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<document id="1C27A89DC2D81D183ED819D213FD778F" ID-CLB-Dataset="169975" ID-DOI="10.11646/zootaxa.5196.2.2" ID-GBIF-Dataset="96e1a27d-a189-4ed0-97e0-c02192b3828d" ID-ISSN="1175-5326" ID-Zenodo-Dep="7224439" ID-ZooBank="8107F44A-C50A-43BB-9D28-0E990E8AA0FC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666167075819" checkinUser="plazi" docAuthor="Sittichaya, Wisut &amp; Smith, Sarah M." docDate="2022" docId="03B387EC1E24FFBEFF54FB78FE13D3CD" docLanguage="en" docName="zootaxa.5196.2.2.pdf" docOrigin="Zootaxa 5196 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amasa macarthorum Sittichaya &amp; Smith 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="205" masterDocId="FF8AFF941E27FFB6FFC3FFB7FFC5D708" masterDocTitle="New species and new records of Amasa Lea, 1894 ambrosia beetles from Thailand (Coleoptera: Curculionidae: Scolytinae: Xyleborini)" masterLastPageNumber="210" masterPageNumber="197" pageNumber="200" updateTime="1699408989087" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="ECE89F8F033984F74822A97301AF06DD">New species and new records of Amasa Lea, 1894 ambrosia beetles from Thailand (Coleoptera: Curculionidae: Scolytinae: Xyleborini)</mods:title>
</mods:titleInfo>
<mods:name id="41AD7B0D39308EAFCEBC1BB0A54DA7C4" type="personal">
<mods:role id="A0540A7B9D7BCA56110C9AD730F146E5">
<mods:roleTerm id="3F7BD342936103359ED98FFE2DA9275D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C54F25E6C8431E9EC305C489ABE0C7A3">Sittichaya, Wisut</mods:namePart>
<mods:affiliation id="8397132E770F8D4CF8077FC7029B8B83">Agricultural Innovation and Management Division, Faculty of Natural Resources, Prince of Songkla University, Songkhla, 90110, Thailand</mods:affiliation>
</mods:name>
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<mods:role id="21A8F36DCBB9DED10AEE92BB50FC19A6">
<mods:roleTerm id="F01B463ED167FAAECF416D174B81B811">Author</mods:roleTerm>
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<mods:namePart id="C470274CCEC9EA4F8C470AA9C04CAAAF">Smith, Sarah M.</mods:namePart>
<mods:nameIdentifier id="039017C96AADDEE62F05561808F1FD95" type="ORCID">0000-0002-5173-3736</mods:nameIdentifier>
<mods:affiliation id="7D581EA7C17B06B3F53154619048966B">Department of Entomology, Michigan State University, 288 Farm Lane, 243 Natural Science Bldg., East Lansing, MI 48824, USA smith 462 @ msu. edu; https: // orcid. org / 0000 - 0002 - 5173 - 3736</mods:affiliation>
<mods:nameIdentifier id="3F369E095C562B8B83D688A39908B3A9" type="email">smith462@msu.edu</mods:nameIdentifier>
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<mods:title id="D170DD24D59A1F870EE97F6DB428060A">Zootaxa</mods:title>
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<mods:part id="A3676AED87F8006ACFB4053269F5ADFA">
<mods:date id="6D8638546868A229C5AEF51425B11CC7">2022</mods:date>
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<mods:number id="E1491A783A26580E6EA500A9CC7F2ABD">2022-10-19</mods:number>
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<treatment id="03B387EC1E24FFBEFF54FB78FE13D3CD" ID-DOI="http://doi.org/10.5281/zenodo.7224467" ID-GBIF-Taxon="203119836" ID-Zenodo-Dep="7224467" LSID="urn:lsid:plazi:treatment:03B387EC1E24FFBEFF54FB78FE13D3CD" httpUri="http://treatment.plazi.org/id/03B387EC1E24FFBEFF54FB78FE13D3CD" lastPageId="8" lastPageNumber="205" pageId="3" pageNumber="200">
<subSubSection id="C30065711E24FFB5FF54FB78FD31D3E1" box="[151,756,1230,1257]" pageId="3" pageNumber="200" type="nomenclature">
<paragraph id="8BA536FA1E24FFB5FF54FB78FD31D3E1" blockId="3.[151,1169,1230,1329]" box="[151,756,1230,1257]" pageId="3" pageNumber="200">
