treatments-xml/data/03/B4/87/03B487A4FFA6FF996CB07D30FDC9FDDE.xml
2024-06-21 12:22:17 +02:00

266 lines
30 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="0A99C90D3F6A82849BFE8AE24453FEB9" ID-DOI="10.5281/zenodo.199004" ID-GBIF-Dataset="1967cb33-23fe-4cfa-9804-ff8420277315" ID-ISSN="1175-5326" ID-Zenodo-Dep="199004" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460320311368" checkinUser="plazi" docAuthor="Kok, Philippe J. R., Macculloch, Ross D., Lathrop, Amy, Willaert, Bert &amp; Bossuyt, Franky" docDate="2010" docId="03B487A4FFA6FF996CB07D30FDC9FDDE" docLanguage="en" docName="zt02660p032.pdf" docOrigin="Zootaxa 2660" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anomaloglossus" docType="treatment" docVersion="9" lastPageNumber="29" masterDocId="FF8DFFDCFFACFF926C27780CFF93FFF9" masterDocTitle="A new species of Anomaloglossus (Anura: Aromobatidae) from the Pakaraima Mountains of Guyana" masterLastPageNumber="32" masterPageNumber="18" pageNumber="28" updateTime="1698256450532" updateUser="plazi">
<mods:mods id="08E947A0E0FC3011ED0913171485C4C0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="04B754C51CE06199CA01925A2699D6D8">
<mods:title id="953F94887D9AB10C5A6D2DFD45FD2DC1">A new species of Anomaloglossus (Anura: Aromobatidae) from the Pakaraima Mountains of Guyana</mods:title>
</mods:titleInfo>
<mods:name id="BE7CD3F2903C31B55AD409B53F113321" type="personal">
<mods:role id="D2453CFEAAB796DE6707BC26A3EDA24F">
<mods:roleTerm id="2E51916894975F7CC0DC33FB13390B2D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="698598C09D2E688FB1566CF5108790D7">Kok, Philippe J. R.</mods:namePart>
</mods:name>
<mods:name id="B427664E5D3A19FEA77410B4250D342E" type="personal">
<mods:role id="47C95B983D8A3C3AA51F231B4EB63871">
<mods:roleTerm id="C505D48FDF3B66DC5FF2268BBAE8F67D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="46723AD5AD7BE305B8D8B8636975C15D">Macculloch, Ross D.</mods:namePart>
</mods:name>
<mods:name id="C158D5853407CE51D2B9827A9B945543" type="personal">
<mods:role id="06120B7750BA457D1DA9827EE1DCA81C">
<mods:roleTerm id="5324F5EFF9E76028216514763394D94A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F76B12281E66D157AEFFE1CEB8E06C3A">Lathrop, Amy</mods:namePart>
</mods:name>
<mods:name id="F103982E177212886709D2A2D898EFF0" type="personal">
<mods:role id="CFB37DF73FD519952EB2633FA9C8B3AC">
<mods:roleTerm id="62FE75BA86F8BBAC43814C0320322F42">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E53746D641D50A121AC43E1A2A5353DE">Willaert, Bert</mods:namePart>
</mods:name>
<mods:name id="662EB8E90EDF27841ED493BB7EB51034" type="personal">
<mods:role id="314CE8DA5D634F6AA9B036E50599F163">
<mods:roleTerm id="CC4247F10BF40A2A51B3ADC6A349F006">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F498B5479CDAD44FBA4C303FA6942741">Bossuyt, Franky</mods:namePart>
</mods:name>
<mods:typeOfResource id="B3C81A2FB2B0D051A004E63A463C69C0">text</mods:typeOfResource>
<mods:relatedItem id="3AA9B255E28D753DC442F2C3676D66EE" type="host">
