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<document id="5BC874CFDF041C329AAA97C116789A68" ID-CLB-Dataset="256481" ID-DOI="10.11646/zootaxa.5240.1.1" ID-GBIF-Dataset="4d650293-3f9a-4bbe-9530-204a3b09ef8b" ID-ISSN="1175-5326" ID-Zenodo-Dep="7639547" ID-ZooBank="2FE467C1-EAC2-4E90-B673-CCE2CCA93C1D" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1676380923589" checkinUser="plazi" docAuthor="Mendonça, Maria Thayane Da Silva, Silva, Valéria Juliete Da &amp; Fernandes, José Antônio Marin" docDate="2023" docId="03B487C0FFDD2508FF47C228FE1FFABB" docLanguage="en" docName="zootaxa.5240.1.1.pdf" docOrigin="Zootaxa 5240 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5240.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Edessa (Edessa) helicoceras Breddin 1905" docType="treatment" docVersion="9" lastPageNumber="26" masterDocId="FF8DFFB8FFCA2511FFD0C712FB1CFFB6" masterDocTitle="Diagnose of the nominal subgenus of Edessa and description of the E. sexdens group (Heteroptera, Pentatomidae, Edessinae)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="24" updateTime="1699475138964" updateUser="ExternalLinkService">
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<mods:title id="5A497A8ED9F1094D3B8AC26EBCB0F4FA">Diagnose of the nominal subgenus of Edessa and description of the E. sexdens group (Heteroptera, Pentatomidae, Edessinae)</mods:title>
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<mods:namePart id="273949B4D450B643EF5ACC126A3F480A">Mendonça, Maria Thayane Da Silva</mods:namePart>
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<mods:namePart id="B96C40420508FF7C9E18E7073437D8DB">Silva, Valéria Juliete Da</mods:namePart>
<mods:nameIdentifier id="B30EC3D022950E087D8F1E21F1AB3036" type="ORCID">0000-0002-1845-5228</mods:nameIdentifier>
<mods:nameIdentifier id="08D55806CB4424550242EDD16F4CF821" type="email">valeriajuliete@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="1981F9FE9C56F2D3F1BB1F7B6C8568AF">Fernandes, José Antônio Marin</mods:namePart>
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<mods:title id="E527B03F2A1B441CECEF063B785AE9A1">Zootaxa</mods:title>
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<mods:part id="229C159CF289AB9620F89C3C9D62D135">
<mods:date id="7CFD1FC89031FFC101979982E360AA49">2023</mods:date>
<mods:detail id="04BF8482248281BEADD522E42BD8B196" type="pubDate">
<mods:number id="6F1C29AB43BFA90F5F3A32C686BF83EE">2023-02-14</mods:number>
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<mods:detail id="FA3FC9EDE545C9B62DAFE91664891791" type="volume">
<mods:number id="673B9D835D487AA7C3E903C30EBE554A">5240</mods:number>
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<mods:number id="8558FEE6A7BAD1926C57AD6842DF8D44">1</mods:number>
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<mods:classification id="B3E7B57CDD1353DE7360D48394DB5C6A">journal article</mods:classification>
<mods:identifier id="DFDB1D7372DCD28675120BC7BC17DE29" type="CLB-Dataset">256481</mods:identifier>
<mods:identifier id="4D3B48E0178D77F7D6060A566219F70A" type="DOI">10.11646/zootaxa.5240.1.1</mods:identifier>
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<treatment id="03B487C0FFDD2508FF47C228FE1FFABB" ID-DOI="http://doi.org/10.5281/zenodo.7639887" ID-GBIF-Taxon="209578832" ID-Zenodo-Dep="7639887" LSID="urn:lsid:plazi:treatment:03B487C0FFDD2508FF47C228FE1FFABB" httpUri="http://treatment.plazi.org/id/03B487C0FFDD2508FF47C228FE1FFABB" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<subSubSection id="C307655DFFDD2506FF47C228F975FAE2" box="[151,617,1338,1365]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BA236D6FFDD2506FF47C228F975FAE2" blockId="23.[151,617,1338,1401]" box="[151,617,1338,1365]" pageId="23" pageNumber="24">
<heading id="D0EA81BAFFDD2506FF47C228F975FAE2" bold="true" box="[151,617,1338,1365]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="4">
<taxonomicName id="4C1D4D55FFDD2506FF47C228F975FAE2" authority="Breddin, 1905" authorityName="Breddin" authorityYear="1905" box="[151,617,1338,1365]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="helicoceras" subGenus="Edessa">
