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<document id="44404DB83B2B107B881E881E7891ABD4" ID-CLB-Dataset="256481" ID-DOI="10.11646/zootaxa.5240.1.1" ID-GBIF-Dataset="4d650293-3f9a-4bbe-9530-204a3b09ef8b" ID-ISSN="1175-5326" ID-Zenodo-Dep="7639547" ID-ZooBank="2FE467C1-EAC2-4E90-B673-CCE2CCA93C1D" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1676380923589" checkinUser="plazi" docAuthor="Mendonça, Maria Thayane Da Silva, Silva, Valéria Juliete Da &amp; Fernandes, José Antônio Marin" docDate="2023" docId="03B487C0FFFA2520FF47C4C0F824FEAB" docLanguage="en" docName="zootaxa.5240.1.1.pdf" docOrigin="Zootaxa 5240 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5240.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Edessa (Edessa) subandina Breddin 1904" docType="treatment" docVersion="9" lastPageNumber="50" masterDocId="FF8DFFB8FFCA2511FFD0C712FB1CFFB6" masterDocTitle="Diagnose of the nominal subgenus of Edessa and description of the E. sexdens group (Heteroptera, Pentatomidae, Edessinae)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="49" updateTime="1699475138964" updateUser="ExternalLinkService">
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<mods:title id="48CE584BB4BED8C91526981FD1B6233F">Diagnose of the nominal subgenus of Edessa and description of the E. sexdens group (Heteroptera, Pentatomidae, Edessinae)</mods:title>
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<mods:namePart id="EA707BFFA3F36FC831B781B08121F5CE">Mendonça, Maria Thayane Da Silva</mods:namePart>
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<mods:namePart id="9417D7FFEB65D9749FFB3388B5E9E23C">Silva, Valéria Juliete Da</mods:namePart>
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<mods:namePart id="99DDC714623A1B086329B4D9732819B5">Fernandes, José Antônio Marin</mods:namePart>
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<treatment id="03B487C0FFFA2520FF47C4C0F824FEAB" ID-DOI="http://doi.org/10.5281/zenodo.7639931" ID-GBIF-Taxon="209578837" ID-Zenodo-Dep="7639931" LSID="urn:lsid:plazi:treatment:03B487C0FFFA2520FF47C4C0F824FEAB" httpUri="http://treatment.plazi.org/id/03B487C0FFFA2520FF47C4C0F824FEAB" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
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<paragraph id="8BA236D6FFFA2521FF47C4C0F943FC5A" blockId="48.[151,607,978,1041]" box="[151,607,978,1005]" pageId="48" pageNumber="49">
<heading id="D0EA81BAFFFA2521FF47C4C0F943FC5A" bold="true" box="[151,607,978,1005]" fontSize="11" level="2" pageId="48" pageNumber="49" reason="4">
<taxonomicName id="4C1D4D55FFFA2521FF47C4C0F943FC5A" authority="Breddin, 1904" authorityName="Breddin" authorityYear="1904" box="[151,607,978,1005]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="subandina" subGenus="Edessa">
<emphasis id="B969EAC4FFFA2521FF47C4C0F943FC5A" bold="true" box="[151,607,978,1005]" pageId="48" pageNumber="49">
<emphasis id="B969EAC4FFFA2521FF47C4C0FAB4FC5B" bold="true" box="[151,424,978,1005]" italics="true" pageId="48" pageNumber="49">Edessa (E.) subandina</emphasis>
<bibRefCitation id="EF8C4B27FFFA2521FE60C4C0F943FC5A" author="Breddin, G." box="[432,607,978,1004]" pageId="48" pageNumber="49" pagination="135 - 154" refId="ref36797" refString="Breddin, G. (1904) Beitrage zur Systematik der Rhynchoten. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 14, 135 - 154. https: // doi. org / 10.5962 / bhl. part. 8958" type="journal article" year="1904">Breddin, 1904</bibRefCitation>