<mods:titleInfo id="B657B0B76C738E81B286BC105E7A58E5">
<mods:title id="B9E742C215DF4B93161DF4F47A6E01E4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="29850B2335B090497C824869F3411637">
<mods:date id="CCD60E24FBC7310FEDE72C2441B00DE5">2010</mods:date>
<mods:detail id="CFB5B1A5220D484DBA9C4B62FF0ABFA7" type="volume">
<mods:number id="81ACE2FE7FCC5CB1395C2F672A7B14D7">2660</mods:number>
</mods:detail>
<mods:extent id="95B8AD656B7CA234F7A9475E189F9C6F" unit="page">
<mods:start id="FD3E8B9E727839FE0AB63584615EC14E">18</mods:start>
<mods:end id="3A418818B141F789C4A753E1D76EEE45">32</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AE3BCEB70236DBAC0F05B6EEF3F1F8B5">journal article</mods:classification>
<mods:identifier id="FC2DABE54BF0286B10C090E82807C0B8" type="DOI">10.5281/zenodo.199004</mods:identifier>
<mods:identifier id="0DC314C573AAAAAB931EBCBD3555141C" type="GBIF-Dataset">1967cb33-23fe-4cfa-9804-ff8420277315</mods:identifier>
<mods:identifier id="ACA4BC86DB5F9933CE6463C505D487B2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EA4B6B39DE823299E42425B47A6628AC" type="Zenodo-Dep">199004</mods:identifier>
</mods:mods>
<treatment id="03B487A4FFA6FF996CB07D30FDC9FDDE" ID-DOI="http://doi.org/10.5281/zenodo.5680005" ID-GBIF-Taxon="119402336" ID-Zenodo-Dep="5680005" LSID="urn:lsid:plazi:treatment:03B487A4FFA6FF996CB07D30FDC9FDDE" httpUri="http://treatment.plazi.org/id/03B487A4FFA6FF996CB07D30FDC9FDDE" lastPageId="11" lastPageNumber="29" pageId="10" pageNumber="28">
<subSubSection id="C3076539FFA6FF986CB07D30FD31FAAF" box="[151,674,1340,1366]" pageId="10" pageNumber="28" type="nomenclature">
<paragraph id="8BA236B2FFA6FF986CB07D30FD31FAAF" blockId="10.[151,1437,1340,2036]" box="[151,674,1340,1366]" pageId="10" pageNumber="28">
<heading id="D0EA81DEFFA6FF986CB07D30FD31FAAF" bold="true" box="[151,674,1340,1366]" fontSize="11" level="1" pageId="10" pageNumber="28" reason="1">
<emphasis id="B969EAA0FFA6FF986CB07D30FD31FAAF" bold="true" box="[151,674,1340,1366]" pageId="10" pageNumber="28">
Western Pantepui
<taxonomicName id="4C1D4D31FFA6FF986D597D30FDD6FAAF" ID-CoL="WS2" box="[382,581,1340,1366]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis id="B969EAA0FFA6FF986D597D30FDD6FAAF" bold="true" box="[382,581,1340,1366]" italics="true" pageId="10" pageNumber="28">Anomaloglossus</emphasis>
</taxonomicName>
species
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3076539FFA6FF986CE27D6EFB67F932" pageId="10" pageNumber="28" type="description">
<paragraph id="8BA236B2FFA6FF986CE27D6EFB67F932" blockId="10.[151,1437,1340,2036]" pageId="10" pageNumber="28">
Among the Western Pantepui
<taxonomicName id="4C1D4D31FFA6FF986E027D6EFD76FA82" box="[549,741,1378,1403]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis id="B969EAA0FFA6FF986E027D6EFD76FA82" box="[549,741,1378,1403]" italics="true" pageId="10" pageNumber="28">Anomaloglossus</emphasis>
</taxonomicName>
species,