<heading id="D0ED81961E24FFB5FF54FB78FD31D3E1" bold="true" box="[151,756,1230,1257]" fontSize="11" level="1" pageId="3" pageNumber="200" reason="1">
<emphasis id="B96EEAE81E24FFB5FF54FB78FD31D3E1" bold="true" box="[151,756,1230,1257]" pageId="3" pageNumber="200">
<taxonomicName id="4C1A4D791E24FFB5FF54FB78FD4AD3E0" authority="Sittichaya &amp; Smith" authorityName="Sittichaya &amp; Smith" authorityYear="2022" box="[151,655,1230,1257]" class="Insecta" family="Curculionidae" genus="Amasa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="species" species="macarthorum" status="sp. nov.">
<emphasis id="B96EEAE81E24FFB5FF54FB78FE5DD3E1" bold="true" box="[151,408,1230,1257]" italics="true" pageId="3" pageNumber="200">Amasa macarthorum</emphasis>
Sittichaya &amp; Smith
</taxonomicName>
<taxonomicNameLabel id="A25D57931E24FFB5FD55FB78FD31D3E1" box="[662,756,1231,1257]" pageId="3" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30065711E24FFB5FF54FB44FB54D239" pageId="3" pageNumber="200" type="description">
<paragraph id="8BA536FA1E24FFB5FF54FB44FF2ED205" blockId="3.[151,1169,1230,1329]" box="[151,235,1267,1293]" pageId="3" pageNumber="200">
(
<figureCitation id="13212A7F1E24FFB5FF5CFB44FF26D205" box="[159,227,1267,1293]" captionStart="FIGURE 4" captionStartId="7.[151,250,1968,1992]" captionTargetBox="[185,1401,181,1938]" captionTargetId="figure-19@7.[185,1401,181,1944]" captionTargetPageId="7" captionText="FIGURE 4. Amasa macarthorum sp. nov. holotype A) dorsal view; B) lateral view; C) postero-lateral view; D) frons; E) venter and antennae." figureDoi="http://doi.org/10.5281/zenodo.7224447" httpUri="https://zenodo.org/record/7224447/files/figure.png" pageId="3" pageNumber="200">Fig. 4</figureCitation>
)
</paragraph>
<paragraph id="8BA536FA1E24FFB5FF54FAA1FB54D239" blockId="3.[151,1169,1230,1329]" box="[151,1169,1302,1329]" pageId="3" pageNumber="200">
http://zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="FFBC0C2F1E24FFB5FD40FAA1FB54D239" box="[643,1169,1302,1329]" pageId="3" pageNumber="200">F6B675A6-F349-4865-B35D-D0E970524B93</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30065711E24FFB5FF54FAEFFEACD142" pageId="3" pageNumber="200" type="materials_examined">
<paragraph id="8BA536FA1E24FFB5FF54FAEFFEACD142" blockId="3.[151,1437,1368,2043]" pageId="3" pageNumber="200">
<emphasis id="B96EEAE81E24FFB5FF54FAEFFE42D27A" bold="true" box="[151,391,1368,1394]" pageId="3" pageNumber="200">Material examined.</emphasis>
<materialsCitation id="3B723CA71E24FFB5FE57FAEFFE7FD2B2" ID-GBIF-Occurrence="3938604305" collectingDate="2019-07-21" collectorName="W. Sittichaya" country="Thailand" latitude="16.394112" location="Dry-dipterocarp forest" longLatPrecision="1" longitude="101.56883" municipality="Phu Khiao Wildlife Sanctuary" pageId="3" pageNumber="200" specimenCount="1" stateProvince="Chaiyaphum" typeStatus="holotype">
<emphasis id="B96EEAE81E24FFB5FE57FAEFFDABD27A" bold="true" box="[404,622,1368,1394]" pageId="3" pageNumber="200">
<typeStatus id="54A188581E24FFB5FE57FAEFFDC6D27A" box="[404,515,1368,1394]" pageId="3" pageNumber="200" type="holotype">Holotype</typeStatus>
, female:
</emphasis>
<collectingCountry id="F30D766A1E24FFB5FDB8FAEFFCCAD27A" box="[635,783,1368,1394]" name="Thailand" pageId="3" pageNumber="200">THAILAND</collectingCountry>
,
<collectingRegion id="49DEF8181E24FFB5FCDCFAEFFBE7D27A" box="[799,1058,1368,1394]" country="Thailand" name="Chaiyaphum" pageId="3" pageNumber="200">Chaiyaphum Province</collectingRegion>
,
<collectingMunicipality id="6BC1AC801E24FFB5FBF7FAEFFA5DD27A" box="[1076,1432,1368,1394]" pageId="3" pageNumber="200">Phu Khiao Wildlife Sanctuary</collectingMunicipality>