</emphasis>
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<subSubSection id="C307655DFFFA2521FF47C4E5FA7EFBA7" box="[151,354,1015,1041]" pageId="48" pageNumber="49" type="description">
<paragraph id="8BA236D6FFFA2521FF47C4E5FA7EFBA7" blockId="48.[151,607,978,1041]" box="[151,354,1015,1041]" pageId="48" pageNumber="49">
(
<figureCitation id="13262A53FFFA2521FF4FC4E5FBE2FBA7" box="[159,254,1015,1041]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Figs. 22</figureCitation>
,
<figureCitation id="13262A53FFFA2521FED9C4E5FA45FBA7" box="[265,345,1015,1041]" captionStart="FIGURE 32" captionStartId="60.[152,255,1984,2008]" captionTargetBox="[189,1399,181,1954]" captionTargetId="figure-31@60.[189,1399,181,1960]" captionTargetPageId="60" captionText="FIGURE 32. AB. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.7639683" httpUri="https://zenodo.org/record/7639683/files/figure.png" pageId="48" pageNumber="49">32 A,B</figureCitation>
)
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<paragraph id="8BA236D6FFFA2521FF47C329F8D1FBE5" blockId="48.[151,973,1083,1107]" box="[151,973,1083,1107]" pageId="48" pageNumber="49">
<treatmentCitationGroup id="AB0D11F8FFFA2521FF47C329F8D1FBE5" box="[151,973,1083,1107]" pageId="48" pageNumber="49">
<taxonomicName id="4C1D4D55FFFA2521FF47C329FA25FBE5" authorityName="Dallas" authorityYear="1851" box="[151,313,1083,1107]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="suturata">
<emphasis id="B969EAC4FFFA2521FF47C329FA25FBE5" box="[151,313,1083,1107]" italics="true" pageId="48" pageNumber="49">Edessa suturata</emphasis>
</taxonomicName>
var. (b)
<treatmentCitation id="0ABC10C7FFFA2521FE40C329F9ECFBE5" author="Breddin, G." box="[400,752,1083,1107]" page="136" pageId="48" pageNumber="49" year="1904">
<taxonomicName id="4C1D4D55FFFA2521FE40C329F9ECFBE5" authority="Breddin 1904: 135 - 136" authorityName="Breddin" authorityPageNumber="135 - 136" authorityYear="1904" box="[400,752,1083,1107]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="subandina">
subandina
<bibRefCitation id="EF8C4B27FFFA2521FE2DC329F9ECFBE5" author="Breddin, G." box="[509,752,1083,1107]" pageId="48" pageNumber="49" pagination="135 - 154" refId="ref36797" refString="Breddin, G. (1904) Beitrage zur Systematik der Rhynchoten. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 14, 135 - 154. https: // doi. org / 10.5962 / bhl. part. 8958" type="journal article" year="1904">Breddin 1904: 135136</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="0ABC10C7FFFA2521FD2BC329F8D5FBE5" author="Kirkaldy, G. W." box="[763,969,1083,1107]" page="165" pageId="48" pageNumber="49" year="1909">
<bibRefCitation id="EF8C4B27FFFA2521FD2BC329F8D5FBE5" author="Kirkaldy, G. W." box="[763,969,1083,1107]" pageId="48" pageNumber="49" refId="ref37776" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) Vol. I: Cimicidae. Felix L. Dames, Berlin, xl + 392 pp." type="book" year="1909">Kirkaldy, 1909: 165</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BA236D6FFFA2521FF47C36EF822FB0C" blockId="48.[151,1437,1148,1210]" pageId="48" pageNumber="49">
<emphasis id="B969EAC4FFFA2521FF47C36EFA76FB20" box="[151,362,1148,1174]" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="4C1D4D55FFFA2521FF47C36EFA7AFB20" authorityName="Breddin" authorityYear="1904" box="[151,358,1148,1174]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="subandina">Edessa subandina</taxonomicName>
.