<taxonomicName id="4C1D4D31FFA6FF986F767D6EFB74FA85" authority="La Marca, 1998" authorityName="La Marca" authorityYear="1998" box="[849,1255,1378,1404]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="ayarzaguenai">
<emphasis id="B969EAA0FFA6FF986F767D6EFB81FA82" box="[849,1042,1378,1403]" italics="true" pageId="10" pageNumber="28">A. ayarzaguenai</emphasis>
(
<bibRefCitation id="EF8C4B43FFA6FF9868067D6EFB4CFA85" author="La" box="[1057,1247,1378,1404]" pageId="10" pageNumber="28" refString="La Marca, E. (1998 &quot; 1996 &quot;) Ranas del genero Colostethus (Amphibia: Anura: Dendrobatidae) de la Guyana Venezolana con la descripcion de siete especies nuevas. Publicaciones de la Asociacion de Amigos de Donana, 9, 1 - 64." type="journal article" year="1998">La Marca, 1998</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C1D4D31FFA6FF9869067D6EFDEFFA58" authority="Barrio-Amoros and Brewer-Carias, 2008" authorityName="Barrio-Amoros and Brewer-Carias" authorityYear="2008" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="moffetti">
<emphasis id="B969EAA0FFA6FF9869067D6EFA0FFA82" box="[1313,1436,1378,1403]" italics="true" pageId="10" pageNumber="28">A. moffetti</emphasis>
<bibRefCitation id="EF8C4B43FFA6FF986CB07D8BFDEFFA58" author="Barrio-Amoros" box="[151,636,1415,1441]" pageId="10" pageNumber="28" refString="Barrio-Amoros, C. L. &amp; Brewer-Carias, C. (2008) Herpetological results of the 2002 expedition to Sarisarinama, a tepui in Venezuelan Guayana, with the description of five new species. Zootaxa, 1942, 1 - 68." type="journal article" year="2008">Barrio-Amorós and Brewer-Carías, 2008</bibRefCitation>
</taxonomicName>
are the geographically closest species to
<taxonomicName id="4C1D4D31FFA6FF98684F7D8BFAA0FA59" box="[1128,1331,1415,1440]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
<emphasis id="B969EAA0FFA6FF98684F7D8BFAA0FA59" box="[1128,1331,1415,1440]" italics="true" pageId="10" pageNumber="28">A. megacephalus</emphasis>
</taxonomicName>
(
<emphasis id="B969EAA0FFA6FF9869637D8BFAFBFA59" box="[1348,1384,1415,1440]" italics="true" pageId="10" pageNumber="28">ca.</emphasis>
<quantity id="4CE59B57FFA6FF9869577D8BFF2CFA3F" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="28" unit="km" value="400.0">400 km</quantity>
airline to the west).
<taxonomicName id="4C1D4D31FFA6FF986DEC7DA0FCDDFA3C" box="[459,846,1452,1477]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="ayarzaguenai">
<emphasis id="B969EAA0FFA6FF986DEC7DA0FCDDFA3C" box="[459,846,1452,1477]" italics="true" pageId="10" pageNumber="28">Anomaloglossus ayarzaguenai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D31FFA6FF986FAA7DA0FB87FA3C" box="[909,1044,1452,1477]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="moffetti">
<emphasis id="B969EAA0FFA6FF986FAA7DA0FB87FA3C" box="[909,1044,1452,1477]" italics="true" pageId="10" pageNumber="28">A. moffetti</emphasis>
</taxonomicName>
are very similar to each other, geographically close (from Cerro Jaua and Cerro Sarisariñama, respectively) and possible synonyms (Kok &amp; Barrio-Amorós unpubl. data).