,
<geoCoordinate id="EE2E503D1E24FFB5FF54FACAFEF4D290" box="[151,305,1405,1432]" degrees="16" direction="north" minutes="23" orientation="latitude" pageId="3" pageNumber="200" precision="1" seconds="38.8" value="16.394112">16°2338.8″N</geoCoordinate>
<geoCoordinate id="EE2E503D1E24FFB5FEFAFACAFE25D290" box="[313,480,1405,1432]" degrees="101" direction="east" minutes="34" orientation="longitude" pageId="3" pageNumber="200" precision="1" seconds="07.8" value="101.56883">101°3407.8″E</geoCoordinate>
,
<location id="8EC560211E24FFB5FE2EFACBFD2DD29E" LSID="urn:lsid:plazi:treatment:03B387EC1E24FFBEFF54FB78FE13D3CD:8EC560211E24FFB5FE2EFACBFD2DD29E" box="[493,744,1404,1430]" country="Thailand" latitude="16.394112" longLatPrecision="1" longitude="101.56883" municipality="Phu Khiao Wildlife Sanctuary" name="Dry-dipterocarp forest" pageId="3" pageNumber="200" stateProvince="Chaiyaphum">Dry-dipterocarp forest</location>
with pine subtype, 870 msl.,
<collectingMethod id="525B4EED1E24FFB5FBF5FACBFACFD29F" box="[1078,1290,1404,1431]" pageId="3" pageNumber="200">ethanol baited trap</collectingMethod>
,
<date id="FFA4103A1E24FFB5FAD5FACBFA5DD29E" box="[1302,1432,1404,1430]" pageId="3" pageNumber="200" value="2019-07-21">
<collectingDate id="EFE0E9D21E24FFB5FAD5FACBFA5DD29E" box="[1302,1432,1404,1430]" pageId="3" pageNumber="200" value="2019-07-21">21.vii.2019</collectingDate>
</date>
,
<collectorName id="26EF532C1E24FFB5FF54FA16FEF2D2B2" box="[151,311,1440,1467]" pageId="3" pageNumber="200">W. Sittichaya</collectorName>
, (NHMW)
</materialsCitation>
;
<materialsCitation id="3B723CA71E24FFB5FE0FFA17FEA0D142" collectingDate="2005-07-05" collectingDateMax="2011-06-01" collectingDateMin="2005-07-05" collectionCode="WSTC, MSUC" collectorName="J. Hulcr &amp; S. Dole" country="Thailand" latitude="17.282804" location="Phu Luang Wildlife Sanctuary" longLatPrecision="1" longitude="101.51956" municipality="Khao Luang National Park" pageId="3" pageNumber="200" specimenCount="1" stateProvince="Loei" typeStatus="paratype">
<emphasis id="B96EEAE81E24FFB5FE0FFA17FD8ED2B2" bold="true" box="[460,587,1440,1466]" pageId="3" pageNumber="200">
<typeStatus id="54A188581E24FFB5FE0FFA17FD80D2B2" box="[460,581,1440,1466]" pageId="3" pageNumber="200" type="paratype">Paratypes</typeStatus>
,
</emphasis>
female, Nakhon Sri Thammarat Province,
<collectingMunicipality id="6BC1AC801E24FFB5FB97FA17FA5DD2B2" box="[1108,1432,1440,1467]" pageId="3" pageNumber="200">Khao Luang National Park</collectingMunicipality>
,
<geoCoordinate id="EE2E503D1E24FFB5FF54FA72FEE6D2E8" box="[151,291,1477,1504]" degrees="8" direction="north" minutes="43" orientation="latitude" pageId="3" pageNumber="200" precision="1" seconds="08.7" value="8.719083">8°4308.7″N</geoCoordinate>
<geoCoordinate id="EE2E503D1E24FFB5FEE8FA72FE00D2E8" box="[299,453,1477,1504]" degrees="99" direction="east" minutes="40" orientation="longitude" pageId="3" pageNumber="200" precision="1" seconds="39.2" value="99.67755">99°4039.2″E</geoCoordinate>
, tropical rainforest,
<collectingMethod id="525B4EED1E24FFB5FD6AFA73FCBBD2D7" box="[681,894,1476,1503]" pageId="3" pageNumber="200">ethanol baited trap</collectingMethod>
,
<date id="FFA4103A1E24FFB5FC49FA73FBC1D2D6" box="[906,1028,1476,1502]" pageId="3" pageNumber="200" value="2011-06-01">
<collectingDate id="EFE0E9D21E24FFB5FC49FA73FBC1D2D6" box="[906,1028,1476,1502]" pageId="3" pageNumber="200" value="2011-06-01">01.vi.2011</collectingDate>
</date>
,
<collectingRegion id="49DEF8181E24FFB5FBD3FA73FB74D2D6" box="[1040,1201,1476,1502]" country="Thailand" name="Loei" pageId="3" pageNumber="200">Loei Province</collectingRegion>