</emphasis>
<materialsCitation id="3B753C8BFFFA2521FEA1C36FF822FB0C" ID-GBIF-Occurrence="4097197356" collectionCode="SDEI" collectorName="Bison Er." country="Venezuela" location="Venezuela" pageId="48" pageNumber="49" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54A68874FFFA2521FEA1C36FFAFDFB21" box="[369,481,1149,1175]" pageId="48" pageNumber="49" type="lectotype">Lectotype</typeStatus>
female. Esmeralda, Venez [
<collectingCountry id="F30A7646FFFA2521FCC9C36EF88DFB20" box="[793,913,1148,1174]" name="Venezuela" pageId="48" pageNumber="49">Venezuela</collectingCountry>
]/
<collectorName id="26E85300FFFA2521FC78C36EFF0DFB20" box="[936,1041,1148,1174]" pageId="48" pageNumber="49">Bison Er.</collectorName>
/
<taxonomicName id="4C1D4D55FFFA2521FBF0C36EFE12FB20" ID-CoL="632LJ" authority="Suturata Dall" authorityName="Suturata Dall" box="[1056,1294,1148,1174]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC4FFFA2521FBF0C36EFFCBFB20" box="[1056,1239,1148,1174]" italics="true" pageId="48" pageNumber="49">Edessa Suturata</emphasis>
Dall
</taxonomicName>
/ DEI Hemimetabola #100292 (
<collectionCode id="ED0CAE13FFFA2521FEA6C3B2FAA6FB0C" box="[374,442,1184,1210]" pageId="48" pageNumber="49">SDEI</collectionCode>
). Examined by photos. (
<figureCitation id="13262A53FFFA2521FD19C3B2F82AFB0C" box="[713,822,1184,1210]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 D</figureCitation>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFFA2521FF47C3FAFE80F818" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="8BA236D6FFFA2521FF47C3FAFF28F8F4" blockId="48.[151,1437,1256,2039]" pageId="48" pageNumber="49">
<emphasis id="B969EAC4FFFA2521FF47C3FAFA0EFAB4" bold="true" box="[151,274,1256,1282]" pageId="48" pageNumber="49">Diagnosis.</emphasis>
Dorsal surface of the body olive green (
<figureCitation id="13262A53FFFA2521FD1FC3FAF820FAB4" box="[719,828,1256,1282]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 A</figureCitation>
). Ventral surface brown with transversal black lines on thorax and abdomen (
<figureCitation id="13262A53FFFA2521FE5EC21EFAE1FA90" box="[398,509,1292,1318]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 B</figureCitation>
). Antennae brown (
<figureCitation id="13262A53FFFA2521FD0DC21EF851FA90" box="[733,845,1292,1318]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 A</figureCitation>
). Pronotum with punctures brown (
<figureCitation id="13262A53FFFA2521FB0FC21EFE53FA90" box="[1247,1359,1292,1318]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 A</figureCitation>
); anterolateral margin and cicatrices with brown punctures (
<figureCitation id="13262A53FFFA2521FD25C222F87EFAFC" box="[757,866,1328,1354]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 A</figureCitation>
). Humeral angles short (1,3 times wider than long) (
<figureCitation id="13262A53FFFA2521FF4FC246FA12FAD8" box="[159,270,1364,1390]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 A</figureCitation>
), apices brown (
<figureCitation id="13262A53FFFA2521FE19C246F925FAD8" box="[457,569,1364,1390]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 A</figureCitation>
); humeral angle brown spot expanding into the pronotum in dorsal view and restricted to the angle in ventral view (
<figureCitation id="13262A53FFFA2521FDEFC26AF9D8FA24" box="[575,708,1400,1426]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 A,B</figureCitation>
). Scutellum with punctures brown (
<figureCitation id="13262A53FFFA2521FB9EC26AFFA6FA24" box="[1102,1210,1400,1426]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 A</figureCitation>
); apex not reach end of corium (
<figureCitation id="13262A53FFFA2521FEC3C28EFA63FA00" box="[275,383,1436,1462]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 A</figureCitation>
). Corium with all veins concolorous with surface (
<figureCitation id="13262A53FFFA2521FC60C28EFF00FA00" box="[944,1052,1436,1462]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 A</figureCitation>
). Posterolateral angles of connexivum with apices black (
<figureCitation id="13262A53FFFA2521FE4EC2D2F917FA6C" box="[414,523,1472,1498]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 A</figureCitation>
); connexival segments with concavities almost entirely covered by subrectangular black spots and separated by a large yellow median spot (
<figureCitation id="13262A53FFFA2521FCC6C2F6F89FFA48" box="[790,899,1508,1534]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 A</figureCitation>
); spots extending ventrally, subrectangular (
<figureCitation id="13262A53FFFA2521FABFC2F6FBD2F994" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 B</figureCitation>
). Ventral surface. Thorax with black stripes (
<figureCitation id="13262A53FFFA2521FD11C11AF837F994" box="[705,811,1544,1570]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 B</figureCitation>
); dark stripe of the propleuron covering 2/3 of the width of the sclerite (
<figureCitation id="13262A53FFFA2521FEECC13EFABBF9F0" box="[316,423,1580,1606]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 B</figureCitation>
). Evaporatorium concolorous with thorax (
<figureCitation id="13262A53FFFA2521FC57C13EF8EDF9F0" box="[903,1009,1580,1606]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 B</figureCitation>
). Metasternal process (
<figureCitation id="13262A53FFFA2521FB22C13EFE47F9F0" box="[1266,1371,1580,1606]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 B</figureCitation>
) with anterior apex straight and laterally little expanded, margin acuminated; anterior face somewhat excavated; anterior bifurcation receiving fourth rostral segment. Legs brown (
<figureCitation id="13262A53FFFA2521FCF1C166F891F938" box="[801,909,1652,1678]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 B</figureCitation>
). Abdomen with spine of third segment acuminated (
<figureCitation id="13262A53FFFA2521FF32C18AFA51F904" box="[226,333,1688,1714]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 B</figureCitation>
). Intersegmental areas solid black reaching ventral spots of connexivum (
<figureCitation id="13262A53FFFA2521FBADC18AFFF4F904" box="[1149,1256,1688,1714]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 B</figureCitation>
). Pseudosutures solid black, continuous, solid and regular border (
<figureCitation id="13262A53FFFA2521FD1AC1AEF825F960" box="[714,825,1724,1750]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 B</figureCitation>
). Median longitudinal black band incomplete (
<figureCitation id="13262A53FFFA2521FA9BC1AEFBB0F94C" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 B</figureCitation>
). Trichobotria parallel to the spiracle. Posterolateral angles of segment VII not reaching level of the apices of laterotergites IX in females (
<figureCitation id="13262A53FFFA2521FE6CC016F935F8A8" box="[444,553,1796,1822]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 C</figureCitation>
). Female genitalia, valvifers VIII with a brown band near the tumescence (
<figureCitation id="13262A53FFFA2521FABFC016FBCCF8F4" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 C</figureCitation>
). Laterotergites VIII with two dark spots on outer lateral margins (
<figureCitation id="13262A53FFFA2521FC69C03AFF3BF8F4" box="[953,1063,1832,1858]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 C</figureCitation>
).