<taxonomicName id="4C1D4D31FFA6FF986E257DFBFCE4F9E9" box="[514,887,1527,1552]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
<emphasis id="B969EAA0FFA6FF986E257DFBFCE4F9E9" box="[514,887,1527,1552]" italics="true" pageId="10" pageNumber="28">Anomaloglossus megacephalus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C1D4D31FFA6FF9868A37DFBFAD4F9E9" box="[1156,1351,1527,1552]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="ayarzaguenai">
<emphasis id="B969EAA0FFA6FF9868A37DFBFAD4F9E9" box="[1156,1351,1527,1552]" italics="true" pageId="10" pageNumber="28">A. ayarzaguenai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D31FFA6FF9869A47DFBFF62F9CC" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="moffetti">
<emphasis id="B969EAA0FFA6FF9869A47DFBFF62F9CC" italics="true" pageId="10" pageNumber="28">A. moffetti</emphasis>
</taxonomicName>
in having a comparatively longer snout, and a larger, somewhat more massive head [compare
<figureCitation id="13262A37FFA6FF98696C7E10FA0FF9CF" box="[1355,1436,1564,1590]" captionStart="FIGURE 1" captionStartId="3.[151,255,1948,1972]" captionTargetBox="[151,1436,1077,1940]" captionTargetId="figure@3.[151,1436,1076,1944]" captionTargetPageId="3" captionText="FIGURE 1. Anomaloglossus megacephalus sp. nov. A: Dorsolateral view of the holotype in life (IRSNB 1986, 27.3 mm SVL). B: Ventral view of the holotype in life. C: Dorsal view of the preserved holotype. D: Ventral view of the preserved holotype. Photos by PJRK." httpUri="https://zenodo.org/record/199005/files/figure.png" pageId="10" pageNumber="28">Figs. 1</figureCitation>
and
<figureCitation id="13262A37FFA6FF986CEF7E4DFF45F9A2" box="[200,214,1601,1627]" captionStart="FIGURE 2" captionStartId="4.[151,255,826,850]" captionTargetBox="[151,1436,186,805]" captionTargetId="figure@4.[151,1436,186,805]" captionTargetPageId="4" captionText="FIGURE 2. Anomaloglossus megacephalus sp. nov. A: Ventral views of left hand (left) and left foot (right) of the female holotype (IRSNB 1986, 27.3 mm SVL). Scale bars are 2 mm. B: ROM 39638, female paratype (27.9 mm SVL), from Mt. Ayanganna. Photos by PJRK (A) and AL (B)." httpUri="https://zenodo.org/record/199006/files/figure.png" pageId="10" pageNumber="28">2</figureCitation>
B with fig. 4 ac illustrating
<taxonomicName id="4C1D4D31FFA6FF986E097E4DFD3BF9A3" box="[558,680,1601,1626]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="moffetti">
<emphasis id="B969EAA0FFA6FF986E097E4DFD3BF9A3" box="[558,680,1601,1626]" italics="true" pageId="10" pageNumber="28">A. moffetti</emphasis>
</taxonomicName>
in
<bibRefCitation id="EF8C4B43FFA6FF986EEA7E4DFB3AF9A2" author="Barrio-Amoros" box="[717,1193,1601,1627]" pageId="10" pageNumber="28" refString="Barrio-Amoros, C. L. &amp; Brewer-Carias, C. (2008) Herpetological results of the 2002 expedition to Sarisarinama, a tepui in Venezuelan Guayana, with the description of five new species. Zootaxa, 1942, 1 - 68." type="journal article" year="2008">Barrio-Amorós &amp; Brewer-Carías (2008)</bibRefCitation>
], in having a distinct tympanum in life (barely distinct in
<taxonomicName id="4C1D4D31FFA6FF986E6D7E6BFC9DF979" box="[586,782,1639,1664]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="ayarzaguenai">
<emphasis id="B969EAA0FFA6FF986E6D7E6BFC9DF979" box="[586,782,1639,1664]" italics="true" pageId="10" pageNumber="28">A. ayarzaguenai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D31FFA6FF986F6D7E6BFC54F979" box="[842,967,1639,1664]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="moffetti">
<emphasis id="B969EAA0FFA6FF986F6D7E6BFC54F979" box="[842,967,1639,1664]" italics="true" pageId="10" pageNumber="28">A. moffetti</emphasis>
</taxonomicName>
), separated from eye by 4260% of its greatest length (25% or less in
<taxonomicName id="4C1D4D31FFA6FF986E257E80FD50F95C" box="[514,707,1676,1701]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="ayarzaguenai">