,
<location id="8EC560211E24FFB5FB7DFA73FECDD10A" LSID="urn:lsid:plazi:treatment:03B387EC1E24FFBEFF54FB78FE13D3CD:8EC560211E24FFB5FB7DFA73FECDD10A" country="Thailand" latitude="17.282804" longLatPrecision="1" longitude="101.51956" municipality="Khao Luang National Park" name="Phu Luang Wildlife Sanctuary" pageId="3" pageNumber="200" stateProvince="Loei">Phu Luang Wildlife Sanctuary</location>
,
<geoCoordinate id="EE2E503D1E24FFB5FED4FA5EFE77D10C" box="[279,434,1513,1540]" degrees="17" direction="north" minutes="16" orientation="latitude" pageId="3" pageNumber="200" precision="1" seconds="58.1" value="17.282804">17°1658.1″N</geoCoordinate>
<geoCoordinate id="EE2E503D1E24FFB5FE7EFA5EFDA1D10C" box="[445,612,1513,1540]" degrees="101" direction="east" minutes="31" orientation="longitude" pageId="3" pageNumber="200" precision="1" seconds="10.4" value="101.51956">101°3110.4″E</geoCoordinate>
, 1,460 msl.,
<collectorName id="26EF532C1E24FFB5FD3CFA5FFCEED10A" box="[767,811,1512,1538]" pageId="3" pageNumber="200">Hill</collectorName>
evergreen forest, ex dry branch, 25.vi.19, (1) (all
<collectorName id="26EF532C1E24FFB5FABEFA5EFECCD12E" pageId="3" pageNumber="200">W. Sittichaya</collectorName>
) (2
<collectionCode id="ED0BAE3F1E24FFB5FEFDF9BBFE55D12E" box="[318,400,1548,1574]" pageId="3" pageNumber="200">WSTC</collectionCode>
),
<collectingRegion id="49DEF8181E24FFB5FE64F9BBFD6DD12E" box="[423,680,1548,1574]" country="Thailand" name="Chaiyaphum" pageId="3" pageNumber="200">Chaiyaphum Province</collectingRegion>
,
<collectorName id="26EF532C1E24FFB5FD74F9BBFBD3D12E" box="[695,1046,1548,1574]" pageId="3" pageNumber="200">Phu Khiao Wildlife Sanctuary</collectorName>
,
<date id="FFA4103A1E24FFB5FBE6F9BBFB5FD12F" box="[1061,1178,1548,1575]" pageId="3" pageNumber="200" value="2005-07-05">
<collectingDate id="EFE0E9D21E24FFB5FBE6F9BBFB5FD12F" box="[1061,1178,1548,1575]" pageId="3" pageNumber="200" value="2005-07-05">5.vii.2005</collectingDate>
</date>
,
<collectorName id="26EF532C1E24FFB5FB6AF9BAFACDD12E" box="[1193,1288,1548,1575]" pageId="3" pageNumber="200">J. Hulcr</collectorName>
,
<collectorName id="26EF532C1E24FFB5FAD4F9BBFAB7D12E" box="[1303,1394,1548,1574]" pageId="3" pageNumber="200">S. Dole</collectorName>
, ex branch (1
<collectionCode id="ED0BAE3F1E24FFB5FECAF987FE98D142" box="[265,349,1584,1610]" pageId="3" pageNumber="200">MSUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30065711E24FFB5FF04F9E3FB0CD02A" pageId="3" pageNumber="200" type="diagnosis">
<paragraph id="8BA536FA1E24FFB5FF04F9E3FEABD1F6" blockId="3.[151,1437,1368,2043]" pageId="3" pageNumber="200">
<emphasis id="B96EEAE81E24FFB5FF04F9E3FE81D166" bold="true" box="[199,324,1620,1646]" pageId="3" pageNumber="200">Diagnosis:</emphasis>
<quantity id="4CE29B1F1E24FFB5FE89F9E3FE29D167" box="[330,492,1620,1647]" metricMagnitude="-3" metricUnit="m" metricValue="2.94" metricValueMax="2.98" metricValueMin="2.9" pageId="3" pageNumber="200" unit="mm" value="2.94" valueMax="2.98" valueMin="2.9">2.902.98 mm</quantity>
long; 2.44× as long as wide. This species is distinguished by the pronotum appearing basic (
<typeStatus id="54A188581E24FFB5FF1DF9CFFED5D19A" box="[222,272,1656,1682]" pageId="3" pageNumber="200">type</typeStatus>
2) when viewed dorsally, anterior margin serrate, pronotal anterior slope shagreened with densely spaced, short fine asperities, strial punctures small, shallow and inconspicuous, elytral discal interstriae densely punctate, punctures small and confused, not seriate, declivital face smooth and shining, punctures on declivity large, much larger then on disc.
