Electrophenacolestes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFEAEFBFEFE511621" box="[354,511,1051,1076]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFEAEFBFEFE511621" box="[354,511,1051,1076]" italics="true" pageId="40" pageNumber="41">Dysagrionites</emphasis>
|
||
</taxonomicName>
|
||
species: not known]; subnodus with normal obliquity (reverse in one specimen of
|
||
<taxonomicName id="4C1D4D57003AFFBFFF5BFBDAFEDD164C" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[151,371,1087,1113]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="hobani">
|
||
<emphasis id="B969EAC6003AFFBFFF5BFBDAFEDD164C" box="[151,371,1087,1113]" italics="true" pageId="40" pageNumber="41">Okanagrion hobani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54A68876003AFFBFFEB3FBDAFE71164C" box="[383,479,1087,1113]" pageId="40" pageNumber="41" type="paratype">paratype</typeStatus>
|
||
1, which we consider adventitious) [separated from
|
||
<taxonomicName id="4C1D4D57003AFFBFFBE9FBDAFB35164D" box="[1061,1179,1087,1112]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFBE9FBDAFB35164D" box="[1061,1179,1087,1112]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
</taxonomicName>
|
||
, reversed obliquity; all others: normal obliquity, slight to notable]; no accessory antenodal crossveins [distinguished from
|
||
<taxonomicName id="4C1D4D57003AFFBFFB39FB86FA3B1668" authorityName="Cockerell" authorityYear="1908" box="[1269,1429,1123,1149]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFB39FB86FA3B1668" box="[1269,1429,1123,1149]" italics="true" pageId="40" pageNumber="41">Phenacolestes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFF5BFB62FE2B16B4" authorityName="Nel & Arillo" authorityYear="2006" box="[151,389,1159,1185]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFF5BFB62FE2B16B4" box="[151,389,1159,1185]" italics="true" pageId="40" pageNumber="41">Electrophenacolestes</emphasis>
|
||
</taxonomicName>
|
||
, some
|
||
<taxonomicName id="4C1D4D57003AFFBFFE19FB62FDE516B5" box="[469,587,1159,1184]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFE19FB62FDE516B5" box="[469,587,1159,1184]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
</taxonomicName>
|
||
species: present; not separated from all others: also none]; IR2 origin rather evenly between RP1-2, RP3-4, connected to RP1-2, RP3-4 at similar level (some species IR2 origin close to or on RP1-2) [
|
||
<taxonomicName id="4C1D4D57003AFFBFFF34FB2AFEC016FD" box="[248,366,1231,1256]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFF34FB2AFEC016FD" box="[248,366,1231,1256]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
</taxonomicName>
|
||
rather evenly between RP1-2, RP3-4, but connected with RP1-2 distinctly more distally than to RP3-4; other genera IR2 origin rather evenly between RP1-2, RP3-4, connected to RP1-2, RP3-4 at similar level]; MA linear from quadrangle to margin [separated from
|
||
<taxonomicName id="4C1D4D57003AFFBFFD3AFAF2FC081724" authority=": MA" authorityName="MA" box="[758,934,1303,1329]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFD3AFAF2FCC21725" box="[758,876,1303,1328]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
: MA
|
||
</taxonomicName>
|
||
linear from quadrangle to zigzagged roughly mid-wing or further, linear again near margin;
|
||
<taxonomicName id="4C1D4D57003AFFBFFD6DFADEFCB81741" box="[673,790,1339,1364]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFD6DFADEFCB81741" box="[673,790,1339,1364]" italics="true" pageId="40" pageNumber="41">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
: similar to
|
||
<taxonomicName id="4C1D4D57003AFFBFFC5AFADEFBA21741" box="[918,1036,1339,1364]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFC5AFADEFBA21741" box="[918,1036,1339,1364]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
</taxonomicName>
|
||
but deeply curved in distal portion;
|
||
<taxonomicName id="4C1D4D57003AFFBFFF5BFABAFE99176C" authorityName="Cockerell" authorityYear="1908" box="[151,311,1375,1401]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFF5BFABAFE99176C" box="[151,311,1375,1401]" italics="true" pageId="40" pageNumber="41">Phenacolestes</emphasis>
|
||
</taxonomicName>
|
||
: very slightly zigzagged or not in mid-portion;
|
||
<taxonomicName id="4C1D4D57003AFFBFFC94FABAFC47176C" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[856,1001,1375,1401]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFC94FABAFC47176C" box="[856,1001,1375,1401]" italics="true" pageId="40" pageNumber="41">Okanopteryx</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFC34FABAFB39176C" authorityName="Archibald & Cannings" authorityYear="2021" box="[1016,1175,1375,1401]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFC34FABAFB39176C" box="[1016,1175,1375,1401]" italics="true" pageId="40" pageNumber="41">Stenodiafanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFB6AFABAFAED176D" box="[1190,1347,1375,1400]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFB6AFABAFAED176D" box="[1190,1347,1375,1400]" italics="true" pageId="40" pageNumber="41">Dysagrionites</emphasis>
|
||
</taxonomicName>
|
||
species (known portions, none known to margin): linear to mid-wing, somewhat to strongly zigzagged to margin; not separated from
|
||
<taxonomicName id="4C1D4D57003AFFBFFEDAFA42FE0317D4" box="[278,429,1447,1473]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFEDAFA42FE0317D4" box="[278,429,1447,1473]" italics="true" pageId="40" pageNumber="41">Primorilestes</emphasis>
|
||
</taxonomicName>
|
||
: linear]; MA, MP far apart at margin [distinct from
|
||
<taxonomicName id="4C1D4D57003AFFBFFC31FA42FB4817D4" authority=": MA, MP" authorityName="MA, MP" box="[1021,1254,1447,1473]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFC31FA42FBDD17D5" box="[1021,1139,1447,1472]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
: MA, MP
|
||
</taxonomicName>
|
||
close, 1–2 cells apart at margin; not clearly distinct from
|
||
<taxonomicName id="4C1D4D57003AFFBFFDAAFA2EFD5917F0" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[614,759,1483,1509]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFDAAFA2EFD5917F0" box="[614,759,1483,1509]" italics="true" pageId="40" pageNumber="41">Okanopteryx</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFCC9FA2EFC0A17F0" authorityName="Archibald & Cannings" authorityYear="2021" box="[773,932,1483,1509]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFCC9FA2EFC0A17F0" box="[773,932,1483,1509]" italics="true" pageId="40" pageNumber="41">Stenodiafanus</emphasis>
|
||
</taxonomicName>
|
||
as difficult to be sure of identity of MA near margin by strong zigzag, but close, probably separated by two cells;
|
||
<taxonomicName id="4C1D4D57003AFFBFFC64FA0AFBEB141D" box="[936,1093,1519,1544]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFC64FA0AFBEB141D" box="[936,1093,1519,1544]" italics="true" pageId="40" pageNumber="41">Dysagrionites</emphasis>
|
||
</taxonomicName>
|
||
species also difficult, perhaps one cell; not separated from
|
||
<taxonomicName id="4C1D4D57003AFFBFFE14F9F6FDE31439" box="[472,589,1555,1580]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFE14F9F6FDE31439" box="[472,589,1555,1580]" italics="true" pageId="40" pageNumber="41">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFD97F9F6FD5C1438" box="[603,754,1555,1581]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFD97F9F6FD5C1438" box="[603,754,1555,1581]" italics="true" pageId="40" pageNumber="41">Primorilestes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFCCCF9F6FBCE1438" authority=": MA, MP" authorityName="MA, MP" box="[768,1120,1555,1581]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFCCCF9F6FC401438" box="[768,1006,1555,1581]" italics="true" pageId="40" pageNumber="41">Electrophenacolestes</emphasis>
|
||
: MA, MP
|
||
</taxonomicName>
|
||
widely separated]; CuA not zigzagged (or only very slightly so), linear to margin [distinct from
|
||
<taxonomicName id="4C1D4D57003AFFBFFC59F9D2FBA51445" box="[917,1035,1591,1616]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFC59F9D2FBA51445" box="[917,1035,1591,1616]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
</taxonomicName>
|
||
: CuA increasingly zigzagged distal to level of nodus;
|
||
<taxonomicName id="4C1D4D57003AFFBFFE91F9BEFE7C1461" box="[349,466,1627,1652]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFE91F9BEFE7C1461" box="[349,466,1627,1652]" italics="true" pageId="40" pageNumber="41">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
: slightly zigzagged distal to level of origin of RP2;
|
||
<taxonomicName id="4C1D4D57003AFFBFFBC0F9BEFB021460" authorityName="Cockerell" authorityYear="1908" box="[1036,1196,1627,1653]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFBC0F9BEFB021460" box="[1036,1196,1627,1653]" italics="true" pageId="40" pageNumber="41">Phenacolestes</emphasis>
|
||
</taxonomicName>
|
||
: may be very slightly zigzagged in parts;
|
||
<emphasis id="B969EAC6003AFFBFFEA2F99AFDA5148C" box="[366,523,1663,1689]" italics="true" pageId="40" pageNumber="41">Okanoptyeryx</emphasis>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFDD4F99AFD19148C" authorityName="Archibald & Cannings" authorityYear="2021" box="[536,695,1663,1689]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFDD4F99AFD19148C" box="[536,695,1663,1689]" italics="true" pageId="40" pageNumber="41">Stenodiafanus</emphasis>
|
||
</taxonomicName>
|
||
: zigzagged to various degrees distal to about mid-wing; not clearly distinct from