<emphasis id="B969EAC4FFDD2506FF47C228F975FAE2" bold="true" box="[151,617,1338,1365]" pageId="23" pageNumber="24">
<emphasis id="B969EAC4FFDD2506FF47C228FAAEFAE3" bold="true" box="[151,434,1338,1365]" italics="true" pageId="23" pageNumber="24">Edessa (E.) helicoceras</emphasis>
<bibRefCitation id="EF8C4B27FFDD2506FE69C228F975FAE2" author="Breddin, G." box="[441,617,1338,1364]" pageId="23" pageNumber="24" pagination="26 - 27" refId="ref36838" refString="Breddin, G. (1905) Neue Rhynchoten (Vorlaufige Diagnosen). Societas entomologica, 20 (4), 26 - 27." type="journal article" year="1905">Breddin, 1905</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFDD2506FF47C24DFA85FACF" box="[151,409,1375,1401]" pageId="23" pageNumber="24" type="description">
<paragraph id="8BA236D6FFDD2506FF47C24DFA85FACF" blockId="23.[151,617,1338,1401]" box="[151,409,1375,1401]" pageId="23" pageNumber="24">
(
<figureCitation id="13262A53FFDD2506FF4FC24DFBF2FACF" box="[159,238,1375,1401]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="23" pageNumber="24">Figs. 8</figureCitation>
,
<figureCitation id="13262A53FFDD2506FF2BC24DFA52FACF" box="[251,334,1375,1401]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="23" pageNumber="24">26 C,D</figureCitation>
,
<figureCitation id="13262A53FFDD2506FE8AC24DFA8DFACF" box="[346,401,1375,1401]" captionStart="FIGURE 31" captionStartId="59.[152,255,1448,1472]" captionTargetBox="[164,1423,181,1414]" captionTargetId="figure-19@59.[164,1423,181,1425]" captionTargetPageId="59" captionText="FIGURE 31. A. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.7639677" httpUri="https://zenodo.org/record/7639677/files/figure.png" pageId="23" pageNumber="24">31 A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFDD2506FF47C2B1F855F9FD" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="8BA236D6FFDD2506FF47C2B1F9EBFA0D" blockId="23.[151,759,1443,1467]" box="[151,759,1443,1467]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="AB0D11F8FFDD2506FF47C2B1F9EBFA0D" box="[151,759,1443,1467]" pageId="23" pageNumber="24">
<treatmentCitation id="0ABC10C7FFDD2506FF47C2B1F907FA0D" author="Breddin, G." box="[151,539,1443,1467]" page="26" pageId="23" pageNumber="24" year="1905">
<taxonomicName id="4C1D4D55FFDD2506FF47C2B1F907FA0D" authority="Breddin, 1905: 26" authorityName="Breddin" authorityPageNumber="26" authorityYear="1905" box="[151,539,1443,1467]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="helicoceras">
<emphasis id="B969EAC4FFDD2506FF47C2B1FA46FA0D" box="[151,346,1443,1467]" italics="true" pageId="23" pageNumber="24">Edessa helicoceras</emphasis>
<bibRefCitation id="EF8C4B27FFDD2506FEB0C2B1F907FA0D" author="Breddin, G." box="[352,539,1443,1467]" pageId="23" pageNumber="24" pagination="26 - 27" refId="ref36838" refString="Breddin, G. (1905) Neue Rhynchoten (Vorlaufige Diagnosen). Societas entomologica, 20 (4), 26 - 27." type="journal article" year="1905">Breddin, 1905: 26</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0ABC10C7FFDD2506FDF5C2B1F9E8FA0D" author="Kirkaldy, G. W." box="[549,756,1443,1467]" page="158" pageId="23" pageNumber="24" year="1909">
<bibRefCitation id="EF8C4B27FFDD2506FDF5C2B1F9E8FA0D" author="Kirkaldy, G. W." box="[549,756,1443,1467]" pageId="23" pageNumber="24" refId="ref37776" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) Vol. I: Cimicidae. Felix L. Dames, Berlin, xl + 392 pp." type="book" year="1909">Kirkaldy, 1909: 158</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA236D6FFDD2506FF47C2FBFA57F991" blockId="23.[151,1436,1513,1611]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="AB0D11F8FFDD2506FF47C2FBFA57F991" pageId="23" pageNumber="24">
<emphasis id="B969EAC4FFDD2506FF47C2FBFA6BF9B5" box="[151,375,1513,1539]" italics="true" pageId="23" pageNumber="24">
<taxonomicName id="4C1D4D55FFDD2506FF47C2FBFA6FF9B5" authorityName="Breddin" authorityYear="1905" box="[151,371,1513,1539]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="helicoceras">Edessa helicoceras</taxonomicName>
.