<emphasis id="B969EAA0FFA6FF986E257E80FD50F95C" box="[514,707,1676,1701]" italics="true" pageId="10" pageNumber="28">A. ayarzaguenai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D31FFA6FF986EDC7E80FCE6F95C" box="[763,885,1676,1701]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="moffetti">
<emphasis id="B969EAA0FFA6FF986EDC7E80FCE6F95C" box="[763,885,1676,1701]" italics="true" pageId="10" pageNumber="28">A. moffetti</emphasis>
</taxonomicName>
), and in lacking a dark area spotted with white ventrolaterally at the level of arm insertion (always present in
<taxonomicName id="4C1D4D31FFA6FF986F577EBDFBA3F933" box="[880,1072,1713,1738]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="ayarzaguenai">
<emphasis id="B969EAA0FFA6FF986F577EBDFBA3F933" box="[880,1072,1713,1738]" italics="true" pageId="10" pageNumber="28">A. ayarzaguenai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D31FFA6FF98684E7EBDFB70F933" box="[1129,1251,1713,1738]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="moffetti">
<emphasis id="B969EAA0FFA6FF98684E7EBDFB70F933" box="[1129,1251,1713,1738]" italics="true" pageId="10" pageNumber="28">A. moffetti</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3076539FFA6FF996CE27EDAFDC9FDDE" lastPageId="11" lastPageNumber="29" pageId="10" pageNumber="28" type="reference_group">
<paragraph id="8BA236B2FFA6FF986CE27EDAFE6EF899" blockId="10.[151,1437,1340,2036]" pageId="10" pageNumber="28">
<taxonomicName id="4C1D4D31FFA6FF986CE27EDAFDA6F916" ID-CoL="6775Q" box="[197,565,1750,1775]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
<emphasis id="B969EAA0FFA6FF986CE27EDAFDA6F916" box="[197,565,1750,1775]" italics="true" pageId="10" pageNumber="28">Anomaloglossus megacephalus</emphasis>
</taxonomicName>
mostly differs from
<taxonomicName id="4C1D4D31FFA6FF986F007EDAFB3DF909" ID-CoL="5V6TQ" authority="La Marca, 1998" authorityName="La Marca" authorityYear="1998" box="[807,1198,1750,1776]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="guanayensis">
<emphasis id="B969EAA0FFA6FF986F007EDAFC4AF916" box="[807,985,1750,1775]" italics="true" pageId="10" pageNumber="28">A. guanayensis</emphasis>
(
<bibRefCitation id="EF8C4B43FFA6FF986FCF7EDAFB35F909" author="La" box="[1000,1190,1750,1776]" pageId="10" pageNumber="28" refString="La Marca, E. (1998 &quot; 1996 &quot;) Ranas del genero Colostethus (Amphibia: Anura: Dendrobatidae) de la Guyana Venezolana con la descripcion de siete especies nuevas. Publicaciones de la Asociacion de Amigos de Donana, 9, 1 - 64." type="journal article" year="1998">La Marca, 1998</bibRefCitation>
)
</taxonomicName>
by (characters of
<taxonomicName id="4C1D4D31FFA6FF9869A47EDAFEBBF8ED" ID-CoL="5V6TQ" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="guanayensis">
<emphasis id="B969EAA0FFA6FF9869A47EDAFEBBF8ED" italics="true" pageId="10" pageNumber="28">A. guanayensis</emphasis>
</taxonomicName>
in parentheses) its larger size, female SVL max
<quantity id="4CE59B57FFA6FF986F447EF7FC5FF8EC" box="[867,972,1787,1813]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="10" pageNumber="28" unit="mm" value="28.3">28.3 mm</quantity>
(n=3) in
<taxonomicName id="4C1D4D31FFA6FF98681C7EF7FA97F8ED" ID-CoL="6775Q" box="[1083,1284,1787,1812]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
<emphasis id="B969EAA0FFA6FF98681C7EF7FA97F8ED" box="[1083,1284,1787,1812]" italics="true" pageId="10" pageNumber="28">A. megacephalus</emphasis>
</taxonomicName>
(
<quantity id="4CE59B57FFA6FF9869347EF7FAEDF8EC" box="[1299,1406,1787,1813]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="10" pageNumber="28" unit="mm" value="23.5">23.5 mm</quantity>
in
<taxonomicName id="4C1D4D31FFA6FF986CB07F2DFED9F8C3" ID-CoL="5V6TQ" box="[151,330,1825,1850]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="guanayensis">
<emphasis id="B969EAA0FFA6FF986CB07F2DFED9F8C3" box="[151,330,1825,1850]" italics="true" pageId="10" pageNumber="28">A. guanayensis</emphasis>
</taxonomicName>
, n=3), and in having less developed keel-like lateral folds on fingers (extensive, present pre- and postaxially on all fingers).