|
||
<taxonomicName id="4C1D4D57003AFFBFFEE3F946FDB314A8" box="[303,541,1699,1725]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFEE3F946FDB314A8" box="[303,541,1699,1725]" italics="true" pageId="40" pageNumber="41">Electrophenacolestes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFDE0F946FD6D14A8" box="[556,707,1699,1725]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFDE0F946FD6D14A8" box="[556,707,1699,1725]" italics="true" pageId="40" pageNumber="41">Primorilestes</emphasis>
|
||
</taxonomicName>
|
||
: linear or very slightly zigzagged]; CuA–A space 4–7 cells wide [separated from
|
||
<taxonomicName id="4C1D4D57003AFFBFFE83F922FE6A14F5" box="[335,452,1735,1760]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFE83F922FE6A14F5" box="[335,452,1735,1760]" italics="true" pageId="40" pageNumber="41">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
: 2;
|
||
<taxonomicName id="4C1D4D57003AFFBFFE3DF922FD3E14F4" authorityName="Archibald & Cannings" authorityYear="2021" box="[497,656,1735,1761]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFE3DF922FD3E14F4" box="[497,656,1735,1761]" italics="true" pageId="40" pageNumber="41">Stenodiafanus</emphasis>
|
||
</taxonomicName>
|
||
: 2 (most likely, wing incomplete);
|
||
<taxonomicName id="4C1D4D57003AFFBFFBD0F922FAA214F4" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[1052,1292,1735,1761]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="delinei">
|
||
<emphasis id="B969EAC6003AFFBFFBD0F922FAA214F4" box="[1052,1292,1735,1761]" italics="true" pageId="40" pageNumber="41">Dysagrionites delinei</emphasis>
|
||
</taxonomicName>
|
||
: 3;
|
||
<taxonomicName id="4C1D4D57003AFFBFFAF5F922FF741511" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFAF5F922FF741511" italics="true" pageId="40" pageNumber="41">Dysagrionites</emphasis>
|
||
</taxonomicName>
|
||
sp. A: 2;
|
||
<taxonomicName id="4C1D4D57003AFFBFFE81F90EFE441511" box="[333,490,1771,1796]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFE81F90EFE441511" box="[333,490,1771,1796]" italics="true" pageId="40" pageNumber="41">Dysagrionites</emphasis>
|
||
</taxonomicName>
|
||
sp. B: probably 3 but unclear; not separated from
|
||
<taxonomicName id="4C1D4D57003AFFBFFBFCF90EFB081511" box="[1072,1190,1771,1796]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFBFCF90EFB081511" box="[1072,1190,1771,1796]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
</taxonomicName>
|
||
: 4;
|
||
<taxonomicName id="4C1D4D57003AFFBFFB1AF90EFAD81510" authorityName="Cockerell" authorityYear="1908" box="[1238,1398,1771,1797]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFB1AF90EFAD81510" box="[1238,1398,1771,1797]" italics="true" pageId="40" pageNumber="41">Phenacolestes</emphasis>
|
||
</taxonomicName>
|
||
: 4;
|
||
<taxonomicName id="4C1D4D57003AFFBFFF5BF8EAFE80153C" box="[151,302,1807,1833]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFF5BF8EAFE80153C" box="[151,302,1807,1833]" italics="true" pageId="40" pageNumber="41">Primorilestes</emphasis>
|
||
</taxonomicName>
|
||
: 3–5;
|
||
<taxonomicName id="4C1D4D57003AFFBFFEB0F8EAFDC4153C" box="[380,618,1807,1833]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFEB0F8EAFDC4153C" box="[380,618,1807,1833]" italics="true" pageId="40" pageNumber="41">Electrophenacolestes</emphasis>
|
||
</taxonomicName>
|
||
: 4 (one column 5);
|
||
<taxonomicName id="4C1D4D57003AFFBFFC9AF8EAFC49153C" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[854,999,1807,1833]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFC9AF8EAFC49153C" box="[854,999,1807,1833]" italics="true" pageId="40" pageNumber="41">Okanopteryx</emphasis>
|
||
</taxonomicName>
|
||
: 2–4]; postnodal, postsubnodal crossveins not aligned at least in proximal half [separated from
|
||
<taxonomicName id="4C1D4D57003AFFBFFC84F8D6FC491558" authorityName="Archibald & Cannings" authorityYear="2021" box="[840,999,1843,1869]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFC84F8D6FC491558" box="[840,999,1843,1869]" italics="true" pageId="40" pageNumber="41">Stenodiafanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFC34F8D6FB271558" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[1016,1161,1843,1869]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFC34F8D6FB271558" box="[1016,1161,1843,1869]" italics="true" pageId="40" pageNumber="41">Okanopteryx</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFB56F8D6FA3B1558" box="[1178,1429,1843,1869]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="violetae">
|
||
<emphasis id="B969EAC6003AFFBFFB56F8D6FA3B1558" box="[1178,1429,1843,1869]" italics="true" pageId="40" pageNumber="41">Primorilestes violetae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1D4D57003AFFBFFF5BF8B2FE2B1564" box="[151,389,1879,1905]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFF5BF8B2FE2B1564" box="[151,389,1879,1905]" italics="true" pageId="40" pageNumber="41">Electrophenacolestes</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1D4D57003AFFBFFE5BF8B2FDA21565" box="[407,524,1879,1904]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFE5BF8B2FDA21565" box="[407,524,1879,1904]" italics="true" pageId="40" pageNumber="41">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
: all aligned or somewhat so in proximal half; not distinct from
|
||
<taxonomicName id="4C1D4D57003AFFBFFB39F8B2FA3B1564" authorityName="Cockerell" authorityYear="1908" box="[1269,1429,1879,1905]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFB39F8B2FA3B1564" box="[1269,1429,1879,1905]" italics="true" pageId="40" pageNumber="41">Phenacolestes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFF5BF89EFE3D1580" box="[151,403,1915,1941]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="madseni">
|
||
<emphasis id="B969EAC6003AFFBFFF5BF89EFE3D1580" box="[151,403,1915,1941]" italics="true" pageId="40" pageNumber="41">Primorilestes madseni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003AFFBFFE6EF89EFDB61581" box="[418,536,1915,1940]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFE6EF89EFDB61581" box="[418,536,1915,1940]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
</taxonomicName>
|
||
: all not]; further from
|
||
<taxonomicName id="4C1D4D57003AFFBFFCD6F89EFC3E1581" box="[794,912,1915,1940]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFCD6F89EFC3E1581" box="[794,912,1915,1940]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
</taxonomicName>
|
||
by nodus at 25–30% wing length [
|
||
<taxonomicName id="4C1D4D57003AFFBFFAD2F89EFA3A1581" box="[1310,1428,1915,1940]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFAD2F89EFA3A1581" box="[1310,1428,1915,1940]" italics="true" pageId="40" pageNumber="41">Dysagrion</emphasis>
|
||
</taxonomicName>
|
||
: Ξ 35%]; further from
|
||
<taxonomicName id="4C1D4D57003AFFBFFE55F87AFD9E15AC" box="[409,560,1951,1977]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFE55F87AFD9E15AC" box="[409,560,1951,1977]" italics="true" pageId="40" pageNumber="41">Primorilestes</emphasis>
|
||
</taxonomicName>
|
||
by no intercalary vein in CuA–A space. Separated from
|
||
<taxonomicName id="4C1D4D57003AFFBFFB1FF87AFA8615AC" box="[1235,1320,1951,1977]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFB1FF87AFA8615AC" box="[1235,1320,1951,1977]" italics="true" pageId="40" pageNumber="41">Valerea</emphasis>
|
||
</taxonomicName>
|
||
(tentative
|
||
<taxonomicName id="4C1D4D57003AFFBFFF5BF826FE9215CB" box="[151,316,1987,2014]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Dysagrioninae">Dysagrioninae</taxonomicName>
|
||
) by no linear supplementary sector between RP1, IR1 [
|
||
<taxonomicName id="4C1D4D57003AFFBFFC77F826FBA115C8" box="[955,1039,1987,2013]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003AFFBFFC77F826FBA115C8" box="[955,1039,1987,2013]" italics="true" pageId="40" pageNumber="41">Valerea</emphasis>
|
||
</taxonomicName>
|
||
: with such sector]; origins of IR1, RP2 further from nodus; origin of IR2 between RP1-2, RP3-4 or near or on RP1-2 [IR2 origin not preserved, only preserved to level of nodus, where it is closer to RP3-4]. Most easily separated from genera regarded here as possible
|
||
<taxonomicName id="4C1D4D57003BFFBEFF1EFF3BFEDB12ED" box="[210,373,222,248]" class="Insecta" family="Dysagrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="family">Dysagrionidae</taxonomicName>
|
||
as follows: from
|
||
<taxonomicName id="4C1D4D57003BFFBEFDFAFF3BFD5412ED" box="[566,762,222,248]" class="Insecta" genus="Thanetophilosina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003BFFBEFDFAFF3BFD5412ED" box="[566,762,222,248]" italics="true" pageId="41" pageNumber="42">Thanetophilosina</emphasis>
|
||
</taxonomicName>
|
||
by pterostigma 3–3.5, rarely 4 times longer than wide [about 6 times width], by no linear supplementary sectors between RP1, IR1, between IR1, RP2 [
|
||
<taxonomicName id="4C1D4D57003BFFBEFB5BFEE7FAF51309" box="[1175,1371,258,284]" class="Insecta" genus="Thanetophilosina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003BFFBEFB5BFEE7FAF51309" box="[1175,1371,258,284]" italics="true" pageId="41" pageNumber="42">Thanetophilosina</emphasis>
|
||
</taxonomicName>
|
||
: with such sectors]; from Chickaloon specimen by MA linear [beginning to zigzag in distal-most portion preserved]; from NHMUK I.9866/I.9718 by origin of RP3-4 [distinctly closer to subnodus]; from
|
||
<emphasis id="B969EAC6003BFFBEFBFEFEACFBE81376" box="[1074,1094,329,355]" italics="true" pageId="41" pageNumber="42">cf</emphasis>
|
||
.