</paragraph>
<paragraph id="8BA536FA1E24FFB5FF04F8BFFB0CD02A" blockId="3.[151,1437,1368,2043]" box="[199,1225,1800,1827]" pageId="3" pageNumber="200">
<emphasis id="B96EEAE81E24FFB5FF04F8BFFE47D02A" bold="true" box="[199,386,1800,1826]" pageId="3" pageNumber="200">Similar species:</emphasis>
<taxonomicName id="4C1A4D791E24FFB5FE4AF8BFFCE9D02A" authority="Smith, Beaver &amp; Cognato" authorityName="Smith, Beaver &amp; Cognato" box="[393,812,1800,1827]" class="Insecta" family="Curculionidae" genus="Amasa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="species" species="lini">
<emphasis id="B96EEAE81E24FFB5FE4AF8BFFDC4D02A" box="[393,513,1800,1826]" italics="true" pageId="3" pageNumber="200">Amasa lini</emphasis>
Smith, Beaver &amp; Cognato
</taxonomicName>
,
<taxonomicName id="4C1A4D791E24FFB5FCFAF8BFFB0CD02A" authority="Smith, Beaver &amp; Cognato." authorityName="Smith, Beaver &amp; Cognato." box="[825,1225,1800,1827]" class="Insecta" family="Curculionidae" genus="Amasa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="species" species="youlii">
<emphasis id="B96EEAE81E24FFB5FCFAF8BFFC5DD02A" box="[825,920,1800,1826]" italics="true" pageId="3" pageNumber="200">A. youlii</emphasis>
Smith, Beaver &amp; Cognato.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30065711E24FFBEFF04F89BFBE8D319" lastPageId="8" lastPageNumber="205" pageId="3" pageNumber="200" type="description">
<paragraph id="8BA536FA1E24FFBEFF04F89BFBE8D319" blockId="3.[151,1437,1368,2043]" lastBlockId="8.[151,1437,151,1221]" lastPageId="8" lastPageNumber="205" pageId="3" pageNumber="200">
<emphasis id="B96EEAE81E24FFB5FF04F89BFE00D04E" bold="true" box="[199,453,1836,1862]" pageId="3" pageNumber="200">Description (female).</emphasis>
<quantity id="4CE29B1F1E24FFB5FE0CF89BFDB3D04F" box="[463,630,1836,1863]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="3" pageNumber="200" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.903.10 mm</quantity>
long (mean =
<quantity id="4CE29B1F1E24FFB5FCE5F89BFC55D04F" box="[806,912,1836,1863]" metricMagnitude="-3" metricUnit="m" metricValue="2.99" pageId="3" pageNumber="200" unit="mm" value="2.99">2.99 mm</quantity>
; n = 3; pronotal anterior margin to declivital summit
<quantity id="4CE29B1F1E24FFB5FF31F8E7FE52D063" box="[242,407,1872,1899]" metricMagnitude="-3" metricUnit="m" metricValue="2.64" metricValueMax="2.7" metricValueMin="2.58" pageId="3" pageNumber="200" unit="mm" value="2.64" valueMax="2.7" valueMin="2.58">2.582.70 mm</quantity>
, mean
<quantity id="4CE29B1F1E24FFB5FE26F8E7FD89D063" box="[485,588,1872,1899]" metricMagnitude="-3" metricUnit="m" metricValue="2.63" pageId="3" pageNumber="200" unit="mm" value="2.63">2.63 mm</quantity>
; n = 2); 2.272.44× as long as wide. Body shining, bicolored: pronotal disc, head, legs, antennae and venter brown to dark brown, anterior slope of pronotum, and elytra, dark reddish brown.