</emphasis>
<materialsCitation id="3B753C8BFFDD2506FEADC2F8FA57F991" ID-GBIF-Occurrence="4097197324" collectionCode="SDEI" collectorName="Schirmer" country="Venezuela" location="Venezuela" pageId="23" pageNumber="24" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54A68874FFDD2506FEADC2F8FAF1F9B2" box="[381,493,1514,1540]" pageId="23" pageNumber="24" type="lectotype">Lectotype</typeStatus>
male. Esmeralda,
<collectingCountry id="F30A7646FFDD2506FD6EC2FBF828F9B5" box="[702,820,1513,1539]" name="Venezuela" pageId="23" pageNumber="24">Venezuela</collectingCountry>
,
<collectorName id="26E85300FFDD2506FCEFC2FBF8BAF9B5" box="[831,934,1513,1539]" pageId="23" pageNumber="24">Schirmer</collectorName>
leg. (
<collectionCode id="ED0CAE13FFDD2506FC36C2FBFF36F9B5" box="[998,1066,1513,1539]" pageId="23" pageNumber="24">SDEI</collectionCode>
). Examined. Designated by
<treatmentCitation id="0ABC10C7FFDD2506FAB6C2FBFA5BF991" author="Gaedike, H." page="87" pageId="23" pageNumber="24" year="1971">
<bibRefCitation id="EF8C4B27FFDD2506FAB6C2FBFA5BF991" author="Gaedike, H." pageId="23" pageNumber="24" pagination="79 - 159" refId="ref37743" refString="Gaedike, H. (1971) Katalog der in Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen - V. Heteroptera. Beitrage zur Entomologie, 21, 79 - 159." type="journal article" year="1971">Gaedike (1971: 87)</bibRefCitation>
</treatmentCitation>
.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA236D6FFDD2506FF47C123F855F9FD" blockId="23.[151,1436,1513,1611]" box="[151,841,1585,1611]" pageId="23" pageNumber="24">
<emphasis id="B969EAC4FFDD2506FF47C123FA6AF9FD" box="[151,374,1585,1611]" italics="true" pageId="23" pageNumber="24">
<taxonomicName id="4C1D4D55FFDD2506FF47C123FA6FF9FD" authorityName="Breddin" authorityYear="1905" box="[151,371,1585,1611]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="helicoceras">Edessa helicoceras</taxonomicName>
.
</emphasis>
<typeStatus id="54A68874FFDD2506FEADC123F909F9FD" box="[381,533,1585,1611]" pageId="23" pageNumber="24" type="paralectotype">Paralectotype</typeStatus>
female. Same data (SDEI).
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFDD2506FF47C16BF950F96D" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BA236D6FFDD2506FF47C16BF950F96D" blockId="23.[151,1437,1656,2008]" pageId="23" pageNumber="24">
<emphasis id="B969EAC4FFDD2506FF47C16BFA98F925" bold="true" box="[151,388,1657,1683]" pageId="23" pageNumber="24">Material examined.</emphasis>
<materialsCitation id="3B753C8BFFDD2506FE5EC16BF855F925" ID-GBIF-Occurrence="4097197340" box="[398,841,1657,1686]" collectionCode="UFPA" country="Colombia" location="COLOMBIA" pageId="23" pageNumber="24" specimenCount="1">
<emphasis id="B969EAC4FFDD2506FE5EC16BF92AF925" bold="true" box="[398,566,1657,1683]" pageId="23" pageNumber="24">
<collectingCountry id="F30A7646FFDD2506FE5EC16BF92DF925" box="[398,561,1657,1683]" name="Colombia" pageId="23" pageNumber="24">COLOMBIA</collectingCountry>
:
</emphasis>
1♁, Colombia (
<collectionCode id="ED0CAE13FFDD2506FD25C16BF85DF925" box="[757,833,1657,1683]" pageId="23" pageNumber="24">UFPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B753C8BFFDD2506FC87C16AF9BBF901" ID-GBIF-Occurrence="4097197322" collectingDate="1925-06-10" collectionCode="USNM" country="Colombia" location="Bogota" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Bogota">
<emphasis id="B969EAC4FFDD2506FC87C16AF8A8F925" bold="true" box="[855,948,1656,1683]" pageId="23" pageNumber="24">
<collectingRegion id="49D9F834FFDD2506FC87C16AF8B3F925" box="[855,943,1656,1683]" country="Colombia" name="Bogota" pageId="23" pageNumber="24">Bogotá</collectingRegion>
:
</emphasis>
<specimenCount id="9D1BFD5FFFDD2506FC6FC16BF8FEF925" box="[959,994,1657,1683]" pageId="23" pageNumber="24" type="female">1♀</specimenCount>
,