|
||
<taxonomicName id="4C1D4D57003BFFBEFB99FEACFB561376" box="[1109,1272,329,355]" class="Insecta" family="Dysagrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="family">Dysagrionidae</taxonomicName>
|
||
genus A, species A (“
|
||
<taxonomicName id="4C1D4D57003BFFBEFF3FFE88FE751392" box="[243,475,365,391]" class="Insecta" family="Megapodagrionidae" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="family">Megapodagrionidae</taxonomicName>
|
||
” genus A, species A of
|
||
<bibRefCitation id="EF8C4B25003BFFBEFD2FFE88FC581392" author="Petrulevicius, J. F. & Wappler, T. & Wedmann, S. & Rust, J. & Nel, A." box="[739,1014,365,392]" pageId="41" pageNumber="42" pagination="1173 - 1181" refId="ref61019" refString="Petrulevicius, J. F., Wappler, T., Wedmann, S., Rust, J. & Nel, A. (2008) New Megapodagrionid Damselflies (Odonata: Zygop- tera) from the Paleogene of Europe. Journal of Paleontology, 82, 1173 - 1181. https: // doi. org / 10.1666 / 07 - 091.1" type="journal article" year="2008">
|
||
Petrulevičius
|
||
<emphasis id="B969EAC6003BFFBEFCB0FE8BFC001392" box="[892,942,365,391]" italics="true" pageId="41" pageNumber="42">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
) by width of CuA–A space [two cells wide, preserved portion, likely maximum], by crossvenation in apical third [not as dense,
|
||
<emphasis id="B969EAC6003BFFBEFBB7FE77FB3313BE" box="[1147,1181,402,427]" italics="true" pageId="41" pageNumber="42">e.g</emphasis>
|
||
., costal space apical to pterostigma two cells wide, RA–RP1, RP1–IR1 spaces one cell wide to apex]; no brace vein [present].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F003BFFBEFF0BFE3DFE4B102F" pageId="41" pageNumber="42" type="type_taxon">
|
||
<paragraph id="8BA236D4003BFFBEFF0BFE3DFE4B102F" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
|
||
<emphasis id="B969EAC6003BFFBEFF0BFE3DFE5413E7" bold="true" box="[199,506,472,498]" pageId="41" pageNumber="42">
|
||
<typeStatus id="54A68876003BFFBEFF0BFE3DFEAC13E7" box="[199,258,472,498]" pageId="41" pageNumber="42">Type</typeStatus>
|
||
and included species
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="54A68876003BFFBEFDC4FE3CFDEE13E6" box="[520,576,473,499]" pageId="41" pageNumber="42">Type</typeStatus>
|
||
species:
|
||
<taxonomicName id="4C1D4D57003BFFBEFD68FE3DFC1413E7" authority="Archibald & Cannings & Erickson & Bybee & Mathewes, 2021" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[676,954,472,498]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="threadgillae" status="sp. nov.">
|
||
<emphasis id="B969EAC6003BFFBEFD68FE3DFC1413E7" box="[676,954,472,498]" italics="true" pageId="41" pageNumber="42">Okanagrion threadgillae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57BD003BFFBEFC0CFE3CFBE713E7" box="[960,1097,472,499]" pageId="41" pageNumber="42" rank="species">new species</taxonomicNameLabel>
|
||
; included:
|
||
<taxonomicName id="4C1D4D57003BFFBEFB0EFE3CFA9C13E7" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[1218,1330,472,499]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hobani">
|
||
<emphasis id="B969EAC6003BFFBEFB0EFE3CFA9C13E7" box="[1218,1330,472,499]" italics="true" pageId="41" pageNumber="42">O. hobani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003BFFBEFAF3FE3CFF031003" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="beardi">
|
||
<emphasis id="B969EAC6003BFFBEFAF3FE3CFF031003" italics="true" pageId="41" pageNumber="42">O. beardi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D57003BFFBEFF71FE19FEE61003" authority="Archibald & Cannings & Erickson & Bybee & Mathewes, 2021" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[189,328,508,535]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="lochmum" status="sp. nov.">
|
||
<emphasis id="B969EAC6003BFFBEFF71FE19FF7F1003" box="[189,209,508,534]" italics="true" pageId="41" pageNumber="42">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC6003BFFBEFF2DFE19FEE61003" box="[225,328,508,534]" italics="true" pageId="41" pageNumber="42">lochmum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57BD003BFFBEFE9CFE18FE751003" box="[336,475,508,535]" pageId="41" pageNumber="42" rank="species">new species</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C1D4D57003BFFBEFE24FE19FDD71003" authority="Archibald & Cannings & Erickson & Bybee & Mathewes, 2021" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[488,633,508,535]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="angustum" status="sp. nov.">
|
||
<emphasis id="B969EAC6003BFFBEFE24FE19FE521003" box="[488,508,508,534]" italics="true" pageId="41" pageNumber="42">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC6003BFFBEFDC0FE18FDD71003" box="[524,633,509,534]" italics="true" pageId="41" pageNumber="42">angustum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57BD003BFFBEFD4DFE18FCA31003" box="[641,781,508,535]" pageId="41" pageNumber="42" rank="species">new species</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C1D4D57003BFFBEFCD5FE19FC0A1003" authority="Archibald & Cannings & Erickson & Bybee & Mathewes, 2021" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[793,932,508,534]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dorrellae" status="sp. nov.">
|
||
<emphasis id="B969EAC6003BFFBEFCD5FE19FC0A1003" box="[793,932,508,534]" italics="true" pageId="41" pageNumber="42">O. dorrellae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57BD003BFFBEFC61FE18FB961003" box="[941,1080,508,535]" pageId="41" pageNumber="42" rank="species">new species</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C1D4D57003BFFBEFB89FE19FAAB1003" authority="Archibald & Cannings & Erickson & Bybee & Mathewes, 2021" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[1093,1285,508,534]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="liquetoalatum" status="sp. nov.">
|
||
<emphasis id="B969EAC6003BFFBEFB89FE19FAAB1003" box="[1093,1285,508,534]" italics="true" pageId="41" pageNumber="42">O. liquetoalatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57BD003BFFBEFAC1FE18FA361003" box="[1293,1432,508,535]" pageId="41" pageNumber="42" rank="species">new species</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C1D4D57003BFFBEFF0BFDC5FEFE102F" authority="Archibald & Cannings & Erickson & Bybee & Mathewes, 2021" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[199,336,544,570]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="worleyae" status="sp. nov.">
|
||
<emphasis id="B969EAC6003BFFBEFF0BFDC5FEFE102F" box="[199,336,544,570]" italics="true" pageId="41" pageNumber="42">O. worleyae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57BD003BFFBEFE9AFDC5FE4E102F" box="[342,480,544,570]" pageId="41" pageNumber="42" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F003BFFBEFF0BFDA6FEE611D6" pageId="41" pageNumber="42" type="description">
|
||
<paragraph id="8BA236D4003BFFBEFF0BFDA6FEE611D6" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
|
||
<emphasis id="B969EAC6003BFFBEFF0BFDA6FEFF1048" bold="true" box="[199,337,579,605]" pageId="41" pageNumber="42">Description</emphasis>
|
||
. As in diagnosis and the following. Colouration of wings varies: broad regions of dark or light fasciae or may be completely hyaline. Pterostigma 3–3.5 (rarely four) times longer than wide; four–seven cells subtend pterostigma (usually six); no accessory crossvein proximal to or distal to CuP; no well-defined supplementary sector between IR1, RP2; CuP proximal to arculus, between levels of Ax1, Ax2; petiole ends proximal to CuA, proximal to CuP; no oblique brace vein; postnodal, postsubnodal crossveins not aligned; CuA–A space maximum four–seven cells wide; origin of RP3-4 proximal to subnodus; no crossvein O; origin of IR2 at (or immediately proximal to) subnodus; IR1 zig-zagged in proximal half, CuA may be slightly zigzagged briefly about nodal level, apart from this all major veins linear (except MP zigzagged distally in
|
||
<taxonomicName id="4C1D4D57003BFFBEFCC6FCDBFC3B1142" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[778,917,829,856]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dorrellae">
|
||
<emphasis id="B969EAC6003BFFBEFCC6FCDBFC3B1142" box="[778,917,829,856]" italics="true" pageId="41" pageNumber="42">O. dorrellae</emphasis>
|
||
</taxonomicName>
|
||
), well defined; usually one–two hyperstigmal crossveins, less often none; two antenodal crossveins (Ax1, 2), rarely Ax0, no accessory antenodal crossveins; Ax2 at or very close to, immediately distal to arculus; CuA long, terminating on posterior margin beyond half length of arculus to apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F003BFFBEFF0BFC28FE611647" pageId="41" pageNumber="42" type="etymology">
|
||
<paragraph id="8BA236D4003BFFBEFF0BFC28FC6C161E" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
|
||
<emphasis id="B969EAC6003BFFBEFF0BFC28FEE611F2" bold="true" box="[199,328,973,999]" pageId="41" pageNumber="42">Etymology</emphasis>
|
||
. The genus name is a toponym derived from “Okanagan Highlands” and -
|
||
<emphasis id="B969EAC6003BFFBEFB4DFC28FB6211F3" box="[1153,1228,973,998]" italics="true" pageId="41" pageNumber="42">agrion</emphasis>
|
||
, the Greek
|
||
<emphasis id="B969EAC6003BFFBEFA81FC2BFA3B11FC" box="[1357,1429,974,1001]" italics="true" pageId="41" pageNumber="42">ἄγρΙΟς</emphasis>
|
||
, often used as a suffix in forming damselfly genus names. Gender neuter.