</paragraph>
<paragraph id="8BA236D6FFFA2521FF17C05EFAE4F8D0" blockId="48.[151,1437,1256,2039]" box="[199,504,1868,1894]" pageId="48" pageNumber="49">
<emphasis id="B969EAC4FFFA2521FF17C05EFA67F8D0" bold="true" box="[199,379,1868,1894]" pageId="48" pageNumber="49">Male genitalia:</emphasis>
Unknown.
</paragraph>
<paragraph id="8BA236D6FFFA2521FF17C062FE80F818" blockId="48.[151,1437,1256,2039]" pageId="48" pageNumber="49">
<emphasis id="B969EAC4FFFA2521FF17C062FA88F83C" bold="true" box="[199,404,1904,1930]" pageId="48" pageNumber="49">Female genitalia:</emphasis>
Valvifers VIII with dark punctures; inner margins contiguous and not divergent; distal margin brown and arched. Laterotergites IX with apices acuminate passing the sclerite uniting laterotergites VIII (
<figureCitation id="13262A53FFFA2521FAF5C086FE93F818" box="[1317,1423,1940,1966]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Fig. 22 C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFFA2520FF17C0AAFFF1FF4F" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="discussion">
<paragraph id="8BA236D6FFFA2520FF17C0AAFFF1FF4F" blockId="48.[151,1437,1256,2039]" lastBlockId="49.[151,1436,151,285]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<emphasis id="B969EAC4FFFA2521FF17C0AAFA53F864" bold="true" box="[199,335,1976,2002]" pageId="48" pageNumber="49">Comments.</emphasis>
<taxonomicName id="4C1D4D55FFFA2521FE86C0AAF94EF864" authorityName="Breddin" authorityYear="1904" box="[342,594,1976,2002]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="subandina" subGenus="Edessa">
<emphasis id="B969EAC4FFFA2521FE86C0AAF94EF864" box="[342,594,1976,2002]" italics="true" pageId="48" pageNumber="49">Edessa (E.) subandina</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C1D4D55FFFA2521FD31C0ABF868F864" authorityName="Walker" authorityYear="1868" box="[737,884,1976,2002]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="electa" subGenus="Edessa">
<emphasis id="B969EAC4FFFA2521FD31C0ABF868F864" box="[737,884,1976,2002]" italics="true" pageId="48" pageNumber="49">E. (E.) electa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D55FFFA2521FC52C0ABFF51F864" authorityName="Stal" authorityYear="1862" box="[898,1101,1976,2002]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="nigricornis" subGenus="Edessa">
<emphasis id="B969EAC4FFFA2521FC52C0ABFF51F864" box="[898,1101,1976,2002]" italics="true" pageId="48" pageNumber="49">E. (E.) nigricornis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D55FFFA2521FB53C0ABFE6EF864" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[1155,1394,1976,2002]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="nigroangulata" subGenus="Edessa">
<emphasis id="B969EAC4FFFA2521FB53C0ABFE6EF864" box="[1155,1394,1976,2002]" italics="true" pageId="48" pageNumber="49">E. (E.) nigroangulata</emphasis>
</taxonomicName>
because of the shape and size of the humeral angle (
<figureCitation id="13262A53FFFA2521FD6EC0CEF829F840" box="[702,821,2012,2038]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="48" pageNumber="49">Figs. 22 A</figureCitation>
,
<figureCitation id="13262A53FFFA2521FC90C0CEF86BF840" box="[832,887,2012,2039]" captionStart="FIGURE 25" captionStartId="53.[152,255,1943,1967]" captionTargetBox="[264,1323,190,1907]" captionTargetId="figure-19@53.[250,1338,181,1919]" captionTargetPageId="53" captionText="FIGURE 25. AF. Dorsal and ventral surface: AB, Edessa (E.) cylindricornis Stål, 1872; CD, Edessa (E.) electa Walker, 1868; EF, Edessa (E.) fallenii Stål, 1872. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639657" httpUri="https://zenodo.org/record/7639657/files/figure.png" pageId="48" pageNumber="49">25 C</figureCitation>