<emphasis id="B96EEAE81E24FFB5FF54F82FFF1BD0BA" bold="true" box="[151,222,1944,1970]" pageId="3" pageNumber="200">Head.</emphasis>
Epistoma entire, transverse, with row of hair-like setae. Frons weakly convex to upper level of eyes. Median impression between eyes triangular and short, surface subshiny shagreen with longitudinal asperities. Frons below eyes alutaceous, subshiny, with moderately large longitudinal asperities, asperities rounder, denser and larger above epistoma, increasing in length dorsally and laterally. Eyes deeply emarginate just above antennal insertion, upper part smaller than lower part. Submentum triangular, deeply impressed. Antennal scape regularly thick, shorter than club (5:8). Pedicel as wide as scape, shorter than funicle. Funicle 4-segmented, segment 1 approximately equal in length with pedicel. Club taller than wide (8:6) and flat,
<typeStatus id="54A188581E2FFFBEFCF8FEB4FCAED615" box="[827,875,259,285]" pageId="8" pageNumber="205">type</typeStatus>
4; segment 1 corneous, small convex on anterior face, occupying approximately basal 1/5; corneous parts of segment 2 and 3 wide nearly as wide as segment 1; segments 13 present on posterior face.
<emphasis id="B96EEAE81E2FFFBEFD98FEFCFD1ED66D" bold="true" box="[603,731,331,357]" pageId="8" pageNumber="205">Pronotum.</emphasis>
1.04× as long as wide. In dorsal view basic and parallel-sided,
<typeStatus id="54A188581E2FFFBEFF54FED8FF02D681" box="[151,199,367,393]" pageId="8" pageNumber="205">type</typeStatus>
2, sides parallel in basal 1/2, rounded anteriorly; anterior margin with a row of seven or eight serrations. In lateral view basic,
<typeStatus id="54A188581E2FFFBEFEB7FE24FE61D6A5" box="[372,420,403,429]" pageId="8" pageNumber="205">type</typeStatus>
0, disc flat, summit at midpoint. Anterior slope shagreened with densely spaced, short fine asperities, becoming lower and more transverse towards summit, bearing long, fine, semi-recumbent hair-like setae. Disc shining, alutaceous, densely minutely punctate, glabrous. Lateral margins sharp obliquely costate. Base transverse, posterior angles narrowly rounded.
<emphasis id="B96EEAE81E2FFFBEFD6CFE48FCC7D511" bold="true" box="[687,770,511,537]" pageId="8" pageNumber="205">Elytra.</emphasis>
1.25× as long as wide, 1.25× as long as pronotum. Elytra subshiny, alutaceous. Scutellum small, narrow, linguiform, flush with elytra, flat, shining, its anterior portion extend forwardly beyond elytral base. Base bisinuate, edge carinate, humeral angles rounded, parallel-sided in basal 4/5, then sharply angulate to apex. Disc flat, shining, glabrous; striae and interstriae laterally slightly diverging from base to declivital summit; striae punctate, punctures small, shallow; interstriae flat, finely punctate, punctures 1/3 size of strial punctures, strongly confused. Declivity truncate, face slightly ascending posteriorly, margin sharp carinated; upper portion of face near lateral margin flattened, lower part on the middle strongly elevated and bearing uniseriate, round apical tubercles on middle of interstria 1. Declivital face strongly shining, smooth; three striae present, strial punctures shining, small, moderately deep, larger than on disc; interstriae slightly inflated, very fine inconspicuous punctate, punctures wide separated; each with very fine very short hair-like setae (visible with high magnitude). Posterolateral margin forming a circumdeclivital carina; carina globous without setae.
<emphasis id="B96EEAE81E2FFFBEFBA7FCD0FB66D489" bold="true" box="[1124,1187,871,897]" pageId="8" pageNumber="205">Legs.</emphasis>
Procoxae contiguous, prosternal coxal piece flat, inconspicuous. Protibiae slender, broadest at apical 1/3; posterior face inflated, coarsely granulate; apical 1/2 of outer margin with six medium size socketed denticles, their length 1/2 time as long as basal width. Meso- and metatibiae broad, flattened, outer margins evenly rounded with 11 and 12 small socketed denticles, respectively; posterior faces unarmed; anterior faces coarsely granulate.