|
||
</paragraph>
|
||
<paragraph id="8BA236D4003BFFBEFF0BFBF1FE611647" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
|
||
<emphasis id="B969EAC6003BFFBEFF0BFBF1FED9163B" bold="true" box="[199,375,1044,1070]" pageId="41" pageNumber="42">Range and age</emphasis>
|
||
. McAbee, BC,
|
||
<collectingCountry id="F30A7644003BFFBEFDE4FBF1FDD2163B" box="[552,636,1044,1070]" name="Canada" pageId="41" pageNumber="42">Canada</collectingCountry>
|
||
and the Klondike Mountain Formation at Republic,
|
||
<collectingRegion id="49D9F836003BFFBEFB05FBF1FAFF163B" box="[1225,1361,1044,1070]" country="United States of America" name="Washington" pageId="41" pageNumber="42">Washington</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30A7644003BFFBEFA90FBF1FA36163B" box="[1372,1432,1044,1070]" name="United States of America" pageId="41" pageNumber="42">USA</collectingCountry>
|
||
; second half of the Ypresian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F003BFFBDFF0BFBBEFDE014A8" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="discussion">
|
||
<paragraph id="8BA236D4003BFFBEFF0BFBBEFCBA17CE" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
|
||
<emphasis id="B969EAC6003BFFBEFF0BFBBEFEE21660" bold="true" box="[199,332,1115,1141]" pageId="41" pageNumber="42">Discussion.</emphasis>
|
||
<taxonomicName id="4C1D4D57003BFFBEFE99FBBEFE071660" box="[341,425,1115,1141]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="133" phylum="Arthropoda" rank="genus">Valerea</taxonomicName>
|
||
<emphasis id="B969EAC6003BFFBEFE7FFBBEFC351660" box="[435,923,1115,1141]" italics="true" pageId="41" pageNumber="42">
|
||
and the Chickaloon specimen.
|
||
<taxonomicName id="4C1D4D57003BFFBEFCD8FBBEFC351660" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[788,923,1115,1141]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">Okanagrion</taxonomicName>
|
||
</emphasis>
|
||
wings share distinctive character states with that of the Thanetian
|
||
<taxonomicName id="4C1D4D57003BFFBEFE46FB9AFD2C168C" box="[394,642,1151,1177]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="multicellulata">
|
||
<emphasis id="B969EAC6003BFFBEFE46FB9AFD2C168C" box="[394,642,1151,1177]" italics="true" pageId="41" pageNumber="42">Valerea multicellulata</emphasis>
|
||
</taxonomicName>
|
||
from Menat,
|
||
<collectingCountry id="F30A7644003BFFBEFCD2FB65FCC4168F" box="[798,874,1152,1178]" name="France" pageId="41" pageNumber="42">France</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8C4B25003BFFBEFCB6FB9AFBDE168C" author="Garrouste, R. & Wedmann, S. & Pouillon, J. - M. & Nel, A." box="[890,1136,1151,1178]" pageId="41" pageNumber="42" pagination="818 - 821" refId="ref57923" refString="Garrouste, R., Wedmann, S., Pouillon, J. - M. & Nel, A. (2017) The oldest ' amphipterygid' damselfly of tropical affinities in the Paleocene of Menat (Zygoptera: Eucaloptera). Historical Biology, 29, 818 - 821. https: // doi. org / 10.1080 / 08912963.2016.1247448" type="journal article" year="2017">
|
||
Garrouste
|
||
<emphasis id="B969EAC6003BFFBEFC3EFB65FB8A168C" box="[1010,1060,1151,1177]" italics="true" pageId="41" pageNumber="42">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
). That insect has a notably large number of small cells in the costal and RA–RP1 spaces distal to the pterostigma, up to five cells wide, a condition that is only approached in the RA–RP1 space by some Mesozoic
|
||
<taxonomicName id="4C1D4D57003BFFBEFC68FB22FBE916F4" baseAuthorityName="Handlirsch" baseAuthorityYear="1906" box="[932,1095,1223,1249]" class="Insecta" family="Aeschnidiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="family">Aeschnidiidae</taxonomicName>
|
||
, in which it is up to three cells wide (
|
||
<bibRefCitation id="EF8C4B25003BFFBEFF17FB0FFE091711" author="Fleck, G. & Nel, A." box="[219,423,1258,1284]" pageId="41" pageNumber="42" pagination="1 - 180" refId="ref57626" refString="Fleck, G. & Nel, A. (2003) Revision of the Mesozoic family Aeschnidiidae (Odonata: Anisoptera). Zoologica, 153, 1 - 180." type="journal article" year="2003">Fleck & Nel 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8C4B25003BFFBEFE7EFB0FFD0E1711" author="Garrouste, R. & Wedmann, S. & Pouillon, J. - M. & Nel, A." box="[434,672,1258,1285]" pageId="41" pageNumber="42" pagination="818 - 821" refId="ref57923" refString="Garrouste, R., Wedmann, S., Pouillon, J. - M. & Nel, A. (2017) The oldest ' amphipterygid' damselfly of tropical affinities in the Paleocene of Menat (Zygoptera: Eucaloptera). Historical Biology, 29, 818 - 821. https: // doi. org / 10.1080 / 08912963.2016.1247448" type="journal article" year="2017">
|
||
Garrouste
|
||
<emphasis id="B969EAC6003BFFBEFDEAFB0EFDF81711" box="[550,598,1258,1284]" italics="true" pageId="41" pageNumber="42">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
). In
|
||
<taxonomicName id="4C1D4D57003BFFBEFD03FB0FFCF81711" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[719,854,1258,1284]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003BFFBEFD03FB0FFCF81711" box="[719,854,1258,1284]" italics="true" pageId="41" pageNumber="42">Okanagrion</emphasis>
|
||
</taxonomicName>
|
||
, there are usually three, and sometimes four, cells in the costal space, but this space does reach five cells wide in
|
||
<taxonomicName id="4C1D4D57003BFFBEFCF7FAEAFC6A173D" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[827,964,1294,1321]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="worleyae">
|
||
<emphasis id="B969EAC6003BFFBEFCF7FAEAFC6A173D" box="[827,964,1294,1321]" italics="true" pageId="41" pageNumber="42">O. worleyae</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<typeStatus id="54A68876003BFFBEFC33FAEAFBF1173C" box="[1023,1119,1295,1321]" pageId="41" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
1, SR 16-006-001,
|
||
<figureCitation id="13262A51003BFFBEFAF0FAEBFA3E173D" box="[1340,1424,1294,1320]" captionStart="FIGURE 40" captionStartId="72.[151,250,2001,2025]" captionTargetBox="[177,1401,921,1972]" captionTargetId="figure-342@72.[176,1411,921,1978]" captionTargetPageId="72" captionText="FIGURE 40. Okanagrion worleyae paratype 1, SR 16-006-001. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558896" httpUri="https://zenodo.org/record/4558896/files/figure.png" pageId="41" pageNumber="42">Fig. 40</figureCitation>
|
||
). The
|
||
<taxonomicName id="4C1D4D57003BFFBEFF04FAD7FE701759" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[200,478,1330,1356]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="threadgillae">
|
||
<emphasis id="B969EAC6003BFFBEFF04FAD7FE701759" box="[200,478,1330,1356]" italics="true" pageId="41" pageNumber="42">Okanagrion threadgillae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13262A51003BFFBEFE27FAD7FDE61759" box="[491,584,1330,1356]" captionStart="FIGURE 19" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[160,1427,181,1912]" captionTargetId="figure-19@44.[160,1427,181,1919]" captionTargetPageId="44" captionText="FIGURE 19. Okanagrion threadgillae holotype, SR 98-12-10. Photograph. A and B refer to drawings in Fig. 20. Scale bar is 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558846" httpUri="https://zenodo.org/record/4558846/files/figure.png" pageId="41" pageNumber="42">Figs. 19</figureCitation>
|
||
,
|
||
<figureCitation id="13262A51003BFFBEFD9EFAD7FDC11759" box="[594,623,1330,1356]" captionStart="FIGURE 20" captionStartId="45.[151,250,1966,1990]" captionTargetBox="[211,1378,198,1934]" captionTargetId="figure-17@45.[196,1392,181,1942]" captionTargetPageId="45" captionText="FIGURE 20. Okanagrion threadgillae holotype, SR 98-12-10. A, drawing of wing A of Fig. 19; B, drawing of wing B of Fig. 19. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558848" httpUri="https://zenodo.org/record/4558848/files/figure.png" pageId="41" pageNumber="42">20</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C1D4D57003BFFBEFD60FAD7FC9A1759" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[684,820,1330,1356]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="lochmum">
|
||