<location id="8EC2600DFFDD2506FC23C16BFF5BF925" LSID="urn:lsid:plazi:treatment:03B487C0FFDD2508FF47C228FE1FFABB:8EC2600DFFDD2506FC23C16BFF5BF925" box="[1011,1095,1657,1683]" country="Colombia" name="Bogota" pageId="23" pageNumber="24" stateProvince="Bogota">Bogotá</location>
,
<date id="FFA31016FFDD2506FB85C16BFFC0F922" box="[1109,1244,1657,1684]" pageId="23" pageNumber="24" value="1925-06-10">
<collectingDate id="EFE7E9FEFFDD2506FB85C16BFFC0F922" box="[1109,1244,1657,1684]" pageId="23" pageNumber="24" value="1925-06-10">10-VI-1925</collectingDate>
</date>
,
<collectorName id="26E85300FFDD2506FB3BC168FE80F925" box="[1259,1436,1657,1684]" pageId="23" pageNumber="24">M. H. Nicefero</collectorName>
(
<collectorName id="26E85300FFDD2506FF4FC18FFA19F90E" box="[159,261,1693,1720]" pageId="23" pageNumber="24">Comp W</collectorName>
<typeStatus id="54A68874FFDD2506FEDCC18FFA20F901" box="[268,316,1693,1719]" pageId="23" pageNumber="24">type</typeStatus>
—
<collectorName id="26E85300FFDD2506FE89C18FFA86F901" box="[345,410,1693,1719]" pageId="23" pageNumber="24">Seiler</collectorName>
&amp;
<collectorName id="26E85300FFDD2506FE6DC18CFAECF90E" box="[445,496,1694,1720]" pageId="23" pageNumber="24">Lutz</collectorName>
1957) (
<collectionCode id="ED0CAE13FFDD2506FD97C18FF982F901" box="[583,670,1693,1719]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="24" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B753C8BFFDD2506FD61C18FF954F96D" ID-GBIF-Occurrence="4097197352" collectingDate="1982-07-18" collectionCode="UFPA" country="Ecuador" location="Tena" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Napo">
<emphasis id="B969EAC4FFDD2506FD61C18FF889F901" bold="true" box="[689,917,1693,1719]" pageId="23" pageNumber="24">
<collectingCountry id="F30A7646FFDD2506FD61C18FF85CF901" box="[689,832,1693,1719]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
,
<collectingRegion id="49D9F834FFDD2506FC9CC18FF88CF901" box="[844,912,1693,1719]" country="Ecuador" name="Napo" pageId="23" pageNumber="24">Napo</collectingRegion>
:
</emphasis>
<specimenCount id="9D1BFD5FFFDD2506FC4CC18FF8A3F901" box="[924,959,1693,1719]" pageId="23" pageNumber="24" type="female">1♀</specimenCount>
,
<location id="8EC2600DFFDD2506FC1DC18CFF1AF90E" LSID="urn:lsid:plazi:treatment:03B487C0FFDD2508FF47C228FE1FFABB:8EC2600DFFDD2506FC1DC18CFF1AF90E" box="[973,1030,1694,1720]" country="Ecuador" name="Tena" pageId="23" pageNumber="24" stateProvince="Napo">Tena</location>
,
<date id="FFA31016FFDD2506FBC0C18FFFBDF901" box="[1040,1185,1693,1719]" pageId="23" pageNumber="24" value="1982-07-18">
<collectingDate id="EFE7E9FEFFDD2506FBC0C18FFFBDF901" box="[1040,1185,1693,1719]" pageId="23" pageNumber="24" value="1982-07-18">18-VII-1982</collectingDate>
</date>
,
<collectorName id="26E85300FFDD2506FB7CC18CFE3CF90E" box="[1196,1312,1694,1720]" pageId="23" pageNumber="24">R. Hensen</collectorName>
(
<collectorName id="26E85300FFDD2506FAFFC18FFE80F90E" box="[1327,1436,1693,1720]" pageId="23" pageNumber="24">Comp. W</collectorName>
<typeStatus id="54A68874FFDD2506FF47C1D3FBD3F96D" box="[151,207,1729,1755]" pageId="23" pageNumber="24">Type</typeStatus>
—
<collectorName id="26E85300FFDD2506FF3CC1D3FA42F96D" box="[236,350,1729,1755]" pageId="23" pageNumber="24">Fernandes</collectorName>
JAM 1990) (
<collectionCode id="ED0CAE13FFDD2506FE24C1D3F95CF96D" box="[500,576,1729,1755]" pageId="23" pageNumber="24">UFPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFDD2506FF17C1F7F81BF8F1" pageId="23" pageNumber="24" type="description">
<paragraph id="8BA236D6FFDD2506FF17C1F7F81BF8F1" blockId="23.[151,1437,1656,2008]" pageId="23" pageNumber="24">
<emphasis id="B969EAC4FFDD2506FF17C1F7FA6AF949" bold="true" box="[199,374,1765,1791]" pageId="23" pageNumber="24">Measurements</emphasis>
(n= 3). Total length: 17.5–19.8; head length: 1.8–2.0; head width: 3.0–3.1; pronotum length: 3.9–4.2; pronotum width: 9.5–10.0; scutellum length: 7.9–8.8; scutellum width: 6.0–6.5; abdominal width: 8.8–9.8; length antennomers: I: 1.0–1.0; II: 1.7–1.8; III: 1.7–1.8.