</paragraph>
<paragraph id="8BA236B2FFA6FF986CE27F67FC9CF80D" blockId="10.[151,1437,1340,2036]" pageId="10" pageNumber="28">
<taxonomicName id="4C1D4D31FFA6FF986CE27F67FDA6F87D" ID-CoL="6775Q" box="[197,565,1899,1924]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
<emphasis id="B969EAA0FFA6FF986CE27F67FDA6F87D" box="[197,565,1899,1924]" italics="true" pageId="10" pageNumber="28">Anomaloglossus megacephalus</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C1D4D31FFA6FF986F527F67FB5DF87C" ID-CoL="5V763" authority="La Marca, 1998" authorityName="La Marca" authorityYear="1998" box="[885,1230,1899,1925]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="parimae">
<emphasis id="B969EAA0FFA6FF986F527F67FC6BF87D" box="[885,1016,1899,1924]" italics="true" pageId="10" pageNumber="28">A. parimae</emphasis>
(
<bibRefCitation id="EF8C4B43FFA6FF9868207F67FB55F87C" author="La" box="[1031,1222,1899,1925]" pageId="10" pageNumber="28" refString="La Marca, E. (1998 &quot; 1996 &quot;) Ranas del genero Colostethus (Amphibia: Anura: Dendrobatidae) de la Guyana Venezolana con la descripcion de siete especies nuevas. Publicaciones de la Asociacion de Amigos de Donana, 9, 1 - 64." type="journal article" year="1998">La Marca, 1998</bibRefCitation>
)
</taxonomicName>
by (characters of
<taxonomicName id="4C1D4D31FFA6FF986CB07F9CFE8AF850" ID-CoL="5V763" box="[151,281,1936,1961]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="parimae">
<emphasis id="B969EAA0FFA6FF986CB07F9CFE8AF850" box="[151,281,1936,1961]" italics="true" pageId="10" pageNumber="28">A. parimae</emphasis>
</taxonomicName>
in parentheses) its larger size, female SVL max
<quantity id="4CE59B57FFA6FF986F6C7F9CFC20F853" box="[843,947,1936,1962]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="10" pageNumber="28" unit="mm" value="28.3">28.3 mm</quantity>
(n=3) in
<taxonomicName id="4C1D4D31FFA6FF9868387F9CFB74F850" ID-CoL="6775Q" box="[1055,1255,1936,1961]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
<emphasis id="B969EAA0FFA6FF9868387F9CFB74F850" box="[1055,1255,1936,1961]" italics="true" pageId="10" pageNumber="28">A. megacephalus</emphasis>
</taxonomicName>
(
<quantity id="4CE59B57FFA6FF9868D27F9CFACDF853" box="[1269,1374,1936,1962]" metricMagnitude="-2" metricUnit="m" metricValue="2.31" pageId="10" pageNumber="28" unit="mm" value="23.1">23.1 mm</quantity>
in
<taxonomicName id="4C1D4D31FFA6FF9869A47F9CFF68F836" ID-CoL="5V763" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="parimae">
<emphasis id="B969EAA0FFA6FF9869A47F9CFF68F836" italics="true" pageId="10" pageNumber="28">A. parimae</emphasis>
</taxonomicName>
, n=2), in having a short, wider than long median lingual process (slender, noticeably longer than wide), and in having distinctly more webbing on toes.
</paragraph>
<paragraph id="8BA236B2FFA7FF996CE27894FB8DFEDB" blockId="11.[151,1437,152,551]" pageId="11" pageNumber="29">
<taxonomicName id="4C1D4D31FFA7FF996CE27894FDA6FF48" ID-CoL="6775Q" box="[197,565,152,177]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="megacephalus">
<emphasis id="B969EAA0FFA7FF996CE27894FDA6FF48" box="[197,565,152,177]" italics="true" pageId="11" pageNumber="29">Anomaloglossus megacephalus</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C1D4D31FFA7FF996F537894FB0DFF4B" ID-CoL="67763" authority="Rivero, 1961" authorityName="Rivero" authorityYear="1961" box="[884,1182,152,178]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="shrevei">
<emphasis id="B969EAA0FFA7FF996F537894FC7AFF48" box="[884,1001,152,177]" italics="true" pageId="11" pageNumber="29">A. shrevei</emphasis>
(
<bibRefCitation id="EF8C4B43FFA7FF996FDE7894FB06FF4B" author="Rivero" box="[1017,1173,152,178]" pageId="11" pageNumber="29" refString="Rivero, J. A. (1961) Salientia of Venezuela. Bulletin of the Museum of Comparative Zoology, 126, 1 - 207." type="journal article" year="1961">Rivero, 1961</bibRefCitation>
)
</taxonomicName>
in having (characters of
<taxonomicName id="4C1D4D31FFA7FF996C9178B1FEB9FF2F" ID-CoL="67763" box="[182,298,189,214]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="shrevei">
<emphasis id="B969EAA0FFA7FF996C9178B1FEB9FF2F" box="[182,298,189,214]" italics="true" pageId="11" pageNumber="29">A. shrevei</emphasis>
</taxonomicName>
in parentheses) a weakly to distinctly curved, slightly tuberclelike tarsal keel, (short, straight, not tuberclelike), less developed keel-like lateral folds on fingers (extensive, present pre- and postaxially on all fingers), and dark blotches on throat and belly (throat and belly immaculate).
</paragraph>
<paragraph id="8BA236B2FFA7FF996CE27921FDC9FDDE" blockId="11.[151,1437,152,551]" pageId="11" pageNumber="29">
<taxonomicName id="4C1D4D31FFA7FF996CE27921FDA5FEBF" ID-CoL="6775Q" box="[197,566,301,326]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="megacephalus">
<emphasis id="B969EAA0FFA7FF996CE27921FDA5FEBF" box="[197,566,301,326]" italics="true" pageId="11" pageNumber="29">Anomaloglossus megacephalus</emphasis>
</taxonomicName>
mostly differs from
<taxonomicName id="4C1D4D31FFA7FF996F0B7921FC6AFEBF" ID-CoL="66V5P" box="[812,1017,301,326]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="tamacuarensis">
<emphasis id="B969EAA0FFA7FF996F0B7921FC6AFEBF" box="[812,1017,301,326]" italics="true" pageId="11" pageNumber="29">A. tamacuarensis</emphasis>
</taxonomicName>
by (characters of
<taxonomicName id="4C1D4D31FFA7FF9968F77921FA0FFEBF" ID-CoL="66V5P" box="[1232,1436,301,326]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="tamacuarensis">
<emphasis id="B969EAA0FFA7FF9968F77921FA0FFEBF" box="[1232,1436,301,326]" italics="true" pageId="11" pageNumber="29">A. tamacuarensis</emphasis>
</taxonomicName>
in parentheses) its larger size, female SVL max
<quantity id="4CE59B57FFA7FF996F2F795EFCEAFE95" box="[776,889,338,364]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="11" pageNumber="29" unit="mm" value="28.3">28.3 mm</quantity>
(n=3) in
<taxonomicName id="4C1D4D31FFA7FF996FD2795EFB43FE92" ID-CoL="6775Q" box="[1013,1232,338,363]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="megacephalus">
<emphasis id="B969EAA0FFA7FF996FD2795EFB43FE92" box="[1013,1232,338,363]" italics="true" pageId="11" pageNumber="29">A. megacephalus</emphasis>
</taxonomicName>
(25.0 mm in
<taxonomicName id="4C1D4D31FFA7FF9969A6795EFED4FE68" ID-CoL="66V5P" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="tamacuarensis">
<emphasis id="B969EAA0FFA7FF9969A6795EFED4FE68" italics="true" pageId="11" pageNumber="29">A. tamacuarensis</emphasis>
</taxonomicName>
, n=2), in having a comparatively longer snout, and a larger, somewhat more massive head [compare
<figureCitation id="13262A37FFA7FF996D2B7991FECFFE4E" box="[268,348,413,439]" captionStart="FIGURE 1" captionStartId="3.[151,255,1948,1972]" captionTargetBox="[151,1436,1077,1940]" captionTargetId="figure@3.[151,1436,1076,1944]" captionTargetPageId="3" captionText="FIGURE 1. Anomaloglossus megacephalus sp. nov. A: Dorsolateral view of the holotype in life (IRSNB 1986, 27.3 mm SVL). B: Ventral view of the holotype in life. C: Dorsal view of the preserved holotype. D: Ventral view of the preserved holotype. Photos by PJRK." httpUri="https://zenodo.org/record/199005/files/figure.png" pageId="11" pageNumber="29">Figs. 1</figureCitation>
and
<figureCitation id="13262A37FFA7FF996DB37991FE30FE4E" box="[404,419,413,439]" captionStart="FIGURE 2" captionStartId="4.[151,255,826,850]" captionTargetBox="[151,1436,186,805]" captionTargetId="figure@4.[151,1436,186,805]" captionTargetPageId="4" captionText="FIGURE 2. Anomaloglossus megacephalus sp. nov. A: Ventral views of left hand (left) and left foot (right) of the female holotype (IRSNB 1986, 27.3 mm SVL). Scale bars are 2 mm. B: ROM 39638, female paratype (27.9 mm SVL), from Mt. Ayanganna. Photos by PJRK (A) and AL (B)." httpUri="https://zenodo.org/record/199006/files/figure.png" pageId="11" pageNumber="29">2</figureCitation>
B with figs. 11 and 13 illustrating
<taxonomicName id="4C1D4D31FFA7FF996F1F7991FB91FE4F" ID-CoL="66V5P" box="[824,1026,413,438]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="tamacuarensis">
<emphasis id="B969EAA0FFA7FF996F1F7991FB91FE4F" box="[824,1026,413,438]" italics="true" pageId="11" pageNumber="29">A. tamacuarensis</emphasis>
</taxonomicName>
in
<bibRefCitation id="EF8C4B43FFA7FF9968017991FACEFE4E" author="Myers" box="[1062,1373,413,439]" pageId="11" pageNumber="29" refString="Myers, C. W. &amp; Donnelly, M. A. (1997) A tepui herpetofauna on a granitic mountain (Tamacuari) in the borderland between Venezuela and Brazil: report from the Phipps Tapirapeco expedition. American Museum Novitates, 3213, 1 - 71." type="journal article" year="1997">Myers &amp; Donnelly (1997)</bibRefCitation>
], less developed keel-like lateral folds on fingers (extensive, present pre- and postaxially on all fingers), a conspicuous tympanum (inconspicuous), and in having a short, wider than long median lingual process (slender, noticeably longer than wide).
</paragraph>
</subSubSection>
</treatment>
</document>