<emphasis id="B969EAC6003BFFBEFD60FAD7FC9A1759" box="[684,820,1330,1356]" italics="true" pageId="41" pageNumber="42">O. lochmum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13262A51003BFFBEFC8DFAD7FC3C1759" box="[833,914,1330,1356]" captionStart="FIGURE 23" captionStartId="50.[151,250,1999,2023]" captionTargetBox="[221,1367,187,1967]" captionTargetId="figure-19@50.[207,1380,181,1975]" captionTargetPageId="50" captionText="FIGURE 23. Okanagrion lochmum holotype, RBCM 11799.001. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558858" httpUri="https://zenodo.org/record/4558858/files/figure.png" pageId="41" pageNumber="42">Fig. 23</figureCitation>
|
||
)
|
||
<typeStatus id="54A68876003BFFBEFC6CFAD7FBA31759" box="[928,1037,1330,1356]" pageId="41" pageNumber="42" type="holotype">holotypes</typeStatus>
|
||
have five cells wide in the RA–RP1 space, and while
|
||
<taxonomicName id="4C1D4D57003BFFBEFE97FAB3FE62177A" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[347,460,1365,1392]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hobani">
|
||
<emphasis id="B969EAC6003BFFBEFE97FAB3FEC11765" box="[347,367,1366,1392]" italics="true" pageId="41" pageNumber="42">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC6003BFFBEFEB2FAB0FE62177A" box="[382,460,1365,1391]" italics="true" pageId="41" pageNumber="42">hobani</emphasis>
|
||
</taxonomicName>
|
||
specimens usually have a maximum of four cells wide in this space, it does reach this remarkable number in
|
||
<typeStatus id="54A68876003BFFBEFE59FA9FFE5B1781" box="[405,501,1402,1428]" pageId="41" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
3 (SR 94-05-22,
|
||
<figureCitation id="13262A51003BFFBEFD74FA9CFCA41786" box="[696,778,1401,1427]" captionStart="FIGURE 27" captionStartId="56.[151,250,1989,2013]" captionTargetBox="[197,1391,187,1959]" captionTargetId="figure-19@56.[185,1403,181,1965]" captionTargetPageId="56" captionText="FIGURE 27. Okanagrion hobani paratype 3, SR 94-05-22. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558870" httpUri="https://zenodo.org/record/4558870/files/figure.png" pageId="41" pageNumber="42">Fig. 27</figureCitation>
|
||
),
|
||
<typeStatus id="54A68876003BFFBEFCD2FA9FFCD01781" box="[798,894,1402,1428]" pageId="41" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
4 (SRUI 99-96-55,
|
||
<figureCitation id="13262A51003BFFBEFB93FA9CFB1F1786" box="[1119,1201,1401,1427]" captionStart="FIGURE 28" captionStartId="57.[151,250,1959,1983]" captionTargetBox="[203,1375,188,1932]" captionTargetId="figure-17@57.[192,1395,181,1935]" captionTargetPageId="57" captionText="FIGURE 28. Okanagrion hobani. Paratype 4, SRUI 99-96-55: A, photograph; B, drawing; SR 13-005-017: C, photograph; D, drawing.All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558872" httpUri="https://zenodo.org/record/4558872/files/figure.png" pageId="41" pageNumber="42">Fig. 28</figureCitation>
|
||
), and
|
||
<typeStatus id="54A68876003BFFBEFB39FA9FFAFB1781" box="[1269,1365,1402,1428]" pageId="41" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
6 (SR 00-04-05,
|
||
<figureCitation id="13262A51003BFFBEFEC1FA78FECF17A2" box="[269,353,1437,1463]" captionStart="FIGURE 30" captionStartId="59.[151,250,1943,1967]" captionTargetBox="[190,1395,184,1909]" captionTargetId="figure-17@59.[190,1398,181,1919]" captionTargetPageId="59" captionText="FIGURE 30. Okanagrion hobani. Paratype 6, SR 00-04-05:A, photograph; B, drawing; SR 06-62-06:A, photograph; B, drawing. Both to scale, 5 mm" figureDoi="http://doi.org/10.5281/zenodo.4558876" httpUri="https://zenodo.org/record/4558876/files/figure.png" pageId="41" pageNumber="42">Fig. 30</figureCitation>
|
||
).
|
||
<bibRefCitation id="EF8C4B25003BFFBEFEB9FA78FDD817A2" author="Garrouste, R. & Wedmann, S. & Pouillon, J. - M. & Nel, A." box="[373,630,1437,1464]" pageId="41" pageNumber="42" pagination="818 - 821" refId="ref57923" refString="Garrouste, R., Wedmann, S., Pouillon, J. - M. & Nel, A. (2017) The oldest ' amphipterygid' damselfly of tropical affinities in the Paleocene of Menat (Zygoptera: Eucaloptera). Historical Biology, 29, 818 - 821. https: // doi. org / 10.1080 / 08912963.2016.1247448" type="journal article" year="2017">
|
||
Garrouste
|
||
<emphasis id="B969EAC6003BFFBEFE27FA7BFDB317A2" box="[491,541,1437,1463]" italics="true" pageId="41" pageNumber="42">et al</emphasis>
|
||
. (2017)
|
||
</bibRefCitation>
|
||
noted that five cells width in these two spaces distal to the pterostigma was then only known in the
|
||
<taxonomicName id="4C1D4D57003BFFBEFE1FFA24FD9C17CE" box="[467,562,1473,1499]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||
in
|
||
<taxonomicName id="4C1D4D57003BFFBEFD9AFA27FCA317CE" box="[598,781,1473,1499]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="multicellulata">
|
||
<emphasis id="B969EAC6003BFFBEFD9AFA27FCA317CE" box="[598,781,1473,1499]" italics="true" pageId="41" pageNumber="42">V. multicellulata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA236D4003BFFBEFF0BFA01FD1D14EC" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
|
||
The pterostigma of
|
||
<taxonomicName id="4C1D4D57003BFFBEFE6DFA00FDF617EB" box="[417,600,1508,1534]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="multicellulata">
|
||
<emphasis id="B969EAC6003BFFBEFE6DFA00FDF617EB" box="[417,600,1508,1534]" italics="true" pageId="41" pageNumber="42">V. multicellulata</emphasis>
|
||
</taxonomicName>
|
||
is also distinct by its shape, the distal side is strongly curved and the proximal side is very long and oblique to C and RA, with two crossveins between the
|
||
<collectingRegion id="49D9F836003BFFBEFBF0F9EDFBD21437" box="[1084,1148,1544,1570]" country="New Zealand" name="West Coast" pageId="41" pageNumber="42">Costa</collectingRegion>
|
||
and proximal side of the pterostigma,
|
||
<emphasis id="B969EAC6003BFFBEFEE6F9C8FEEB1453" box="[298,325,1581,1606]" italics="true" pageId="41" pageNumber="42">i.e</emphasis>
|
||
., hyperstigmal crossveins. Hyperstigmal crossveins are uncommon in extant
|
||
<taxonomicName id="4C1D4D57003BFFBEFB66F9C8FAB31452" box="[1194,1309,1581,1607]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="124" phylum="Arthropoda" rank="subOrder" subOrder="Zygoptera">Zygoptera</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF8C4B25003BFFBEFAE0F9C9FEB6147F" author="Garrouste, R. & Wedmann, S. & Pouillon, J. - M. & Nel, A." pageId="41" pageNumber="42" pagination="818 - 821" refId="ref57923" refString="Garrouste, R., Wedmann, S., Pouillon, J. - M. & Nel, A. (2017) The oldest ' amphipterygid' damselfly of tropical affinities in the Paleocene of Menat (Zygoptera: Eucaloptera). Historical Biology, 29, 818 - 821. https: // doi. org / 10.1080 / 08912963.2016.1247448" type="journal article" year="2017">
|
||
Garrouste
|
||
<emphasis id="B969EAC6003BFFBEFF5BF9B5FF7C147F" box="[151,210,1616,1642]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
). The pterostigmata of
|
||
<taxonomicName id="4C1D4D57003BFFBEFDEFF9B5FD04147F" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[547,682,1616,1642]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003BFFBEFDEFF9B5FD04147F" box="[547,682,1616,1642]" italics="true" pageId="41" pageNumber="42">Okanagrion</emphasis>
|
||
</taxonomicName>
|
||
species are shaped like this, and the majority of specimens of
|
||
<taxonomicName id="4C1D4D57003BFFBEFA4DF9B5FF4B1498" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="hobani">
|
||
<emphasis id="B969EAC6003BFFBEFA4DF9B5FF4B1498" italics="true" pageId="41" pageNumber="42">O. hobani</emphasis>
|
||
</taxonomicName>
|
||
, and one each of
|
||
<taxonomicName id="4C1D4D57003BFFBEFE63F991FD971498" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[431,569,1651,1678]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="worleyae">
|
||
<emphasis id="B969EAC6003BFFBEFE63F991FE6D149B" box="[431,451,1652,1678]" italics="true" pageId="41" pageNumber="42">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC6003BFFBEFE1EF996FD971498" box="[466,569,1651,1677]" italics="true" pageId="41" pageNumber="42">worleyae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D57003BFFBEFDBDF991FD721498" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[625,732,1651,1678]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="beardi">
|
||