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFDD2508FF17C043F899FBA7" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BA236D6FFDD2508FF17C043FA77FC8F" blockId="23.[151,1437,1656,2008]" lastBlockId="25.[151,1437,151,1293]" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<emphasis id="B969EAC4FFDD2506FF17C043FA5EF8DD" bold="true" box="[199,322,1873,1899]" pageId="23" pageNumber="24">Diagnosis.</emphasis>
Specimens large (
<quantity id="4CE59B33FFDD2506FDC0C043F9A5F8DD" box="[528,697,1873,1900]" metricMagnitude="-2" metricUnit="m" metricValue="1.8649999999999998" metricValueMax="1.98" metricValueMin="1.75" pageId="23" pageNumber="24" unit="mm" value="18.65" valueMax="19.8" valueMin="17.5">17.5–19.8 mm</quantity>
). Dorsal surface olive green (
<figureCitation id="13262A53FFDD2506FBD5C043FF6FF8DD" box="[1029,1139,1873,1899]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="23" pageNumber="24">Fig. 26 C</figureCitation>
). Ventral surface dark yellow to brown with transversal black lines on thorax and abdomen (
<figureCitation id="13262A53FFDD2506FC50C067F8F3F839" box="[896,1007,1909,1935]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="23" pageNumber="24">Fig. 26 D</figureCitation>
). Antennae reddish brown (
<figureCitation id="13262A53FFDD2506FAF2C067FE93F839" box="[1314,1423,1909,1935]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="23" pageNumber="24">Fig. 26 C</figureCitation>
). Pronotum with punctures concolorous with surface to brown (
<figureCitation id="13262A53FFDD2506FC86C08BF8D9F805" box="[854,965,1945,1971]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="23" pageNumber="24">Fig. 26 C</figureCitation>
). Humeral angles long (1,75 times longer than wide) (
<figureCitation id="13262A53FFDD2506FEF2C0AFFA88F861" box="[290,404,1981,2007]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="23" pageNumber="24">Fig. 26 C</figureCitation>
), apices black (
<figureCitation id="13262A53FFDD2506FD99C0AFF9A7F861" box="[585,699,1981,2007]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="23" pageNumber="24">Fig. 26 C</figureCitation>
); humeral angle dark spot restricted to the angle in dorsal view, but in ventral view spot expanding into the propleuron (
<figureCitation id="13262A53FFD32508FCC2C785F880FF07" box="[786,924,151,177]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 C,D</figureCitation>
). Scutellum with punctures concolorous with surface to brown (
<figureCitation id="13262A53FFD32508FEB8C7A9FAC5FF63" box="[360,473,187,213]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 C</figureCitation>
); apex not reach end of corium (
<figureCitation id="13262A53FFD32508FC81C7A9F8DEFF63" box="[849,962,187,213]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 C</figureCitation>
). Corium with all veins concolorous with surface (
<figureCitation id="13262A53FFD32508FF27C7CDFA7FFF4F" box="[247,355,223,249]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 C</figureCitation>
). Posterolateral angles of connexivum with apices brown (
<figureCitation id="13262A53FFD32508FC20C7CDFF40FF4F" box="[1008,1116,223,249]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 C</figureCitation>
); connexival segments without spot and flat (
<figureCitation id="13262A53FFD32508FEB1C611FACCFEAB" box="[353,464,259,285]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 C</figureCitation>
). Ventral surface. Thorax with black stripes (
<figureCitation id="13262A53FFD32508FC00C611FF5CFEAB" box="[976,1088,259,285]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 D</figureCitation>
); dark stripe of the propleuron connected to the dark spot of the humeral angle (
<figureCitation id="13262A53FFD32508FD10C635F82DFEF7" box="[704,817,295,321]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 D</figureCitation>
). Proepisternum with dark stripe. Evaporatorium black (
<figureCitation id="13262A53FFD32508FF4FC659FA11FED3" box="[159,269,331,357]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 D</figureCitation>
). Metasternal process (
<figureCitation id="13262A53FFD32508FDDDC659F975FED3" box="[525,617,331,357]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 G</figureCitation>
) with anterior apex straight and laterally well expanded, margin rounded; anterior face somewhat excavated; anterior bifurcation receiving fourth rostral segment; with black central spot. Legs with tibiae and tarsi darker than the rest (
<figureCitation id="13262A53FFD32508FD77C681F804FE1B" box="[679,792,403,429]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 D</figureCitation>
). Abdomen with spine of third segment acuminated (
<figureCitation id="13262A53FFD32508FABFC681FBD2FE67" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 F</figureCitation>
). Intersegmental areas covered by wide black band reaching lateral margin; band connected to the pseudosutures by black spots that also cover spiracles (
<figureCitation id="13262A53FFD32508FD4FC6C9F80CFE43" box="[671,784,475,501]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 D</figureCitation>
). Pseudosutures solid black, margin irregular (
<figureCitation id="13262A53FFD32508FACEC6C9FE8CFE43" box="[1310,1424,475,501]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 D</figureCitation>
). Median longitudinal black band complete (
<figureCitation id="13262A53FFD32508FDABC6EDF9F7FDAF" box="[635,747,511,537]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 D</figureCitation>
). Trichobotria lateral to the spiracles. Posterolateral angles of segment VII not reaching the level of apices of laterotergites IX in females (
<figureCitation id="13262A53FFD32508FC2DC531FF40FD8B" box="[1021,1116,547,573]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 F</figureCitation>
). Male genitalia, dorsal side of the pygophore with a suffused brown area occupying 1/3 of the surface (
<figureCitation id="13262A53FFD32508FC31C555FF90FDD7" box="[993,1164,583,609]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 A,B,D,F</figureCitation>
). Posterolateral angle of pygophore slightly developed (
<figureCitation id="13262A53FFD32508FE27C579F946FD33" box="[503,602,619,645]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 A</figureCitation>
). Superior process of genital cup little developed, fixed to the wall of the tenth segment, rectangular, thick, tumid, not continuing ventrally in a carina (
<figureCitation id="13262A53FFD32508FC28C59DFF73FD1F" box="[1016,1135,655,681]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 B,E</figureCitation>
). Anterior half of proctiger black (
<figureCitation id="13262A53FFD32508FF35C5A1FA88FD7B" box="[229,404,691,717]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 A,B,D,E</figureCitation>
). Ventral rim not setose (
<figureCitation id="13262A53FFD32508FD64C5A1F80AFD7B" box="[692,790,691,717]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 C</figureCitation>
). Female genitalia, valvifers VIII with a small dentiform projection on the valvifers IX; tumescence in the central region of the valvifers VIII with black spot; valvifers IX with black spot; laterotergites VIII with dark band on outer lateral margins; laterotergites IX with black spot in dug the base (
<figureCitation id="13262A53FFD32508FED1C40DFA42FC8F" box="[257,350,799,825]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 F</figureCitation>
).