</paragraph>
</subSubSection>
<caption id="DF6566721E23FFB2FF54F830FBF3D0EF" ID-DOI="http://doi.org/10.5281/zenodo.7224441" ID-Zenodo-Dep="7224441" httpUri="https://zenodo.org/record/7224441/files/figure.png" pageId="4" pageNumber="201" startId="4.[151,250,1927,1951]" targetBox="[258,1324,197,1892]" targetPageId="4">
<paragraph id="8BA536FA1E23FFB2FF54F830FBF3D0EF" blockId="4.[151,1437,1927,2023]" pageId="4" pageNumber="201">
<emphasis id="B96EEAE81E23FFB2FF54F830FED3D097" bold="true" box="[151,278,1927,1951]" pageId="4" pageNumber="201">FIGURE 1.</emphasis>
Insect survey localities in Thailand. Red stars indicate surveyed areas, blue stars indicate locations where new species and newly recorded
<taxonomicName id="4C1A4D791E23FFB2FE03F81BFD92D0CB" box="[448,599,1963,1987]" pageId="4" pageNumber="201">
<emphasis id="B96EEAE81E23FFB2FE03F81BFDC0D0CB" box="[448,517,1964,1987]" italics="true" pageId="4" pageNumber="201">Amasa</emphasis>
species
</taxonomicName>
were captured. Yellow stars indicate localities of previously recorded
<taxonomicName id="4C1A4D791E23FFB2FAF9F81BFABAD0CB" authorityName="Lea" authorityYear="1894" box="[1338,1407,1964,1987]" class="Insecta" family="Curculionidae" genus="Amasa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="201" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAE81E23FFB2FAF9F81BFABAD0CB" box="[1338,1407,1964,1987]" italics="true" pageId="4" pageNumber="201">Amasa</emphasis>
</taxonomicName>
in Thailand. Source: the map was modified from a Royal Forest Department of Thailand map.
</paragraph>
</caption>
<caption id="DF6566721E22FFB3FF54F804FEDDD0E6" ID-DOI="http://doi.org/10.5281/zenodo.7224443" ID-Zenodo-Dep="7224443" httpUri="https://zenodo.org/record/7224443/files/figure.png" pageId="5" pageNumber="202" startId="5.[151,250,1971,1995]" targetBox="[166,1420,181,1934]" targetPageId="5">
<paragraph id="8BA536FA1E22FFB3FF54F804FEDDD0E6" blockId="5.[151,1436,1971,2031]" pageId="5" pageNumber="202">
<emphasis id="B96EEAE81E22FFB3FF54F804FED6D0C3" bold="true" box="[151,275,1971,1995]" pageId="5" pageNumber="202">FIGURE 2.</emphasis>
<taxonomicName id="4C1A4D791E22FFB3FEDAF804FE75D0C3" authority="Sittichaya &amp; Smith, 2022" authorityName="Sittichaya &amp; Smith" authorityYear="2022" box="[281,432,1971,1995]" class="Insecta" family="Curculionidae" genus="Amasa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="202" phylum="Arthropoda" rank="species" species="beaveri" status="sp. nov.">
<emphasis id="B96EEAE81E22FFB3FEDAF804FE75D0C3" box="[281,432,1971,1995]" italics="true" pageId="5" pageNumber="202">Amasa beaveri</emphasis>
</taxonomicName>
<emphasis id="B96EEAE81E22FFB3FE75F804FDC3D0C3" bold="true" box="[438,518,1971,1995]" pageId="5" pageNumber="202">
<taxonomicNameLabel id="A25D57931E22FFB3FE75F804FDC3D0C3" box="[438,518,1971,1995]" pageId="5" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype A) dorsal view; B) lateral view; C) postero-lateral view; D) frons; E) head lateral and antenna.
</paragraph>
</caption>
<caption id="DF6566721E21FFB0FF54F803FF3CD0F8" ID-DOI="http://doi.org/10.5281/zenodo.7224445" ID-Zenodo-Dep="7224445" httpUri="https://zenodo.org/record/7224445/files/figure.png" pageId="6" pageNumber="203" startId="6.[151,250,1972,1996]" targetBox="[214,1374,181,1948]" targetPageId="6">
<paragraph id="8BA536FA1E21FFB0FF54F803FF3CD0F8" blockId="6.[151,1437,1972,2032]" pageId="6" pageNumber="203">
<emphasis id="B96EEAE81E21FFB0FF54F803FED7D0C3" bold="true" box="[151,274,1972,1996]" pageId="6" pageNumber="203">FIGURE 3.</emphasis>
<taxonomicName id="4C1A4D791E21FFB0FEDAF803FE78D0C4" authority="Sittichaya &amp; Smith, 2022" authorityName="Sittichaya &amp; Smith" authorityYear="2022" box="[281,445,1972,1996]" class="Insecta" family="Curculionidae" genus="Amasa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="203" phylum="Arthropoda" rank="species" species="cognatoi" status="sp. nov.">
<emphasis id="B96EEAE81E21FFB0FEDAF803FE78D0C4" box="[281,445,1972,1996]" italics="true" pageId="6" pageNumber="203">Amasa cognatoi</emphasis>
</taxonomicName>
<emphasis id="B96EEAE81E21FFB0FE01F803FDD6D0C4" bold="true" box="[450,531,1972,1996]" pageId="6" pageNumber="203">
<taxonomicNameLabel id="A25D57931E21FFB0FE01F803FDD6D0C4" box="[450,531,1972,1996]" pageId="6" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype A) dorsal view; B) lateral view; C) postero-lateral view; D) frons; E) venter and antennae.