<emphasis id="B969EAC6003BFFBEFDBDF991FD2B149B" box="[625,645,1652,1678]" italics="true" pageId="41" pageNumber="42">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC6003BFFBEFD58F996FD721498" box="[660,732,1651,1677]" italics="true" pageId="41" pageNumber="42">beardi</emphasis>
|
||
</taxonomicName>
|
||
have a hyperstigmal crossvein, and the
|
||
<typeStatus id="54A68876003BFFBEFB6EF996FAAA1498" box="[1186,1284,1651,1677]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
hind wing of
|
||
<taxonomicName id="4C1D4D57003BFFBEFF5BF97DFEEC14A4" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[151,322,1687,1714]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="threadgillae">
|
||
<emphasis id="B969EAC6003BFFBEFF5BF97DFEEC14A4" box="[151,322,1687,1714]" italics="true" pageId="41" pageNumber="42">O. threadgillae</emphasis>
|
||
</taxonomicName>
|
||
bears two (
|
||
<figureCitation id="13262A51003BFFBEFE0AF972FD8314A4" box="[454,557,1687,1713]" captionStart="FIGURE 20" captionStartId="45.[151,250,1966,1990]" captionTargetBox="[211,1378,198,1934]" captionTargetId="figure-17@45.[196,1392,181,1942]" captionTargetPageId="45" captionText="FIGURE 20. Okanagrion threadgillae holotype, SR 98-12-10. A, drawing of wing A of Fig. 19; B, drawing of wing B of Fig. 19. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558848" httpUri="https://zenodo.org/record/4558848/files/figure.png" pageId="41" pageNumber="42">Fig. 20B</figureCitation>
|
||
). The pterostigma of
|
||
<taxonomicName id="4C1D4D57003BFFBEFCD3F972FCDA14A4" box="[799,884,1687,1713]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003BFFBEFCD3F972FCDA14A4" box="[799,884,1687,1713]" italics="true" pageId="41" pageNumber="42">Valerea</emphasis>
|
||
</taxonomicName>
|
||
is about three times longer than wide, within the lower range of
|
||
<taxonomicName id="4C1D4D57003BFFBEFE89F95EFE6214C0" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[325,460,1723,1749]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003BFFBEFE89F95EFE6214C0" box="[325,460,1723,1749]" italics="true" pageId="41" pageNumber="42">Okanagrion</emphasis>
|
||
</taxonomicName>
|
||
. The vein fragment of
|
||
<taxonomicName id="4C1D4D57003BFFBEFD18F959FC2314C0" box="[724,909,1723,1749]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="multicellulata">
|
||
<emphasis id="B969EAC6003BFFBEFD18F959FC2314C0" box="[724,909,1723,1749]" italics="true" pageId="41" pageNumber="42">V. multicellulata</emphasis>
|
||
</taxonomicName>
|
||
that we interpret as CuA is linear at least well proximal to the origin of IR1, as in
|
||
<taxonomicName id="4C1D4D57003BFFBEFDE9F93AFD0214EC" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[549,684,1759,1785]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003BFFBEFDE9F93AFD0214EC" box="[549,684,1759,1785]" italics="true" pageId="41" pageNumber="42">Okanagrion</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA236D4003BFFBEFF0BF8E7FABC159D" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
|
||
The proximal portion of IR
|
||
<quantity id="4CE59B31003BFFBEFDCDF8E7FD831509" box="[513,557,1794,1820]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="41" pageNumber="42" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="4C1D4D57003BFFBEFDFAF8E6FD411509" box="[566,751,1794,1820]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="multicellulata">
|
||
<emphasis id="B969EAC6003BFFBEFDFAF8E6FD411509" box="[566,751,1794,1820]" italics="true" pageId="41" pageNumber="42">V. multicellulata</emphasis>
|
||
</taxonomicName>
|
||
is not preserved from immediately distal to the level of the nodus. Unlike the condition in
|
||
<taxonomicName id="4C1D4D57003BFFBEFE34F8C3FDD11555" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[504,639,1830,1856]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6003BFFBEFE34F8C3FDD11555" box="[504,639,1830,1856]" italics="true" pageId="41" pageNumber="42">Okanagrion</emphasis>
|
||
</taxonomicName>
|
||
species where IR2 originates at this level either closer to RP1-2 or on RP1-2, the proximal-most preserved portion in
|
||
<taxonomicName id="4C1D4D57003BFFBEFD7DF8AEFCC41571" box="[689,874,1866,1892]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="multicellulata">
|
||
<emphasis id="B969EAC6003BFFBEFD7DF8AEFCC41571" box="[689,874,1866,1892]" italics="true" pageId="41" pageNumber="42">V. multicellulata</emphasis>
|
||
</taxonomicName>
|
||
is relatively far from RP1-2 and closer to RP3-4, indicating that it likely originates well proximal to the nodus, or if at that level, then on or near RP3-4.
|
||
</paragraph>
|
||
<paragraph id="8BA236D4003BFFBEFF0BF874FAA015E1" blockId="41.[151,1437,151,2036]" pageId="41" pageNumber="42">
|
||
The Thanetian of Menat and the Ypresian Okanagan Highlands are probably about three to six million years apart, and would have been connected by continuous land at that time, with mild climates reaching high latitudes facilitating intercontinental dispersal (reviewed:
|
||
<bibRefCitation id="EF8C4B25003BFFBEFD7AF83CFC5715E6" author="Archibald, S. B. & Makarkin, V. N." box="[694,1017,2009,2035]" pageId="41" pageNumber="42" pagination="119 - 155" refId="ref55867" refString="Archibald, S. B. & Makarkin, V. N. (2006) Tertiary Giant Lacewings (Neuroptera: Polystoechotidae) revision and description of new taxa from western North America and Denmark. Journal of Systematic Paleontology, 4, 119 - 155, 307 (errata). https: // doi. org / 10.1017 / S 1477201906001817" type="journal article" year="2006">Archibald & Makarkin 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8C4B25003BFFBEFBCFF83CFAAD15E1" author="Archibald, S. B. & Johnson, K. R. & Mathewes, R. W. & Greenwood, D. R." box="[1027,1283,2009,2036]" pageId="41" pageNumber="42" pagination="3679 - 3686" refId="ref55797" refString="Archibald, S. B., Johnson, K. R., Mathewes, R. W. & Greenwood, D. R. (2011 b) Intercontinental dispersal of giant thermophilic ants across the Arctic during early Eocene hyperthermals. Proceedings of the Royal Society B, 278, 3679 - 3686. https: // doi. org / 10.1098 / rspb. 2011.0729" type="journal article" year="2011">
|
||
Archibald
|
||
<emphasis id="B969EAC6003BFFBEFBB6F83FFB0515E6" box="[1146,1195,2009,2035]" italics="true" pageId="41" pageNumber="42">et al</emphasis>
|
||
. 2011b
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA236D40038FFBDFF0BFF72FC91139C" blockId="42.[151,1437,151,1725]" pageId="42" pageNumber="43">
|
||
The specimen of undetermined genus and species from the Thanetian or Ypresian Chickaloon Formation of
|
||
<collectingRegion id="49D9F8360038FFBDFF5BFF5EFF4B12C0" box="[151,229,187,213]" country="United States of America" name="Alaska" pageId="42" pageNumber="43">Alaska</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EF8C4B250038FFBDFF38FF5EFE5D12C0" author="Garrouste, R. & Nel, A." box="[244,499,187,213]" pageId="42" pageNumber="43" pagination="1939 - 1946" refId="ref57867" refString="Garrouste, R. & Nel, A. (2019) Alaskan Palaeogene insects: a challenge for a better knowledge of the Beringian ' route' (Odonata: Aeshnidae, Dysagrionidae). Journal of Systematic Palaeontology, 17, 1939 - 1946. https: // doi. org / 10.1080 / 14772019.2019.1572235" type="journal article" year="2019">Garrouste & Nel 2019</bibRefCitation>
|
||
) appears quite close to
|
||
<taxonomicName id="4C1D4D570038FFBDFD31FF5EFC2A12C0" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[765,900,187,213]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC60038FFBDFD31FF5EFC2A12C0" box="[765,900,187,213]" italics="true" pageId="42" pageNumber="43">Okanagrion</emphasis>
|
||
</taxonomicName>
|
||
with its dense venation, including an expanded CuA–A space four cells wide at its widest, between the levels of origins of
|
||
<collectingRegion id="49D9F8360038FFBDFC15FF3AFC5212EC" box="[985,1020,223,249]" country="Bulgaria" name="Ruse" pageId="42" pageNumber="43">RS</collectingRegion>
|
||
2 and IR1. In its diagnosis, above, we cite the MA beginning to zigzag in the distal-most portion of preserved in the Chickaloon specimen as distinguishing it from
|
||
<taxonomicName id="4C1D4D570038FFBDFED8FEC2FE351354" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[276,411,295,321]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC60038FFBDFED8FEC2FE351354" box="[276,411,295,321]" italics="true" pageId="42" pageNumber="43">Okanagrion</emphasis>