</paragraph>
<caption id="DF62665EFFD22509FF48C085F8D6F865" ID-DOI="http://doi.org/10.5281/zenodo.7639581" ID-Zenodo-Dep="7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="24" pageNumber="25" startId="24.[152,255,1943,1967]" targetBox="[186,1401,193,1904]" targetPageId="24">
<paragraph id="8BA236D6FFD22509FF48C085F8D6F865" blockId="24.[151,1436,1943,2004]" pageId="24" pageNumber="25">
<emphasis id="B969EAC4FFD22509FF48C085FA05F819" bold="true" box="[152,281,1943,1967]" pageId="24" pageNumber="25">FIGURE 8.</emphasis>
A–G.
<taxonomicName id="4C1D4D55FFD22509FE8EC08AF9F7F819" authority="Breddin, 1905" authorityName="Breddin" authorityYear="1905" box="[350,747,1943,1968]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="helicoceras" subGenus="Edessa">
<emphasis id="B969EAC4FFD22509FE8EC08AFABAF806" box="[350,422,1944,1968]" italics="true" pageId="24" pageNumber="25">Edessa</emphasis>
(
<emphasis id="B969EAC4FFD22509FE65C08AFAD7F819" box="[437,459,1944,1967]" italics="true" pageId="24" pageNumber="25">E.</emphasis>
)
<emphasis id="B969EAC4FFD22509FE0AC08AF953F806" box="[474,591,1944,1968]" italics="true" pageId="24" pageNumber="25">helicoceras</emphasis>
<bibRefCitation id="EF8C4B27FFD22509FD86C08AF9F7F819" author="Breddin, G." box="[598,747,1943,1968]" pageId="24" pageNumber="25" pagination="26 - 27" refId="ref36838" refString="Breddin, G. (1905) Neue Rhynchoten (Vorlaufige Diagnosen). Societas entomologica, 20 (4), 26 - 27." type="journal article" year="1905">Breddin, 1905</bibRefCitation>
</taxonomicName>
: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="8BA236D6FFD32508FF17C451F869FC13" blockId="25.[151,1437,151,1293]" pageId="25" pageNumber="26">
<emphasis id="B969EAC4FFD32508FF17C451FA68FCEA" bold="true" box="[199,372,834,861]" pageId="25" pageNumber="26">Male genitalia</emphasis>
(
<figureCitation id="13262A53FFD32508FE56C451F91BFCEB" box="[390,519,835,861]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 A–E</figureCitation>
): Parameres with brown margin; anterior lobe round and subrectangular; dorsal and posterior lobe rounded (
<figureCitation id="13262A53FFD32508FE01C475F97EFC37" box="[465,610,871,897]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 B,D,E</figureCitation>
). Proctiger with subtriangular posterior face (
<figureCitation id="13262A53FFD32508FB8CC475FFCFFC37" box="[1116,1235,871,897]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 D,E</figureCitation>
). Ventral rim with expansions undeveloped and concolorous with surface (
<figureCitation id="13262A53FFD32508FCD9C499F874FC13" box="[777,872,907,933]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 C</figureCitation>
).
</paragraph>
<paragraph id="8BA236D6FFD32508FF17C4BDF899FBA7" blockId="25.[151,1437,151,1293]" pageId="25" pageNumber="26">
<emphasis id="B969EAC4FFD32508FF17C4BDFA89FC7E" bold="true" box="[199,405,942,969]" pageId="25" pageNumber="26">Female genitalia:</emphasis>
Valvifers VIII with dark punctures and setulose; inner margins black, contiguous and divergent; distal margin forming distal U-shaped excavation, with black band and arched. Laterotergites IX with apices acuminate passing the sclerite uniting laterotergites VIII (
<figureCitation id="13262A53FFD32508FCCBC4E5F864FBA7" box="[795,888,1015,1041]" captionStart="FIGURE 8" captionStartId="24.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1904]" captionTargetId="figure-31@24.[186,1401,179,1917]" captionTargetPageId="24" captionText="FIGURE 8. A–G. Edessa (E.) helicoceras Breddin, 1905: A pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639581" httpUri="https://zenodo.org/record/7639581/files/figure.png" pageId="25" pageNumber="26">Fig. 8 F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFD32508FF17C308FE8DFB5F" pageId="25" pageNumber="26" type="discussion">
<paragraph id="8BA236D6FFD32508FF17C308FE8DFB5F" blockId="25.[151,1437,151,1293]" pageId="25" pageNumber="26">
<emphasis id="B969EAC4FFD32508FF17C308FA53FB82" bold="true" box="[199,335,1050,1076]" pageId="25" pageNumber="26">Comments.</emphasis>
The
<typeStatus id="54A68874FFD32508FE5EC309FAA2FB83" box="[398,446,1051,1077]" pageId="25" pageNumber="26">type</typeStatus>
locality of this species, Esmeralda, was considered
<collectingCountry id="F30A7646FFD32508FBC9C309FF90FB83" box="[1049,1164,1051,1077]" name="Venezuela" pageId="25" pageNumber="26">Venezuela</collectingCountry>
by
<bibRefCitation id="EF8C4B27FFD32508FB6BC309FE76FB83" author="Breddin, G." box="[1211,1386,1050,1077]" pageId="25" pageNumber="26" pagination="26 - 27" refId="ref36838" refString="Breddin, G. (1905) Neue Rhynchoten (Vorlaufige Diagnosen). Societas entomologica, 20 (4), 26 - 27." type="journal article" year="1905">Breddin (1905)</bibRefCitation>
and
<bibRefCitation id="EF8C4B27FFD32508FF47C32DFA54FBEF" author="Gaedike, H." box="[151,328,1087,1113]" pageId="25" pageNumber="26" pagination="79 - 159" refId="ref37743" refString="Gaedike, H. (1971) Katalog der in Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen - V. Heteroptera. Beitrage zur Entomologie, 21, 79 - 159." type="journal article" year="1971">Gaedike (1971)</bibRefCitation>
, but Esmeralda was considered
<collectingCountry id="F30A7646FFD32508FD77C32DF809FBEF" box="[679,789,1087,1113]" name="Colombia" pageId="25" pageNumber="26">Colombia</collectingCountry>
by Breddin in his description of
<taxonomicName id="4C1D4D55FFD32508FBADC32DFE72FBEF" authorityName="Breddin" authorityYear="1904" box="[1149,1390,1087,1113]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="schirmeri" subGenus="Edessa">
<emphasis id="B969EAC4FFD32508FBADC32DFE72FBEF" box="[1149,1390,1087,1113]" italics="true" pageId="25" pageNumber="26">Edessa (E.) schirmeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D55FFD32508FF47C371FA43FBCB" authorityName="Breddin" authorityYear="1904" box="[151,351,1123,1149]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="subandina" subGenus="Edessa">
<emphasis id="B969EAC4FFD32508FF47C371FA43FBCB" box="[151,351,1123,1149]" italics="true" pageId="25" pageNumber="26">E. (E.) subandina</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C4B27FFD32508FEBEC371F911FBCB" author="Breddin, G." box="[366,525,1123,1149]" pageId="25" pageNumber="26" pagination="135 - 154" refId="ref36797" refString="Breddin, G. (1904) Beitrage zur Systematik der Rhynchoten. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 14, 135 - 154. https: // doi. org / 10.5962 / bhl. part. 8958" type="journal article" year="1904">Breddin 1904</bibRefCitation>
,
<bibRefCitation id="EF8C4B27FFD32508FDC8C371F9A6FBCB" author="Gaedike, H." box="[536,698,1123,1149]" pageId="25" pageNumber="26" pagination="79 - 159" refId="ref37743" refString="Gaedike, H. (1971) Katalog der in Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen - V. Heteroptera. Beitrage zur Entomologie, 21, 79 - 159." type="journal article" year="1971">Gaedike 1971</bibRefCitation>
). But in one of the
<taxonomicName id="4C1D4D55FFD32508FC45C371FF41FBCB" authorityName="Breddin" authorityYear="1904" box="[917,1117,1123,1149]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="subandina" subGenus="Edessa">