</paragraph>
</caption>
<caption id="DF6566721E20FFB1FF54F807FEE6D0E4" ID-DOI="http://doi.org/10.5281/zenodo.7224447" ID-Zenodo-Dep="7224447" httpUri="https://zenodo.org/record/7224447/files/figure.png" pageId="7" pageNumber="204" startId="7.[151,250,1968,1992]" targetBox="[185,1401,181,1938]" targetPageId="7">
<paragraph id="8BA536FA1E20FFB1FF54F807FEE6D0E4" blockId="7.[151,1436,1968,2028]" pageId="7" pageNumber="204">
<emphasis id="B96EEAE81E20FFB1FF54F807FED7D0C0" bold="true" box="[151,274,1968,1992]" pageId="7" pageNumber="204">FIGURE 4.</emphasis>
<taxonomicName id="4C1A4D791E20FFB1FEDBF806FE29D0C0" authority="Sittichaya &amp; Smith, 2022" authorityName="Sittichaya &amp; Smith" authorityYear="2022" box="[280,492,1968,1992]" class="Insecta" family="Curculionidae" genus="Amasa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="macarthorum" status="sp. nov.">
<emphasis id="B96EEAE81E20FFB1FEDBF806FE29D0C0" box="[280,492,1968,1992]" italics="true" pageId="7" pageNumber="204">Amasa macarthorum</emphasis>
</taxonomicName>
<emphasis id="B96EEAE81E20FFB1FE31F807FD87D0C0" bold="true" box="[498,578,1968,1992]" pageId="7" pageNumber="204">
<taxonomicNameLabel id="A25D57931E20FFB1FE31F807FD87D0C0" box="[498,578,1968,1992]" pageId="7" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype A) dorsal view; B) lateral view; C) postero-lateral view; D) frons; E) venter and antennae.
</paragraph>
</caption>
<subSubSection id="C30065711E2FFFBEFF04FBACFD2ED375" pageId="8" pageNumber="205" type="etymology">
<paragraph id="8BA536FA1E2FFFBEFF04FBACFD2ED375" blockId="8.[151,1437,151,1221]" pageId="8" pageNumber="205">
<emphasis id="B96EEAE81E2FFFBEFF04FBACFE97D33D" bold="true" box="[199,338,1051,1077]" pageId="8" pageNumber="205">Etymology:</emphasis>
The name honors the John D. and Catherine T. MacArthur Foundation, the primary funding source for the Biodiversity Heritage Library, which “serves to improve research methodology by collaboratively making biodiversity literature openly available to the world”.
</paragraph>
</subSubSection>
<subSubSection id="C30065711E2FFFBEFF04FB31FAF3D3A8" box="[199,1334,1158,1185]" pageId="8" pageNumber="205" type="distribution">
<paragraph id="8BA536FA1E2FFFBEFF04FB31FAF3D3A8" blockId="8.[151,1437,151,1221]" box="[199,1334,1158,1185]" pageId="8" pageNumber="205">
<emphasis id="B96EEAE81E2FFFBEFF04FB31FEA6D3A8" bold="true" box="[199,355,1158,1184]" pageId="8" pageNumber="205">Distribution:</emphasis>
<collectingCountry id="F30D766A1E2FFFBEFEA9FB30FE3AD3A9" box="[362,511,1159,1185]" name="Thailand" pageId="8" pageNumber="205">THAILAND</collectingCountry>
:
<collectingRegion id="49DEF8181E2FFFBEFDCAFB30FCC0D3A9" box="[521,773,1159,1185]" country="Thailand" name="Chaiyaphum" pageId="8" pageNumber="205">Chaiyaphum Province</collectingRegion>
,
<collectingRegion id="49DEF8181E2FFFBEFCD2FB30FC74D3A9" box="[785,945,1159,1185]" country="Thailand" name="Loei" pageId="8" pageNumber="205">Loei Province</collectingRegion>
, Nakhon Sri Thammarat Province.
</paragraph>
</subSubSection>
<subSubSection id="C30065711E2FFFBEFF04FB1CFE13D3CD" box="[199,470,1195,1221]" pageId="8" pageNumber="205" type="biology_ecology">
<paragraph id="8BA536FA1E2FFFBEFF04FB1CFE13D3CD" blockId="8.[151,1437,151,1221]" box="[199,470,1195,1221]" pageId="8" pageNumber="205">
<emphasis id="B96EEAE81E2FFFBEFF04FB1CFE9CD3CD" bold="true" box="[199,345,1195,1221]" pageId="8" pageNumber="205">Host plants:</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>