|
||
</taxonomicName>
|
||
(where this is linear), although this is not a great difference, and they might be congeners or closely related genera. It is clearly not a species of
|
||
<taxonomicName id="4C1D4D570038FFBDFD17FEAEFC9E1370" box="[731,816,331,357]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC60038FFBDFD17FEAEFC9E1370" box="[731,816,331,357]" italics="true" pageId="42" pageNumber="43">Valerea</emphasis>
|
||
</taxonomicName>
|
||
as,
|
||
<emphasis id="B969EAC60038FFBDFC91FEAEFCC71371" box="[861,873,331,356]" italics="true" pageId="42" pageNumber="43">e</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60038FFBDFCBDFEAEFCD11371" box="[881,895,331,356]" italics="true" pageId="42" pageNumber="43">g</emphasis>
|
||
., the crossvenation is much more dense and the origins of IR1 and RP2 are nearer to the nodus in that genus.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40038FFBDFF0BFE76FECF1090" blockId="42.[151,1437,151,1725]" pageId="42" pageNumber="43">
|
||
<emphasis id="B969EAC60038FFBDFF0BFE76FE2F13B8" box="[199,385,403,429]" italics="true" pageId="42" pageNumber="43">Species richness</emphasis>
|
||
. It is not surprising that there is high species richness of
|
||
<taxonomicName id="4C1D4D570038FFBDFBDCFE76FB3913B8" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[1040,1175,403,429]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC60038FFBDFBDCFE76FB3913B8" box="[1040,1175,403,429]" italics="true" pageId="42" pageNumber="43">Okanagrion</emphasis>
|
||
</taxonomicName>
|
||
at two localities: eight in total, with four at McAbee and five at Republic (one of these shared between them). High alpha diversity was established at McAbee by
|
||
<bibRefCitation id="EF8C4B250038FFBDFE0CFE3EFD6B13E0" author="Archibald, S. B. & Bossert, W. H. & Greenwood, D. R. & Farrell, B. D." box="[448,709,475,501]" pageId="42" pageNumber="43" pagination="374 - 398" refId="ref55509" refString="Archibald, S. B., Bossert, W. H., Greenwood, D. R. & Farrell, B. D. (2010) Seasonality, the latitudinal gradient of diversity, and Eocene insects. Paleobiology, 36, 374 - 398. https: // doi. org / 10.1666 / 09021.1" type="journal article" year="2010">
|
||
Archibald
|
||
<emphasis id="B969EAC60038FFBDFDF4FE3EFDC713E0" box="[568,617,475,501]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. (2010)
|
||
</bibRefCitation>
|
||
, who found insect and woody dicotyledon species richness there to be equivalent to that of a modern lowland tropical rainforest in
|
||
<collectingCountry id="F30A76440038FFBDFCB8FE1AFC5C100C" box="[884,1010,511,537]" name="Costa Rica" pageId="42" pageNumber="43">Costa Rica</collectingCountry>
|
||
. That this richness is not an artefact of splitting was confirmed by the independent assessments of experts in individual insect orders (see
|
||
<bibRefCitation id="EF8C4B250038FFBDFB20FDC6FF7A1074" author="Archibald, S. B. & Bossert, W. H. & Greenwood, D. R. & Farrell, B. D." pageId="42" pageNumber="43" pagination="374 - 398" refId="ref55509" refString="Archibald, S. B., Bossert, W. H., Greenwood, D. R. & Farrell, B. D. (2010) Seasonality, the latitudinal gradient of diversity, and Eocene insects. Paleobiology, 36, 374 - 398. https: // doi. org / 10.1666 / 09021.1" type="journal article" year="2010">
|
||
Archibald
|
||
<emphasis id="B969EAC60038FFBDFAA8FDC6FA3B1028" box="[1380,1429,547,573]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
). Such high, tropical levels of alpha diversity for woody dicotyledons have also been established at Republic (
|
||
<bibRefCitation id="EF8C4B250038FFBDFF52FD8EFEF81090" author="Wilf, P. & Cuneo, N. R. & Johnson, K. R. & Hicks, J. F. & Wing, S. L. & Obradovich, J. D." box="[158,342,619,645]" pageId="42" pageNumber="43" pagination="122 - 125" refId="ref62279" refString="Wilf, P., Cuneo, N. R. Johnson, K. R., Hicks, J. F., Wing, S. L. & Obradovich, J. D. (2003) High plant diversity in Eocene South America: evidence from Patagonia. Science, 300, 122 - 125. https: // doi. org / 10.1126 / science. 1080475" type="journal article" year="2003">
|
||
Wilf
|
||
<emphasis id="B969EAC60038FFBDFF16FD8EFEA51090" box="[218,267,619,645]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. 2003
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA236D40038FFBDFF0BFD6AFCD8112C" blockId="42.[151,1437,151,1725]" pageId="42" pageNumber="43">
|
||
This is also consistent with the findings of
|
||
<bibRefCitation id="EF8C4B250038FFBDFD6FFD6AFC0D10BC" author="Archibald, S. B. & Greenwood, D. R. & Mathewes, R. W." box="[675,931,655,681]" pageId="42" pageNumber="43" pagination="1 - 8" refId="ref55667" refString="Archibald, S. B., Greenwood, D. R. & Mathewes, R. W. (2013) Seasonality, montane beta diversity, and Eocene insects: testing Janzen's dispersal hypothesis in an equable world. Palaeogeography, Palaeoclimatology, Palaeoecology, 371, 1 - 8. https: // doi. org / 10.1016 / j. palaeo. 2012.10.043" type="journal article" year="2013">
|
||
Archibald
|
||
<emphasis id="B969EAC60038FFBDFCD6FD6AFCE510BC" box="[794,843,655,681]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. (2013)
|
||
</bibRefCitation>
|
||
that insect beta diversity is high between site pairs in the Okanagan Highlands. They extended Janzen’s hypothesis “Mountain passes are higher in the tropics” to mid-latitude mountainous regions of the Eocene by their “greenhouse world” extra-tropical climatic equability. This diversity pattern is then expected with climate-driven high alpha diversity, and climate and topography-driven high beta diversity. Other genera described here are less speciose.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40038FFBDFF0BFCA6FB8011DC" blockId="42.[151,1437,151,1725]" pageId="42" pageNumber="43">
|
||
<emphasis id="B969EAC60038FFBDFF0BFCA6FE611149" box="[199,463,835,861]" italics="true" pageId="42" pageNumber="43">Wing colour patterning</emphasis>
|
||
. There is sexual dimorphism in wing colouration in a variety of extant
|
||
<taxonomicName id="4C1D4D570038FFBDFB35FCA6FAC21148" box="[1273,1388,835,861]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="124" phylum="Arthropoda" rank="subOrder" subOrder="Zygoptera">Zygoptera</taxonomicName>
|
||
and this may also be true in
|
||
<taxonomicName id="4C1D4D570038FFBDFE69FC82FDDA1194" box="[421,628,871,897]" kingdom="Animalia" pageId="42" pageNumber="27" rank="subOrder" subOrder="Cephalozygoptera">Cephalozygoptera</taxonomicName>
|
||
. Further, there may be differences in patterning between the hind wings and forewings. In some
|
||
<taxonomicName id="4C1D4D570038FFBDFE6EFC6EFDBB11B0" box="[418,533,907,933]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="124" phylum="Arthropoda" rank="subOrder" subOrder="Zygoptera">Zygoptera</taxonomicName>
|
||
species, there is also colour polymorphism in a single sex (
|
||
<emphasis id="B969EAC60038FFBDFB6AFC6EFB6611B1" box="[1190,1224,907,932]" italics="true" pageId="42" pageNumber="43">e.g</emphasis>
|
||
., clear-winged and pattern-winged males of
|
||
<taxonomicName id="4C1D4D570038FFBDFE62FC4AFC8611DF" box="[430,808,943,970]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC60038FFBDFE62FC4AFD9411DC" box="[430,570,943,969]" italics="true" pageId="42" pageNumber="43">Paraphlebia</emphasis>
|
||
(Thaumatoneuridae)
|
||
</taxonomicName>
|
||
, but this is uncommon.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40038FFBDFF0BFC36FD5616B4" blockId="42.[151,1437,151,1725]" pageId="42" pageNumber="43">
|
||
There are
|
||
<specimenCount id="9D1BFD5D0038FFBDFEF4FC36FE7E11F8" box="[312,464,979,1005]" pageId="42" pageNumber="43" type="generic">17 specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="4C1D4D570038FFBDFE39FC36FD7F11F8" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[501,721,979,1005]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="hobani">
|
||
<emphasis id="B969EAC60038FFBDFE39FC36FD7F11F8" box="[501,721,979,1005]" italics="true" pageId="42" pageNumber="43">Okanagrion hobani</emphasis>
|
||
</taxonomicName>
|
||
illustrated and discussed here, one of which we know is a male. While it seems unlikely that all 17 would be males, still, this is possible.