<emphasis id="B969EAC4FFD32508FC45C371FF41FBCB" box="[917,1117,1123,1149]" italics="true" pageId="25" pageNumber="26">E. (E.) subandina</emphasis>
</taxonomicName>
labels is written Esmeralda,
<collectingCountry id="F30A7646FFD32508FF47C395FA16FB17" box="[151,266,1159,1185]" name="Venezuela" pageId="25" pageNumber="26">Venezuela</collectingCountry>
(
<figureCitation id="13262A53FFD32508FEC9C395FA9BFB17" box="[281,391,1159,1185]" captionStart="FIGURE 22" captionStartId="49.[152,255,1924,1948]" captionTargetBox="[153,1426,317,1879]" captionTargetId="figure-83@49.[151,1436,306,1897]" captionTargetPageId="49" captionText="FIGURE 22. A–D. Edessa (E.) subandina Breddin, 1904: A–B, dorsal and ventral surface of the lectotype; C, genital plates; D, lectotype labels. Scale 1 mm for genitalia and 5 mm for dorsal and ventral surfaces. The photos were taken by Dr. Stephan Blank (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.7639640" httpUri="https://zenodo.org/record/7639640/files/figure.png" pageId="25" pageNumber="26">Fig. 22 D</figureCitation>
). Therefore, we were unable to correctly place this location in any of the mentioned countries. The
<taxonomicName id="4C1D4D55FFD32508FF18C3B9FA84FB73" authorityName="Breddin" authorityYear="1905" box="[200,408,1195,1221]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="helicoceras" subGenus="Edessa">
<emphasis id="B969EAC4FFD32508FF18C3B9FA84FB73" box="[200,408,1195,1221]" italics="true" pageId="25" pageNumber="26">E. (E.) helicoceras</emphasis>
</taxonomicName>
can be easily separated from the other species of the group by the humeral angles narrow and long (
<figureCitation id="13262A53FFD32508FF08C3DDFA7DFB5F" box="[216,353,1231,1257]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 C,D</figureCitation>
); connexivum without spots (
<figureCitation id="13262A53FFD32508FD7DC3DDF804FB5F" box="[685,792,1231,1257]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 C</figureCitation>
) and ventral surface with wide black stripes (
<figureCitation id="13262A53FFD32508FAC5C3DDFE99FB5F" box="[1301,1413,1231,1257]" captionStart="FIGURE 26" captionStartId="54.[152,255,1943,1967]" captionTargetBox="[264,1322,194,1904]" captionTargetId="figure-31@54.[251,1337,181,1919]" captionTargetPageId="54" captionText="FIGURE 26. A–F. Dorsal and ventral surface: A–B, Edessa (E.) gentilitia Distant, 1890; C–D, Edessa (E.) helicoceras Breddin, 1905; E–F, Edessa (E.) helix Erichson, 1848. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639659" httpUri="https://zenodo.org/record/7639659/files/figure.png" pageId="25" pageNumber="26">Fig. 26 D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFD32508FF17C3E0FE1FFABB" box="[199,1283,1266,1293]" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BA236D6FFD32508FF17C3E0FE1FFABB" blockId="25.[151,1437,151,1293]" box="[199,1283,1266,1293]" pageId="25" pageNumber="26">
<emphasis id="B969EAC4FFD32508FF17C3E0FA45FABA" bold="true" box="[199,345,1266,1292]" pageId="25" pageNumber="26">Distribution</emphasis>
(
<figureCitation id="13262A53FFD32508FEB9C3E1FAC9FABB" box="[361,469,1267,1293]" captionStart="FIGURE 31" captionStartId="59.[152,255,1448,1472]" captionTargetBox="[164,1423,181,1414]" captionTargetId="figure-19@59.[164,1423,181,1425]" captionTargetPageId="59" captionText="FIGURE 31. A. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.7639677" httpUri="https://zenodo.org/record/7639677/files/figure.png" pageId="25" pageNumber="26">Fig. 31 A</figureCitation>
):
<collectingCountry id="F30A7646FFD32508FE3CC3E1F99AFABB" box="[492,646,1267,1293]" name="Colombia" pageId="25" pageNumber="26">COLOMBIA</collectingCountry>
:
<collectingRegion id="49D9F834FFD32508FD41C3E1F9F9FABB" box="[657,741,1267,1293]" country="Colombia" name="Bogota" pageId="25" pageNumber="26">Bogotá</collectingRegion>
;
<collectingCountry id="F30A7646FFD32508FD20C3E1F887FABB" box="[752,923,1267,1293]" name="Venezuela" pageId="25" pageNumber="26">VENEZUELA</collectingCountry>
:
<collectingRegion id="49D9F834FFD32508FC73C3E1FF01FABB" box="[931,1053,1267,1293]" country="Venezuela" name="Amazonas" pageId="25" pageNumber="26">Amazonas</collectingRegion>
;
<collectingCountry id="F30A7646FFD32508FBF8C3E1FFA8FABB" box="[1064,1204,1267,1293]" name="Ecuador" pageId="25" pageNumber="26">ECUADOR</collectingCountry>
:
<collectingRegion id="49D9F834FFD32508FB6EC3E1FFE2FABB" box="[1214,1278,1267,1293]" country="Ecuador" name="Napo" pageId="25" pageNumber="26">Napo</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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