,
<figureCitation id="13262A53FFFA2521FC52C0CEF8A7F840" box="[898,955,2012,2038]" captionStart="FIGURE 28" captionStartId="56.[152,255,1943,1967]" captionTargetBox="[272,1317,184,1915]" captionTargetId="figure-31@56.[256,1332,181,1919]" captionTargetPageId="56" captionText="FIGURE 28. AF. Dorsal and ventral surface: AB, Edessa (E.) nigriclava Walker, 1868; CD, Edessa (E.) nigricornis Stål, 1862; EF, Edessa (E.) nigroangulata Fernandes &amp; Silva, 2015. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639667" httpUri="https://zenodo.org/record/7639667/files/figure.png" pageId="48" pageNumber="49">28 C</figureCitation>
). However,
<taxonomicName id="4C1D4D55FFFA2521FB91C0CFFE1AF840" authorityName="Breddin" authorityYear="1904" box="[1089,1286,2012,2038]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="subandina" subGenus="Edessa">
<emphasis id="B969EAC4FFFA2521FB91C0CFFE1AF840" box="[1089,1286,2012,2038]" italics="true" pageId="48" pageNumber="49">E. (E.) subandina</emphasis>
</taxonomicName>
can be easily separated from the others by the large spots on connexival segments (
<figureCitation id="13262A53FFFB2520FC76C785FF08FF07" box="[934,1044,151,177]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="49" pageNumber="50">Fig. 22 A</figureCitation>
), complete black stripes on ventral side of abdomen (
<figureCitation id="13262A53FFFB2520FEB2C7A9FACCFF63" box="[354,464,187,213]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="49" pageNumber="50">Fig. 22 B</figureCitation>
), and a small stripe on valvifers VIII close to distal margin (
<figureCitation id="13262A53FFFB2520FBACC7A9FFF7FF63" box="[1148,1259,187,213]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. AD. Edessa (E.) subandina Breddin, 1904: AB, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="49" pageNumber="50">Fig. 22 C</figureCitation>
). The specimen was examined by photos of the
<typeStatus id="54A68874FFFB2520FE2AC7CDF97DFF4F" box="[506,609,223,249]" pageId="49" pageNumber="50" type="lectotype">lectotype</typeStatus>
female. The photos were taken by Stephan Blank (SDEI).
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFFB2520FF17C610F824FEAB" box="[199,824,258,285]" pageId="49" pageNumber="50" type="distribution">
<paragraph id="8BA236D6FFFB2520FF17C610F824FEAB" blockId="49.[151,1436,151,285]" box="[199,824,258,285]" pageId="49" pageNumber="50">
<emphasis id="B969EAC4FFFB2520FF17C610FA45FEAA" bold="true" box="[199,345,258,284]" pageId="49" pageNumber="50">Distribution</emphasis>
(
<figureCitation id="13262A53FFFB2520FEB9C611FAF1FEAB" box="[361,493,259,285]" captionStart="FIGURE 32" captionStartId="60.[152,255,1984,2008]" captionTargetBox="[189,1399,181,1954]" captionTargetId="figure-31@60.[189,1399,181,1960]" captionTargetPageId="60" captionText="FIGURE 32. AB. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.7639683" httpUri="https://zenodo.org/record/7639683/files/figure.png" pageId="49" pageNumber="50">Fig. 32 A,B</figureCitation>
):
<collectingCountry id="F30A7646FFFB2520FDD6C611F9ADFEAB" box="[518,689,259,285]" name="Venezuela" pageId="49" pageNumber="50">VENEZUELA</collectingCountry>
:
<collectingRegion id="49D9F834FFFB2520FD6AC611F828FEAB" box="[698,820,259,285]" country="Venezuela" name="Amazonas" pageId="49" pageNumber="50">Amazonas</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>