|
||
<taxonomicName id="4C1D4D570038FFBDFC02FC12FBFB1604" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[974,1109,1015,1041]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC60038FFBDFC02FC12FBFB1604" box="[974,1109,1015,1041]" italics="true" pageId="42" pageNumber="43">Okanagrion</emphasis>
|
||
</taxonomicName>
|
||
species concepts are defined by other wing traits, with wing colouration as supplementary. Taking a conservative stance, we identify wing colouration as male in the
|
||
<taxonomicName id="4C1D4D570038FFBDFE4BFBDAFE56164C" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[391,504,1087,1113]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="hobani">
|
||
<emphasis id="B969EAC60038FFBDFE4BFBDAFE56164C" box="[391,504,1087,1113]" italics="true" pageId="42" pageNumber="43">O. hobani</emphasis>
|
||
</taxonomicName>
|
||
diagnosis, with female colouration unknown, and in other species specify that the sex is unknown. Colouration is consistent between wings identified as fore- and hind wings, and in the articulated fore- and hind wings of
|
||
<taxonomicName id="4C1D4D570038FFBDFE69FB62FDBB16B4" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[421,533,1159,1185]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="hobani">
|
||
<emphasis id="B969EAC60038FFBDFE69FB62FDBB16B4" box="[421,533,1159,1185]" italics="true" pageId="42" pageNumber="43">O. hobani</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54A688760038FFBDFDD0FB62FDD216B4" box="[540,636,1159,1185]" pageId="42" pageNumber="43" type="paratype">paratype</typeStatus>
|
||
7, F-1044.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40038FFBDFF0BFB4EFDE014A8" blockId="42.[151,1437,151,1725]" pageId="42" pageNumber="43">
|
||
<emphasis id="B969EAC60038FFBDFF0BFB4EFE5C16D1" box="[199,498,1195,1221]" italics="true" pageId="42" pageNumber="43">Forewings and hind wings</emphasis>
|
||
. Although forewings are close in shape to hind wings in
|
||
<taxonomicName id="4C1D4D570038FFBDFBBBFB4EFAE816D0" box="[1143,1350,1195,1221]" kingdom="Animalia" pageId="42" pageNumber="27" rank="subOrder" subOrder="Cephalozygoptera">Cephalozygoptera</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D570038FFBDFA9CFB4EFEAB16FC" authorityName="Handlirsch" authorityYear="1906" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="124" phylum="Arthropoda" rank="subOrder" subOrder="Anisozygoptera">Anisozygoptera</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D570038FFBDFEF6FB2AFE1E16FC" box="[314,432,1231,1257]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="124" phylum="Arthropoda" rank="subOrder" subOrder="Zygoptera">Zygoptera</taxonomicName>
|
||
, they may in cases be somewhat different (
|
||
<emphasis id="B969EAC60038FFBDFC47FB2AFC0316FD" box="[907,941,1231,1256]" italics="true" pageId="42" pageNumber="43">e.g</emphasis>
|
||
., slightly in
|
||
<taxonomicName id="4C1D4D570038FFBDFBFBFB2AFA9F16FC" authority="Selys" authorityName="Selys" authorityYear="1861" box="[1079,1329,1231,1257]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="zoe">
|
||
<emphasis id="B969EAC60038FFBDFBFBFB2AFB4016FD" box="[1079,1262,1231,1257]" italics="true" pageId="42" pageNumber="43">Paraphlebia zoe</emphasis>
|
||
Selys
|
||
</taxonomicName>
|
||
in Hagen see Figs. 470 and 436 and 437 of
|
||
<bibRefCitation id="EF8C4B250038FFBDFDC0FB16FD431718" author="Garrison, R. W. & von Ellenrieder, N. & Louton, J. A." box="[524,749,1267,1293]" pageId="42" pageNumber="43" refId="ref57766" refString="Garrison, R. W., von Ellenrieder, N. & Louton, J. A. (2010) Damselfly genera of the New World: an illustrated and annotated key to the Zygoptera. Johns Hopkins University Press, Baltimore, 490 pp." type="book" year="2010">
|
||
Garrison
|
||
<emphasis id="B969EAC60038FFBDFDB8FB16FD0B1718" box="[628,677,1267,1293]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
, more so in
|
||
<taxonomicName id="4C1D4D570038FFBDFCBFFB16FA8E1718" authority="McLachlan" authorityName="McLachlan" authorityYear="1897" box="[883,1312,1267,1293]" class="Insecta" family="Megapodagrionidae" genus="Thaumatoneura" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="inopinata">
|
||
<emphasis id="B969EAC60038FFBDFCBFFB16FB391719" box="[883,1175,1267,1293]" italics="true" pageId="42" pageNumber="43">Thaumatoneura inopinata</emphasis>
|
||
McLachlan
|
||
</taxonomicName>
|
||
, their Figs. 470 and 471). We base our assignment of fore- and hind wings on those of the
|
||
<typeStatus id="54A688760038FFBDFBC1FAF2FBC11724" box="[1037,1135,1303,1329]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
|
||
(and only specimen) of
|
||
<taxonomicName id="4C1D4D570038FFBDFA4DFAF2FEB11740" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="threadgillae">
|
||
<emphasis id="B969EAC60038FFBDFA4DFAF2FEB11740" italics="true" pageId="42" pageNumber="43">O. threadgillae</emphasis>
|
||
</taxonomicName>
|
||
, where two wings that conform to the differences seen in some modern species are in close association: the wing that we presume to be a forewing is slightly wider at the base, with the posterior margin slightly convex, widened at the level of the distal portion of the quadrangle (
|
||
<figureCitation id="13262A510038FFBDFCF9FA66FC261788" box="[821,904,1410,1437]" captionStart="FIGURE 35" captionStartId="65.[151,250,1490,1514]" captionTargetBox="[157,1426,185,1456]" captionTargetId="figure-17@65.[151,1436,181,1466]" captionTargetPageId="65" captionText="FIGURE 35. Okanagrion hobani relatively complete wings, comparative forewings (FW) and hind wings (HW). Red, blue arrows, see text. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558886" httpUri="https://zenodo.org/record/4558886/files/figure.png" pageId="42" pageNumber="43">Fig. 35</figureCitation>
|
||
, red arrows), and the anterior margin is slightly curved between the nodus and the pterostigma. In the presumptive hind wing, the proximal posterior margin is smoothly curved through the level of the quadrangle (
|
||
<figureCitation id="13262A510038FFBDFD25FA2EFC9417F0" box="[745,826,1482,1509]" captionStart="FIGURE 35" captionStartId="65.[151,250,1490,1514]" captionTargetBox="[157,1426,185,1456]" captionTargetId="figure-17@65.[151,1436,181,1466]" captionTargetPageId="65" captionText="FIGURE 35. Okanagrion hobani relatively complete wings, comparative forewings (FW) and hind wings (HW). Red, blue arrows, see text. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558886" httpUri="https://zenodo.org/record/4558886/files/figure.png" pageId="42" pageNumber="43">Fig. 35</figureCitation>
|
||
, blue arrow), the anterior margin is rather straight from the nodus to the pterostigma, and the posterior margin is more deeply curved. We compare this difference to that of the fore- and hind wings of
|
||
<taxonomicName id="4C1D4D570038FFBDFE05F9F6FDD21438" authorityName="McLachlan" authorityYear="1897" box="[457,636,1555,1581]" class="Insecta" family="Megapodagrionidae" genus="Thaumatoneura" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC60038FFBDFE05F9F6FDD21438" box="[457,636,1555,1581]" italics="true" pageId="42" pageNumber="43">Thaumatoneura</emphasis>
|
||
</taxonomicName>
|
||
species, although not as pronounced as in some of those. Unfortunately, in the male specimen F-1044 of
|
||
<taxonomicName id="4C1D4D570038FFBDFDCEF9D2FDDC1444" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[514,626,1591,1617]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="hobani">
|
||
<emphasis id="B969EAC60038FFBDFDCEF9D2FDB81444" box="[514,534,1591,1617]" italics="true" pageId="42" pageNumber="43">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60038FFBDFDE8F9D2FDDC1444" box="[548,626,1591,1617]" italics="true" pageId="42" pageNumber="43">hobani</emphasis>
|
||
</taxonomicName>
|
||
with all four wings preserved, these are somewhat damaged, precluding meaningful comparison of details of shape. Rather complete and undamaged wings of
|
||
<taxonomicName id="4C1D4D570038FFBDFBA6F9BEFB5F1460" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[1130,1265,1627,1653]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC60038FFBDFBA6F9BEFB5F1460" box="[1130,1265,1627,1653]" italics="true" pageId="42" pageNumber="43">Okanagrion</emphasis>
|
||
</taxonomicName>
|
||
species appear to vary in the same manner as do those that we presume to be the fore- and hind wing of
|
||
<taxonomicName id="4C1D4D570038FFBDFBB9F99AFAB1148C" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[1141,1311,1663,1689]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="threadgillae">
|
||
<emphasis id="B969EAC60038FFBDFBB9F99AFAB1148C" box="[1141,1311,1663,1689]" italics="true" pageId="42" pageNumber="43">O. threadgillae</emphasis>
|
||
</taxonomicName>
|
||
, and so we assume these identities where possible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |